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<mods:title id="4D44CBAA06002F4DE5017BCF2B3D7D5D">A survey of Brignoliolus Ovchinnikov, 1999 stat. rev. (Araneae, Agelenidae) with the description of a new species from Israel</mods:title>
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<mods:namePart id="F4CAB51B608C6E322A3F471F3397A3FA">Zonstein, Sergei</mods:namePart>
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<taxonomicName id="4C404D23FFCCFF892BF5FC9FFD2FB155" authority="(Ovchinnikov, 1999)" baseAuthorityName="Ovchinnikov" baseAuthorityYear="1999" box="[151,664,855,882]" class="Arachnida" family="Agelenidae" genus="Brignoliolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="nenilini" status="comb. nov.">
<emphasis id="B934EAB2FFCCFF892BF5FC9FFE22B156" bold="true" box="[151,405,855,882]" italics="true" pageId="4" pageNumber="219">Brignoliolus nenilini</emphasis>
(
<bibRefCitation id="EFD14B51FFCCFF892AC6FC9FFD27B155" author="Ovchinnikov, S. V." box="[420,656,855,881]" pageId="4" pageNumber="219" pagination="63 - 80" refId="ref7423" refString="Ovchinnikov, S. V. (1999) On the supraspecific systematics of the subfamily Coelotinae (Araneae, Amaurobiidae) in the former USSR fauna. Tethys Entomological Research, 1, 63 - 80. [in Russian]" type="journal article" year="1999">Ovchinnikov, 1999</bibRefCitation>
)
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<figureCitation id="137B2A25FFCCFF892BF5FCB3FEDEB1B1" box="[151,361,891,917]" captionStart="FIGURES 914" captionStartId="5.[151,264,1237,1261]" captionTargetBox="[151,1435,489,1214]" captionTargetId="figure-202@5.[151,1435,489,1214]" captionTargetPageId="5" captionText="FIGURES 914. Brignoliolus juglandicola (Ovchinnikov, 1984) (9, 12), and B. nenilini (Ovchinnikov, 1999) (10, 11, 13, 14), females from Arslanbob and Sidjak, respectively. 9, 10 epigyne, ventral; 11 vulva, dorsal; 12, 14 spinnerets, ventral; 13 same, caudal. Scale bars = 0.2 mm. Abbreviations: Am anterior margin of fovea; Cl colulus; Et tooth of epigyne; Fo fovea." figureDoi="http://doi.org/10.5281/zenodo.14240974" httpUri="https://zenodo.org/record/14240974/files/figure.png" pageId="4" pageNumber="219">Figs 10, 11, 13, 14</figureCitation>
,
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(
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)
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<bibRefCitation id="EFD14B51FFCCFF892A82FC08FD7CB1FC" author="Ovchinnikov, S. V." box="[480,715,960,984]" pageId="4" pageNumber="219" pagination="63 - 80" refId="ref7423" refString="Ovchinnikov, S. V. (1999) On the supraspecific systematics of the subfamily Coelotinae (Araneae, Amaurobiidae) in the former USSR fauna. Tethys Entomological Research, 1, 63 - 80. [in Russian]" type="journal article" year="1999">Ovchinnikov, 1999: 78</bibRefCitation>
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, figs 4044 (
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).
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<taxonomicName id="4C404D23FFCCFF892BF5FC16FEF4B1D2" ID-CoL="WTQM" authorityName="Ovchinnikov" authorityYear="1999" box="[151,323,990,1014]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="B934EAB2FFCCFF892BF5FC16FEF4B1D2" box="[151,323,990,1014]" italics="true" pageId="4" pageNumber="219">Coelotes nenilini</emphasis>
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:
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<bibRefCitation id="EFD14B51FFCCFF892A32FC17FDF3B1D2" author="Wang, X. P. &amp; Zhu, M. S." box="[336,580,990,1015]" pageId="4" pageNumber="219" pagination="272 - 281" refId="ref7657" refString="Wang, X. P. &amp; Zhu, M. S. (2009) On the charitonovi species group of the spider genus Coelotes (Araneae: Amaurobiidae). Journal of Arachnology, 37, 272 - 281. https: // doi. org / 10.1636 / a 08 - 105.1" type="journal article" year="2009">Wang &amp; Zhu 2009: 280</bibRefCitation>
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:
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(
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Ridge,
<collectingMunicipality id="6B9BACDAFFCCFF892F16FBECFA90B61B" box="[1140,1319,1060,1087]" pageId="4" pageNumber="219">Kansai Canyon</collectingMunicipality>
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W Khumsan
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a.s.l.
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(
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)
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;
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<specimenCount id="9D46FD29FFCCFF8928F2FB81FC04B647" box="[912,947,1097,1123]" count="4" pageId="4" pageNumber="219" type="female">4♀</specimenCount>
(
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), same collection data as preceding but
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N Handailyk Vill.
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,
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,
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a.s.l.
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,
<date id="FFFE1060FFCCFF892F28FBA5FB7FB6A3" box="[1098,1224,1133,1159]" pageId="4" pageNumber="219" value="2023-04-23">
<collectingDate id="EFBAE988FFCCFF892F28FBA5FB7FB6A3" box="[1098,1224,1133,1159]" pageId="4" pageNumber="219" value="2023-04-23">23.04.2023</collectingDate>
</date>
(
<collectorName id="26B55376FFCCFF892FBEFBA5FAD4B6A2" box="[1244,1379,1132,1159]" pageId="4" pageNumber="219">S. Zonstein</collectorName>
)
</materialsCitation>
;
<materialsCitation id="3B283CFDFFCCFF892E1BFBA5FE87B6EA" collectingDate="2023-05-09" collectionCode="SMNH" collectorName="S. Zonstein" country="Uzbekistan" elevation="1000" latitude="41.683334" location="Ugam Mt." longLatPrecision="1155" longitude="69.916664" pageId="4" pageNumber="219" specimenCount="2" specimenCount-female="2">
<specimenCount id="9D46FD29FFCCFF892E1BFBA5FA2BB6A3" box="[1401,1436,1133,1159]" count="2" pageId="4" pageNumber="219" type="female">2♀</specimenCount>
(
<collectionCode id="ED51AE65FFCCFF892BFDFB59FF42B68F" box="[159,245,1169,1195]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="219">SMNH</collectionCode>
), same region,
<location id="8E9F607BFFCCFF892AC6FB59FDAEB68F" LSID="urn:lsid:plazi:treatment:03E987B6FFCCFF892BF5FC9FFE87B47E:8E9F607BFFCCFF892AC6FB59FDAEB68F" box="[420,537,1169,1195]" country="Uzbekistan" latitude="41.683334" longLatPrecision="1155" longitude="69.916664" name="Ugam Mt." pageId="4" pageNumber="219">Ugam Mt.</location>
Ridge,
<locationDeviation id="9A4D2F82FFCCFF892911FB59FCCBB68E" box="[627,892,1168,1195]" location="Sidjak Vill." pageId="4" pageNumber="219">
<quantity id="4CB89B45FFCCFF892911FB59FD19B68E" box="[627,686,1168,1195]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="219" unit="km" value="4.0">4 km</quantity>
SSW Sidjak Vill.
</locationDeviation>
,
<geoCoordinate id="EE745067FFCCFF8928EAFB5AFC5EB689" box="[904,1001,1170,1197]" degrees="41" direction="north" minutes="41" orientation="latitude" pageId="4" pageNumber="219" precision="925" value="41.683334">41°41N</geoCoordinate>
,
<geoCoordinate id="EE745067FFCCFF892897FB5AFBE4B689" box="[1013,1107,1170,1197]" degrees="69" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="219" precision="925" value="69.916664">69°55E</geoCoordinate>
,
<emphasis id="B934EAB2FFCCFF892F3DFB59FBCEB68E" box="[1119,1145,1169,1194]" italics="true" pageId="4" pageNumber="219">ca</emphasis>
.
<elevation id="006DD193FFCCFF892FEBFB59FAA9B68E" box="[1161,1310,1168,1195]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="219" unit="m" value="1000.0">
<quantity id="4CB89B45FFCCFF892FEBFB59FB57B68F" box="[1161,1248,1169,1195]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="219" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
,
<date id="FFFE1060FFCCFF892E4EFB58FA2BB68F" box="[1324,1436,1168,1195]" pageId="4" pageNumber="219" value="2023-05-09">
<collectingDate id="EFBAE988FFCCFF892E4EFB58FA2BB68F" box="[1324,1436,1168,1195]" pageId="4" pageNumber="219" value="2023-05-09">9.05.2023</collectingDate>
</date>
(
<collectorName id="26B55376FFCCFF892BC3FB7DFE9FB6EA" box="[161,296,1204,1231]" pageId="4" pageNumber="219">S. Zonstein</collectorName>
)
</materialsCitation>
;
<materialsCitation id="3B283CFDFFCCFF892A5CFB7DFE13B6D6" collectingDate="2023-05-08" collectionCode="SMNH" collectorName="S. Zonstein" country="Uzbekistan" elevation="1300" latitude="41.55" location="Chimgan Mts." longLatPrecision="1155" longitude="70.066666" municipality="Gulikamsai Canyon" pageId="4" pageNumber="219" specimenCount="4" specimenCount-female="4">
<specimenCount id="9D46FD29FFCCFF892A5CFB7DFED5B6EB" box="[318,354,1205,1231]" count="4" pageId="4" pageNumber="219" type="female">4♀</specimenCount>
(
<collectionCode id="ED51AE65FFCCFF892A16FB7DFE7DB6EB" box="[372,458,1205,1231]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="4" pageNumber="219">SMNH</collectionCode>
), same region,
<location id="8E9F607BFFCCFF89291FFB7CFC94B6EB" LSID="urn:lsid:plazi:treatment:03E987B6FFCCFF892BF5FC9FFE87B47E:8E9F607BFFCCFF89291FFB7CFC94B6EB" box="[637,803,1204,1231]" country="Uzbekistan" latitude="41.55" longLatPrecision="1155" longitude="70.066666" municipality="Gulikamsai Canyon" name="Chimgan Mts." pageId="4" pageNumber="219">Chimgan Mts.</location>
,
<collectingMunicipality id="6B9BACDAFFCCFF892853FB7CFBA1B6EB" box="[817,1046,1204,1231]" pageId="4" pageNumber="219">Gulikamsai Canyon</collectingMunicipality>
,
<geoCoordinate id="EE745067FFCCFF892F47FB7EFB32B6F5" box="[1061,1157,1206,1233]" degrees="41" direction="north" minutes="33" orientation="latitude" pageId="4" pageNumber="219" precision="925" value="41.55">41°33N</geoCoordinate>
,
<geoCoordinate id="EE745067FFCCFF892FF1FB7EFB47B6F5" box="[1171,1264,1206,1233]" degrees="70" direction="east" minutes="04" orientation="longitude" pageId="4" pageNumber="219" precision="925" value="70.066666">70°04E</geoCoordinate>
,
<elevation id="006DD193FFCCFF892F9DFB7DFA20B6EA" box="[1279,1431,1204,1231]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="219" unit="m" value="1300.0">
<quantity id="4CB89B45FFCCFF892F9DFB7DFAE0B6EB" box="[1279,1367,1205,1231]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="219" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</elevation>
,
<date id="FFFE1060FFCCFF892BF5FB10FEB0B6D7" box="[151,263,1240,1267]" pageId="4" pageNumber="219" value="2023-05-08">
<collectingDate id="EFBAE988FFCCFF892BF5FB10FEB0B6D7" box="[151,263,1240,1267]" pageId="4" pageNumber="219" value="2023-05-08">8.05.2023</collectingDate>
</date>
(
<collectorName id="26B55376FFCCFF892A7AFB11FE2BB6D6" box="[280,412,1240,1267]" pageId="4" pageNumber="219">S. Zonstein</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A652BFFCCFF892BA5FB34FCB3B437" pageId="4" pageNumber="219" type="discussion">
<paragraph id="8BFF36A0FFCCFF892BA5FB34FCB3B437" blockId="4.[151,1437,1060,1627]" pageId="4" pageNumber="219">
<emphasis id="B934EAB2FFCCFF892BA5FB34FEF8B732" bold="true" box="[199,335,1276,1302]" pageId="4" pageNumber="219">Comments.</emphasis>
In the structure of the epigyne (with tightly clustered epigynal teeth), females of
<taxonomicName id="4C404D23FFCCFF892FA2FB35FA84B732" authorityName="Ovchinnikov" authorityYear="1999" box="[1216,1331,1276,1303]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="B934EAB2FFCCFF892FA2FB35FB66B732" box="[1216,1233,1277,1302]" italics="true" pageId="4" pageNumber="219">B</emphasis>
.
<emphasis id="B934EAB2FFCCFF892FBFFB34FA84B732" box="[1245,1331,1276,1302]" italics="true" pageId="4" pageNumber="219">nenilini</emphasis>
</taxonomicName>
resemble those of
<taxonomicName id="4C404D23FFCCFF892B9DFAE9FE0FB71E" authorityName="Zonstein &amp; Marusik" authorityYear="2024" baseAuthorityName="Ovchinnikov" baseAuthorityYear="1999" box="[255,440,1312,1339]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="turkestanicus">
<emphasis id="B934EAB2FFCCFF892B9DFAE9FEA7B71E" box="[255,272,1313,1338]" italics="true" pageId="4" pageNumber="219">B</emphasis>
.
<emphasis id="B934EAB2FFCCFF892A41FAE8FE0FB71E" box="[291,440,1312,1338]" italics="true" pageId="4" pageNumber="219">turkestanicus</emphasis>
</taxonomicName>
. They differ by the shape of the epigynal fovea (
<emphasis id="B934EAB2FFCCFF892F60FAE9FB96B71E" box="[1026,1057,1313,1338]" italics="true" pageId="4" pageNumber="219">Fo</emphasis>
) with wide (
<emphasis id="B934EAB2FFCCFF892FDFFAE9FB63B71E" box="[1213,1236,1313,1338]" italics="true" pageId="4" pageNumber="219">vs</emphasis>
. narrow) median constriction, as well as by the shape and arrangement of the receptacles which are narrower and more widely spaced in the middle part
<emphasis id="B934EAB2FFCCFF892A06FAA1FECCB7A6" box="[356,379,1385,1410]" italics="true" pageId="4" pageNumber="219">vs</emphasis>
. narrowly spaced (
<figureCitation id="137B2A25FFCCFF89293AFAA0FD63B7A7" box="[600,724,1384,1411]" captionStart="FIGURES 914" captionStartId="5.[151,264,1237,1261]" captionTargetBox="[151,1435,489,1214]" captionTargetId="figure-202@5.[151,1435,489,1214]" captionTargetPageId="5" captionText="FIGURES 914. Brignoliolus juglandicola (Ovchinnikov, 1984) (9, 12), and B. nenilini (Ovchinnikov, 1999) (10, 11, 13, 14), females from Arslanbob and Sidjak, respectively. 9, 10 epigyne, ventral; 11 vulva, dorsal; 12, 14 spinnerets, ventral; 13 same, caudal. Scale bars = 0.2 mm. Abbreviations: Am anterior margin of fovea; Cl colulus; Et tooth of epigyne; Fo fovea." figureDoi="http://doi.org/10.5281/zenodo.14240974" httpUri="https://zenodo.org/record/14240974/files/figure.png" pageId="4" pageNumber="219">Figs 10, 11</figureCitation>
<emphasis id="B934EAB2FFCCFF8929B8FAA1FD59B7A7" box="[730,750,1385,1411]" italics="true" pageId="4" pageNumber="219">cf</emphasis>
.
<bibRefCitation id="EFD14B51FFCCFF89299EFAA0FC19B7A7" author="Fomichev, A. A." box="[764,942,1384,1411]" pageId="4" pageNumber="219" pagination="691 - 694" refId="ref6826" refString="Fomichev, A. A. (2021) The north-easternmost record of a Central Asian funnelweb spider, Coelotes turkestanicus Ovtchinnikov, 1999 (Araneae, Agelenidae). Check List, 17 (2), 691 - 694. https: // doi. org / 10.15560 / 17.2.691" type="journal article" year="2021">Fomichev 2021</bibRefCitation>
, fig. 1DF). Males of
<taxonomicName id="4C404D23FFCCFF892FC4FAA1FAACB7A6" authorityName="Ovchinnikov" authorityYear="1999" box="[1190,1307,1384,1411]" class="Arachnida" family="Agelenidae" genus="Coelotes" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="nenilini">
<emphasis id="B934EAB2FFCCFF892FC4FAA1FB00B7A6" box="[1190,1207,1385,1410]" italics="true" pageId="4" pageNumber="219">B</emphasis>
.
<emphasis id="B934EAB2FFCCFF892FA7FAA0FAACB7A6" box="[1221,1307,1384,1410]" italics="true" pageId="4" pageNumber="219">nenilini</emphasis>
</taxonomicName>
differ from other known male congeners, except for
<taxonomicName id="4C404D23FFCCFF892909FA45FCADB782" authorityName="Zonstein &amp; Marusik" authorityYear="2024" baseAuthorityName="Ovchinnikov" baseAuthorityYear="1984" box="[619,794,1420,1447]" class="Arachnida" family="Agelenidae" genus="Brignoliolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="219" phylum="Arthropoda" rank="species" species="juglandicola">
<emphasis id="B934EAB2FFCCFF892909FA45FDCBB782" box="[619,636,1421,1446]" italics="true" pageId="4" pageNumber="219">B</emphasis>
.
<emphasis id="B934EAB2FFCCFF8929EEFA44FCADB782" box="[652,794,1420,1446]" italics="true" pageId="4" pageNumber="219">juglandicola</emphasis>
</taxonomicName>
, in possessing the bifurcated (
<emphasis id="B934EAB2FFCCFF892F1AFA45FB38B782" box="[1144,1167,1421,1446]" italics="true" pageId="4" pageNumber="219">vs</emphasis>
. undivided) retrolateral patellar apophysis of the male palp; they can be distinguished from the latter species by the different shape of the conductor and median apophysis of the tegulum (Ovtchinnikov 1984, fig. 1; 1999, figs 35, 37, 4042;
<bibRefCitation id="EFD14B51FFCCFF892E69FA1DFF65B437" author="Wang, X. P. &amp; Zhu, M. S." pageId="4" pageNumber="219" pagination="272 - 281" refId="ref7657" refString="Wang, X. P. &amp; Zhu, M. S. (2009) On the charitonovi species group of the spider genus Coelotes (Araneae: Amaurobiidae). Journal of Arachnology, 37, 272 - 281. https: // doi. org / 10.1636 / a 08 - 105.1" type="journal article" year="2009">Wang &amp; Zhu 2009</bibRefCitation>
, figs 10, 22,30;
<bibRefCitation id="EFD14B51FFCCFF892AE4FA30FD39B437" author="Dimitrov, D. K. &amp; Jager, P." box="[390,654,1528,1555]" pageId="4" pageNumber="219" pagination="1 - 7" refId="ref6703" refString="Dimitrov, D. K. &amp; Jager, P. (2021) Coelotes vignai Brignoli, 1978 (Araneae: Agelenidae) from Turkey: first description of male and annotations on terminology of copulatory organs. Biodiversity Data Journal, 9 (e 73127), 1 - 7. https: // doi. org / 10.3897 / BDJ. 9. e 73127" type="book chapter" year="2021">Dimitrov &amp; Jager 2021</bibRefCitation>
, figs 1, 2).
</paragraph>
</subSubSection>
<subSubSection id="C35A652BFFCCFF892BA5F9D4FE87B47E" pageId="4" pageNumber="219" type="distribution">
<paragraph id="8BFF36A0FFCCFF892BA5F9D4FE87B47E" blockId="4.[151,1437,1060,1627]" pageId="4" pageNumber="219">
<emphasis id="B934EAB2FFCCFF892BA5F9D4FED6B412" bold="true" box="[199,353,1564,1590]" pageId="4" pageNumber="219">Distribution.</emphasis>
The species is known from several localities in the forested midlands of Western Tien-Shan (
<collectingCountry id="F3577630FFCCFF892BC3F988FE93B47E" box="[161,292,1600,1626]" name="Uzbekistan" pageId="4" pageNumber="219">Uzbekistan</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>