treatments-xml/data/03/DA/9D/03DA9D6FFF9AFFFF92EBD6FEFA82FC6E.xml
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<mods:title id="8B35527E2CCDB5856BC37F13969F53E5">The Larvae of European Ascalaphidae (Neuroptera)</mods:title>
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<mods:namePart id="A523B03A443123821220269556C09811">Badano, Davide</mods:namePart>
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<treatment id="03DA9D6FFF9AFFFF92EBD6FEFA82FC6E" ID-DOI="http://doi.org/10.5281/zenodo.4915189" ID-GBIF-Taxon="183403925" ID-Zenodo-Dep="4915189" LSID="urn:lsid:plazi:treatment:03DA9D6FFF9AFFFF92EBD6FEFA82FC6E" httpUri="http://treatment.plazi.org/id/03DA9D6FFF9AFFFF92EBD6FEFA82FC6E" lastPageId="10" lastPageNumber="297" pageId="8" pageNumber="295">
<subSubSection id="C3697FF2FF9AFFFD92EBD6FEFDCEF92A" box="[151,527,1737,1763]" pageId="8" pageNumber="295" type="nomenclature">
<paragraph id="8BCC2C79FF9AFFFD92EBD6FEFDCEF92A" blockId="8.[151,527,1737,1797]" box="[151,527,1737,1763]" pageId="8" pageNumber="295">
<heading id="D0849B15FF9AFFFD92EBD6FEFDCEF92A" bold="true" box="[151,527,1737,1763]" fontSize="11" level="1" pageId="8" pageNumber="295" reason="1">
<taxonomicName id="4C7357FAFF9AFFFD92EBD6FEFDCEF92A" ID-CoL="78Q35" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[151,527,1737,1763]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="295" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF9AFFFD92EBD6FEFDCEF92A" bold="true" box="[151,527,1737,1763]" pageId="8" pageNumber="295">
<emphasis id="B907F06BFF9AFFFD92EBD6FEFE97F92A" bold="true" box="[151,342,1737,1763]" italics="true" pageId="8" pageNumber="295">Puer maculatus</emphasis>
(Olivier, 1789)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3697FF2FF9AFFFD92EBD6DBFF1EF85C" pageId="8" pageNumber="295" type="description">
<paragraph id="8BCC2C79FF9AFFFD92EBD6DBFE43F8CC" blockId="8.[151,527,1737,1797]" box="[151,386,1772,1797]" pageId="8" pageNumber="295">
(
<figureCitation id="134830FCFF9AFFFD92E3D6DBFEC1F8CC" box="[159,256,1772,1797]" captionStart="FIGURE 1" captionStartId="2.[151,249,1953,1975]" captionTargetBox="[260,1345,203,1915]" captionTargetId="figure-58@2.[232,1354,193,1931]" captionTargetPageId="2" captionText="FIGURE 1. Taxonomic characters of Ascalaphidae larvae. A, Deleproctophylla australis dorsal and ventral view; B, head of Ascalaphus festivus; C, head of Puer maculatus; D, VIII and IX sternites of Libelloides siculus." figureDoi="http://doi.org/10.5281/zenodo.4915175" httpUri="https://zenodo.org/record/4915175/files/figure.png" pageId="8" pageNumber="295">Figs. 1C</figureCitation>
,
<figureCitation id="134830FCFF9AFFFD9371D6DBFEF1F8CC" box="[269,304,1772,1797]" captionStart="FIGURE 2" captionStartId="5.[151,249,1866,1888]" captionTargetBox="[214,1374,688,1811]" captionTargetId="figure-236@5.[204,1382,673,1817]" captionTargetPageId="5" captionText="FIGURE 2. VIII and IX abdominal sternites of 3rd instar larva of Ascalaphidae. A, Ascalaphus festivus; B, Bubopsis agrionoides; C, Puer maculatus; D, Deleproctophylla australis; E, Libelloides macaronius. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4915181" httpUri="https://zenodo.org/record/4915181/files/figure.png" pageId="8" pageNumber="295">2C</figureCitation>
,
<figureCitation id="134830FCFF9AFFFD9340D6DBFE9FF8CC" box="[316,350,1772,1797]" captionStart="FIGURE 3" captionStartId="6.[151,249,1300,1322]" captionTargetBox="[194,1362,201,1269]" captionTargetId="figure-247@6.[184,1403,193,1276]" captionTargetPageId="6" captionText="FIGURE 3. Chaetotaxy of the VIII abdominal sternite of 3rd instar larva of Ascalaphidae. A, Ascalaphus festivus; B, Bubopsis agrionoides; C, Puer maculatus; D, Deleproctophylla australis; E, Libelloides longicornis. Scale bar: 100 µm." figureDoi="http://doi.org/10.5281/zenodo.4915185" httpUri="https://zenodo.org/record/4915185/files/figure.png" pageId="8" pageNumber="295">3C</figureCitation>
,
<figureCitation id="134830FCFF9AFFFD9316D6DBFEB9F8CC" box="[362,376,1772,1797]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="8" pageNumber="295">5</figureCitation>
)
</paragraph>
<paragraph id="8BCC2C79FF9AFFFD92EBD703FF1EF85C" blockId="8.[151,1436,1844,2013]" pageId="8" pageNumber="295">
Description of
<bibRefCitation id="EFE25188FF9AFFFD9334D703FE26F884" author="Hagen, H. A." box="[328,487,1844,1869]" pageId="8" pageNumber="295" pagination="33 - 62" refId="ref16310" refString="Hagen, H. A. (1873) Die Larven von Ascalaphus. Stettiner Entomologische Zeitung, 34, 33 - 62." type="journal article" year="1873">Hagen (1873)</bibRefCitation>
effectively refers to this peculiar and rare species, despite his identification was mainly based on geographical assumptions thus it was considered doubtful by later authors (H.
<bibRefCitation id="EFE25188FF9AFFFD9683D76FFF12F85C" author="Aspock, H. &amp; Aspock, U. &amp; Holzel, H." pageId="8" pageNumber="295" refId="ref15298" refString="Aspock, H., Aspock, U. &amp; Holzel, H. (1980) Die Neuropteren Europas. Eine zusammenfassende Darstellung der Systematik, Okologie und Chorologie der Neuropteroidea (Megaloptera, Raphidioptera, Planipennia) Europas. Goecke and Evers, Krefeld, Germany, 495 + 355 pp." type="book" year="1980">
Aspöck
<emphasis id="B907F06BFF9AFFFD971CD76EFA5DF8B8" box="[1376,1436,1880,1905]" italics="true" pageId="8" pageNumber="295">et al.</emphasis>
1980
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3697FF2FF9AFFFD92BBD797FD6AF814" pageId="8" pageNumber="295" type="materials_examined">
<paragraph id="8BCC2C79FF9AFFFD92BBD797FD6AF814" blockId="8.[151,1436,1844,2013]" pageId="8" pageNumber="295">
Examined specimens.
<materialsCitation id="3B1B2624FF9AFFFD93AAD797FD69F814" ID-GBIF-Occurrence="3319797311" collectingDate="2011-06" collectorName="B. Michel &amp; D. Badano" country="France" county="Les Hauts de Boscorre" location="Herault" municipality="Argelliers" pageId="8" pageNumber="295" specimenCount="1">
<emphasis id="B907F06BFF9AFFFD93AAD797FDF0F870" bold="true" box="[470,561,1952,1977]" pageId="8" pageNumber="295">
<collectingCountry id="F3646CE9FF9AFFFD93AAD797FDEDF870" box="[470,556,1952,1977]" name="France" pageId="8" pageNumber="295">France</collectingCountry>
.
</emphasis>
<location id="8EAC7AA2FF9AFFFD9040D797FD52F870" LSID="urn:lsid:plazi:treatment:03DA9D6FFF9AFFFF92EBD6FEFA82FC6E:8EAC7AA2FF9AFFFD9040D797FD52F870" box="[572,659,1952,1977]" country="France" county="Les Hauts de Boscorre" municipality="Argelliers" name="Herault" pageId="8" pageNumber="295">Hérault</location>
,
<collectingMunicipality id="6BA8B603FF9AFFFD90DED797FCD5F870" box="[674,788,1952,1977]" pageId="8" pageNumber="295">Argelliers</collectingMunicipality>
,
<collectingCounty id="62AD54F5FF9AFFFD915FD796FBEEF871" box="[803,1071,1952,1977]" pageId="8" pageNumber="295">Les Hauts de Boscorre</collectingCounty>
, holm oak wood on calcareous rocks,
<date id="FFCD0AB9FF9AFFFD929DD7F3FEFCF814" box="[225,317,1988,2013]" pageId="8" pageNumber="295" value="2011-06">
<collectingDate id="EF89F351FF9AFFFD929DD7F3FEFCF814" box="[225,317,1988,2013]" pageId="8" pageNumber="295" value="2011-06">VI.2011</collectingDate>
</date>
(
<collectorName id="268649AFFF9AFFFD9331D7F3FE7CF814" box="[333,445,1988,2013]" pageId="8" pageNumber="295">B. Michel</collectorName>
&amp;
<collectorName id="268649AFFF9AFFFD939DD7F3FD9CF814" box="[481,605,1988,2013]" pageId="8" pageNumber="295">D. Badano</collectorName>
), 2 L3
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0C7CF1FF9BFFFC92EBD792FCACF813" ID-DOI="http://doi.org/10.5281/zenodo.4915191" ID-Zenodo-Dep="4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="9" pageNumber="296" startId="9.[151,249,1957,1979]" targetBox="[254,1327,215,1914]" targetPageId="9">
<paragraph id="8BCC2C79FF9BFFFC92EBD792FCACF813" blockId="9.[151,1435,1954,2010]" pageId="9" pageNumber="296">
<emphasis id="B907F06BFF9BFFFC92EBD792FED3F873" bold="true" box="[151,274,1957,1979]" pageId="9" pageNumber="296">FIGURE 5.</emphasis>
<taxonomicName id="4C7357FAFF9BFFFC9365D791FD99F872" authority="(Olivier, 1789)" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[281,600,1957,1979]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="296" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF9BFFFC9365D791FE78F872" box="[281,441,1957,1979]" italics="true" pageId="9" pageNumber="296">Puer maculatus</emphasis>
(Olivier, 1789)
</taxonomicName>
, 3
<superScript id="7C068131FF9BFFFC900CD795FDBDF866" attach="left" box="[624,636,1954,1967]" fontSize="5" pageId="9" pageNumber="296">rd</superScript>
instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description.
</paragraph>
</caption>
<subSubSection id="C3697FF2FF98FFFF92BBD0ADFAAAFCF2" pageId="10" pageNumber="297" type="description">
<paragraph id="8BCC2C79FF98FFFF92BBD0ADFE09FDAA" blockId="10.[151,1437,151,935]" pageId="10" pageNumber="297">
<emphasis id="B907F06BFF98FFFF92BBD0ADFDF2FF7A" bold="true" box="[199,563,151,179]" pageId="10" pageNumber="297">
Description of 3
<superScript id="7C068131FF98FFFF93F4D0A0FE56FF6C" attach="left" box="[392,407,151,165]" fontSize="6" pageId="10" pageNumber="297">rd</superScript>
instar larva.
</emphasis>
Size (based on
<specimenCount id="9D75E7F0FF98FFFF9097D0ADFCBAFF7A" box="[747,891,154,179]" pageId="10" pageNumber="297" type="generic">2 specimens</specimenCount>
): BL
<quantity id="4C8B819CFF98FFFF91C0D0ADFBF2FF7B" box="[956,1075,154,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.043" pageId="10" pageNumber="297" unit="mm" value="10.43">10.43 mm</quantity>
; HL
<quantity id="4C8B819CFF98FFFF9612D0ADFB13FF7B" box="[1134,1234,154,179]" metricMagnitude="-3" metricUnit="m" metricValue="2.21" pageId="10" pageNumber="297" unit="mm" value="2.21">2.21 mm</quantity>
(2.122.30), HW
<quantity id="4C8B819CFF98FFFF92EBD089FF3CFF1F" box="[151,253,190,215]" metricMagnitude="-3" metricUnit="m" metricValue="3.24" pageId="10" pageNumber="297" unit="mm" value="3.24">3.24 mm</quantity>
(3.233.25), ML
<quantity id="4C8B819CFF98FFFF93B7D089FDF0FF1F" box="[459,561,190,215]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="10" pageNumber="297" unit="mm" value="2.6">2.60 mm</quantity>
(2.592.62), HW/HL 1.47, ML/HL 1.17. General colouring ochre with grey markings and spots, ventral side noticeably paler, mottled with grey; dorsal side of the head capsule light brown, ventral side of the head light brown with a dark area on the mouthparts, mandibles brown; legs ochre; setae black, most dolichasters grey. Head rectangular, dorsal side with a narrow posterior depression; long setae are present at the base of the forward-placed ocular tubercles (
<figureCitation id="134830FCFF98FFFF90C8D179FCC6FEAE" box="[692,775,334,359]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="10" pageNumber="297">Fig. 5b</figureCitation>
); mandibles narrow, close at the base (
<figureCitation id="134830FCFF98FFFF96CBD179FAC9FEAE" box="[1207,1288,334,359]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="10" pageNumber="297">Fig. 5c</figureCitation>
), longer than the head capsule (
<figureCitation id="134830FCFF98FFFF931DD145FE75FE42" box="[353,436,370,395]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="10" pageNumber="297">Fig. 5a</figureCitation>
); basal tooth equidistant between the base of the mandible and median tooth; interdental pseudo-teeth (43)(43)(0); thin and hair-like dolichasters are present at the base of the mandible; external margin of mandible equipped with short setae, longer toward the base. Pronotum covered with short dolichasters; mesothoracic spiracles black; anterior pair of mesothoracic scoli-like processes very large, bent at 90° (
<figureCitation id="134830FCFF98FFFF9745D1E9FA4EFE3E" box="[1337,1423,478,503]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="10" pageNumber="297">Fig. 5d</figureCitation>
). First pair of abdominal spiracles dorsally disposed, black in colour (
<figureCitation id="134830FCFF98FFFF91DAD235FC38FDD2" box="[934,1017,514,539]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="10" pageNumber="297">Fig. 5e</figureCitation>
); the following seven pairs disposed ventrally; dorsal series of abdominal scolus-like processes positioned on the lower sides of the body, in contact with the substrate (
<figureCitation id="134830FCFF98FFFF9310D27DFE7AFDAA" box="[364,443,586,611]" captionStart="FIGURE 5" captionStartId="9.[151,249,1957,1979]" captionTargetBox="[254,1327,215,1914]" captionTargetId="figure-49@9.[240,1347,193,1933]" captionTargetPageId="9" captionText="FIGURE 5. Puer maculatus (Olivier, 1789), 3rd instar larva (France: Hérault, Argelliers). Dorsal (above), ventral (middle) and lateral (below) view; af: diagnostic characters, see species description." figureDoi="http://doi.org/10.5281/zenodo.4915191" httpUri="https://zenodo.org/record/4915191/files/figure.png" pageId="10" pageNumber="297">Fig. 5f</figureCitation>
).
</paragraph>
<paragraph id="8BCC2C79FF98FFFF92BBD259FAAAFCF2" blockId="10.[151,1437,151,935]" pageId="10" pageNumber="297">
<emphasis id="B907F06BFF98FFFF92BBD259FE96FD4E" bold="true" box="[199,343,622,647]" pageId="10" pageNumber="297">Bio-ecology.</emphasis>
<taxonomicName id="4C7357FAFF98FFFF931DD247FE32FD4E" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[353,499,622,647]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="297" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF98FFFF931DD247FE32FD4E" box="[353,499,622,647]" italics="true" pageId="10" pageNumber="297">P. maculatus</emphasis>
</taxonomicName>
is associated with open Mediterranean biotopes with isolated trees or scrubs on extensive barren rocky areas. The investigated site is characterized by thick hedges of holm oaks interspersed with wide surfaces of cracked limestone. The larvae live adhering to the inferior surface of flat stones, normally disposed under larger rocks. The cocoon is spun and fixed in the same condition. The larva is extremely motionless and it is camouflaged by the fine rock dust retained by the dolichasters. The larval and pupal stages were both collected during the first decade of June, just before the known flight period in the area (
<bibRefCitation id="EFE25188FF98FFFF9608D315FA9EFCF2" author="Michel, B. &amp; Kral, K." box="[1140,1375,802,827]" pageId="10" pageNumber="297" pagination="127 - 134" refId="ref16761" refString="Michel, B. &amp; Kral, K. (2008) Ecology and eye morphology in Bubopsis agrionoides, Puer maculatus and Deleproctophylla dusmeti (Neuroptera, Ascalaphidae). Annales, Anali za Istrske in Mediteranske Studije, Series Historia Naturalis, 18, 127 - 134." type="journal article" year="2008">Michel &amp; Kral 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3697FF2FF98FFFF92BBD371FB33FC96" box="[199,1266,838,863]" pageId="10" pageNumber="297" type="distribution">
<paragraph id="8BCC2C79FF98FFFF92BBD371FB33FC96" blockId="10.[151,1437,151,935]" box="[199,1266,838,863]" pageId="10" pageNumber="297">
<emphasis id="B907F06BFF98FFFF92BBD371FEA1FC96" bold="true" box="[199,352,838,863]" inLineHeading="true" pageId="10" pageNumber="297" reason="1">Distribution.</emphasis>
<taxonomicName id="4C7357FAFF98FFFF931BD37FFE37FC96" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[359,502,838,863]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="297" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF98FFFF931BD37FFE37FC96" box="[359,502,838,863]" italics="true" pageId="10" pageNumber="297">P. maculatus</emphasis>
</taxonomicName>
has been reported for southern
<collectingCountry id="F3646CE9FF98FFFF9124D370FC66FC97" box="[856,935,839,862]" name="France" pageId="10" pageNumber="297">France</collectingCountry>
, Iberian Peninsula and
<collectingCountry id="F3646CE9FF98FFFF96D0D371FB2FFC96" box="[1196,1262,838,863]" name="Israel" pageId="10" pageNumber="297">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3697FF2FF98FFFF92BBD35DFA82FC6E" pageId="10" pageNumber="297" type="discussion">
<paragraph id="8BCC2C79FF98FFFF92BBD35DFA82FC6E" blockId="10.[151,1437,151,935]" pageId="10" pageNumber="297">
<emphasis id="B907F06BFF98FFFF92BBD35DFEFAFC4A" bold="true" box="[199,315,874,899]" inLineHeading="true" pageId="10" pageNumber="297" reason="1">Remarks.</emphasis>
The larva of
<taxonomicName id="4C7357FAFF98FFFF93B3D35CFD9CFC4A" baseAuthorityName="Olivier" baseAuthorityYear="1790" box="[463,605,874,899]" class="Insecta" family="Ascalaphidae" genus="Puer" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="297" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B907F06BFF98FFFF93B3D35CFD9CFC4A" box="[463,605,874,899]" italics="true" pageId="10" pageNumber="297">P. maculatus</emphasis>
</taxonomicName>
, the only member of the genus whose larva is known, is remarkable due to the flattened habitus and peculiar morphology, noticeably differing from the other European
<taxonomicName id="4C7357FAFF98FFFF96D8D3B9FAFEFC6E" box="[1188,1343,910,935]" class="Insecta" family="Ascalaphidae" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="297" phylum="Arthropoda" rank="family">Ascalaphidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>