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<document id="418AA55C577F2A4EC2C80F17BF74E035" ID-DOI="10.11646/zootaxa.4098.3.3" ID-GBIF-Dataset="d63adea2-3c11-47ac-9de7-0ec4542a65db" ID-ISSN="1175-5326" ID-Zenodo-Dep="265642" ID-ZooBank="1ACE4A63-6A0A-4138-BB56-08FC5AEF9F2D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460010950499" checkinUser="plazi" docAuthor="Bidzilya, Oleksiy &amp; Li, Houhun" docDate="2016" docId="03E9725F3F55FFCD74FEFE2DFCD1F77E" docLanguage="en" docName="zootaxa.4098.3.3.pdf.imf" docOrigin="Zootaxa 4098 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Kiwaia (Empista) spinosa Povolny 1976" docType="treatment" docVersion="9" lastPageNumber="482" masterDocId="FFD00A273F5EFFC67469FF95FFFCF210" masterDocTitle="A review of the genus Kiwaia Philpott, 1930 (Lepidoptera, Gelechiidae) in the Palaearctic Region" masterLastPageNumber="497" masterPageNumber="471" pageNumber="482" updateTime="1698233421406" updateUser="plazi">
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<mods:title id="B6B7A00B891D316353435E48091A854B">A review of the genus Kiwaia Philpott, 1930 (Lepidoptera, Gelechiidae) in the Palaearctic Region</mods:title>
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<mods:namePart id="1853B055E53DF766EC6AC80671AFA0B5">Bidzilya, Oleksiy</mods:namePart>
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<mods:namePart id="5370F6F75C53A78BB99D8DC1F158EED7">Li, Houhun</mods:namePart>
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<mods:date id="1C787001E02568CA4A18FBF3CCF89A48">2016</mods:date>
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<treatment id="03E9725F3F55FFCD74FEFE2DFCD1F77E" ID-DOI="http://doi.org/10.5281/zenodo.6062811" ID-GBIF-Taxon="119381419" ID-Zenodo-Dep="6062811" LSID="urn:lsid:plazi:treatment:03E9725F3F55FFCD74FEFE2DFCD1F77E" httpUri="http://treatment.plazi.org/id/03E9725F3F55FFCD74FEFE2DFCD1F77E" lastPageNumber="482" pageId="11" pageNumber="482">
<subSubSection id="C35A90C23F55FFCD74FEFE2DFEDBF3E4" pageId="11" pageNumber="482" type="nomenclature">
<paragraph id="8BFFC3493F55FFCD74FEFE2DFD82F3C0" blockId="11.[151,638,439,501]" box="[151,638,439,464]" pageId="11" pageNumber="482">
<heading id="D0B774253F55FFCD74FEFE2DFD82F3C0" bold="true" box="[151,638,439,464]" fontSize="10" level="2" pageId="11" pageNumber="482" reason="2">
<taxonomicName id="4C40B8CA3F55FFCD74FEFE2DFD82F3C0" ID-CoL="935VW" authority="Povolny, 1976" authorityName="Povolny" authorityYear="1976" box="[151,638,439,464]" class="Insecta" family="Gelechiidae" genus="Kiwaia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="spinosa" subGenus="Empista">
<emphasis id="B9341F5B3F55FFCD74FEFE2DFD82F3C0" bold="true" box="[151,638,439,464]" pageId="11" pageNumber="482">
<emphasis id="B9341F5B3F55FFCD74FEFE2DFF14F3C0" bold="true" box="[151,232,440,464]" italics="true" pageId="11" pageNumber="482">Kiwaia</emphasis>
(
<emphasis id="B9341F5B3F55FFCD7491FE2DFEAAF3C0" bold="true" box="[248,342,440,464]" italics="true" pageId="11" pageNumber="482">Empista</emphasis>
)
<emphasis id="B9341F5B3F55FFCD750FFE2DFE41F3C0" bold="true" box="[358,445,440,464]" italics="true" pageId="11" pageNumber="482">spinosa</emphasis>
(
<bibRefCitation id="EFD1BEB83F55FFCD75A4FE22FD8AF3C0" author="Povolny" box="[461,630,439,464]" pageId="11" pageNumber="482" refString="Povolny, D. (1976). Two new species of the genus Empista Povolny from Nepal (Lepidoptera, Gelechiidae). Acta entomologica bohemoslovaca, 73, 174 - 182." type="journal article" year="1976">Povolný, 1976</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFFC3493F55FFCD74FEFE49FEDBF3E4" blockId="11.[151,638,439,501]" box="[151,295,476,501]" pageId="11" pageNumber="482">
(
<figureCitation id="137BDFCC3F55FFCD74F6FE49FF09F3E4" box="[159,245,476,501]" captionStart="FIGURES 11 – 20" captionStartId="17.[151,264,1874,1897]" captionTargetBox="[187,1351,203,1823]" captionTargetId="figure@17.[171,1415,193,1836]" captionTargetPageId="17" captionText="FIGURES 11 – 20. Adults of Kiwaia (Empista) spp. 11, K. (E.) septentrionala sp. n., PT, ♀ (China: Shanxi); 12, K. (E.) septentrionala sp. n., PT, ♀ (China: Shanxi); 13, K. (E.) septentrionala sp. n., PT, ♂ (China: Gansu); 14, K. (E.) centrica sp. n., HT, ♂ (China: Hunan); 15, K. (E.) centrica sp. n., PT, ♀ (China: Hunan); 16, K. (E.) centrica sp. n., ♀ (China: Gansu); 17, K. (E.) spinosa (Povolný), HT, ♂ (Nepal); 18, K. (E.) montana sp. n., HT, ♂ (China: Xizang); 19, K. (E.) montana sp. n., PT, ♂ (China: Xizang); 20, K. (E.) medogia sp. n., HT, ♂ (China: Xizang)." httpUri="https://zenodo.org/record/265644/files/figure.png" pageId="11" pageNumber="482">Figs 17</figureCitation>
,
<figureCitation id="137BDFCC3F55FFCD7568FE49FEE3F3E4" box="[257,287,476,500]" captionStart="FIGURES 42 – 47" captionStartId="21.[151,264,1919,1942]" captionTargetBox="[163,1418,213,1873]" captionTargetId="figure@21.[151,1436,193,1898]" captionTargetPageId="21" captionText="FIGURES 42 – 47. Male genitalia of Kiwaia (Empista) spp. 42, K. (E.) centrica sp. n., HT (China: Hunan), slide no. SYW 05204; 43, K. (E.) centrica sp. n., PT (China: Hunan), slide no. SYW 05224; 44, K. (E.) spinosa (Povolný), HT (Nepal), slide Jp. 4833; 45, K. (E.) montana sp. n., HT (China: Xizang), slide no. L 07029; 46, K. (E.) montana sp. n., PT (China: Xizang), slide no. L 13017; 47 K. (E.) medogia sp. n., HT (China: Xizang), slide no. L 07038." httpUri="https://zenodo.org/record/265648/files/figure.png" pageId="11" pageNumber="482">44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35A90C23F55FFCD74FEFDB1FC19F048" pageId="11" pageNumber="482" type="reference_group">
<paragraph id="8BFFC3493F55FFCD74FEFDB1FDEEF02A" blockId="11.[151,997,548,600]" box="[151,530,548,570]" pageId="11" pageNumber="482">
<treatmentCitationGroup id="AB50E4673F55FFCD74FEFDB1FDEEF02A" box="[151,530,548,570]" pageId="11" pageNumber="482">
<taxonomicName id="4C40B8CA3F55FFCD74FEFDB1FDF2F02A" ID-CoL="93G9Q" authority="Povolny, 1976: 179" authorityName="Povolny" authorityPageNumber="179" authorityYear="1976" box="[151,526,548,570]" class="Insecta" family="Gelechiidae" genus="Empista" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="spinosa">
<emphasis id="B9341F5B3F55FFCD74FEFDB1FEC1F029" box="[151,317,548,569]" italics="true" pageId="11" pageNumber="482">Empista spinosa</emphasis>
<treatmentCitation id="0AE1E5583F55FFCD752DFDB1FDF2F02A" author="Povolny" box="[324,526,548,570]" page="179" pageId="11" pageNumber="482" year="1976">
<bibRefCitation id="EFD1BEB83F55FFCD752DFDB1FE27F02A" author="Povolny" box="[324,475,548,570]" pageId="11" pageNumber="482" refString="Povolny, D. (1976). Two new species of the genus Empista Povolny from Nepal (Lepidoptera, Gelechiidae). Acta entomologica bohemoslovaca, 73, 174 - 182." type="journal article" year="1976">Povolný, 1976</bibRefCitation>
: 179
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFFC3493F55FFCD74FEFDD6FC19F048" blockId="11.[151,997,548,600]" box="[151,997,578,600]" pageId="11" pageNumber="482">
<treatmentCitationGroup id="AB50E4673F55FFCD74FEFDD6FC19F048" box="[151,997,578,600]" pageId="11" pageNumber="482">
<taxonomicName id="4C40B8CA3F55FFCD74FEFDD6FDB1F048" ID-CoL="935VW" authority="Povolny, 1976" authorityName="Povolny" authorityYear="1976" box="[151,589,578,600]" class="Insecta" family="Gelechiidae" genus="Kiwaia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="spinosa" subGenus="Empista">
<emphasis id="B9341F5B3F55FFCD74FEFDD6FF1CF048" box="[151,224,579,600]" italics="true" pageId="11" pageNumber="482">Kiwaia</emphasis>
(
<emphasis id="B9341F5B3F55FFCD7486FDD6FEBEF048" box="[239,322,579,600]" italics="true" pageId="11" pageNumber="482">Empista</emphasis>
)
<emphasis id="B9341F5B3F55FFCD7539FDD6FE61F048" box="[336,413,579,600]" italics="true" pageId="11" pageNumber="482">spinosa</emphasis>
(
<bibRefCitation id="EFD1BEB83F55FFCD75C2FDD7FDB8F048" author="Povolny" box="[427,580,578,600]" pageId="11" pageNumber="482" refString="Povolny, D. (1976). Two new species of the genus Empista Povolny from Nepal (Lepidoptera, Gelechiidae). Acta entomologica bohemoslovaca, 73, 174 - 182." type="journal article" year="1976">Povolný, 1976</bibRefCitation>
)
</taxonomicName>
:
<treatmentCitation id="0AE1E5583F55FFCD763FFDD7FCF1F048" author="Sattler" box="[598,781,578,600]" page="234" pageId="11" pageNumber="482" year="1988">
<bibRefCitation id="EFD1BEB83F55FFCD763FFDD7FD26F048" author="Sattler" box="[598,730,578,600]" pageId="11" pageNumber="482" refString="Sattler, K. (1988) The systematic status of the genera Ilseopsis Povolny, 1965, and Empista Povolny, 1968 (Lepidoptera: Gelechiidae: Gnorimoschemini). Nota lepidopterologica, 10 (4), 224 - 235." type="journal article" year="1988">Sattler, 1988</bibRefCitation>
: 234
</treatmentCitation>
;
<treatmentCitation id="0AE1E5583F55FFCD777EFDD7FC1DF048" author="Povolny" box="[791,993,578,600]" page="101" pageId="11" pageNumber="482" year="2002">
<bibRefCitation id="EFD1BEB83F55FFCD777EFDD7FC53F048" author="Povolny" box="[791,943,578,600]" pageId="11" pageNumber="482" refString="Povolny, D. (2002) Iconographia tribus Gnorimoschemini (Lepidoptera, Gelechiidae) Regionis Palaearcticae. Verlag F. Slamka, Bratislava, 110 pp." type="book" year="2002">Povolný, 2002</bibRefCitation>
: 101
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35A90C23F55FFCD74FEFD10FE94F0D2" pageId="11" pageNumber="482" type="materials_examined">
<paragraph id="8BFFC3493F55FFCD74FEFD10FE94F0D2" blockId="11.[151,1437,645,1390]" pageId="11" pageNumber="482">
<emphasis id="B9341F5B3F55FFCD74FEFD10FE7DF08E" bold="true" box="[151,385,645,670]" pageId="11" pageNumber="482">Material examined.</emphasis>
<typeStatus id="54FB7DEB3F55FFCD75E0FD10FE0DF08E" box="[393,497,645,670]" pageId="11" pageNumber="482" type="holotype">Holotype</typeStatus>
♂ &quot;Tukucha, Palpa,
<date id="FFFEE5893F55FFCD76B7FD10FCBCF08E" box="[734,832,645,670]" pageId="11" pageNumber="482" value="1968-05-06">6.v.1968</date>
,
<quantity id="4CB86EAC3F55FFCD7724FD10FC5BF08E" box="[845,935,645,670]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="11" pageNumber="482" unit="m" value="2600.0">2600 m</quantity>
&quot;, &quot;
<collectingCountry id="F35783D93F55FFCD77AFFD10FBF2F08E" box="[966,1038,645,670]" name="Nepal" pageId="11" pageNumber="482">Nepal</collectingCountry>
, T. Kumata&quot;, &quot;Jp. 4833&quot;, gen. slide Jp. 4833 (
<collectionCode id="ED515B8C3F55FFCD756CFD3FFEA0F0D2" box="[261,348,682,706]" httpUri="http://grbio.org/cool/tc3a-fmpc" name="Staatliches Museum fuer Naturkunde Karlsruhe (State Museum of Natural History)" pageId="11" pageNumber="482">SMNK</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35A90C23F55FFCD74AEFD58FE6CF6CE" pageId="11" pageNumber="482" type="description">
<paragraph id="8BFFC3493F55FFCD74AEFD58FCFAF1AE" blockId="11.[151,1437,645,1390]" pageId="11" pageNumber="482">
<emphasis id="B9341F5B3F55FFCD74AEFD58FEA5F0F6" bold="true" box="[199,345,717,742]" pageId="11" pageNumber="482">Description.</emphasis>
Adult (
<figureCitation id="137BDFCC3F55FFCD75DFFD58FDF1F0F6" box="[438,525,717,742]" captionStart="FIGURES 11 – 20" captionStartId="17.[151,264,1874,1897]" captionTargetBox="[187,1351,203,1823]" captionTargetId="figure@17.[171,1415,193,1836]" captionTargetPageId="17" captionText="FIGURES 11 – 20. Adults of Kiwaia (Empista) spp. 11, K. (E.) septentrionala sp. n., PT, ♀ (China: Shanxi); 12, K. (E.) septentrionala sp. n., PT, ♀ (China: Shanxi); 13, K. (E.) septentrionala sp. n., PT, ♂ (China: Gansu); 14, K. (E.) centrica sp. n., HT, ♂ (China: Hunan); 15, K. (E.) centrica sp. n., PT, ♀ (China: Hunan); 16, K. (E.) centrica sp. n., ♀ (China: Gansu); 17, K. (E.) spinosa (Povolný), HT, ♂ (Nepal); 18, K. (E.) montana sp. n., HT, ♂ (China: Xizang); 19, K. (E.) montana sp. n., PT, ♂ (China: Xizang); 20, K. (E.) medogia sp. n., HT, ♂ (China: Xizang)." httpUri="https://zenodo.org/record/265644/files/figure.png" pageId="11" pageNumber="482">Fig. 17</figureCitation>
): Wingspan 14.0 mm. Head, thorax and tegulae grey mixed with black, frons lighter, grey; labial palpus up-curved, brown with white medial and apical belts, inner surface of segment 2 light grey, segment 3 about 1.5 times shorter and 1.5 times narrower than segment 2; scapus brown, other antennal segments brown with white rings; forewing grey, costal margin black, black dash under 1/4 of costal margin, two diffuse black spots and black dash near base in mid-width, black spot near base of dorsal margin, diffuse black pattern from 1/3 to about one half length in cell, two black dots in the middle of cell, apical area weakly mottled with black; cilia grey black tipped; hindwing light grey.
</paragraph>
<paragraph id="8BFFC3493F55FFCD74AEFC5CFEEEF6AA" blockId="11.[151,1437,645,1390]" pageId="11" pageNumber="482">
Male genitalia (
<figureCitation id="137BDFCC3F55FFCD7515FC5CFE2FF1F2" box="[380,467,969,994]" captionStart="FIGURES 42 – 47" captionStartId="21.[151,264,1919,1942]" captionTargetBox="[163,1418,213,1873]" captionTargetId="figure@21.[151,1436,193,1898]" captionTargetPageId="21" captionText="FIGURES 42 – 47. Male genitalia of Kiwaia (Empista) spp. 42, K. (E.) centrica sp. n., HT (China: Hunan), slide no. SYW 05204; 43, K. (E.) centrica sp. n., PT (China: Hunan), slide no. SYW 05224; 44, K. (E.) spinosa (Povolný), HT (Nepal), slide Jp. 4833; 45, K. (E.) montana sp. n., HT (China: Xizang), slide no. L 07029; 46, K. (E.) montana sp. n., PT (China: Xizang), slide no. L 13017; 47 K. (E.) medogia sp. n., HT (China: Xizang), slide no. L 07038." httpUri="https://zenodo.org/record/265648/files/figure.png" pageId="11" pageNumber="482">Fig. 44</figureCitation>
): Uncus twice longer than broad on base, gradually narrowed, posterior margin with small medial incision; gnathos weakly curved, rather stout; tegumen nearly parallel-sided, anteromedial emargination deeply rounded; valva moderately broad, about of equal width, tapers into long thorn, not reaching the middle of uncus; sacculus broad, weakly curved inwards, 1/3 as long as valva; vinculum slightly broader than long, posteromedial emargination broad, vincular processes broadly rounded, covered apically with dense hairs and rare spines; saccus stout, broad; phallus nearly straight, caecum weakly inflated, top rounded, apical arm well developed.
</paragraph>
<paragraph id="8BFFC3493F55FFCD74AEFB50FE6CF6CE" blockId="11.[151,1437,645,1390]" box="[199,400,1221,1246]" pageId="11" pageNumber="482">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C35A90C23F55FFCD74AEFB7CFE4EF712" box="[199,434,1257,1282]" pageId="11" pageNumber="482" type="distribution">
<paragraph id="8BFFC3493F55FFCD74AEFB7CFE4EF712" blockId="11.[151,1437,645,1390]" box="[199,434,1257,1282]" pageId="11" pageNumber="482">
<emphasis id="B9341F5B3F55FFCD74AEFB7CFE9CF712" bold="true" box="[199,352,1257,1282]" pageId="11" pageNumber="482">Distribution.</emphasis>
<collectingCountry id="F35783D93F55FFCD750EFB7CFE52F712" box="[359,430,1257,1282]" name="Nepal" pageId="11" pageNumber="482">Nepal</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35A90C23F55FFCD74AEFA98FCD1F77E" pageId="11" pageNumber="482" type="discussion">
<paragraph id="8BFFC3493F55FFCD74AEFA98FCD1F77E" blockId="11.[151,1437,645,1390]" pageId="11" pageNumber="482">
<emphasis id="B9341F5B3F55FFCD74AEFA98FEC7F736" bold="true" box="[199,315,1293,1318]" pageId="11" pageNumber="482">Remarks.</emphasis>
As far as one can judge after the slightly worn
<typeStatus id="54FB7DEB3F55FFCD770BFA98FC38F736" box="[866,964,1293,1318]" pageId="11" pageNumber="482" type="holotype">holotype</typeStatus>
<taxonomicName id="4C40B8CA3F55FFCD77A4FA9BFB85F735" box="[973,1145,1294,1318]" class="Insecta" family="Gelechiidae" genus="Kiwaia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="spinosa" subGenus="Empista">
<emphasis id="B9341F5B3F55FFCD77A4FA9BFC1BF735" box="[973,999,1294,1317]" italics="true" pageId="11" pageNumber="482">K.</emphasis>
(
<emphasis id="B9341F5B3F55FFCD7790FA9BFBEDF735" box="[1017,1041,1294,1317]" italics="true" pageId="11" pageNumber="482">E.</emphasis>
)
<emphasis id="B9341F5B3F55FFCD704AFA9BFB85F735" box="[1059,1145,1294,1317]" italics="true" pageId="11" pageNumber="482">spinosa</emphasis>
</taxonomicName>
seems to be most related externally to
<taxonomicName id="4C40B8CA3F55FFCD7540FAA7FE2EF75A" box="[297,466,1329,1354]" class="Insecta" family="Gelechiidae" genus="Kiwaia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="482" phylum="Arthropoda" rank="species" species="kostjuki" subGenus="Empista">
<emphasis id="B9341F5B3F55FFCD7540FAA7FEBFF759" box="[297,323,1330,1353]" italics="true" pageId="11" pageNumber="482">K.</emphasis>
(
<emphasis id="B9341F5B3F55FFCD753AFAA7FE97F759" box="[339,363,1330,1353]" italics="true" pageId="11" pageNumber="482">E.</emphasis>
)
<emphasis id="B9341F5B3F55FFCD7512FAA4FE2EF75A" box="[379,466,1329,1354]" italics="true" pageId="11" pageNumber="482">kostjuki</emphasis>
</taxonomicName>
(for differences see under that species). The male genitalia can easily be recognized by the narrow prolonged uncus and the apically tapered valva.
</paragraph>
</subSubSection>
</treatment>
</document>
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