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<document id="315A8E7D3DB4E77C678C9A82F3C0B839" ID-DOI="10.11646/phytotaxa.515.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8061143" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="jonas" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="jonas" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="jonas" IM.treatments_approvedBy="jonas" checkinTime="1687282820830" checkinUser="plazi" docAuthor="Beek, Abraham Van De" docDate="2021" docId="03DE3646FF82FFD3FF43FD3FFCF9FACF" docLanguage="en" docName="Phytotaxa.515.1.1.pdf" docOrigin="Phytotaxa 515 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.515.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Rubus pappei Ecklon &amp; Zeyher 1836" docType="treatment" docVersion="6" lastPageNumber="15" masterDocId="FFE74E3EFF8FFFDDFFCBFFE1FF96FF93" masterDocTitle="Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa" masterLastPageNumber="71" masterPageNumber="1" pageNumber="14" updateTime="1699527950928" updateUser="plazi">
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<mods:title id="6018396D304C8064E3CBCAF68DA17FF1">Rubi Capenses: a further contribution to the knowledge of the genus Rubus (Rosaceae) in South Africa</mods:title>
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<mods:namePart id="B0603B4F736BC0772FD3CF536313B293">Beek, Abraham Van De</mods:namePart>
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<mods:title id="3E9C156D20438D9A662C3E6D4F6BB289">Phytotaxa</mods:title>
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<mods:date id="7002F088375C40EBF03CC2DB24DE378F">2021</mods:date>
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<treatment id="03DE3646FF82FFD3FF43FD3FFCF9FACF" ID-DOI="http://doi.org/10.5281/zenodo.8071552" ID-Zenodo-Dep="8071552" LSID="urn:lsid:plazi:treatment:03DE3646FF82FFD3FF43FD3FFCF9FACF" httpUri="http://treatment.plazi.org/id/03DE3646FF82FFD3FF43FD3FFCF9FACF" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<subSubSection id="C36DD4DBFF82FFD0FF43FD3FFD34FD6B" box="[136,674,734,761]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BC88750FF82FFD0FF43FD3FFD34FD6B" blockId="13.[136,1452,734,797]" box="[136,674,734,761]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF43FD3FFF0BFD6B" bold="true" box="[136,157,734,760]" pageId="13" pageNumber="14">3.</emphasis>
<taxonomicName id="4C77FCD3FF82FFD0FF6EFD3FFD0BFD6B" ID-CoL="4TL66" ID-ENA="2904924" authority="Ecklon &amp; Zeyher (1836: 263)" authorityName="Ecklon &amp; Zeyher" authorityPageNumber="263" authorityYear="1836" box="[165,669,734,761]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF82FFD0FF6EFD3FFED5FD6B" bold="true" box="[165,323,734,761]" italics="true" pageId="13" pageNumber="14">Rubus pappei</emphasis>
<bibRefCitation id="EFE6FAA1FF82FFD0FE80FD3FFD0BFD6B" author="Ecklon, C. F. &amp; Zeyher, C. L. P." box="[331,669,734,760]" pageId="13" pageNumber="14" refId="ref46945" refString="Ecklon, C. F. &amp; Zeyher, C. L. P. (1836) Enumeratio plantarum Africae australis extratropicae: quae collectae, determinatae et exposita, Vol. 2. Perthes &amp; Besser, Hamburg, 144 pp. https: // doi. org / 10.5962 / bhl. title. 48" type="book" year="1836">Ecklon &amp; Zeyher (1836: 263)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD0FD62FD3EFCF2FC8F" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BC88750FF82FFD0FD62FD3EFCF2FC8F" blockId="13.[136,1452,734,797]" pageId="13" pageNumber="14">
<materialsCitation id="3B1F8D0DFF82FFD0FD62FD3EFCF2FC8F" pageId="13" pageNumber="14" typeStatus="lectotype">
<typeStatus id="54CC39F2FF82FFD0FD62FD3EFC8FFD6A" box="[681,793,735,761]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F360C7C0FF82FFD0FBCFFD3FFB42FD6B" box="[1028,1236,734,760]" name="South Africa" pageId="13" pageNumber="14">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF82FFD0FB14FD3EFA16FD6B" box="[1247,1408,734,761]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Western Cape</collectingRegion>
:
<collectingCounty id="62A9FFDCFF82FFD0FA5BFD3FFA3AFD6B" box="[1424,1452,734,760]" pageId="13" pageNumber="14">In</collectingCounty>
dumetis prope
<collectingMunicipality id="6BAC1D2AFF82FFD0FEE5FCE3FE53FC8F" box="[302,453,770,796]" pageId="13" pageNumber="14">Rondebosch</collectingMunicipality>
, 1833,
<emphasis id="B9035B42FF82FFD0FDDDFCE2FD1DFC8F" box="[534,651,771,796]" italics="true" pageId="13" pageNumber="14">
<location id="8EA8D18BFF82FFD0FDDDFCE2FDC8FC8F" LSID="urn:lsid:plazi:treatment:03DE3646FF82FFD3FF43FD3FFCF9FACF:8EA8D18BFF82FFD0FDDDFCE2FDC8FC8F" box="[534,606,771,796]" country="South Africa" county="In" municipality="Rondebosch'" name="Pappe" pageId="13" pageNumber="14" stateProvince="Western Cape">Pappe</location>
s.n.
</emphasis>
(BREM_0002212)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD0FF43FCABFDEBFB8B" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BC88750FF82FFD0FF43FCABFDEBFB8B" blockId="13.[136,1452,842,1048]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF43FCABFEF0FCF7" bold="true" box="[136,358,842,868]" pageId="13" pageNumber="14">
Note on
<typeStatus id="54CC39F2FF82FFD0FF3AFCABFEF7FCF7" box="[241,353,842,868]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
:
</emphasis>
—A specimen collected by
<collectorName id="2682E286FF82FFD0FD6DFCAAFD7CFCF6" box="[678,746,843,869]" pageId="13" pageNumber="14">Pappe</collectorName>
was selected as the
<typeStatus id="54CC39F2FF82FFD0FC11FCABFBD2FCF7" box="[986,1092,842,868]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
. The protologue mentions only localities in the
<collectingRegion id="49B349B2FF82FFD0FEFDFC8EFE5AFC1B" box="[310,460,878,905]" country="South Africa" name="Eastern Cape" pageId="13" pageNumber="14">Eastern Cape</collectingRegion>
, but the labels of
<collectorName id="2682E286FF82FFD0FD47FC8FFD4DFC1B" box="[652,731,878,904]" pageId="13" pageNumber="14">Ecklon</collectorName>
&amp;
<collectorName id="2682E286FF82FFD0FD36FC8FFCDAFC1B" box="[765,844,878,904]" pageId="13" pageNumber="14">Zeyher</collectorName>
are notoriously imprecise. Because Zeyher cooperated with Pappe in the time when both were in
<collectingRegion id="49B349B2FF82FFD0FDBDFC73FD26FC3F" box="[630,688,914,940]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Cape</collectingRegion>
Town, he would probably have seen Pappes specimen, which the epithet also indicates. This specimen is well defined, with a precise locality and corresponds with other specimens that were collected in that region. The localities on the label of Ecklon &amp; Zeyher might refer to their own collections, which they considered identical with Pappes plant.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD0FF43FBABFB30FA13" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BC88750FF82FFD0FF43FBABFB37FB17" blockId="13.[136,1453,1098,1408]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF43FBABFEF6FBF3" box="[136,352,1098,1120]" italics="true" pageId="13" pageNumber="14">Heterotypic synonyms</emphasis>
:—
<taxonomicName id="4C77FCD3FF82FFD0FEB5FBABFD60FBCC" ID-CoL="4TJVM" authority="Gandoger (1918: 24)" authorityName="Gandoger" authorityPageNumber="24" authorityYear="1918" box="[382,758,1098,1120]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="cumingii">
<emphasis id="B9035B42FF82FFD0FEB5FBABFD8FFBCC" box="[382,537,1098,1119]" italics="true" pageId="13" pageNumber="14">Rubus cumingii</emphasis>
<bibRefCitation id="EFE6FAA1FF82FFD0FDE8FBABFD60FBCC" author="Gandoger, M." box="[547,758,1098,1120]" pageId="13" pageNumber="14" pagination="24 - 69" refId="ref47605" refString="Gandoger, M. (1918). Sertum plantarum novarum. Pars prima. Bulletin de la Societe Botanique de France 65: 24 - 69. https: // doi. org / 10.1080 / 00378941.1918.10836046" type="journal article" year="1918">Gandoger (1918: 24)</bibRefCitation>
</taxonomicName>
. Type:
<emphasis id="B9035B42FF82FFD0FC8FFBABFC5CFBCC" box="[836,970,1098,1119]" italics="true" pageId="13" pageNumber="14">Cuming 2455</emphasis>
(
<typeStatus id="54CC39F2FF82FFD0FC11FBABFBA6FBF3" box="[986,1072,1098,1120]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
LY [https://herbier2014.univ-lyon1.fr/ fiche/8574], isotypes BREM_00002217, L.1918961, L.1918962,
<specimenCode id="DBD12F2BFF82FFD0FCF9FB8FFC34FB17" box="[818,930,1134,1156]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="13" pageNumber="14" type="Herbarium">P00682403</specimenCode>
,
<specimenCode id="DBD12F2BFF82FFD0FC60FB8FFB8AFB17" box="[939,1052,1134,1156]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="13" pageNumber="14" type="Herbarium">P00682406</specimenCode>
,
<specimenCode id="DBD12F2BFF82FFD0FBEEFB8FFB01FB17" box="[1061,1175,1134,1156]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="13" pageNumber="14" type="Herbarium">P04203582</specimenCode>
).
</paragraph>
<paragraph id="8BC88750FF82FFD0FF43FB73FCECFAAB" blockId="13.[136,1453,1098,1408]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB67A07EFF82FFD0FF43FB73FC57FB34" box="[136,961,1170,1192]" pageId="13" pageNumber="14">
?
<treatmentCitation id="0AD6A141FF82FFD0FF50FB73FD09FB3B" author="Meisner, C. F." box="[155,671,1170,1192]" page=")" pageId="13" pageNumber="14" year="1843">
<taxonomicName id="4C77FCD3FF82FFD0FF50FB73FD09FB3B" ID-CoL="4TKD8" authority="E. Mey. ex Meisner (1843: 527)" authorityName="Meisner" authorityPageNumber="527" authorityYear="1843" box="[155,671,1170,1192]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF82FFD0FF50FB73FEF7FB34" box="[155,353,1170,1191]" italics="true" pageId="13" pageNumber="14">Rubus heterophyllus</emphasis>
E.Mey. ex
<bibRefCitation id="EFE6FAA1FF82FFD0FE1FFB73FD09FB3B" author="Meisner, C. F." box="[468,671,1170,1192]" pageId="13" pageNumber="14" pagination="53 - 105" refId="ref48333" refString="Meisner, C. F. (1843) Contributions towards a Flora of South Africa. London Journal of Botany 2: 53 - 105; 527 - 559. [https: // www. biodiversitylibrary. org / item / 6313 # page / 56 / mode / 1 up]" type="journal article" year="1843">Meisner (1843: 527)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, non
<treatmentCitation id="0AD6A141FF82FFD0FD1CFB73FC2BFB34" author="Willdenow, C. L." box="[727,957,1170,1192]" page="413" pageId="13" pageNumber="14" year="1811">
<bibRefCitation id="EFE6FAA1FF82FFD0FD1CFB73FC2BFB34" author="Willdenow, C. L." box="[727,957,1170,1192]" pageId="13" pageNumber="14" refId="ref50580" refString="Willdenow, C. L. (1811) Die wilde Baumzucht oder Beschreibung aller im K ˆ nigl. Botan Garten etc. bey Berlin im Freien ausdauernden Baume und Straucher. Nauck, Berlin, 204 pp." type="book" year="1811">Willdenow (1811: 413)</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
<materialsCitation id="3B1F8D0DFF82FFD0FC03FB73FE2BFB63" collectionCode="LY" country="South Africa" location="Waldern im Voormansbosch" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Western Cape" typeStatus="lectotype">
<typeStatus id="54CC39F2FF82FFD0FC03FB73FBBEFB34" box="[968,1064,1170,1191]" pageId="13" pageNumber="14" type="lectotype">Lectotype</typeStatus>
(designated here):—
<collectingCountry id="F360C7C0FF82FFD0FB3FFB73FA3EFB34" box="[1268,1448,1170,1192]" name="South Africa" pageId="13" pageNumber="14">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF82FFD0FF76FB57FED3FB58" box="[189,325,1206,1227]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Western Cape</collectingRegion>
: 2452. …
<collectorName id="2682E286FF82FFD0FE67FB57FDA9FB5F" box="[428,575,1206,1228]" pageId="13" pageNumber="14">
In den
<location id="8EA8D18BFF82FFD0FE27FB57FDA9FB5F" LSID="urn:lsid:plazi:treatment:03DE3646FF82FFD3FF43FD3FFCF9FACF:8EA8D18BFF82FFD0FE27FB57FDA9FB5F" box="[492,575,1206,1228]" country="South Africa" name="Waldern im Voormansbosch" pageId="13" pageNumber="14" stateProvince="Western Cape">Wäldern</location>
</collectorName>
im Voormansbosch und
<collectorName id="2682E286FF82FFD0FCECFB57FC4EFB5F" box="[807,984,1206,1228]" pageId="13" pageNumber="14">Grootvadersbosch</collectorName>
,
<collectorName id="2682E286FF82FFD0FC2BFB57FBAFFB5F" box="[992,1081,1206,1228]" pageId="13" pageNumber="14">October</collectorName>
,
<emphasis id="B9035B42FF82FFD0FB8AFB57FB2FFB58" box="[1089,1209,1206,1227]" italics="true" pageId="13" pageNumber="14">
<collectorName id="2682E286FF82FFD0FB8AFB57FB12FB58" box="[1089,1156,1206,1227]" pageId="13" pageNumber="14">Zeyher</collectorName>
2452
</emphasis>
(
<collectionCode id="ED661F95FF82FFD0FB08FB57FB74FB58" box="[1219,1250,1206,1227]" country="France" name="Laboratoire de Mycologie associe au CNRS" pageId="13" pageNumber="14">LY</collectionCode>
[https://herbier2014. univ-lyon1.fr/fiche/8576])
</materialsCitation>
.
<materialsCitation id="3B1F8D0DFF82FFD0FE0DFB3BFCE0FAAB" collectionCode="P" country="South Africa" location="Zeyher" municipality="Puspasvalei" pageId="13" pageNumber="14" specimenCode="P02579472" specimenCount="1" stateProvince="Western Cape" typeStatus="syntype">
<typeStatus id="54CC39F2FF82FFD0FE0DFB3BFD83FB7C" box="[454,533,1242,1263]" pageId="13" pageNumber="14" type="syntype">Syntype</typeStatus>
:—
<collectingCountry id="F360C7C0FF82FFD0FDFAFB3BFD74FB7C" box="[561,738,1242,1263]" name="South Africa" pageId="13" pageNumber="14">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF82FFD0FD20FB3BFCE5FB7C" box="[747,883,1242,1263]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Western Cape</collectingRegion>
: 2452. Flora Capensis.
<taxonomicName id="4C77FCD3FF82FFD0FB91FB3BFF47FA80" ID-CoL="6X9P4" authority="Wild. Swellendam. In" authorityName="Wild. Swellendam. In" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF82FFD0FB91FB3BFB79FB7C" box="[1114,1263,1242,1263]" italics="true" pageId="13" pageNumber="14">Rubus pinnatus</emphasis>
Wild. Swellendam. In
</taxonomicName>
sylvaticis montium ad
<collectingMunicipality id="6BAC1D2AFF82FFD0FE74FB1FFDA5FA87" box="[447,563,1278,1300]" pageId="13" pageNumber="14">Puspasvalei</collectingMunicipality>
, Duyvelsbosch. Octobris. Legit Zeyher. Communicavit Drège,
<emphasis id="B9035B42FF82FFD0FB7AFB1FFABBFA80" box="[1201,1325,1278,1299]" italics="true" pageId="13" pageNumber="14">
<location id="8EA8D18BFF82FFD0FB7AFB1FFB62FA80" LSID="urn:lsid:plazi:treatment:03DE3646FF82FFD3FF43FD3FFCF9FACF:8EA8D18BFF82FFD0FB7AFB1FFB62FA80" box="[1201,1268,1278,1299]" country="South Africa" municipality="Puspasvalei" name="Zeyher" pageId="13" pageNumber="14" stateProvince="Western Cape">Zeyher</location>
2452
</emphasis>
(
<specimenCode id="DBD12F2BFF82FFD0FAF7FB1FFA3DFA87" box="[1340,1451,1278,1300]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="13" pageNumber="14" type="Herbarium">P02579472</specimenCode>
[https://science.mnhn.fr/institution/mnhn/collection/p/item/p02579472])
</materialsCitation>
.
</paragraph>
<paragraph id="8BC88750FF82FFD0FF43FAA7FB30FA13" blockId="13.[136,1453,1098,1408]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB67A07EFF82FFD0FF43FAA7FB30FA13" pageId="13" pageNumber="14">
?
<treatmentCitation id="0AD6A141FF82FFD0FF57FAA7FD95FACF" author="Gandoger, M." box="[156,515,1350,1372]" page=")" pageId="13" pageNumber="14" year="1918">
<taxonomicName id="4C77FCD3FF82FFD0FF57FAA7FD95FACF" authority="Gandoger (1918: 25)" authorityName="Gandoger" authorityPageNumber="25" authorityYear="1918" box="[156,515,1350,1372]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="B9035B42FF82FFD0FF57FAA7FEB0FAC8" box="[156,294,1350,1371]" italics="true" pageId="13" pageNumber="14">Rubus zeyheri</emphasis>
<materialsCitation id="3B1F8D0DFF82FFD0FEFBFAA7FD95FACF" box="[304,515,1350,1372]" collectionCode="LY" collectorName="In den Waldern" country="South Africa" location="Waldern im Voormansbosch" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Western Cape" typeStatus="holotype">
<bibRefCitation id="EFE6FAA1FF82FFD0FEFBFAA7FD95FACF" author="Gandoger, M." box="[304,515,1350,1372]" pageId="13" pageNumber="14" pagination="24 - 69" refId="ref47605" refString="Gandoger, M. (1918). Sertum plantarum novarum. Pars prima. Bulletin de la Societe Botanique de France 65: 24 - 69. https: // doi. org / 10.1080 / 00378941.1918.10836046" type="journal article" year="1918">Gandoger (1918: 25)</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
. Type:—
<collectingCountry id="F360C7C0FF82FFD0FD97FAA7FC86FAC8" box="[604,784,1350,1371]" name="South Africa" pageId="13" pageNumber="14">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF82FFD0FCD7FAA7FC3FFAC8" box="[796,937,1350,1371]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Western Cape</collectingRegion>
: 2452. …
<collectorName id="2682E286FF82FFD0FBD5FAA7FB2CFACF" box="[1054,1210,1350,1372]" pageId="13" pageNumber="14">
In den
<location id="8EA8D18BFF82FFD0FBACFAA7FB2CFACF" LSID="urn:lsid:plazi:treatment:03DE3646FF82FFD3FF43FD3FFCF9FACF:8EA8D18BFF82FFD0FBACFAA7FB2CFACF" box="[1127,1210,1350,1372]" country="South Africa" name="Waldern im Voormansbosch" pageId="13" pageNumber="14" stateProvince="Western Cape">Wäldern</location>
</collectorName>
im Voormansbosch und Grootvadersbosch. October,
<emphasis id="B9035B42FF82FFD0FE11FA8BFDC5FAEC" box="[474,595,1386,1407]" italics="true" pageId="13" pageNumber="14">Zeyher 2452</emphasis>
(
<typeStatus id="54CC39F2FF82FFD0FD94FA8BFD23FA13" box="[607,693,1386,1408]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
LY [https://herbier2014.univ-lyon1.fr/fiche/8576]).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD0FF43FA4FFC3DF8A3" pageId="13" pageNumber="14" type="description">
<paragraph id="8BC88750FF82FFD0FF43FA4FFC3DF8A3" blockId="13.[136,1452,1454,2093]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF43FA4FFEC4FA5B" box="[136,338,1454,1480]" italics="true" pageId="13" pageNumber="14">Flowering branch</emphasis>
bluntly angular, densely very short hairy; prickles 414 per
<quantity id="4C8F2AB5FF82FFD0FC26FA4FFBB0FA5A" box="[1005,1062,1454,1481]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="14" unit="cm" value="5.0">5 cm</quantity>
, unequal, from
<quantity id="4C8F2AB5FF82FFD0FB19FA4FFAB8FA5A" box="[1234,1326,1454,1481]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="13" pageNumber="14" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">15 mm</quantity>
broad base compressed, curved, up to
<quantity id="4C8F2AB5FF82FFD0FE7FFA33FD84FA7E" box="[436,530,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
long; stipules lanceolate,
<quantity id="4C8F2AB5FF82FFD0FCFEFA33FC03FA7E" box="[821,917,1490,1517]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="13" pageNumber="14" unit="mm" value="8.0" valueMax="9.0" valueMin="7.0">79 mm</quantity>
.
<emphasis id="B9035B42FF82FFD0FC6BFA32FC78FA7F" box="[928,1006,1491,1516]" italics="true" pageId="13" pageNumber="14">Leaves</emphasis>
pinnate 5- to 7-foliolate, adaxially with strigose hairs, sometimes almost glabrous, abaxially with long white hairs on nerves; serrature narrow, irregular, deep, very sharp, with long more or less upward directed mucros; central leaflet
<quantity id="4C8F2AB5FF82FFD0FC00F9FBFBD2F9A6" box="[971,1092,1562,1589]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="5.5" metricValueMin="2.9" pageId="13" pageNumber="14" unit="mm" value="42.0" valueMax="55.0" valueMin="29.0">2955 mm</quantity>
long, ovate or rhombic, with an attenuate, rounded or subemarginate base, acute or gradually attenuate into short to moderate tip, widthlength index 0.540.71, length of petiolule (12)1728% of length of leaflet; petiolule of lowest leaflets 1(4) mm.
<emphasis id="B9035B42FF82FFD0FADDF983FA3AF9EF" box="[1302,1452,1634,1660]" italics="true" pageId="13" pageNumber="14">Inflorescence</emphasis>
narrow cylindrical, with long appressed and stellate hairs, with rather many declining or curved prickles; peduncles with sharp angle, with up to 4 flowers; pedicels
<quantity id="4C8F2AB5FF82FFD0FD6CF94BFC91F956" box="[679,775,1706,1733]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="13" pageNumber="14" unit="mm" value="3.5" valueMax="6.0" valueMin="1.0">16 mm</quantity>
, short tomentose and with long hairs, with few to numerous prickles and some sessile glands.
<emphasis id="B9035B42FF82FFD0FDD8F92FFDF9F97B" box="[531,623,1742,1768]" italics="true" pageId="13" pageNumber="14">Flowers</emphasis>
: sepals narrow, long pointed, 24 ×
<quantity id="4C8F2AB5FF82FFD0FBE9F92FFB04F97A" box="[1058,1170,1742,1769]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="13" pageNumber="14" unit="mm" value="8.0" valueMax="11.0" valueMin="5.0">511 mm</quantity>
, loosely patent or erect, greyish, tomentose and hairy, sometimes with few prickles and subsessile glands; petals
<quantity id="4C8F2AB5FF82FFD0FBB0F913FB4BF89E" box="[1147,1245,1778,1805]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="14" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">34 mm</quantity>
; anthers glabrous; ovaries with a few hairs; receptacle almost glabrous or with some hairs.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD0FF76F8DBFE55F8EB" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BC88750FF82FFD0FF76F8DBFE55F8EB" blockId="13.[136,1452,1454,2093]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF76F8DBFECFF8C7" bold="true" box="[189,345,1850,1876]" pageId="13" pageNumber="14">Distribution:</emphasis>
—Common on
<collectingCountry id="F360C7C0FF82FFD0FDC2F8DBFD15F8C7" box="[521,643,1850,1876]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="13" pageNumber="14">St. Helena</collectingCountry>
; in
<collectingCountry id="F360C7C0FF82FFD0FD66F8DBFCABF8C7" box="[685,829,1850,1876]" name="South Africa" pageId="13" pageNumber="14">South Africa</collectingCountry>
only some old collections near
<collectingRegion id="49B349B2FF82FFD0FB6CF8DBFB77F8C7" box="[1191,1249,1850,1876]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Cape</collectingRegion>
Town and maybe Swellendam,
<collectingRegion id="49B349B2FF82FFD0FEEBF8BEFE29F8EB" box="[288,447,1886,1913]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Western Cape</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD0FF76F863FB88F79B" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BC88750FF82FFD0FF76F863FAD8F80E" blockId="13.[136,1452,1454,2093]" box="[189,1358,1922,1949]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF76F863FDACF80F" bold="true" box="[189,570,1922,1949]" pageId="13" pageNumber="14">
Specimens:—
<collectingCountry id="F360C7C0FF82FFD0FEAAF863FDA3F80F" box="[353,565,1922,1948]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="13" pageNumber="14">SAINT HELENA</collectingCountry>
.
</emphasis>
<emphasis id="B9035B42FF82FFD0FD8AF862FD4FF80F" box="[577,729,1922,1949]" italics="true" pageId="13" pageNumber="14">Cuming 2453</emphasis>
(BREM_00002220); 1841,
<emphasis id="B9035B42FF82FFD0FBDEF863FB30F80F" box="[1045,1190,1922,1948]" italics="true" pageId="13" pageNumber="14">Hombron s.n</emphasis>
. (P02579475).
</paragraph>
<paragraph id="8BC88750FF82FFD0FF76F846FB88F79B" blockId="13.[136,1452,1454,2093]" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF12F847FDFEF853" bold="true" box="[217,616,1958,1984]" pageId="13" pageNumber="14">
<collectingCountry id="F360C7C0FF82FFD0FF12F847FE27F853" box="[217,433,1958,1984]" name="South Africa" pageId="13" pageNumber="14">SOUTH AFRICA</collectingCountry>
.
<collectingRegion id="49B349B2FF82FFD0FE75F847FDFEF853" box="[446,616,1958,1984]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Western Cape</collectingRegion>
</emphasis>
:
<collectingRegion id="49B349B2FF82FFD0FDB7F847FD2CF853" box="[636,698,1958,1984]" country="South Africa" name="Western Cape" pageId="13" pageNumber="14">Cape</collectingRegion>
pen. July [?] 1924,
<emphasis id="B9035B42FF82FFD0FC62F846FBC6F853" box="[937,1104,1959,1985]" italics="true" pageId="13" pageNumber="14">J.C. Smuts s.n.</emphasis>
(NBG). Inter frutices (altit. I, II) nemorum in “Krakakamma et Adow,” tum ad “Zwartehoogdes” et “Katrivier” (Uytenhage, Albany), s.d.,
<emphasis id="B9035B42FF82FFD0FA95F82BFEA4F79B" italics="true" pageId="13" pageNumber="14">Ecklon &amp; Zeyher 1706</emphasis>
(P02579473) [it is not certain where this specimen was collected].
</paragraph>
</subSubSection>
<subSubSection id="C36DD4DBFF82FFD3FF76F7F3FCF9FACF" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BC88750FF82FFD3FF76F7F3FE44FEDB" blockId="13.[136,1452,1454,2093]" lastBlockId="14.[136,1452,158,1373]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="B9035B42FF82FFD0FF76F7F3FE9FF7BF" bold="true" box="[189,265,2066,2092]" pageId="13" pageNumber="14">Notes:</emphasis>
—The material of
<taxonomicName id="4C77FCD3FF82FFD0FE1CF7F2FDD7F7BF" authorityName="Ecklon &amp; Zeyher" authorityYear="1836" box="[471,577,2067,2092]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF82FFD0FE1CF7F2FDD7F7BF" box="[471,577,2067,2092]" italics="true" pageId="13" pageNumber="14">R. pappei</emphasis>
</taxonomicName>
that Ecklon &amp; Zeyher distributed under the number 1706 is not homogeneous. The specimens in P (P02579473), BREM (BREM-0002214) and the short inflorescences in L (L 0019831), are identical with the
<taxonomicName id="4C77FCD3FF81FFD3FE9CFF22FE41FF4F" box="[343,471,195,220]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF81FFD3FE9CFF22FE41FF4F" box="[343,471,195,220]" italics="true" pageId="14" pageNumber="15">R. pinnatus</emphasis>
</taxonomicName>
in B-W. The long inflorescences in L are
<taxonomicName id="4C77FCD3FF81FFD3FC77FF22FAA3FF4F" authority="Cordemoy (1895: 407)" authorityName="Cordemoy" authorityPageNumber="407" authorityYear="1895" box="[956,1333,194,221]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="glaber">
<emphasis id="B9035B42FF81FFD3FC77FF22FBB0FF4F" box="[956,1062,194,220]" italics="true" pageId="14" pageNumber="15">R. glaber</emphasis>
<bibRefCitation id="EFE6FAA1FF81FFD3FBE5FF23FAA3FF4F" author="Cordemoy, E. J. de" box="[1070,1333,194,221]" pageId="14" pageNumber="15" refId="ref46635" refString="Cordemoy, E. J. de (1895) Flore de l'ile de la Reunion. Klincksieck, Paris, 602 pp. https: // doi. org / 10.5962 / bhl. title. 28055" type="book" year="1895">Cordemoy (1895: 407)</bibRefCitation>
</taxonomicName>
, while the sample in B (with handwritten label by Zeyher,) represents
<taxonomicName id="4C77FCD3FF81FFD3FCE1FF06FC3FFE93" box="[810,937,231,256]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF81FFD3FCE1FF06FC3FFE93" box="[810,937,231,256]" italics="true" pageId="14" pageNumber="15">R. pinnatus</emphasis>
</taxonomicName>
. Because the first
<taxonomicName id="4C77FCD3FF81FFD3FBB2FF07FAA8FE93" box="[1145,1342,230,256]" form="corresponds" pageId="14" pageNumber="15" rank="form">form corresponds</taxonomicName>
best with the protologue (ramis …cano-tomentosis …foliis … subtus breve pilosis rugoso-nervosis), the specimen in BREM was selected as the
<typeStatus id="54CC39F2FF81FFD3FEA8FECFFE5BFEDB" box="[355,461,302,328]" pageId="14" pageNumber="15" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BC88750FF81FFD3FF76FEB3FCA5FE4B" blockId="14.[136,1452,158,1373]" pageId="14" pageNumber="15">
<bibRefCitation id="EFE6FAA1FF81FFD3FF76FEB3FE34FEFE" author="Meisner, C. F." box="[189,418,338,365]" pageId="14" pageNumber="15" pagination="53 - 105" refId="ref48333" refString="Meisner, C. F. (1843) Contributions towards a Flora of South Africa. London Journal of Botany 2: 53 - 105; 527 - 559. [https: // www. biodiversitylibrary. org / item / 6313 # page / 56 / mode / 1 up]" type="journal article" year="1843">Meisner (1843: 527)</bibRefCitation>
published another species from
<collectingCountry id="F360C7C0FF81FFD3FCC2FEB3FC01FEFF" box="[777,919,338,364]" name="South Africa" pageId="14" pageNumber="15">South Africa</collectingCountry>
under the name
<taxonomicName id="4C77FCD3FF81FFD3FB9CFEB3FA3EFEFF" authority="E. Meyer" authorityName="E. Meyer" box="[1111,1448,338,364]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FB9CFEB3FAACFEFF" box="[1111,1338,338,364]" italics="true" pageId="14" pageNumber="15">Rubus heterophyllus</emphasis>
E.Meyer
</taxonomicName>
! in herb. Drège. There is no valid publication of this name by Meyer. The name is also not in
<bibRefCitation id="EFE6FAA1FF81FFD3FB0BFE97FACCFE03" author="Drege, J. F." box="[1216,1370,374,400]" pageId="14" pageNumber="15" refId="ref46761" refString="Drege, J. F. (1839) Catalogus plantarum exsiccatarum Africae Australioris. Ab auctore, Hamburg, 32 pp." type="book" year="1839">Drège (1839)</bibRefCitation>
, nor in
<bibRefCitation id="EFE6FAA1FF81FFD3FF43FE7BFEB4FE27" author="Drege, J. F." box="[136,290,410,436]" pageId="14" pageNumber="15" refId="ref46784" refString="Drege, J. F. (1843) Zwei pflanzengeographische Documente. Besondere Beigabe zur Flora Band II, Regensburg, 230 pp. https: // doi. org / 10.5962 / bhl. title. 87612" type="book" year="1843">Drège (1843)</bibRefCitation>
. Consequently, the comparison with
<taxonomicName id="4C77FCD3FF81FFD3FD75FE7AFCABFE27" box="[702,829,411,436]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF81FFD3FD75FE7AFCABFE27" box="[702,829,411,436]" italics="true" pageId="14" pageNumber="15">R. pinnatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE6FAA1FF81FFD3FCA3FE7BFB83FE27" author="Meisner, C. F." box="[872,1045,410,436]" pageId="14" pageNumber="15" pagination="53 - 105" refId="ref48333" refString="Meisner, C. F. (1843) Contributions towards a Flora of South Africa. London Journal of Botany 2: 53 - 105; 527 - 559. [https: // www. biodiversitylibrary. org / item / 6313 # page / 56 / mode / 1 up]" type="journal article" year="1843">Meisner (1843)</bibRefCitation>
is the validating description and the species must be cited as
<taxonomicName id="4C77FCD3FF81FFD3FE50FE5EFCA5FE4B" authority="E. Mey. ex Meisner." box="[411,819,446,473]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FE50FE5EFDC7FE4B" box="[411,593,446,472]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
E.Mey. ex Meisner.
</taxonomicName>
</paragraph>
<paragraph id="8BC88750FF81FFD3FF76FE02FADBFCF7" blockId="14.[136,1452,158,1373]" pageId="14" pageNumber="15">
In his description,
<bibRefCitation id="EFE6FAA1FF81FFD3FE52FE03FDDFFE6F" author="Meisner, C. F." box="[409,585,482,508]" pageId="14" pageNumber="15" pagination="53 - 105" refId="ref48333" refString="Meisner, C. F. (1843) Contributions towards a Flora of South Africa. London Journal of Botany 2: 53 - 105; 527 - 559. [https: // www. biodiversitylibrary. org / item / 6313 # page / 56 / mode / 1 up]" type="journal article" year="1843">Meisner (1843)</bibRefCitation>
states that
<taxonomicName id="4C77FCD3FF81FFD3FD19FE02FC1DFE6F" box="[722,907,482,508]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FD19FE02FC1DFE6F" box="[722,907,482,508]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C77FCD3FF81FFD3FBE1FE02FB3BFE6F" box="[1066,1197,483,508]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF81FFD3FBE1FE02FB3BFE6F" box="[1066,1197,483,508]" italics="true" pageId="14" pageNumber="15">R. pinnatus</emphasis>
</taxonomicName>
in having the leaves pubescent underneath. There are specimens determined as
<taxonomicName id="4C77FCD3FF81FFD3FCD1FDE6FBB0FDB2" authority="E. Mey." authorityName="E. Mey." box="[794,1062,518,545]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FCD1FDE6FC59FDB3" box="[794,975,518,544]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
E.Mey.
</taxonomicName>
in the Drège collections in P and in K (P02579458, K000312557 R. heterophylla). Both are samples of
<taxonomicName id="4C77FCD3FF81FFD3FC4CFDCAFC7DFDD7" authorityName="Weihe &amp; Nees" authorityYear="1822" box="[903,1003,554,580]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B9035B42FF81FFD3FC4CFDCAFC7DFDD7" box="[903,1003,554,580]" italics="true" pageId="14" pageNumber="15">R. affinis</emphasis>
</taxonomicName>
and have no similarity with
<taxonomicName id="4C77FCD3FF81FFD3FAEDFDCAFA32FDD7" box="[1318,1444,555,580]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF81FFD3FAEDFDCAFA32FDD7" box="[1318,1444,555,580]" italics="true" pageId="14" pageNumber="15">R. pinnatus</emphasis>
</taxonomicName>
. Thus, these cannot be the plant that Meisner had on his desk. Fortunately, there is another reference to
<taxonomicName id="4C77FCD3FF81FFD3FB3DFDAEFA3DFDFB" box="[1270,1451,590,616]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FB3DFDAEFA3DFDFB" box="[1270,1451,590,616]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE6FAA1FF81FFD3FF67FD93FE18FD1F" author="Drege, J. F." box="[172,398,626,652]" pageId="14" pageNumber="15" pagination="599 - 680" refId="ref46878" refString="Drege, J. F. (1847 b) Vergleichungen der von Ecklon und Zeyher und von Drege gesammelten susdafrikanischen Pflanzen (so weit dieselbe noch vorhanden) mit den Exemplaren von Zeyher's neuesten Sammlungen. Linnaea 19: 599 - 680. [https: // www. biodiversitylibrary. org / item / 10863 # page / 602 / mode / 1 up]" type="journal article" year="1847">Drège (1847b: 648)</bibRefCitation>
: E.Z.
<taxonomicName id="4C77FCD3FF81FFD3FE11FD93FD11FD1F" box="[474,647,626,652]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pinnatus">
<emphasis id="B9035B42FF81FFD3FE11FD93FD11FD1F" box="[474,647,626,652]" italics="true" pageId="14" pageNumber="15">Rubus pinnatus</emphasis>
</taxonomicName>
W. = D.
<taxonomicName id="4C77FCD3FF81FFD3FD24FD92FC30FD1F" box="[751,934,626,652]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FD24FD92FC30FD1F" box="[751,934,626,652]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
E.M. a = Z. I, 2452 (70.10). Specimens with the number 2452 are in LY and P. They are inflorescences with pinnate and abaxially hairy leaves, thus corresponding well with the description of
<taxonomicName id="4C77FCD3FF81FFD3FE74FD5AFDE2FD47" box="[447,628,698,724]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FE74FD5AFDE2FD47" box="[447,628,698,724]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE6FAA1FF81FFD3FD56FD5BFCDCFD47" author="Meisner, C. F." box="[669,842,698,724]" pageId="14" pageNumber="15" pagination="53 - 105" refId="ref48333" refString="Meisner, C. F. (1843) Contributions towards a Flora of South Africa. London Journal of Botany 2: 53 - 105; 527 - 559. [https: // www. biodiversitylibrary. org / item / 6313 # page / 56 / mode / 1 up]" type="journal article" year="1843">Meisner (1843)</bibRefCitation>
and can be considered as the taxon, which he meant. It was collected by Zeyher near Swellendam in October. The year is not mentioned (as is usual with Zeyhers collections), but this must be 1831 when Zeyher collected at that location in October. Another sample with the same number 2452 is also conserved in P (P02579478). It does not have an original label, but only a printed tag with the name Coll. De Zeyher 1847. This is obviously not the collection date, but the date that Drège revisited Zeyhers herbarium.
</paragraph>
<paragraph id="8BC88750FF81FFD3FF76FC8FFC5FFC3E" blockId="14.[136,1452,158,1373]" pageId="14" pageNumber="15">
<bibRefCitation id="EFE6FAA1FF81FFD3FF76FC8FFE36FC1B" author="Drege, J. F." box="[189,416,878,904]" pageId="14" pageNumber="15" pagination="599 - 680" refId="ref46878" refString="Drege, J. F. (1847 b) Vergleichungen der von Ecklon und Zeyher und von Drege gesammelten susdafrikanischen Pflanzen (so weit dieselbe noch vorhanden) mit den Exemplaren von Zeyher's neuesten Sammlungen. Linnaea 19: 599 - 680. [https: // www. biodiversitylibrary. org / item / 10863 # page / 602 / mode / 1 up]" type="journal article" year="1847">Drège (1847b: 648)</bibRefCitation>
continues his comments on
<taxonomicName id="4C77FCD3FF81FFD3FD20FC8EFC32FC1B" box="[747,932,878,904]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FD20FC8EFC32FC1B" box="[747,932,878,904]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
with: Hierzu auch E.Z.
<taxonomicName id="4C77FCD3FF81FFD3FB03FC8EFAA0FC1B" authorityName="Ecklon &amp; Zeyher" authorityYear="1836" box="[1224,1334,879,904]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF81FFD3FB03FC8EFAA0FC1B" box="[1224,1334,879,904]" italics="true" pageId="14" pageNumber="15">R. pappei</emphasis>
</taxonomicName>
E.Z. =
<taxonomicName id="4C77FCD3FF81FFD3FA58FC8EFE89FC3F" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FA58FC8EFE89FC3F" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
E.M. b. Thus, he identifies
<taxonomicName id="4C77FCD3FF81FFD3FDAAFC72FC81FC3F" box="[609,791,914,940]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FDAAFC72FC81FC3F" box="[609,791,914,940]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
with
<taxonomicName id="4C77FCD3FF81FFD3FC9CFC72FC54FC3F" authorityName="Ecklon &amp; Zeyher" authorityYear="1836" box="[855,962,915,940]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF81FFD3FC9CFC72FC54FC3F" box="[855,962,915,940]" italics="true" pageId="14" pageNumber="15">R. pappei</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC88750FF81FFD3FF76FC56FAC8FB8B" blockId="14.[136,1452,158,1373]" pageId="14" pageNumber="15">
Because the protologue of
<taxonomicName id="4C77FCD3FF81FFD3FE2DFC56FD0DFC43" box="[486,667,950,976]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FE2DFC56FD0DFC43" box="[486,667,950,976]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
is earlier than Drèges comments, the type of
<taxonomicName id="4C77FCD3FF81FFD3FB53FC56FA94FC43" authorityName="Ecklon &amp; Zeyher" authorityYear="1836" box="[1176,1282,951,976]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF81FFD3FB53FC56FA94FC43" box="[1176,1282,951,976]" italics="true" pageId="14" pageNumber="15">R. pappei</emphasis>
</taxonomicName>
is not included in it. Consequently, both names are heterotypic and a separate type of
<taxonomicName id="4C77FCD3FF81FFD3FC5BFC3AFBD3FC67" box="[912,1093,986,1012]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FC5BFC3AFBD3FC67" box="[912,1093,986,1012]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
must be selected. The specimen of
<emphasis id="B9035B42FF81FFD3FF6DFC1FFEA5FB8B" box="[166,307,1022,1048]" italics="true" pageId="14" pageNumber="15">
Zeyher
<quantity id="4C8F2AB5FF81FFD3FF30FC1FFEA5FB8B" box="[251,307,1022,1048]" metricMagnitude="1" metricUnit="m" metricValue="6.22808" pageId="14" pageNumber="15" unit="in" value="2452.0">2452</quantity>
</emphasis>
in LY with the original label was chosen for this, while the other specimens are
<typeStatus id="54CC39F2FF81FFD3FB7DFC1EFA8DFB8A" box="[1206,1307,1023,1049]" pageId="14" pageNumber="15" type="syntype">syntypes</typeStatus>
, then.
</paragraph>
<paragraph id="8BC88750FF81FFD3FF76FBC3FDF0FB17" blockId="14.[136,1452,158,1373]" pageId="14" pageNumber="15">
<taxonomicName id="4C77FCD3FF81FFD3FF76FBC3FD1BFBAF" ID-CoL="4TKD8" authority="E. Mey. ex Meisner" authorityName="Meisner" box="[189,653,1058,1085]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FF76FBC3FE33FBAF" box="[189,421,1058,1084]" italics="true" pageId="14" pageNumber="15">Rubus heterophyllus</emphasis>
E.Mey. ex Meisner
</taxonomicName>
is a later homonym of
<taxonomicName id="4C77FCD3FF81FFD3FC6CFBC2FAF8FBAF" ID-CoL="4TKD8" authority="Willdenow [1811: 413" authorityName="Willdenow" authorityYear="1811" box="[935,1390,1058,1084]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FC6CFBC2FBF7FBAF" box="[935,1121,1058,1084]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
<bibRefCitation id="EFE6FAA1FF81FFD3FBA0FBC3FAF8FBAF" author="Willdenow, C. L." box="[1131,1390,1058,1084]" pageId="14" pageNumber="15" refId="ref50580" refString="Willdenow, C. L. (1811) Die wilde Baumzucht oder Beschreibung aller im K ˆ nigl. Botan Garten etc. bey Berlin im Freien ausdauernden Baume und Straucher. Nauck, Berlin, 204 pp." type="book" year="1811">Willdenow [1811: 413</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C77FCD3FF81FFD3FA58FBC2FD8EFBF3" ID-CoL="87C95" authority="Blanchard (1906: 149)" authorityName="Blanchard" authorityPageNumber="149" authorityYear="1906" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="plicatifolius">
<emphasis id="B9035B42FF81FFD3FA58FBC2FE98FBF3" italics="true" pageId="14" pageNumber="15">R. plicatifolius</emphasis>
<bibRefCitation id="EFE6FAA1FF81FFD3FEDDFBA7FD8EFBF3" author="Blanchard, W. H." box="[278,536,1094,1120]" pageId="14" pageNumber="15" pagination="146 - 157" refId="ref46352" refString="Blanchard, W. H. (1906) Some Maine Rubi. The blackberries of the Kennebunks and Wells 1. Rhodora 8: 146 - 157. [https: // www. jstor. org / stable / 23293880]" type="journal article" year="1906">Blanchard (1906: 149)</bibRefCitation>
</taxonomicName>
, see
<bibRefCitation id="EFE6FAA1FF81FFD3FD84FBA7FC06FBF3" author="Beek, A. van de &amp; Widrlechner, M." box="[591,912,1094,1120]" pageId="14" pageNumber="15" pagination="67 - 98" refId="ref46292" refString="Beek, A. van de &amp; Widrlechner, M. (2021) North American species of Rubus L. (Rosaceae) described from European botanical gardens (1789 - 1823). Adansonia 43 (8): 67 - 98. https: // doi. org / 10.5252 / adansonia 2021 v 43 a 8" type="journal article" year="2021">Beek &amp; Widrlechner (2021)</bibRefCitation>
], of
<taxonomicName id="4C77FCD3FF81FFD3FC08FBA6FAAEFBF3" ID-CoL="4TKD8" authority="Host (1831: 32)" authorityName="Host" authorityPageNumber="32" authorityYear="1831" box="[963,1336,1094,1121]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FC08FBA6FBECFBF3" box="[963,1146,1094,1120]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
<bibRefCitation id="EFE6FAA1FF81FFD3FB49FBA6FAAEFBF3" author="Host, N. T." box="[1154,1336,1094,1121]" pageId="14" pageNumber="15" refId="ref47966" refString="Host, N. T. (1831) Flora Austriaca, Vol. 2. C. F. Beck, Viennae, 814 pp." type="book" year="1831">Host (1831: 32)</bibRefCitation>
</taxonomicName>
, and of
<taxonomicName id="4C77FCD3FF81FFD3FA58FBA6FDF4FB17" ID-CoL="4TKD8" authority="Hegetschweiler (1839: 486)" authorityName="Hegetschweiler" authorityPageNumber="486" authorityYear="1839" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FA58FBA6FE89FB17" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
<bibRefCitation id="EFE6FAA1FF81FFD3FEEDFB8BFDF4FB17" author="Hegetschweiler, J." box="[294,610,1130,1156]" pageId="14" pageNumber="15" refId="ref47792" refString="Hegetschweiler, J. (1839 [' 1840 ']) Flora der Schweiz. F. Schulthess, Zurich, xxviii + 1135 pp. https: // doi. org / 10.5962 / bhl. title. 6350" type="book" year="1839">Hegetschweiler (1839: 486)</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC88750FF81FFD3FF76FB6FFCF9FACF" blockId="14.[136,1452,158,1373]" pageId="14" pageNumber="15">
<bibRefCitation id="EFE6FAA1FF81FFD3FF76FB6FFE35FB3A" author="Gandoger, M." box="[189,419,1166,1193]" pageId="14" pageNumber="15" pagination="24 - 69" refId="ref47605" refString="Gandoger, M. (1918). Sertum plantarum novarum. Pars prima. Bulletin de la Societe Botanique de France 65: 24 - 69. https: // doi. org / 10.1080 / 00378941.1918.10836046" type="journal article" year="1918">Gandoger (1918: 25)</bibRefCitation>
published the specimen
<emphasis id="B9035B42FF81FFD3FD65FB6FFCAFFB3B" box="[686,825,1166,1192]" italics="true" pageId="14" pageNumber="15">Zeyher 2452</emphasis>
once again under the name
<taxonomicName id="4C77FCD3FF81FFD3FBAFFB6EFB44FB3B" authorityName="Gandoger" authorityYear="1918" box="[1124,1234,1166,1192]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="B9035B42FF81FFD3FBAFFB6EFB44FB3B" box="[1124,1234,1166,1192]" italics="true" pageId="14" pageNumber="15">R. zeyheri</emphasis>
</taxonomicName>
. This is a legitimate name for the taxon. That makes the remark by
<bibRefCitation id="EFE6FAA1FF81FFD3FD43FB53FCBDFB5F" author="Drege, J. F." box="[648,811,1202,1228]" pageId="14" pageNumber="15" pagination="599 - 680" refId="ref46878" refString="Drege, J. F. (1847 b) Vergleichungen der von Ecklon und Zeyher und von Drege gesammelten susdafrikanischen Pflanzen (so weit dieselbe noch vorhanden) mit den Exemplaren von Zeyher's neuesten Sammlungen. Linnaea 19: 599 - 680. [https: // www. biodiversitylibrary. org / item / 10863 # page / 602 / mode / 1 up]" type="journal article" year="1847">Drège (1847b)</bibRefCitation>
that
<taxonomicName id="4C77FCD3FF81FFD3FCABFB52FB83FB5F" authorityName="Hegetschweiler" authorityYear="1839" box="[864,1045,1202,1228]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FCABFB52FB83FB5F" box="[864,1045,1202,1228]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
is identical with
<taxonomicName id="4C77FCD3FF81FFD3FB1AFB52FAADFB5F" authorityName="Ecklon &amp; Zeyher" authorityYear="1836" box="[1233,1339,1203,1228]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF81FFD3FB1AFB52FAADFB5F" box="[1233,1339,1203,1228]" italics="true" pageId="14" pageNumber="15">R. pappei</emphasis>
</taxonomicName>
of special interest, for the question is now whether
<taxonomicName id="4C77FCD3FF81FFD3FD9DFB36FD51FB63" authorityName="Gandoger" authorityYear="1918" box="[598,711,1238,1264]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="zeyheri">
<emphasis id="B9035B42FF81FFD3FD9DFB36FD51FB63" box="[598,711,1238,1264]" italics="true" pageId="14" pageNumber="15">R. zeyheri</emphasis>
</taxonomicName>
, which is identical with
<taxonomicName id="4C77FCD3FF81FFD3FC16FB36FB02FB63" authorityName="Hegetschweiler" authorityYear="1839" box="[989,1172,1238,1264]" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="heterophyllus">
<emphasis id="B9035B42FF81FFD3FC16FB36FB02FB63" box="[989,1172,1238,1264]" italics="true" pageId="14" pageNumber="15">R. heterophyllus</emphasis>
</taxonomicName>
, is also identical with
<taxonomicName id="4C77FCD3FF81FFD3FA58FB36FF42FA87" authorityName="Ecklon &amp; Zeyher" authorityYear="1836" class="Magnoliopsida" family="Rosaceae" genus="Rubus" kingdom="Plantae" order="Rosales" pageId="14" pageNumber="15" phylum="Tracheophyta" rank="species" species="pappei">
<emphasis id="B9035B42FF81FFD3FA58FB36FF42FA87" italics="true" pageId="14" pageNumber="15">R. pappei</emphasis>
</taxonomicName>
or not. Both have abaxially hairy leaves. However, the prickles are less numerous and the leaves weaker and less hairy, but this may be due to a shady location. As long as no other material is found, the decision must be left open. Maybe field research around Swellendam might give more clarity.
</paragraph>
</subSubSection>
</treatment>
</document>