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<document id="981F14AD3D9C1C9EA731BAEB05BEB6B1" ID-DOI="10.11646/phytotaxa.515.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8061143" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1687282820830" checkinUser="plazi" docAuthor="Beek, Abraham Van De" docDate="2021" docId="03DE3646FFAFFFFFFF43FA4FFCBEFD6B" docLanguage="en" docName="Phytotaxa.515.1.1.pdf" docOrigin="Phytotaxa 515 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.515.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Rubus mundtii Chamisso &amp; Schlechtendal 1827" docType="treatment" docVersion="7" lastPageNumber="35" masterDocId="FFE74E3EFF8FFFDDFFCBFFE1FF96FF93" masterDocTitle="Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa" masterLastPageNumber="71" masterPageNumber="1" pageNumber="33" updateTime="1699527950928" updateUser="plazi">
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<mods:title id="15A73F3F9E37F1CE7ECA3A071E225FF7">Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa</mods:title>
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<mods:title id="30FA5C18BD3A673AB8C14C94B102626F">Phytotaxa</mods:title>
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<subSubSection id="C36DD4DBFFAFFFFDFF43FA4FFC87FA5B" box="[136,785,1454,1481]" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="8BC88750FFAFFFFDFF43FA4FFC87FA5B" blockId="32.[136,785,1454,1481]" box="[136,785,1454,1481]" pageId="32" pageNumber="33">
<emphasis id="B9035B42FFAFFFFDFF43FA4FFF3DFA5B" bold="true" box="[136,171,1454,1480]" pageId="32" pageNumber="33">12.</emphasis>
<taxonomicName id="4C77FCD3FFAFFFFDFF79FA4FFC87FA5B" ID-CoL="4TKXZ" authority="Chamisso &amp; Schlechtendal (1827: 18)" authorityName="Chamisso &amp; Schlechtendal" authorityYear="1827" box="[178,785,1454,1481]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="species" species="mundtii">
<emphasis id="B9035B42FFAFFFFDFF79FA4FFEC8FA5A" bold="true" box="[178,350,1454,1481]" italics="true" pageId="32" pageNumber="33">Rubus mundtii</emphasis>
<bibRefCitation id="EFE6FAA1FFAFFFFDFEAEFA4FFC87FA5B" author="Chamisso, A. de &amp; Schlechtendal, D. de" box="[357,785,1454,1480]" pageId="32" pageNumber="33" pagination="1 - 20" refId="ref46583" refString="Chamisso, A. de &amp; Schlechtendal, D. de (1827) De plantis in expeditione speculatoria romanzoffiana observatis. Linnaea 2: 1 - 20. [https: // www. biodiversitylibrary. org / item / 10846 # page / 5 / mode / 1 up]" type="journal article" year="1827">Chamisso &amp; Schlechtendal (1827: 18)</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAFFFFDFF43FA1BFBC4F99C" box="[136,1106,1530,1552]" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="8BC88750FFAFFFFDFF43FA1BFBC4F99C" blockId="32.[136,1451,1530,1624]" box="[136,1106,1530,1552]" pageId="32" pageNumber="33">
<emphasis id="B9035B42FFAFFFFDFF43FA1BFEDDF99C" box="[136,331,1530,1551]" italics="true" pageId="32" pageNumber="33">Homotypic synonym</emphasis>
:—
<taxonomicName id="4C77FCD3FFAFFFFDFEA1FA1BFBD8F99C" ID-CoL="4TKXZ" authority="(Cham. &amp; Schltdl.) Harvey &amp; Sonder (1862: 287)" authorityName="Harvey &amp; Sonder" authorityPageNumber="287" authorityYear="1862" baseAuthorityName="Cham. &amp; Schltdl." box="[362,1102,1530,1552]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="32" pageNumber="33" phylum="Tracheophyta" rank="variety" species="rigidus" variety="mundtii">
<emphasis id="B9035B42FFAFFFFDFEA1FA1BFE66F99C" box="[362,496,1530,1551]" italics="true" pageId="32" pageNumber="33">Rubus rigidus</emphasis>
var.
<emphasis id="B9035B42FFAFFFFDFDD4FA1BFDFEF99C" box="[543,616,1530,1551]" italics="true" pageId="32" pageNumber="33">mundtii</emphasis>
(Cham. &amp; Schltdl.)
<bibRefCitation id="EFE6FAA1FFAFFFFDFCE7FA1BFBD8F99C" author="Harvey, W. H. &amp; Sonder, O. W." box="[812,1102,1530,1552]" pageId="32" pageNumber="33" refId="ref47716" refString="Harvey, W. H. &amp; Sonder, O. W. (1862) Flora Capensis. Being a systematic description of the plants of the Cape Colony, Cafraria, &amp; Port Natal, Vol. 2. Hodges, Smith &amp; Co Dublin / A. S. Robertson, Cape Town, ix + 621 pp. https: // doi. org / 10.5962 / bhl. title. 821" type="book" year="1862">Harvey &amp; Sonder (1862: 287)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAFFFFDFB9CFA1BFBD8F9CB" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BC88750FFAFFFFDFB9CFA1BFBD8F9CB" blockId="32.[136,1451,1530,1624]" pageId="32" pageNumber="33">
<materialsCitation id="3B1F8D0DFFAFFFFDFB9CFA1BFBDDF9CB" collectingDate="1821-07" collectionCode="ID" collectorName="Mundt &amp; Maire" country="South Africa" location="Hangklipp" pageId="32" pageNumber="33" specimenCode="B 10 1146998" specimenCount="1" stateProvince="Western Cape" typeStatus="type">
<typeStatus id="54CC39F2FFAFFFFDFB9CFA1BFB1FF99C" box="[1111,1161,1530,1551]" pageId="32" pageNumber="33">Type</typeStatus>
:—
<collectingCountry id="F360C7C0FFAFFFFDFB6FFA1BFAC3F99C" box="[1188,1365,1530,1551]" name="South Africa" pageId="32" pageNumber="33">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FFAFFFFDFA96FA1BFF64F9A0" country="South Africa" name="Western Cape" pageId="32" pageNumber="33">Western Cape</collectingRegion>
: In promontorio bonae spei ad
<location id="8EA8D18BFFAFFFFDFDEAF9FFFD10F9A7" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFAFFFFDFDEAF9FFFD10F9A7" box="[545,646,1566,1588]" country="South Africa" municipality="In" name="Hangklipp" pageId="32" pageNumber="33" stateProvince="Western Cape">Hangklipp</location>
legerunt
<date id="FFC9A190FFAFFFFDFD2BF9FFFCD2F9A0" box="[736,836,1566,1588]" pageId="32" pageNumber="33" value="1821-07">
<collectingDate id="EF8D5878FFAFFFFDFD2BF9FFFCD2F9A0" box="[736,836,1566,1588]" pageId="32" pageNumber="33" value="1821-07">Julio 1821</collectingDate>
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Mund et Maire,
<emphasis id="B9035B42FFAFFFFDFC27F9FFFB3CF9A0" box="[1004,1194,1566,1587]" italics="true" pageId="32" pageNumber="33">
<collectorName id="2682E286FFAFFFFDFC27F9FFFBBDF9A0" box="[1004,1067,1566,1587]" pageId="32" pageNumber="33">Mundt</collectorName>
&amp;
<collectorName id="2682E286FFAFFFFDFB82F9FFFB15F9A0" box="[1097,1155,1566,1587]" pageId="32" pageNumber="33">Maire</collectorName>
s.n.
</emphasis>
(the number 224 is written on the label by Schlechtendal) (
<specimenCode id="DBD12F2BFFAFFFFDFE26F9A3FDE3F9CB" box="[493,629,1602,1624]" pageId="32" pageNumber="33">B 10 1146998</specimenCode>
[https://www.jacq.org/detail.php?
<collectionCode id="ED661F95FFAFFFFDFC0AF9A3FC4DF9CB" box="[961,987,1602,1624]" pageId="32" pageNumber="33">ID</collectionCode>
=1627315])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAFFFFDFF43F967FEF8F956" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BC88750FFAFFFFDFF43F967FEF8F956" blockId="32.[136,1452,1670,2093]" pageId="32" pageNumber="33">
<emphasis id="B9035B42FFAFFFFDFF43F967FEA5F933" bold="true" box="[136,307,1670,1696]" pageId="32" pageNumber="33">Note on name:</emphasis>
—The correct spelling of the epithet is mundtii. The family name is Mundt, as Dr. Uwe Braun in Halle kindly informed me.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAFFFFCFF76F92EFA96FE27" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
<paragraph id="8BC88750FFAFFFFCFF76F92EFA96FE27" blockId="32.[136,1452,1670,2093]" lastBlockId="33.[136,1452,158,436]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B9035B42FFAFFFFDFF76F92EFEAEF97B" box="[189,312,1743,1768]" italics="true" pageId="32" pageNumber="33">Primocane</emphasis>
(
<figureCitation id="134C9BD5FFAFFFFDFE82F92FFE41F97B" box="[329,471,1742,1768]" captionStart="FIGURE 10" captionStartId="33.[136,229,1424,1446]" captionTargetBox="[151,1436,461,1399]" captionTargetId="figure-163@33.[151,1436,461,1399]" captionTargetPageId="33" captionText="FIGURE 10. Rubus mundtii. A. Primocane with leaves. B. Primocane. C. Primocane. D. Leaf, abaxially. E. Flowering branch. F. Flowering branch. G. Inflorescence. H. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061171" httpUri="https://zenodo.org/record/8061171/files/figure.png" pageId="32" pageNumber="33">Figs 10AC</figureCitation>
) arching, diam.
<quantity id="4C8F2AB5FFAFFFFDFD5EF92FFD60F97A" box="[661,758,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="32" pageNumber="33" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">47 mm</quantity>
, angular and furrowed, with dense short grey hairs; prickles 515 per
<quantity id="4C8F2AB5FFAFFFFDFF3AF913FEBEF89E" box="[241,296,1778,1805]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="32" pageNumber="33" unit="cm" value="5.0">5 cm</quantity>
from
<quantity id="4C8F2AB5FFAFFFFDFEA7F913FE5FF89E" box="[364,457,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="32" pageNumber="33" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">48 mm</quantity>
broad base compressed, hooked or with geniculate tip, up to
<quantity id="4C8F2AB5FFAFFFFDFBBFF913FB47F89E" box="[1140,1233,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="32" pageNumber="33" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">48 mm</quantity>
long, densely short hairy; stipules
<quantity id="4C8F2AB5FFAFFFFDFEE5F8F7FE0DF8A2" box="[302,411,1814,1841]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.5" metricValueMin="0.7" pageId="32" pageNumber="33" unit="mm" value="11.0" valueMax="15.0" valueMin="7.0">715 mm</quantity>
, linear-lanceolate, hairy.
<emphasis id="B9035B42FFAFFFFDFD7EF8F6FC95F8A3" box="[693,771,1815,1840]" italics="true" pageId="32" pageNumber="33">Leaves</emphasis>
with flat margin, 3-foliolate or bi-jugate pinnate, sometimes with lobate or divided basal leaflets, adaxially with some to moderately dense short appressed hairs, abaxially (
<figureCitation id="134C9BD5FFAFFFFDFAB5F8DBFF2CF8EB" captionStart="FIGURE 10" captionStartId="33.[136,229,1424,1446]" captionTargetBox="[151,1436,461,1399]" captionTargetId="figure-163@33.[151,1436,461,1399]" captionTargetPageId="33" captionText="FIGURE 10. Rubus mundtii. A. Primocane with leaves. B. Primocane. C. Primocane. D. Leaf, abaxially. E. Flowering branch. F. Flowering branch. G. Inflorescence. H. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061171" httpUri="https://zenodo.org/record/8061171/files/figure.png" pageId="32" pageNumber="33">Fig. 10D</figureCitation>
) densely grey to white tomentose; serrature triangular, hardly to short mucronate, clearly short periodical; petiole
<quantity id="4C8F2AB5FFAFFFFDFF43F863FF4CF80E" box="[136,218,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="32" pageNumber="33" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">57 cm</quantity>
long, densely hairy, with 1013 curved or hooked prickles; central leaflet of 3-foliolate leaves
<quantity id="4C8F2AB5FFAFFFFDFB3EF863FAFBF80E" box="[1269,1389,1922,1949]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="32" pageNumber="33" unit="mm" value="57.5" valueMax="60.0" valueMin="55.0">5560 mm</quantity>
long, broad ovate with emarginated or subcordate base, gradually short pointed, widthlength index 0.780.87, length of petiolule 3843% of length of leaflet; central leaflet of 5-foliolate leaves
<quantity id="4C8F2AB5FFAFFFFDFC06F82BFBD1F876" box="[973,1095,1994,2021]" metricMagnitude="-2" metricUnit="m" metricValue="4.95" metricValueMax="6.0" metricValueMin="3.9" pageId="32" pageNumber="33" unit="mm" value="49.5" valueMax="60.0" valueMin="39.0">3960 mm</quantity>
long, (ob-)ovate-elliptical with rounded base, acute or sometimes emarginated at tip, widthlength index 0.570.71, length of petiolule 1430% of length of leaflet; petiolule of lateral leaves
<quantity id="4C8F2AB5FFAFFFFDFDAFF7F3FD55F7BE" box="[612,707,2066,2093]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="4.0" metricValueMin="0.0" pageId="32" pageNumber="33" unit="mm" value="2.0" valueMax="4.0" valueMin="0.0">04 mm</quantity>
.
<emphasis id="B9035B42FFAFFFFDFD06F7F3FC01F7BF" box="[717,919,2066,2092]" italics="true" pageId="32" pageNumber="33">Flowering branch</emphasis>
(
<figureCitation id="134C9BD5FFAFFFFDFC6EF7F3FBBFF7BF" box="[933,1065,2066,2092]" captionStart="FIGURE 10" captionStartId="33.[136,229,1424,1446]" captionTargetBox="[151,1436,461,1399]" captionTargetId="figure-163@33.[151,1436,461,1399]" captionTargetPageId="33" captionText="FIGURE 10. Rubus mundtii. A. Primocane with leaves. B. Primocane. C. Primocane. D. Leaf, abaxially. E. Flowering branch. F. Flowering branch. G. Inflorescence. H. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061171" httpUri="https://zenodo.org/record/8061171/files/figure.png" pageId="32" pageNumber="33">Figs 10EF</figureCitation>
) angular and furrowed, with dense short hairs and sessile glands; prickles 420 per
<quantity id="4C8F2AB5FFAEFFFCFD7EFF7FFD66FF2A" box="[693,752,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="34" unit="cm" value="5.0">5 cm</quantity>
, unequal, from
<quantity id="4C8F2AB5FFAEFFFCFC6EFF7FFB92FF2A" box="[933,1028,158,185]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="33" pageNumber="34" unit="mm" value="4.0" valueMax="6.0" valueMin="2.0">26 mm</quantity>
base compressed to almost subulate, curved and hooked, up to
<quantity id="4C8F2AB5FFAEFFFCFE64FF23FD9BFF4E" box="[431,525,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="33" pageNumber="34" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">46 mm</quantity>
long.
<emphasis id="B9035B42FFAEFFFCFD9EFF22FD35FF4F" box="[597,675,195,220]" italics="true" pageId="33" pageNumber="34">Leaves</emphasis>
3- to 5-foliolate or upper simple, adaxially with some to dense short appressed hairs, abaxially densely white tomentose; serrature sharper than at primocane leaves.
<emphasis id="B9035B42FFAEFFFCFB1DFF07FAFAFE93" box="[1238,1388,230,256]" italics="true" pageId="33" pageNumber="34">Inflorescence</emphasis>
(
<figureCitation id="134C9BD5FFAEFFFCFAB5FF07FF2CFEB7" captionStart="FIGURE 10" captionStartId="33.[136,229,1424,1446]" captionTargetBox="[151,1436,461,1399]" captionTargetId="figure-163@33.[151,1436,461,1399]" captionTargetPageId="33" captionText="FIGURE 10. Rubus mundtii. A. Primocane with leaves. B. Primocane. C. Primocane. D. Leaf, abaxially. E. Flowering branch. F. Flowering branch. G. Inflorescence. H. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061171" httpUri="https://zenodo.org/record/8061171/files/figure.png" pageId="33" pageNumber="34">Fig. 10G</figureCitation>
) with rigid axis, with 28 axillary branches or fully leafy, with dense, narrow cylindric apex, densely hairy with patent stellate hairs and with curved and hooked prickles; peduncles divided from base or in middle, longest ones with 58 flowers; pedicels
<quantity id="4C8F2AB5FFAEFFFCFE48FEB3FE75FEFE" box="[387,483,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="8.0" metricValueMin="0.0" pageId="33" pageNumber="34" unit="mm" value="4.0" valueMax="8.0" valueMin="0.0">08 mm</quantity>
, densely hairy, often with some prickles.
<emphasis id="B9035B42FFAEFFFCFC76FEB3FB8FFEFF" box="[957,1049,338,364]" italics="true" pageId="33" pageNumber="34">Flowers</emphasis>
: sepals erect, 34 ×
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, short pointed, densely grey tomentose; petals pink or white, small, roundish; stamens shorter than styles; anthers glabrous; styles red; ovaries with few short stellate hairs; receptacle with long hairs. Ripe
<emphasis id="B9035B42FFAEFFFCFBC3FE7BFBAEFE27" box="[1032,1080,410,436]" italics="true" pageId="33" pageNumber="34">fruit</emphasis>
black (
<figureCitation id="134C9BD5FFAEFFFCFB40FE7BFB62FE27" box="[1163,1268,410,436]" captionStart="FIGURE 10" captionStartId="33.[136,229,1424,1446]" captionTargetBox="[151,1436,461,1399]" captionTargetId="figure-163@33.[151,1436,461,1399]" captionTargetPageId="33" captionText="FIGURE 10. Rubus mundtii. A. Primocane with leaves. B. Primocane. C. Primocane. D. Leaf, abaxially. E. Flowering branch. F. Flowering branch. G. Inflorescence. H. Fruits. Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061171" httpUri="https://zenodo.org/record/8061171/files/figure.png" pageId="33" pageNumber="34">Fig. 10H</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF08D7D8FFAEFFFCFF43FA71FCFDFA59" ID-DOI="http://doi.org/10.5281/zenodo.8061171" ID-Zenodo-Dep="8061171" httpUri="https://zenodo.org/record/8061171/files/figure.png" pageId="33" pageNumber="34" startId="33.[136,229,1424,1446]" targetBox="[151,1436,461,1399]" targetPageId="33" targetType="figure">
<paragraph id="8BC88750FFAEFFFCFF43FA71FCFDFA59" blockId="33.[136,1451,1424,1482]" pageId="33" pageNumber="34">
<emphasis id="B9035B42FFAEFFFCFF43FA71FE98FA36" bold="true" box="[136,270,1424,1446]" pageId="33" pageNumber="34">FIGURE 10.</emphasis>
<taxonomicName id="4C77FCD3FFAEFFFCFEDCFA71FE30FA36" authorityName="Chamisso &amp; Schlechtendal" authorityYear="1827" box="[279,422,1424,1445]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="mundtii">
<emphasis id="B9035B42FFAEFFFCFEDCFA71FE30FA36" box="[279,422,1424,1445]" italics="true" pageId="33" pageNumber="34">Rubus mundtii</emphasis>
</taxonomicName>
.
<emphasis id="B9035B42FFAEFFFCFE7CFA71FE58FA36" bold="true" box="[439,462,1424,1445]" pageId="33" pageNumber="34">A.</emphasis>
Primocane with leaves.
<emphasis id="B9035B42FFAEFFFCFD02FA71FD49FA36" bold="true" box="[713,735,1424,1445]" pageId="33" pageNumber="34">B.</emphasis>
Primocane.
<emphasis id="B9035B42FFAEFFFCFCABFA71FCE1FA35" bold="true" box="[864,887,1424,1446]" pageId="33" pageNumber="34">C.</emphasis>
Primocane.
<emphasis id="B9035B42FFAEFFFCFC3CFA71FB99FA36" bold="true" box="[1015,1039,1424,1445]" pageId="33" pageNumber="34">D.</emphasis>
Leaf, abaxially.
<emphasis id="B9035B42FFAEFFFCFB70FA71FB47FA36" bold="true" box="[1211,1233,1424,1445]" pageId="33" pageNumber="34">E.</emphasis>
Flowering branch.
<emphasis id="B9035B42FFAEFFFCFA52FA71FA3DFA36" bold="true" box="[1433,1451,1424,1445]" pageId="33" pageNumber="34">F.</emphasis>
Flowering branch.
<emphasis id="B9035B42FFAEFFFCFEF5FA55FEC1FA5A" bold="true" box="[318,343,1460,1482]" pageId="33" pageNumber="34">G.</emphasis>
Inflorescence.
<emphasis id="B9035B42FFAEFFFCFE21FA55FD95FA5A" bold="true" box="[490,515,1460,1481]" pageId="33" pageNumber="34">H.</emphasis>
Fruits. Photographs: A. van de Beek.
</paragraph>
</caption>
<subSubSection id="C36DD4DBFFAEFFFCFF76F9FDFDD0F9A5" box="[189,582,1564,1591]" pageId="33" pageNumber="34" type="biology_ecology">
<paragraph id="8BC88750FFAEFFFCFF76F9FDFDD0F9A5" blockId="33.[136,1452,1564,2059]" box="[189,582,1564,1591]" pageId="33" pageNumber="34">
<emphasis id="B9035B42FFAEFFFCFF76F9FDFEB0F9A5" bold="true" box="[189,294,1564,1590]" pageId="33" pageNumber="34">Ecology:</emphasis>
—Roadsides, open fields.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAEFFFCFF76F9A1FBFCF954" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BC88750FFAEFFFCFF76F9A1FBFCF954" blockId="33.[136,1452,1564,2059]" pageId="33" pageNumber="34">
<emphasis id="B9035B42FFAEFFFCFF76F9A1FECFF9C9" bold="true" box="[189,345,1600,1626]" pageId="33" pageNumber="34">Distribution:</emphasis>
—Widespread in
<collectingCountry id="F360C7C0FFAEFFFCFDEAF9A1FD27F9C9" box="[545,689,1600,1626]" name="South Africa" pageId="33" pageNumber="34">South Africa</collectingCountry>
and
<collectingCountry id="F360C7C0FFAEFFFCFD21F9A1FCDEF9C9" box="[746,840,1600,1626]" name="Lesotho" pageId="33" pageNumber="34">Lesotho</collectingCountry>
. The typical
<taxonomicName id="4C77FCD3FFAEFFFCFC17F9A1FB54F9C9" authority="Hangklip" authorityName="Hangklip" box="[988,1218,1600,1626]" form="from" pageId="33" pageNumber="34" rank="form">form from Hangklip</taxonomicName>
eastward to the Van Stadens Pass. Similar forms at many places in the eastern and northern provinces. In spite of extensive searches in the region of Hangklip, the species could not be found there again. A good match was found in hedges along the road from Swellendam to Suurbraak, where it grows together with other forms of
<taxonomicName id="4C77FCD3FFAEFFFCFC6CF94DFBF5F955" authorityName="A. Beek" authorityYear="2021" box="[935,1123,1708,1735]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="series" series="Scleri">
<emphasis id="B9035B42FFAEFFFCFC6CF94DFC7BF955" box="[935,1005,1708,1734]" italics="true" pageId="33" pageNumber="34">Rubus</emphasis>
ser.
<emphasis id="B9035B42FFAEFFFCFBE9F94DFBF5F955" box="[1058,1123,1708,1734]" italics="true" pageId="33" pageNumber="34">Scleri</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAEFFFCFF76F931FAABF8E9" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BC88750FFAEFFFCFF76F931FAABF8E9" blockId="33.[136,1452,1564,2059]" pageId="33" pageNumber="34">
<emphasis id="B9035B42FFAEFFFCFF76F931FD62F979" bold="true" box="[189,756,1744,1771]" pageId="33" pageNumber="34">
<collectingRegion id="49B349B2FFAEFFFCFF76F931FEA9F979" box="[189,319,1744,1770]" name="Specimens" pageId="33" pageNumber="34">Specimens</collectingRegion>
:—
<collectingCountry id="F360C7C0FFAEFFFCFEAAF931FDACF979" box="[353,570,1744,1770]" name="South Africa" pageId="33" pageNumber="34">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FFAEFFFCFD83F931FD62F979" box="[584,756,1744,1770]" country="South Africa" name="Western Cape" pageId="33" pageNumber="34">Western Cape</collectingRegion>
</emphasis>
: along the road from Swellendam to Suurbraak,
<date id="FFC9A190FFAEFFFCFAE5F931FF55F89D" pageId="33" pageNumber="34" value="2020-01-13">13 January 2020</date>
,
<emphasis id="B9035B42FFAEFFFCFF1BF915FEFDF89D" box="[208,363,1780,1806]" italics="true" pageId="33" pageNumber="34">Beek 2020.05</emphasis>
(L); Swellendam, in utroque litore Buffelsjagrivier ab Swellendam ad Rietkuil,
<emphasis id="B9035B42FFAEFFFCFACCF914FAD0F89D" box="[1287,1350,1780,1806]" italics="true" pageId="33" pageNumber="34">s. dat</emphasis>
.,
<emphasis id="B9035B42FFAEFFFCFA95F915FF56F8A1" italics="true" pageId="33" pageNumber="34">Zeyher 2450</emphasis>
(PRE,
<taxonomicName id="4C77FCD3FFAEFFFCFEDDF8F9FEF5F8A1" authorityName="Linnaeus" authorityYear="1753" box="[278,355,1816,1842]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
chrysocarpus, not all specimens under this number are identical); Wilderness, Fairy Know, at the Ebb and Flow River,
<date id="FFC9A190FFAEFFFCFE4BF8DDFDD7F8C5" box="[384,577,1852,1879]" pageId="33" pageNumber="34" value="1929-01-13">13 January 1929</date>
,
<emphasis id="B9035B42FFAEFFFCFD85F8DCFD4BF8C5" box="[590,733,1852,1878]" italics="true" pageId="33" pageNumber="34">Mogg 11740</emphasis>
(PRE).
<emphasis id="B9035B42FFAEFFFCFCF6F8DDFC77F8C5" bold="true" box="[829,993,1852,1878]" pageId="33" pageNumber="34">
<collectingRegion id="49B349B2FFAEFFFCFCF6F8DDFC77F8C5" box="[829,993,1852,1878]" country="South Africa" name="Eastern Cape" pageId="33" pageNumber="34">Eastern Cape</collectingRegion>
</emphasis>
: Humansdorp, Clarkson,
<date id="FFC9A190FFAEFFFCFAC2F8DDFA31F8C5" box="[1289,1447,1852,1878]" pageId="33" pageNumber="34" value="1926-10">October 1926</date>
,
<emphasis id="B9035B42FFAEFFFCFF43F881FE99F8E9" box="[136,271,1888,1914]" italics="true" pageId="33" pageNumber="34">Thode A822</emphasis>
(PRE,
<taxonomicName id="4C77FCD3FFAEFFFCFEACF881FE6FF8E9" authorityName="Linnaeus" authorityYear="1753" box="[359,505,1888,1914]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="fruticosus">R. fruticosus</taxonomicName>
var. bergii); Near Van Stadens Pass,
<date id="FFC9A190FFAEFFFCFC6AF881FBCBF8E9" box="[929,1117,1888,1915]" pageId="33" pageNumber="34" value="1949-01-14">14 January 1949</date>
,
<emphasis id="B9035B42FFAEFFFCFBA3F881FB7EF8E9" box="[1128,1256,1888,1914]" italics="true" pageId="33" pageNumber="34">
<taxonomicName id="4C77FCD3FFAEFFFCFBA3F881FB2EF8E9" authorityName="Eyton" authorityYear="1845" box="[1128,1208,1888,1914]" class="Aves" genus="Theron" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">Theron</taxonomicName>
568
</emphasis>
(PRE).
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFAEFFFFFF76F865FBA8FDD7" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8BC88750FFAEFFFFFF76F865FEA1FE92" blockId="33.[136,1452,1564,2059]" lastBlockId="34.[136,1452,158,761]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B9035B42FFAEFFFCFF76F865FED2F80D" bold="true" box="[189,324,1924,1950]" pageId="33" pageNumber="34">Taxonomy:</emphasis>
—The typical
<taxonomicName id="4C77FCD3FFAEFFFCFE2DF865FDDFF80D" box="[486,585,1924,1950]" form="has" pageId="33" pageNumber="34" rank="form">form has</taxonomicName>
very strong, compressed, hooked prickles, adaxially moderately hairy leaves and straight teeth with short mucros. From the Van Stadens Pass eastward plants occur with less compressed only curved prickles (
<figureCitation id="134C9BD5FFAEFFFCFF3AF82DFECEF875" box="[241,344,1996,2022]" captionStart="FIGURE 11" captionStartId="34.[136,229,1715,1737]" captionTargetBox="[151,1436,785,1690]" captionTargetId="figure-286@34.[151,1436,785,1690]" captionTargetPageId="34" captionText="FIGURE 11. Rubus sclerus. A. Primocane. B. Pedicels. C. Herbarium specimen (Beek 2018.18). Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061173" httpUri="https://zenodo.org/record/8061173/files/figure.png" pageId="33" pageNumber="34">Fig. 11A</figureCitation>
), adaxially denser hairy leaves and with very fine almost fringed serrature with long mucros (
<figureCitation id="134C9BD5FFAEFFFCFAB5F82DFF2FF799" captionStart="FIGURE 11" captionStartId="34.[136,229,1715,1737]" captionTargetBox="[151,1436,785,1690]" captionTargetId="figure-286@34.[151,1436,785,1690]" captionTargetPageId="34" captionText="FIGURE 11. Rubus sclerus. A. Primocane. B. Pedicels. C. Herbarium specimen (Beek 2018.18). Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061173" httpUri="https://zenodo.org/record/8061173/files/figure.png" pageId="33" pageNumber="34">Fig. 11B</figureCitation>
). It was given the provisional name
<taxonomicName id="4C77FCD3FFAEFFFCFD9DF810FD52F799" box="[598,708,2032,2058]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="33" pageNumber="34" phylum="Tracheophyta" rank="species" species="sclerus">
<emphasis id="B9035B42FFAEFFFCFD9DF810FD52F799" box="[598,708,2032,2058]" italics="true" pageId="33" pageNumber="34">R. sclerus</emphasis>
</taxonomicName>
(
<figureCitation id="134C9BD5FFAEFFFCFD11F811FCA9F799" box="[730,831,2032,2058]" captionStart="FIGURE 11" captionStartId="34.[136,229,1715,1737]" captionTargetBox="[151,1436,785,1690]" captionTargetId="figure-286@34.[151,1436,785,1690]" captionTargetPageId="34" captionText="FIGURE 11. Rubus sclerus. A. Primocane. B. Pedicels. C. Herbarium specimen (Beek 2018.18). Photographs: A. van de Beek." figureDoi="http://doi.org/10.5281/zenodo.8061173" httpUri="https://zenodo.org/record/8061173/files/figure.png" pageId="33" pageNumber="34">Fig. 11C</figureCitation>
). However, many intermediate plants were found and it was not possible to find consistent combinations of characteristics for separating both forms. Different characteristics were sometimes found at the same plant, so that more research is needed before these plants can be conceived as an accepted taxon.
</paragraph>
<paragraph id="8BC88750FFADFFFFFF76FEEBFBA8FDD7" blockId="34.[136,1452,158,761]" pageId="34" pageNumber="35">
<emphasis id="B9035B42FFADFFFFFF76FEEBFDD3FEB6" bold="true" box="[189,581,266,293]" pageId="34" pageNumber="35">Specimens of the eastern form:—</emphasis>
<materialsCitation id="3B1F8D0DFFADFFFFFD8EFEEBFE8EFEDB" collectingDate="1907-12-26" collectionCode="L" collectorName="Heidelberg" country="South Africa" location="Leendertz" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Gauteng">
<emphasis id="B9035B42FFADFFFFFD8EFEEBFC1AFEB7" bold="true" box="[581,908,266,292]" pageId="34" pageNumber="35">
<collectingCountry id="F360C7C0FFADFFFFFD8EFEEBFC8CFEB7" box="[581,794,266,292]" name="South Africa" pageId="34" pageNumber="35">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FFADFFFFFCEEFEEBFC1AFEB7" box="[805,908,266,292]" country="South Africa" name="Gauteng" pageId="34" pageNumber="35">Gauteng</collectingRegion>
</emphasis>
:
<collectorName id="2682E286FFADFFFFFC52FEEBFB8EFEB7" box="[921,1048,266,292]" pageId="34" pageNumber="35">Heidelberg</collectorName>
,
<date id="FFC9A190FFADFFFFFBEAFEEBFB60FEB7" box="[1057,1270,266,292]" pageId="34" pageNumber="35" value="1907-12-26">
<collectingDate id="EF8D5878FFADFFFFFBEAFEEBFB60FEB7" box="[1057,1270,266,292]" pageId="34" pageNumber="35" value="1907-12-26">26 December 1907</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFB34FEEBFA3DFEB7" box="[1279,1451,266,292]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFB34FEEBFAFBFEB7" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFB34FEEBFAFBFEB7" box="[1279,1389,266,292]" country="South Africa" name="Leendertz" pageId="34" pageNumber="35" stateProvince="Gauteng">Leendertz</location>
1069
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFF5BFECFFF35FEDB" box="[144,163,302,328]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="34" pageNumber="35" type="Herbarium">L</collectionCode>
.1913163)
</materialsCitation>
.
<materialsCitation id="3B1F8D0DFFADFFFFFEEEFECFFB1DFEDB" box="[293,1163,302,329]" collectingDate="1972-04" collectionCode="PRE" collectorName="Golden Gate" country="South Africa" location="Jacobs" municipality="Wodehouse Kop" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Free State">
<emphasis id="B9035B42FFADFFFFFEEEFECFFE37FEDB" bold="true" box="[293,417,302,328]" pageId="34" pageNumber="35">
<collectingRegion id="49B349B2FFADFFFFFEEEFECFFE37FEDB" box="[293,417,302,328]" country="South Africa" name="Free State" pageId="34" pageNumber="35">Free State</collectingRegion>
</emphasis>
:
<collectorName id="2682E286FFADFFFFFE79FECFFDD0FEDB" box="[434,582,302,328]" pageId="34" pageNumber="35">Golden Gate</collectorName>
,
<collectingMunicipality id="6BAC1D2AFFADFFFFFD99FECFFC87FEDA" box="[594,785,302,329]" pageId="34" pageNumber="35">Wodehouse Kop</collectingMunicipality>
,
<date id="FFC9A190FFADFFFFFCD6FECFFC0AFEDB" box="[797,924,302,328]" pageId="34" pageNumber="35" value="1972-04">
<collectingDate id="EF8D5878FFADFFFFFCD6FECFFC0AFEDB" box="[797,924,302,328]" pageId="34" pageNumber="35" value="1972-04">April 1972</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFC62FECFFBAFFEDB" box="[937,1081,302,328]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFC62FECFFC61FEDB" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFC62FECFFC61FEDB" box="[937,1015,302,328]" country="South Africa" municipality="Wodehouse Kop" name="Jacobs" pageId="34" pageNumber="35" stateProvince="Free State">Jacobs</location>
8557
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFB81FECFFB15FEDB" box="[1098,1155,302,328]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="34" pageNumber="35" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFADFFFFFB52FECFFA99FEFF" collectingDate="2019-02-17" collectionCode="L" collectorName="Clarens &amp; Golden Gate" country="South Africa" location="Beek" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Free State">
<collectorName id="2682E286FFADFFFFFB52FECFFB67FEDB" box="[1177,1265,302,328]" pageId="34" pageNumber="35">Clarens</collectorName>
, road to
<collectorName id="2682E286FFADFFFFFA92FECFFF56FEFF" pageId="34" pageNumber="35">Golden Gate</collectorName>
, short before entrance of the park, south side of the road,
<date id="FFC9A190FFADFFFFFCADFEB3FBA0FEFF" box="[870,1078,338,364]" pageId="34" pageNumber="35" value="2019-02-17">
<collectingDate id="EF8D5878FFADFFFFFCADFEB3FBA0FEFF" box="[870,1078,338,364]" pageId="34" pageNumber="35" value="2019-02-17">17 February 2019</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFB88FEB3FB48FEFF" box="[1091,1246,338,364]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFB88FEB3FBEFFEFF" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFB88FEB3FBEFFEFF" box="[1091,1145,338,364]" country="South Africa" name="Beek" pageId="34" pageNumber="35" stateProvince="Free State">Beek</location>
2019.37
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFB3AFEB3FA90FEFF" box="[1265,1286,338,364]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="34" pageNumber="35" type="Herbarium">L</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFADFFFFFAD5FEB3FDE7FE03" collectingDate="1918-01-08" collectionCode="PRE" collectorName="Fouriesburg &amp; Dunelm" country="South Africa" location="Potts" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Free State">
<collectorName id="2682E286FFADFFFFFAD5FEB3FA3EFEFF" box="[1310,1448,338,364]" pageId="34" pageNumber="35">Fouriesburg</collectorName>
,
<collectorName id="2682E286FFADFFFFFF43FE97FF73FE03" box="[136,229,374,400]" pageId="34" pageNumber="35">Dunelm</collectorName>
,
<date id="FFC9A190FFADFFFFFF3BFE97FE08FE03" box="[240,414,374,401]" pageId="34" pageNumber="35" value="1918-01-08">
<collectingDate id="EF8D5878FFADFFFFFF3BFE97FE08FE03" box="[240,414,374,401]" pageId="34" pageNumber="35" value="1918-01-08">8 January 1918</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFE63FE96FDB7FE03" box="[424,545,374,400]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFE63FE96FE74FE03" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFE63FE96FE74FE03" box="[424,482,375,400]" country="South Africa" name="Potts" pageId="34" pageNumber="35" stateProvince="Free State">Potts</location>
6683
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFDE4FE97FDFEFE03" box="[559,616,374,400]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="34" pageNumber="35" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1F8D0DFFADFFFFFDB0FE97FE87FE27" collectingDate="2018-02-16" collectionCode="L" collectorName="Spioenkop" country="South Africa" location="Beek" pageId="34" pageNumber="35" specimenCount="1" stateProvince="KwaZulu-Natal">
<emphasis id="B9035B42FFADFFFFFDB0FE97FCA1FE03" bold="true" box="[635,823,374,400]" pageId="34" pageNumber="35">
<collectingRegion id="49B349B2FFADFFFFFDB0FE97FCA1FE03" box="[635,823,374,400]" country="South Africa" name="KwaZulu-Natal" pageId="34" pageNumber="35">KwaZulu-Natal</collectingRegion>
</emphasis>
:
<collectorName id="2682E286FFADFFFFFC8DFE97FC28FE03" box="[838,958,374,400]" pageId="34" pageNumber="35">Spioenkop</collectorName>
memorial, parking,
<date id="FFC9A190FFADFFFFFB6AFE97FAFCFE03" box="[1185,1386,374,400]" pageId="34" pageNumber="35" value="2018-02-16">
<collectingDate id="EF8D5878FFADFFFFFB6AFE97FAFCFE03" box="[1185,1386,374,400]" pageId="34" pageNumber="35" value="2018-02-16">16 February 2018</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFABEFE97FF75FE27" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFABEFE97FA3DFE03" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFABEFE97FA3DFE03" box="[1397,1451,374,400]" country="South Africa" name="Beek" pageId="34" pageNumber="35" stateProvince="KwaZulu-Natal">Beek</location>
2018.18
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFF38FE7BFE9EFE27" box="[243,264,410,436]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="34" pageNumber="35" type="Herbarium">L</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B1F8D0DFFADFFFFFED7FE7BFDD8FE4B" collectionCode="NBG" collectorName="Zeyher" country="South Africa" county="On" location="Adow" municipality="Zwartkops River" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Eastern Cape">
<emphasis id="B9035B42FFADFFFFFED7FE7BFE28FE27" bold="true" box="[284,446,410,436]" pageId="34" pageNumber="35">
<collectingRegion id="49B349B2FFADFFFFFED7FE7BFE28FE27" box="[284,446,410,436]" country="South Africa" name="Eastern Cape" pageId="34" pageNumber="35">Eastern Cape</collectingRegion>
</emphasis>
:
<collectorName id="2682E286FFADFFFFFE19FE7BFE78FE27" box="[466,494,410,436]" pageId="34" pageNumber="35">In</collectorName>
mont. supra
<collectorName id="2682E286FFADFFFFFDB4FE7AFCDCFE27" box="[639,842,410,437]" pageId="34" pageNumber="35">Van Stadensriver</collectorName>
,
<emphasis id="B9035B42FFADFFFFFC9EFE7BFC50FE27" box="[853,966,410,436]" italics="true" pageId="34" pageNumber="35">
<collectorName id="2682E286FFADFFFFFC9EFE7BFC35FE27" box="[853,931,410,436]" pageId="34" pageNumber="35">Zeyher</collectorName>
66
</emphasis>
=
<collectingCounty id="62A9FFDCFFADFFFFFC2FFE7BFB90FE27" box="[996,1030,410,436]" pageId="34" pageNumber="35">On</collectingCounty>
the banks of
<collectingMunicipality id="6BAC1D2AFFADFFFFFB57FE7BFACFFE27" box="[1180,1369,410,436]" pageId="34" pageNumber="35">Zwartkops River</collectingMunicipality>
and on the hills of
<location id="8EA8D18BFFADFFFFFECEFE5FFEDDFE4B" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFECEFE5FFEDDFE4B" box="[261,331,446,472]" country="South Africa" county="On" municipality="Zwartkops River" name="Adow" pageId="34" pageNumber="35" stateProvince="Eastern Cape">Adow</location>
,
<location id="8EA8D18BFFADFFFFFE9DFE5FFE1EFE4B" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFE9DFE5FFE1EFE4B" box="[342,392,446,472]" country="South Africa" county="On" municipality="Zwartkops River" name="Oct." pageId="34" pageNumber="35" stateProvince="Eastern Cape">Oct.</location>
,
<emphasis id="B9035B42FFADFFFFFE58FE5FFE60FE4B" box="[403,502,446,472]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFE58FE5FFE77FE4B" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFE58FE5FFE77FE4B" box="[403,481,446,472]" country="South Africa" county="On" municipality="Zwartkops River" name="Zeyher" pageId="34" pageNumber="35" stateProvince="Eastern Cape">Zeyher</location>
4
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFDCEFE5FFDD0FE4B" box="[517,582,446,472]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="34" pageNumber="35" type="Herbarium">NBG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFADFFFFFD95FE5FFB2FFDB3" collectingDate="2020-01-15" collectionCode="L, PRE" collectorName="Ecklon &amp; Zeyher" country="South Africa" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Eastern Cape">
<collectorName id="2682E286FFADFFFFFD95FE5FFD13FE4B" box="[606,645,446,472]" pageId="34" pageNumber="35">Ad</collectorName>
ripam fluminis
<collectorName id="2682E286FFADFFFFFCF6FE5FFC69FE4B" box="[829,1023,446,472]" pageId="34" pageNumber="35">Zwartkopsrivier</collectorName>
et in collibus (alt. I-III) terrae
<collectorName id="2682E286FFADFFFFFA90FE5FFA3AFE4B" box="[1371,1452,446,472]" pageId="34" pageNumber="35">Adow</collectorName>
(
<collectorName id="2682E286FFADFFFFFF5BFE03FE9FFE6F" box="[144,265,482,508]" pageId="34" pageNumber="35">Uitenhage</collectorName>
), tum in
<collectorName id="2682E286FFADFFFFFEBEFE03FE51FE6F" box="[373,455,482,508]" pageId="34" pageNumber="35">Albany</collectorName>
et
<collectorName id="2682E286FFADFFFFFE3BFE03FDE3FE6F" box="[496,629,482,508]" pageId="34" pageNumber="35">Kafferland</collectorName>
,
<emphasis id="B9035B42FFADFFFFFD4FFE03FC1BFE6F" box="[644,909,482,509]" italics="true" pageId="34" pageNumber="35">
<collectorName id="2682E286FFADFFFFFD4FFE03FD44FE6F" box="[644,722,482,508]" pageId="34" pageNumber="35">Ecklon</collectorName>
&amp;
<collectorName id="2682E286FFADFFFFFD36FE03FCDDFE6F" box="[765,843,482,508]" pageId="34" pageNumber="35">Zeyher</collectorName>
1709
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFC6BFE03FC24FE6F" box="[928,946,482,508]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="34" pageNumber="35" type="Herbarium">L</collectionCode>
.1913162,
<collectionCode id="ED661F95FFADFFFFFBE6FE02FBF5FE6E" box="[1069,1123,483,509]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="34" pageNumber="35" type="Herbarium">PRE</collectionCode>
, other samples of
<emphasis id="B9035B42FFADFFFFFAF6FE03FE82FDB3" italics="true" pageId="34" pageNumber="35">
<collectorName id="2682E286FFADFFFFFAF6FE03FA1DFE6F" box="[1341,1419,482,508]" pageId="34" pageNumber="35">Ecklon</collectorName>
&amp;
<collectorName id="2682E286FFADFFFFFF43FDE7FF40FDB3" box="[136,214,518,544]" pageId="34" pageNumber="35">Zeyher</collectorName>
<date id="FFC9A190FFADFFFFFF17FDE7FE82FDB3" box="[220,276,518,544]" pageId="34" pageNumber="35" value="1709-05">1709</date>
</emphasis>
may be different);
<collectorName id="2682E286FFADFFFFFE2DFDE7FDCAFDB3" box="[486,604,518,544]" pageId="34" pageNumber="35">Uitenhage</collectorName>
,
<collectorName id="2682E286FFADFFFFFDADFDE6FCBEFDB2" box="[614,808,518,545]" pageId="34" pageNumber="35">Van Stadens Pass</collectorName>
,
<date id="FFC9A190FFADFFFFFCF9FDE7FC7AFDB3" box="[818,1004,518,545]" pageId="34" pageNumber="35" value="2020-01-15">
<collectingDate id="EF8D5878FFADFFFFFCF9FDE7FC7AFDB3" box="[818,1004,518,545]" pageId="34" pageNumber="35" value="2020-01-15">15 January 2020</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFC3DFDE7FB1BFDB3" box="[1014,1165,518,544]" italics="true" pageId="34" pageNumber="35">Beek 2020.10</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFB57FDE7FB27FDB3" box="[1180,1201,518,544]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="34" pageNumber="35" type="Herbarium">L</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFADFFFFFB0FFDE7FDF4FDD6" collectionCode="C" collectorName="Galgebosch" country="South Africa" location="Drege" pageId="34" pageNumber="35" specimenCode="HBG 503845" specimenCount="1" stateProvince="Eastern Cape">
<collectorName id="2682E286FFADFFFFFB0FFDE7FADEFDB3" box="[1220,1352,518,544]" pageId="34" pageNumber="35">Galgebosch</collectorName>
(locality code IV
<collectionCode id="ED661F95FFADFFFFFF23FDCBFF6DFDD7" box="[232,251,554,580]" country="Denmark" name="University of Copenhagen" pageId="34" pageNumber="35" type="Herbarium">C</collectionCode>
c 9),
<emphasis id="B9035B42FFADFFFFFEF1FDCAFE38FDD7" box="[314,430,555,581]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFEF1FDCAFE16FDD6" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFEF1FDCAFE16FDD6" box="[314,384,555,581]" country="South Africa" name="Drege" pageId="34" pageNumber="35" stateProvince="Eastern Cape">Drège</location>
s.n.
</emphasis>
(
<specimenCode id="DBD12F2BFFADFFFFFE76FDCBFDCCFDD6" box="[445,602,554,581]" collectionCode="HBG" country="Japan" httpUri="http://grbio.org/cool/fv5j-intd" name="Hiroshima Botanical Garden" pageId="34" pageNumber="35">HBG 503845</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B1F8D0DFFADFFFFFDA6FDCAFBACFDD7" box="[621,1082,554,581]" collectionCode="NBG" country="South Africa" location="Thode" municipality="East London" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Eastern Cape">
<collectingMunicipality id="6BAC1D2AFFADFFFFFDA6FDCAFD68FDD7" box="[621,766,554,581]" pageId="34" pageNumber="35">East London</collectingMunicipality>
, 1888,
<emphasis id="B9035B42FFADFFFFFC84FDCBFC74FDD7" box="[847,994,554,580]" italics="true" pageId="34" pageNumber="35">
<location id="8EA8D18BFFADFFFFFC84FDCBFC03FDD7" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFC84FDCBFC03FDD7" box="[847,917,554,580]" country="South Africa" municipality="East London" name="Thode" pageId="34" pageNumber="35" stateProvince="Eastern Cape">Thode</location>
10440
</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFC3AFDCBFBA4FDD7" box="[1009,1074,554,580]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15565" lsid="urn:lsid:biocol.org:col:15565" name="South African National Biodiversity Institute" pageId="34" pageNumber="35" type="Herbarium">NBG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFADFFFFFF76FDAEFC31FDFB" box="[189,935,590,617]" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BC88750FFADFFFFFF76FDAEFC31FDFB" blockId="34.[136,1452,158,761]" box="[189,935,590,617]" pageId="34" pageNumber="35">
<materialsCitation id="3B1F8D0DFFADFFFFFF12FDAFFC35FDFB" box="[217,931,590,617]" collectingDate="1949-04-15" collectionCode="PRE" country="Lesotho" location="Mamathes" pageId="34" pageNumber="35" specimenCount="1">
<emphasis id="B9035B42FFADFFFFFF12FDAFFEFAFDFB" bold="true" box="[217,364,590,616]" pageId="34" pageNumber="35">
<collectingCountry id="F360C7C0FFADFFFFFF12FDAFFEF0FDFB" box="[217,358,590,616]" name="Lesotho" pageId="34" pageNumber="35">LESOTHO</collectingCountry>
:
</emphasis>
<location id="8EA8D18BFFADFFFFFEB8FDAFFE7DFDFB" LSID="urn:lsid:plazi:treatment:03DE3646FFAFFFFFFF43FA4FFCBEFD6B:8EA8D18BFFADFFFFFEB8FDAFFE7DFDFB" box="[371,491,590,616]" country="Lesotho" name="Mamathes" pageId="34" pageNumber="35">Mamathes</location>
,
<date id="FFC9A190FFADFFFFFE3EFDAFFD02FDFB" box="[501,660,590,617]" pageId="34" pageNumber="35" value="1949-04-15">
<collectingDate id="EF8D5878FFADFFFFFE3EFDAFFD02FDFB" box="[501,660,590,617]" pageId="34" pageNumber="35" value="1949-04-15">15 April 1949</collectingDate>
</date>
,
<emphasis id="B9035B42FFADFFFFFD54FDAFFCC5FDFB" box="[671,851,590,616]" italics="true" pageId="34" pageNumber="35">Guillarmod 868</emphasis>
(
<collectionCode id="ED661F95FFADFFFFFCA9FDAFFC0DFDFB" box="[866,923,590,616]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15547" lsid="urn:lsid:biocol.org:col:15547" name="South African National Biodiversity Institute (SANBI)" pageId="34" pageNumber="35" type="Herbarium">PRE</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFFADFFFFFF76FD93FCBEFD6B" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BC88750FFADFFFFFF76FD93FCBEFD6B" blockId="34.[136,1452,158,761]" pageId="34" pageNumber="35">
<emphasis id="B9035B42FFADFFFFFF76FD93FF69FD1F" bold="true" box="[189,255,626,652]" pageId="34" pageNumber="35">Note:</emphasis>
—Also plants with other different characteristics, such as patent straight prickles, white petals, abaxially glabrous leaves, or incised leave margins, were found. In the studied region such plants are rare, but their occurrence increase to the east and north, especially in northern
<collectingRegion id="49B349B2FFADFFFFFD18FD5BFC10FD47" box="[723,902,698,724]" country="South Africa" name="KwaZulu-Natal" pageId="34" pageNumber="35">KwaZulu-Natal</collectingRegion>
, where they are abundant. The precise relation of all these diverging forms must be subject of future research.
</paragraph>
</subSubSection>
</treatment>
</document>