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<mods:title id="A31743959213FB4428FE2A1F5F3DC8C3">On the land snail Rhagada Albers, 1860 (Gastropoda: Camaenidae) from across the Lesser Sunda Islands</mods:title>
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<treatment id="03F3175FEF6B5E41FF213ED4FEFAF9B3" ID-DOI="http://doi.org/10.5281/zenodo.4775640" ID-GBIF-Taxon="182237994" ID-Zenodo-Dep="4775640" LSID="urn:lsid:plazi:treatment:03F3175FEF6B5E41FF213ED4FEFAF9B3" httpUri="http://treatment.plazi.org/id/03F3175FEF6B5E41FF213ED4FEFAF9B3" lastPageNumber="119" pageId="4" pageNumber="119">
<subSubSection id="C340F5C2EF6B5E41FF213ED4FD42FCE9" box="[255,656,767,791]" pageId="4" pageNumber="119" type="nomenclature">
<paragraph id="8BE5A649EF6B5E41FF213ED4FD42FCE9" blockId="4.[255,656,767,791]" box="[255,656,767,791]" pageId="4" pageNumber="119">
<heading id="D0AD1125EF6B5E41FF213ED4FD42FCE9" bold="true" box="[255,656,767,791]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="119" reason="2">
<taxonomicName id="4C5ADDCAEF6B5E41FF213ED4FD42FCE9" ID-CoL="4RYBY" authority="(Martens, 1891)" baseAuthorityName="Martens" baseAuthorityYear="1891" box="[255,656,767,791]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="floresiana">
<emphasis id="B92E7A5BEF6B5E41FF213ED4FE0BFCE9" bold="true" box="[255,473,767,791]" italics="true" pageId="4" pageNumber="119">Rhagada floresiana</emphasis>
(
<bibRefCitation id="EFCBDBB8EF6B5E41FE373F2BFD55FCE9" author="Martens E von" box="[489,647,768,791]" pageId="4" pageNumber="119" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C340F5C2EF6B5E41FF563F15FEC7FAE7" pageId="4" pageNumber="119" type="reference_group">
<paragraph id="8BE5A649EF6B5E41FF563F15FD8FFC73" blockId="4.[136,776,830,967]" pageId="4" pageNumber="119">
<treatmentCitationGroup id="AB4A8167EF6B5E41FF563F15FD8FFC73" pageId="4" pageNumber="119">
<treatmentCitation id="0AFB8058EF6B5E41FF563F15FD5DFCAD" author="Martens E von" box="[136,655,830,851]" page="236" pageId="4" pageNumber="119" year="1891">
<taxonomicName id="4C5ADDCAEF6B5E41FF563F15FD5DFCAD" ID-CoL="3KDR7" authority="Martens, 1891: 235 - 236" authorityName="Martens" authorityPageNumber="235 - 236" authorityYear="1891" box="[136,655,830,851]" class="Gastropoda" family="Camaenidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="floresiana" subGenus="Rhagada">
<emphasis id="B92E7A5BEF6B5E41FF563F15FF6EFCAD" box="[136,188,830,851]" italics="true" pageId="4" pageNumber="119">Helix</emphasis>
(
<emphasis id="B92E7A5BEF6B5E41FF133F15FEF6FCAD" box="[205,292,830,851]" italics="true" pageId="4" pageNumber="119">Rhagada</emphasis>
)
<emphasis id="B92E7A5BEF6B5E41FEEA3F15FE41FCAD" box="[308,403,830,851]" italics="true" pageId="4" pageNumber="119">floresiana</emphasis>
<bibRefCitation id="EFCBDBB8EF6B5E41FE423F15FD5DFCAD" author="Martens E von" box="[412,655,830,851]" pageId="4" pageNumber="119" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891: 235236</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 14, figs 13, 14 (from
<collectingCountry id="F34DE6D9EF6B5E41FEE43F70FEAFFC8E" box="[314,381,859,880]" name="Indonesia" pageId="4" pageNumber="119">Flores</collectingCountry>
,
<location id="8E85F092EF6B5E41FE573F70FE22FC8E" LSID="urn:lsid:plazi:treatment:03F3175FEF6B5E41FF213ED4FEFAF9B3:8E85F092EF6B5E41FE573F70FE22FC8E" box="[393,496,859,880]" country="Indonesia" latitude="-8.62" longLatPrecision="21" longitude="122.22" name="Maumere'" pageId="4" pageNumber="119">Maumere</location>
,
<geoCoordinate id="EE6EC08EEF6B5E41FE233F70FDA9FC8E" box="[509,635,859,880]" degrees="8" direction="south" minutes="37" orientation="latitude" pageId="4" pageNumber="119" precision="15" seconds="12" value="-8.62">8° 37' 12&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6EC08EEF6B5E41FD563F70FF6DFC73" degrees="122" direction="east" minutes="13" orientation="longitude" pageId="4" pageNumber="119" precision="15" seconds="12" value="122.22">122° 13' 12&quot; E</geoCoordinate>
;
<specimenCount id="9D5C6DC0EF6B5E41FF153F53FEE1FC73" box="[203,307,888,909]" pageId="4" pageNumber="119" type="generic" typeStatus="syntypes">3 syntypes</specimenCount>
ZMB Moll. 44732, not seen).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE5A649EF6B5E41FF563FBEFEF1FC39" blockId="4.[136,776,830,967]" pageId="4" pageNumber="119">
<treatmentCitationGroup id="AB4A8167EF6B5E41FF563FBEFEF1FC39" pageId="4" pageNumber="119">
<taxonomicName id="4C5ADDCAEF6B5E41FF563FBEFE7BFC54" ID-CoL="4RYBY" baseAuthorityName="Martens" baseAuthorityYear="1891" box="[136,425,917,938]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="subSpecies" species="solorensis" subSpecies="floresiana">
<emphasis id="B92E7A5BEF6B5E41FF563FBEFE7BFC54" box="[136,425,917,938]" italics="true" pageId="4" pageNumber="119">Rhagada solorensis floresiana</emphasis>
</taxonomicName>
<treatmentCitation id="0AFB8058EF6B5E41FE613FBEFDB5FC54" author="Rensch B" box="[447,615,917,938]" page="95" pageId="4" pageNumber="119" year="1931">
<bibRefCitation id="EFCBDBB8EF6B5E41FE613FBEFDB5FC54" author="Rensch B" box="[447,615,917,938]" pageId="4" pageNumber="119" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch, 1931: 95</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0AFB8058EF6B5E41FDAF3FBEFF14FC39" author="Maassen WJM" page="66" pageId="4" pageNumber="119" year="2009">
<bibRefCitation id="EFCBDBB8EF6B5E41FDAF3FBEFF14FC39" author="Maassen WJM" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen, 2009: 66</bibRefCitation>
</treatmentCitation>
(partim).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE5A649EF6B5E41FF563FC5FEC7FAE7" blockId="4.[136,776,1006,1305]" pageId="4" pageNumber="119">
<emphasis id="B92E7A5BEF6B5E41FF563FC5FE8CFBFB" bold="true" box="[136,350,1006,1029]" pageId="4" pageNumber="119">Taxonomic history.</emphasis>
<bibRefCitation id="EFCBDBB8EF6B5E41FEBC3FC5FDD7FBFB" author="Martens E von" box="[354,517,1006,1029]" pageId="4" pageNumber="119" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens (1891)</bibRefCitation>
recognised
<taxonomicName id="4C5ADDCAEF6B5E41FD5F3FC5FCD5FBFB" ID-CoL="4RYCR" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[641,775,1006,1029]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FD5F3FC5FCD5FBFB" box="[641,775,1006,1029]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
as distinct by its thinner, more turreted shell, less thickened columellar lip and slightly different colour. This taxon has subsequently been treated as a subspecies of
<taxonomicName id="4C5ADDCAEF6B5E41FD813861FD34FB9F" ID-CoL="4RYCR" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[607,742,1098,1121]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FD813861FD34FB9F" box="[607,742,1098,1121]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCBDBB8EF6B5E41FF563842FEF7FB7E" author="Rensch B" box="[136,293,1129,1152]" pageId="4" pageNumber="119" pagination="361 - 396" refId="ref6470" refString="Rensch B (1931) Die Molluskenfauna der Kleinen Sunda-Inseln Bali, Lombok, Sumbawa, Flores und Sumba. I. Zoologische Jahrbucher. Abteilung fur Systematik Okologie und Geographie der Tiere, 61: 361 - 396." type="journal article" year="1931">Rensch (1931)</bibRefCitation>
and
<bibRefCitation id="EFCBDBB8EF6B5E41FE843842FDD7FB7E" author="Maassen WJM" box="[346,517,1129,1152]" pageId="4" pageNumber="119" pagination="65 - 69" refId="ref6197" refString="Maassen WJM (2009) A new Rhagada species from Alor Island, Lesser Sunda Islands, Indonesia (Gastropoda, Pulmonata, Camaenidae). Miscellanea Malacologica, 3: 65 - 69." type="journal article" year="2009">Maassen (2009)</bibRefCitation>
based on shell features. I prefer following the original concept and maintaining
<taxonomicName id="4C5ADDCAEF6B5E41FD2E38A3FF23FB43" ID-CoL="3KDR7" authority="(Martens, 1891)" authorityName="Martens" authorityYear="1891" baseAuthorityName="Martens" baseAuthorityYear="1891" class="Gastropoda" family="Camaenidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="floresiana">
<emphasis id="B92E7A5BEF6B5E41FD2E38A3FF23FB43" italics="true" pageId="4" pageNumber="119">R. floresiana</emphasis>
</taxonomicName>
as a distinct species (see also under
<taxonomicName id="4C5ADDCAEF6B5E41FDB1388DFD25FB43" ID-CoL="4RYCR" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[623,759,1190,1213]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FDB1388DFD25FB43" box="[623,759,1190,1213]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
). Genitalia were figured by
<bibRefCitation id="EFCBDBB8EF6B5E41FE6B38EEFD53FB22" author="Wiegmann F" box="[437,641,1221,1244]" pageId="4" pageNumber="119" pagination="112 - 259" refId="ref7036" refString="Wiegmann F (1893) Beitrage zur Anatomie der Landschnecken des Indischen Archipels. In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 3. Brill, Leiden. Pp. 112 - 259." type="book chapter" year="1893">Wiegmann (1893)</bibRefCitation>
, who noted that in this illustration the insertion of the penial retractor was missing.
</paragraph>
</subSubSection>
<subSubSection id="C340F5C2EF6B5E41FF56396BFEFAF9B3" pageId="4" pageNumber="119" type="diagnosis">
<paragraph id="8BE5A649EF6B5E41FF56396BFE61FA4D" blockId="4.[136,775,1344,1459]" pageId="4" pageNumber="119">
<emphasis id="B92E7A5BEF6B5E41FF56396BFED3FAA9" bold="true" box="[136,257,1344,1367]" pageId="4" pageNumber="119">Diagnosis.</emphasis>
Shell comparatively large (D =
<quantity id="4CA20BACEF6B5E41FDA3396BFD28FAA9" box="[637,762,1344,1367]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="4" pageNumber="119" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">1619 mm</quantity>
), smooth or with fine axial ribs only, umbilicus forming a chink or narrowly open, peripheral band particularly welldeveloped (
<bibRefCitation id="EFCBDBB8EF6B5E41FED839B7FE75FA4D" author="Martens E von" box="[262,423,1436,1459]" pageId="4" pageNumber="119" pagination="209 - 263" refId="ref6232" refString="Martens E von (1891) Landschnecken des Indischen Archipels In: M. Weber (ed.) Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien. Band 2. Brill, Leiden. Pp. 209 - 263, pls. XII-XIV." type="book chapter" year="1891">Martens, 1891</bibRefCitation>
).
</paragraph>
<paragraph id="8BE5A649EF6B5E41FF5639F2FEFAF9B3" blockId="4.[136,775,1497,1613]" pageId="4" pageNumber="119">
<emphasis id="B92E7A5BEF6B5E41FF5639F2FE4FFA0E" bold="true" box="[136,413,1497,1520]" pageId="4" pageNumber="119">Comparative remarks.</emphasis>
Shell similar to
<taxonomicName id="4C5ADDCAEF6B5E41FDB739F2FD2CFA0E" baseAuthorityName="Martens" baseAuthorityYear="1863" box="[617,766,1497,1520]" class="Gastropoda" family="Camaenidae" genus="Rhagada" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="119" phylum="Mollusca" rank="species" species="solorensis">
<emphasis id="B92E7A5BEF6B5E41FDB739F2FD2CFA0E" box="[617,766,1497,1520]" italics="true" pageId="4" pageNumber="119">R. solorensis</emphasis>
</taxonomicName>
; species poorly defined by shell characters only; comparative study of anatomical and molecular characters required to confirm status.
</paragraph>
</subSubSection>
</treatment>
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