252 lines
33 KiB
XML
252 lines
33 KiB
XML
<document id="E0287F43838D82404305F6CB7FA1CA4C" ID-DOI="10.5281/zenodo.178756" ID-GBIF-Dataset="f9ea8775-9b2d-4364-b20a-4738aab6a54c" ID-ISSN="1175-5326" ID-Zenodo-Dep="178756" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459920987951" checkinUser="plazi" docAuthor="Kataev, Boris M. & Liang, Hongbin" docDate="2007" docId="03E987D1FFB4FFF7FF178AD23713FA59" docLanguage="en" docName="zt01604p020.pdf" docOrigin="Zootaxa 1604" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Harpalus vernicosus Kataev & Liang, 2007, sp. n." docType="treatment" docVersion="9" lastPageNumber="4" masterDocId="FFD0FFA9FFB5FFF4FF8088153517FFD3" masterDocTitle="Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)" masterLastPageNumber="20" masterPageNumber="1" pageNumber="2" updateTime="1698229235843" updateUser="plazi">
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<mods:mods id="4F026EBCF568CF2BC324CCF920595050" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="003183E1541ADFFA302FD5822B47F2F2">
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<mods:title id="E3A14C0B0605F1EB05F13F8496AE72AF">Contribution to the knowledge of the genus Harpalus in China, with description of new taxa (Coleoptera: Carabidae: Harpalini)</mods:title>
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</mods:titleInfo>
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<mods:name id="AE90CFE325F0565773A10E4BCC2048CF" type="personal">
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<mods:role id="D5FCA83D35D91F2AD2EB885C35D253B4">
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<mods:roleTerm id="DF15D241A88983069CEEEDF43EC35A5E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B0EE143C0DC407BCA437CB6C262B7DF1">Kataev, Boris M.</mods:namePart>
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</mods:name>
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<mods:name id="E423DCA58058F3F97FE4983CCE190635" type="personal">
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<mods:role id="463F6530F05E034EA16EF77A1C2ED458">
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<mods:roleTerm id="E7FB8BFA0765D9EE83DCDB269ACA2D75">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="16BC451DAF703FC9C04BFF95C8427C9B">Liang, Hongbin</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="92DDBF4CB8E6BCE59914B0C8E8CF225F">text</mods:typeOfResource>
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<mods:titleInfo id="BAFE35DDECC17374812618A34848240B">
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<mods:title id="EC5F22051238D84221EC0462F126DF0A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="3B512D8CC815DE0757873178CE787844">
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<mods:date id="839C4D41D12DE11A2B4FB1CC0138E6CA">2007</mods:date>
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<mods:detail id="4E219A7487A026FA8726BCAAD401F48C" type="volume">
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<mods:number id="16AFC97D913D850AEA0CBDAF178D31CE">1604</mods:number>
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<mods:start id="C64DFD963045467F1C5F1AA5FE060CB6">1</mods:start>
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<mods:end id="D3A4BA50B8DFDA27905BDBA2121C91BE">20</mods:end>
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</mods:extent>
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<mods:classification id="1F9F1DD6EDBE8797CA066A849F7B91A6">journal article</mods:classification>
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<mods:identifier id="00CDEF3747986465AAA876D2297D2523" type="DOI">10.5281/zenodo.178756</mods:identifier>
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<mods:identifier id="ACA22CF17F6167484600D1E83D86E021" type="GBIF-Dataset">f9ea8775-9b2d-4364-b20a-4738aab6a54c</mods:identifier>
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<mods:identifier id="D5634D61CF22BF86D6628C896EAD7184" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A2DE5EB8DE445FD75FDD05C4F2094889" type="Zenodo-Dep">178756</mods:identifier>
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<treatment id="03E987D1FFB4FFF7FF178AD23713FA59" ID-DOI="http://doi.org/10.5281/zenodo.6250224" ID-GBIF-Taxon="119373721" ID-Zenodo-Dep="6250224" LSID="urn:lsid:plazi:treatment:03E987D1FFB4FFF7FF178AD23713FA59" httpUri="http://treatment.plazi.org/id/03E987D1FFB4FFF7FF178AD23713FA59" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="C35A654CFFB4FFF5FF178AD23407FCD7" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8BFF36C7FFB4FFF5FF178AD234CDFD32" blockId="1.[151,474,711,772]" box="[151,474,711,737]" pageId="1" pageNumber="2">
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<heading id="D0B781ABFFB4FFF5FF178AD234CDFD32" bold="true" box="[151,474,711,737]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="B934EAD5FFB4FFF5FF178AD234CDFD32" bold="true" box="[151,474,711,737]" pageId="1" pageNumber="2">
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<taxonomicName id="4C404D44FFB4FFF5FF178AD23487FD32" ID-CoL="3JSSM" box="[151,400,711,737]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="vernicosus" status="sp. nov.">
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<emphasis id="B934EAD5FFB4FFF5FF178AD23487FD32" bold="true" box="[151,400,711,737]" italics="true" pageId="1" pageNumber="2">Harpalus vernicosus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20757AEFFB4FFF5FE188AD234CDFD32" box="[408,474,711,737]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BFF36C7FFB4FFF5FF178AFF3407FCD7" blockId="1.[151,474,711,772]" box="[151,272,746,772]" pageId="1" pageNumber="2">
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(
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<figureCitation id="137B2A42FFB4FFF5FF1F8AFF3410FCD7" box="[159,263,746,772]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1499,1523]" captionTargetBox="[164,1336,380,1498]" captionTargetId="figure@2.[151,1436,380,1551]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Harpalus vernicosus sp. n. (Sichuan, Xiangcheng Co.: 1 – 3, paratype; 4 – 6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1 – 2, 5 – 6), C = 1.0 mm (Fig. 3)." httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="1" pageNumber="2">Figs 1–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35A654CFFB4FFF5FF178B2F34D9FC27" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="8BFF36C7FFB4FFF5FF178B2F35F6FCAF" blockId="1.[151,1436,826,2012]" pageId="1" pageNumber="2">
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<emphasis id="B934EAD5FFB4FFF5FF178B2F345FFC87" bold="true" box="[151,328,826,852]" pageId="1" pageNumber="2">
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<typeStatus id="54FB8865FFB4FFF5FF178B2F35C3FC87" box="[151,212,826,852]" pageId="1" pageNumber="2">Type</typeStatus>
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material
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</emphasis>
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.
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<typeStatus id="54FB8865FFB4FFF5FED98B2F34DCFC87" box="[345,459,826,852]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
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: ɗ,
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<collectingCountry id="F3577657FFB4FFF5FE7D8B2F3751FC87" box="[509,582,826,852]" name="China" pageId="1" pageNumber="2">China</collectingCountry>
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, Sichuan, Xiangcheng Co.,
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<quantity id="4CB89B22FFB4FFF5FC168B2F36E2FC87" box="[918,1013,826,852]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="2" unit="m" value="3000.0">3000 m</quantity>
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,
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<date id="FFFE1007FFB4FFF5FB838B2F319CFC87" box="[1027,1163,826,852]" pageId="1" pageNumber="2" value="1982-06-21">21.VI.1982</date>
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, Zhang Xuezhong leg. (
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<collectionCode id="ED51AE02FFB4FFF5FF208B7735C2FCAF" box="[160,213,866,892]" pageId="1" pageNumber="2">IZB</collectionCode>
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||
).
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</paragraph>
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<paragraph id="8BFF36C7FFB4FFF5FF478B9F34D9FC27" blockId="1.[151,1436,826,2012]" pageId="1" pageNumber="2">
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<typeStatus id="54FB8865FFB4FFF5FF478B9F3429FC77" box="[199,318,906,932]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
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: 3 ɗ, 2 Ψ, same data as
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<typeStatus id="54FB8865FFB4FFF5FDE98B9F37D8FC77" box="[617,719,906,932]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED51AE02FFB4FFF5FD628B9F3602FC77" box="[738,789,906,932]" pageId="1" pageNumber="2">IZB</collectionCode>
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,
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<collectionCode id="ED51AE02FFB4FFF5FCA48B9F3672FC77" box="[804,869,906,932]" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="1" pageNumber="2">ZISP</collectionCode>
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); 1 ɗ, 2 Ψ, Sichuan, Xiangcheng Co.,
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<quantity id="4CB89B22FFB4FFF5FAB98B9F308FFC77" box="[1337,1432,906,932]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="1" pageNumber="2" unit="m" value="2900.0">2900 m</quantity>
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,
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<date id="FFFE1007FFB4FFF5FF178BA73408FC1F" box="[151,287,946,972]" pageId="1" pageNumber="2" value="1982-06-18">18.VI.1982</date>
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, Zhang Xuezhong & Wang Shuyong leg. (
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<collectionCode id="ED51AE02FFB4FFF5FC968BA7365DFC1F" box="[790,842,946,972]" pageId="1" pageNumber="2">IZB</collectionCode>
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||
); 6 ɗ, 3 Ψ, Tibet, Markam,
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<quantity id="4CB89B22FFB4FFF5FB138BA731F8FC1F" box="[1171,1263,946,972]" metricMagnitude="3" metricUnit="m" metricValue="3.25" pageId="1" pageNumber="2" unit="m" value="3250.0">3250 m</quantity>
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,
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<date id="FFFE1007FFB4FFF5FB7C8BA73080FC1F" box="[1276,1431,946,972]" pageId="1" pageNumber="2" value="1982-08-12">12.VIII.1982</date>
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, Wang Shuyong leg. (
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<collectionCode id="ED51AE02FFB4FFF5FE0D8BCF34D6FC27" box="[397,449,986,1012]" pageId="1" pageNumber="2">IZB</collectionCode>
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||
).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35A654CFFB4FFF7FF478C173069FDC9" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
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<paragraph id="8BFF36C7FFB4FFF5FF478C17366FFBCF" blockId="1.[151,1436,826,2012]" box="[199,888,1026,1052]" pageId="1" pageNumber="2">
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<emphasis id="B934EAD5FFB4FFF5FF478C17344FFBCF" bold="true" box="[199,344,1026,1052]" pageId="1" pageNumber="2">Description</emphasis>
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. Body length
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<quantity id="4CB89B22FFB4FFF5FE7E8C173799FBCF" box="[510,654,1026,1052]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="7.1" metricValueMin="5.4" pageId="1" pageNumber="2" unit="mm" value="6.25" valueMax="7.1" valueMin="5.4">5.4–7.1 mm</quantity>
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, width
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<quantity id="4CB89B22FFB4FFF5FD638C173664FBCF" box="[739,883,1026,1052]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.1" metricValueMin="2.4" pageId="1" pageNumber="2" unit="mm" value="2.75" valueMax="3.1" valueMin="2.4">2.4–3.1 mm</quantity>
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||
.
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||
</paragraph>
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||
<paragraph id="8BFF36C7FFB4FFF5FF478C3F346AFB97" blockId="1.[151,1436,826,2012]" box="[199,381,1066,1092]" pageId="1" pageNumber="2">
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Habitus:
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<figureCitation id="137B2A42FFB4FFF5FEB18C3F346EFB97" box="[305,377,1066,1092]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1499,1523]" captionTargetBox="[164,1336,380,1498]" captionTargetId="figure@2.[151,1436,380,1551]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Harpalus vernicosus sp. n. (Sichuan, Xiangcheng Co.: 1 – 3, paratype; 4 – 6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1 – 2, 5 – 6), C = 1.0 mm (Fig. 3)." httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="1" pageNumber="2">Fig. 4</figureCitation>
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||
.
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</paragraph>
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||
<paragraph id="8BFF36C7FFB4FFF5FF478C473071FB6F" blockId="1.[151,1436,826,2012]" pageId="1" pageNumber="2">Colour: Body dark brown to black, shiny on dorsum; middle portion of mandibles, labrum externally, very narrow margins of pronotum and elytra often reddish brown; femora dark brown, tibiae and tarsi much paler, mid- to light brown, but tibiae usually infuscated apically; palpi and antennae brownish yellow to brown.</paragraph>
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<paragraph id="8BFF36C7FFB4FFF5FF478CDF3115F9F7" blockId="1.[151,1436,826,2012]" pageId="1" pageNumber="2">
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Head: Medium-sized (WHmax/WPmax =
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||
<quantity id="4CB89B22FFB4FFF5FD388CDF365BFB37" box="[696,844,1226,1252]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" metricValueMax="1.7018" metricValueMin="1.651" pageId="1" pageNumber="2" unit="in" value="0.66" valueMax="0.67" valueMin="0.65">0.65–0.67 in</quantity>
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male and
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<quantity id="4CB89B22FFB4FFF5FC458CDF314EFB37" box="[965,1113,1226,1252]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" metricValueMax="1.7272" metricValueMin="1.6256" pageId="1" pageNumber="2" unit="in" value="0.66" valueMax="0.68" valueMin="0.64">0.64–0.68 in</quantity>
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female; WHmin/WPmax =
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<quantity id="4CB89B22FFB4FFF5FF178CE7343BFADF" box="[151,300,1266,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.3716" metricValueMax="1.4224" metricValueMin="1.3208" pageId="1" pageNumber="2" unit="in" value="0.54" valueMax="0.56" valueMin="0.52">0.52–0.56 in</quantity>
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male and
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||
<quantity id="4CB89B22FFB4FFF5FE288CE7372AFADF" box="[424,573,1266,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.4350999999999998" metricValueMax="1.4731999999999998" metricValueMin="1.3970000000000002" pageId="1" pageNumber="2" unit="in" value="0.565" valueMax="0.58" valueMin="0.55">0.55–0.58 in</quantity>
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||
female), with moderately convex eyes (WHmax/WHmin =
|
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<quantity id="4CB89B22FFB4FFF5FA878CE7308BFADF" box="[1287,1436,1266,1292]" metricMagnitude="-2" metricUnit="m" metricValue="3.0734" metricValueMax="3.175" metricValueMin="2.9718" pageId="1" pageNumber="2" unit="in" value="1.21" valueMax="1.25" valueMin="1.17">1.17–1.25 in</quantity>
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male and
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||
<quantity id="4CB89B22FFB4FFF5FE898D0F348AFAE7" box="[265,413,1306,1332]" metricMagnitude="-2" metricUnit="m" metricValue="2.9972" metricValueMax="3.0734" metricValueMin="2.921" pageId="1" pageNumber="2" unit="in" value="1.18" valueMax="1.21" valueMin="1.15">1.15–1.21 in</quantity>
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||
female), separated from buccal fissure ventrally by distance slightly shorter than width of first antennomere. Tempora rather short, flat or weakly convex. Clypeus with one lateral setigerous pore on each side (one of
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<typeStatus id="54FB8865FFB4FFF5FEE78D7F34C0FA57" box="[359,471,1386,1412]" pageId="1" pageNumber="2" type="paratype">paratypes</typeStatus>
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||
with an anomalous additional setigerous pore on left side). Dorsal microsculpture visible on areas around supraorbital pores and behind eyes, usually also on vertex laterally, more distinct in female than in male; meshes fine, more or less isodiametric. Labium (
|
||
<figureCitation id="137B2A42FFB4FFF5FC638DAF313EFA07" box="[995,1065,1466,1492]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1499,1523]" captionTargetBox="[164,1336,380,1498]" captionTargetId="figure@2.[151,1436,380,1551]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Harpalus vernicosus sp. n. (Sichuan, Xiangcheng Co.: 1 – 3, paratype; 4 – 6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1 – 2, 5 – 6), C = 1.0 mm (Fig. 3)." httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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||
) with prominent mental tooth; submentum with one seta on each side; ligular sclerite narrow, weakly widened apically. Antennae short, not reaching pronotal base; antennomeres 5–10 only slightly longer than wide.
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</paragraph>
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||
<paragraph id="8BFF36C7FFB4FFF5FF478E273610F8B7" blockId="1.[151,1436,826,2012]" pageId="1" pageNumber="2">Pronotum: Moderately wide, 1.44–1.51 times as wide as long, widest before middle, with unisetose sides roundly or almost rectilinearly converging basad. Apical margin slightly, arcuately emarginate, bordered only laterally. Apical angles weakly protruding, widely rounded. Basal margin usually widely rounded, sometimes almost straight, bordered throughout, wider than anterior margin; basal edge glabrous. Border along sides narrow throughout. Basal angles obtuse, blunt or rounded. Pronotal disc moderately and rather evenly convex, occasionally slightly flattened at basal angles, impunctate. Median line superficial, anterior transverse depression indistinct. Basal foveae very small and shallow. Microsculpture visible throughout, meshes fine, more or less isodiametric, slightly obliterate on disc medially.</paragraph>
|
||
<paragraph id="8BFF36C7FFB4FFF6FF478F67316CFEA9" blockId="1.[151,1436,826,2012]" lastBlockId="2.[151,1436,152,378]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">Elytra: Moderately convex, 1.34–1.44 times as long as wide, 2.35–2.55 times as long and 1.15–1.23 times as wide as pronotum, widest just behind middle, widely rounded at sides. Base of elytra slightly wider than pronotal base. Humeri angulate, obtuse, each with a small denticle visible dorsally. Preapical sinuation distinct, deeper in female than in male. Sutural angle acute, slightly blunt at apex. Basal border glabrous, weakly sinuate, forming a distinct obtuse angle with lateral margin. Striae impunctate, largely superficial, slightly impressed apically. Scutellar striole long, with a basal pore. Intervals flat, occasionally slightly convex apically. Third interval with a small discal pore in apical third. Fifth and seventh intervals lacking rows of pores before apices. Series umbilicate irregular and not interrupted medially. Microsculpture visible throughout, consisting of fine meshes, isodiametric along margins and weakly transverse on disc.</paragraph>
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||
<caption id="DF3F664FFFB7FFF6FF178DCE3125F9EB" httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="2" pageNumber="3" targetBox="[164,1336,380,1498]" targetPageId="2">
|
||
<paragraph id="8BFF36C7FFB7FFF6FF178DCE3125F9EB" blockId="2.[151,1436,1499,1592]" pageId="2" pageNumber="3">
|
||
<emphasis id="B934EAD5FFB7FFF6FF178DCE3451FA20" bold="true" box="[151,326,1499,1523]" pageId="2" pageNumber="3">FIGURES 1–6.</emphasis>
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||
<taxonomicName id="4C404D44FFB7FFF6FED08DC93726FA20" box="[336,561,1500,1523]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="vernicosus" status="sp. nov.">
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<emphasis id="B934EAD5FFB7FFF6FED08DC93726FA20" box="[336,561,1500,1523]" italics="true" pageId="2" pageNumber="3">Harpalus vernicosus</emphasis>
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</taxonomicName>
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||
<emphasis id="B934EAD5FFB7FFF6FDBC8DCE376CFA20" bold="true" box="[572,635,1499,1523]" pageId="2" pageNumber="3">
|
||
<taxonomicNameLabel id="A20757AEFFB7FFF6FDBC8DCE376CFA20" box="[572,635,1499,1523]" pageId="2" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
|
||
(Sichuan, Xiangcheng Co.: 1–3, paratype; 4–6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1–2, 5–6), C = 1.0 mm (Fig. 3).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFF36C7FFB7FFF6FF478E7E34F0F956" blockId="2.[151,1436,1643,2029]" box="[199,487,1643,1669]" pageId="2" pageNumber="3">Wings: Fully developed.</paragraph>
|
||
<paragraph id="8BFF36C7FFB7FFF6FF478E8631D5F92E" blockId="2.[151,1436,1643,2029]" pageId="2" pageNumber="3">
|
||
Ventral surface: Prosternum glabrous medially. Proepisterna impunctate. Metepisterna (
|
||
<figureCitation id="137B2A42FFB7FFF6FB668E863027F97E" box="[1254,1328,1683,1709]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1499,1523]" captionTargetBox="[164,1336,380,1498]" captionTargetId="figure@2.[151,1436,380,1551]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Harpalus vernicosus sp. n. (Sichuan, Xiangcheng Co.: 1 – 3, paratype; 4 – 6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1 – 2, 5 – 6), C = 1.0 mm (Fig. 3)." httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="2" pageNumber="3">Fig. 3</figureCitation>
|
||
) notably longer than wide, narrowed posteriad. Three apical abdominal sternites glabrous, only with obligatory fixed setae. Anal sternite rounded at apex in both sexes, without pronounced sexual dimorphism.
|
||
</paragraph>
|
||
<paragraph id="8BFF36C7FFB7FFF7FF478F1E3612FF61" blockId="2.[151,1436,1643,2029]" lastBlockId="3.[151,1437,152,1418]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">Legs: Metacoxa without additional setae. Protibia markedly widened apically, without prominent ventroapical tubercle in male, with one ventroapical spine and with three preapical spines at outer margin isolated from spines on ventral surface. Metafemur ventrally with 4–5 setigerous pores along posterior margin and occasionally one pore at anterior margin closer to apex (in addition to an apical pore situated just on anterior margin). Tarsi impunctate and glabrous dorsally. First mesotarsomere in male rather narrow, with two tiny scales apically. Metatarsomeres with sides slightly diverging apicad; first metatarsomere longer than second and shorter than second and third combined together.</paragraph>
|
||
<paragraph id="8BFF36C7FFB6FFF7FF4788D535D3FEF9" blockId="3.[151,1437,152,1418]" pageId="3" pageNumber="4">
|
||
Female genitalia (
|
||
<figureCitation id="137B2A42FFB6FFF7FE1988D534F6FF09" box="[409,481,192,218]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1499,1523]" captionTargetBox="[164,1336,380,1498]" captionTargetId="figure@2.[151,1436,380,1551]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Harpalus vernicosus sp. n. (Sichuan, Xiangcheng Co.: 1 – 3, paratype; 4 – 6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1 – 2, 5 – 6), C = 1.0 mm (Fig. 3)." httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
): Hemisternite comparatively wide, with 3–4 short setae distally. Basal stylomere with two setae distally. Apical stylomere arcuate, relatively wide, with 2–5 basal setae on external dorsal margin.
|
||
</paragraph>
|
||
<paragraph id="8BFF36C7FFB6FFF7FF47892D3069FDC9" blockId="3.[151,1437,152,1418]" pageId="3" pageNumber="4">
|
||
Male genitalia: Median lobe (
|
||
<figureCitation id="137B2A42FFB6FFF7FDA3892D379BFE81" box="[547,652,312,338]" captionStart="FIGURES 1 – 6" captionStartId="2.[151,269,1499,1523]" captionTargetBox="[164,1336,380,1498]" captionTargetId="figure@2.[151,1436,380,1551]" captionTargetPageId="2" captionText="FIGURES 1 – 6. Harpalus vernicosus sp. n. (Sichuan, Xiangcheng Co.: 1 – 3, paratype; 4 – 6, holotype). 1, labium; 2, hemisternite and stylus; 3, left metepisternum; 4, habitus; 5, median lobe of aedeagus, dorsal aspect; 6, same, lateral aspect. Scales: A = 1.0 mm (Fig. 4), B = 0.5 mm (Figs 1 – 2, 5 – 6), C = 1.0 mm (Fig. 3)." httpUri="https://zenodo.org/record/178757/files/figure.png" pageId="3" pageNumber="4">Figs 5–6</figureCitation>
|
||
) bent before middle, with almost straight ventral margin in its apical half (lateral aspect). Terminal lamella slightly directed dorsad and weakly bent ventrad just at apex, it moderately long (notably longer than wide), almost parallel-sided, narrowly rounded at apex and with very small dorsoapical marginal flange forming an oblique horseshoe–shaped apical capitulum. Apical orifice shifted to left, extending to middle of median lobe. Internal sac with two groups of small spines: a larger dorsolateral group on left side and a smaller ventrolateral group on right side, without any separate large teeth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A654CFFB6FFF7FF478A323015FD91" box="[199,1282,551,578]" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="8BFF36C7FFB6FFF7FF478A323015FD91" blockId="3.[151,1437,152,1418]" box="[199,1282,551,578]" pageId="3" pageNumber="4">
|
||
<emphasis id="B934EAD5FFB6FFF7FF478A323477FD92" bold="true" box="[199,352,551,577]" pageId="3" pageNumber="4">Distribution</emphasis>
|
||
. This new species is known only from northwestern Sichuan and eastern Tibet.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A654CFFB6FFF7FF478A5A314CFDB9" box="[199,1115,591,618]" pageId="3" pageNumber="4" type="etymology">
|
||
<paragraph id="8BFF36C7FFB6FFF7FF478A5A314CFDB9" blockId="3.[151,1437,152,1418]" box="[199,1115,591,618]" pageId="3" pageNumber="4">
|
||
<emphasis id="B934EAD5FFB6FFF7FF478A5A345BFDBA" bold="true" box="[199,332,591,617]" pageId="3" pageNumber="4">Etymology</emphasis>
|
||
. The species epithet refers to the shiny dorsum of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35A654CFFB6FFF7FF478A623713FA59" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="8BFF36C7FFB6FFF7FF478A623713FA59" blockId="3.[151,1437,152,1418]" pageId="3" pageNumber="4">
|
||
<emphasis id="B934EAD5FFB6FFF7FF478A62342EFD42" bold="true" box="[199,313,631,657]" pageId="3" pageNumber="4">Remarks</emphasis>
|
||
. The taxonomic position of this new species is not absolutely clear. In appearance, it is very similar to representatives of the
|
||
<emphasis id="B934EAD5FFB6FFF7FD928AB53761FD6A" box="[530,630,672,697]" italics="true" pageId="3" pageNumber="4">tinctulus</emphasis>
|
||
group sensu Kataev 1997 (=
|
||
<emphasis id="B934EAD5FFB6FFF7FC548AB53190FD6A" box="[980,1159,672,697]" italics="true" pageId="3" pageNumber="4">Zangoharpalus</emphasis>
|
||
Huang), particularly to some specimens of
|
||
<taxonomicName id="4C404D44FFB6FFF7FE078ADD3614FD31" authority="Schauberger" authorityName="Schauberger" box="[391,771,712,738]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="pseudotinctulus">
|
||
<emphasis id="B934EAD5FFB6FFF7FE078ADD3770FD32" box="[391,615,712,737]" italics="true" pageId="3" pageNumber="4">H. pseudotinctulus</emphasis>
|
||
Schauberger
|
||
</taxonomicName>
|
||
with impunctate base of pronotum, but differs sharply from them in such important characters as the structure of the aedeagus and the presence of ventral adhesive scales at apex of the fist mesotarsomere of the male. In addition, unlike all known members of the
|
||
<emphasis id="B934EAD5FFB6FFF7FAB78B0D308BFCE2" box="[1335,1436,792,817]" italics="true" pageId="3" pageNumber="4">tinctulus</emphasis>
|
||
group,
|
||
<taxonomicName id="4C404D44FFB6FFF7FF6D8B553486FC8A" box="[237,401,832,857]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vernicosus" status="sp. nov.">
|
||
<emphasis id="B934EAD5FFB6FFF7FF6D8B553486FC8A" box="[237,401,832,857]" italics="true" pageId="3" pageNumber="4">H. vernicosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B934EAD5FFB6FFF7FE1C8B2A34F6FC8A" bold="true" box="[412,481,831,857]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A20757AEFFB6FFF7FE1C8B2A34F6FC8A" box="[412,481,831,857]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has the median lobe of aedeagus with horseshoe–shaped apical capitulum and without any individual teeth in the internal sac. Based on total combination of the characters of the male genitalia and external morphology,
|
||
<taxonomicName id="4C404D44FFB6FFF7FD918B8537A2FC7A" box="[529,693,912,937]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vernicosus" status="sp. nov.">
|
||
<emphasis id="B934EAD5FFB6FFF7FD918B8537A2FC7A" box="[529,693,912,937]" italics="true" pageId="3" pageNumber="4">H. vernicosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B934EAD5FFB6FFF7FD408B9A3611FC7A" bold="true" box="[704,774,911,937]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A20757AEFFB6FFF7FD408B9A3611FC7A" box="[704,774,911,937]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
probably belongs to the
|
||
<emphasis id="B934EAD5FFB6FFF7FBB88B853195FC7A" box="[1080,1154,912,937]" italics="true" pageId="3" pageNumber="4">tardus</emphasis>
|
||
group sensu Kataev in
|
||
<bibRefCitation id="EFD14B36FFB6FFF7FF178BAD34D0FC01" author="Kryzhanovskij" box="[151,455,952,978]" pageId="3" pageNumber="20" refString="Kryzhanovskij, O. L., Belousov, I. A., Kabak, I. I., Kataev, B. M., Makarov, K. V., Shilenkov, V. G. (1995) A checklist of the ground-beetles of Russia and adjacent lands (Insecta, Coleoptera, Carabidae). Pensoft Publishers: Sofia - Moscow, 272 pp." type="book" year="1995">Kryzhanovskij et al. 1995</bibRefCitation>
|
||
, and is most closely related, in our opinion, to
|
||
<taxonomicName id="4C404D44FFB6FFF7FC678BAD31C3FC01" authority="Dejean" authorityName="Dejean" box="[999,1236,952,978]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="modestus">
|
||
<emphasis id="B934EAD5FFB6FFF7FC678BAD3160FC02" box="[999,1143,952,977]" italics="true" pageId="3" pageNumber="4">H. modestus</emphasis>
|
||
Dejean
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D44FFB6FFF7FB5F8BAD35DFFC29" authority="Chaudoir" authorityName="Chaudoir" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="bungii">
|
||
<emphasis id="B934EAD5FFB6FFF7FB5F8BAD3059FC02" box="[1247,1358,952,977]" italics="true" pageId="3" pageNumber="4">H. bungii</emphasis>
|
||
Chaudoir
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D44FFB6FFF7FF558BF534C6FC29" authority="Lafer" authorityName="Lafer" box="[213,465,992,1018]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="chasanensis">
|
||
<emphasis id="B934EAD5FFB6FFF7FF558BF5349FFC2A" box="[213,392,992,1017]" italics="true" pageId="3" pageNumber="4">H. chasanensis</emphasis>
|
||
Lafer
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C404D44FFB6FFF7FE5E8BF537FEFC29" authority="Kataev" authorityName="Kataev" box="[478,745,992,1018]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="tangutorum">
|
||
<emphasis id="B934EAD5FFB6FFF7FE5E8BF5379AFC2A" box="[478,653,992,1017]" italics="true" pageId="3" pageNumber="4">H. tangutorum</emphasis>
|
||
Kataev
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C404D44FFB6FFF7FCA98BF531A4FC29" authority="Schauberger. The" authorityName="Schauberger. The" box="[809,1203,992,1018]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="praecurrens" status="sp. nov.">
|
||
<emphasis id="B934EAD5FFB6FFF7FCA98BF536CAFC2A" box="[809,989,992,1017]" italics="true" pageId="3" pageNumber="4">H. praecurrens</emphasis>
|
||
Schauberger. The
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20757AEFFB6FFF7FB3C8BF5305EFC29" box="[1212,1353,992,1018]" pageId="3" pageNumber="4" rank="species">new species</taxonomicNameLabel>
|
||
shares with these species the following distinctive character states: antennae pale, pronotum with glabrous basal edge, protibia with one ventroapical spine and without prominent ventroapical tubercle in male, mesotibia and anal sternite without pronounced sexual dimorphism, three last abdominal sternites glabrous, elytra with only one discal setigerous pore on each third interval and without rows of pores at apex of fifth and seventh intervals, and internal sac of median lobe with several spiny patches medially and without any large separate teeth.
|
||
<taxonomicName id="4C404D44FFB6FFF7FF178CC5342FFB3A" box="[151,312,1232,1257]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vernicosus" status="sp. nov.">
|
||
<emphasis id="B934EAD5FFB6FFF7FF178CC5342FFB3A" box="[151,312,1232,1257]" italics="true" pageId="3" pageNumber="4">H. vernicosus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B934EAD5FFB6FFF7FEC08CDA3494FB3A" bold="true" box="[320,387,1231,1257]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="A20757AEFFB6FFF7FEC08CDA3494FB3A" box="[320,387,1231,1257]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is rather distinct within this group and clearly recognized by its small and relatively narrower body, impunctate base of pronotum, finer elytral striae and specific pattern of spiny patches in the internal sac of the aedeagus. A similar pattern of spiny patches is characteristic also of the species of the
|
||
<taxonomicName id="4C404D44FFB6FFF7FAC58D35308CFAEA" box="[1349,1435,1312,1337]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="vittatus">
|
||
<emphasis id="B934EAD5FFB6FFF7FAC58D35308CFAEA" box="[1349,1435,1312,1337]" italics="true" pageId="3" pageNumber="4">vittatus</emphasis>
|
||
</taxonomicName>
|
||
group sensu
|
||
<bibRefCitation id="EFD14B36FFB6FFF7FEB38D5D34DBFAB1" author="Kataev" box="[307,460,1352,1378]" pageId="3" pageNumber="20" refString="Kataev, B. M. (1990) Ground-beetles of the Harpalus vittatus group (Coleoptera, Carabidae). Entomologicheskoe obozrenie, 69 (2), 391 - 400 (in Russian)." type="journal article" year="1990">Kataev 1990</bibRefCitation>
|
||
which differ, however, in the dark antennae and clearly prominent ventroapical tuberculum of protibia in male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |