1189 lines
119 KiB
XML
1189 lines
119 KiB
XML
<document ID-DOI="10.11646/zootaxa.5263.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7800918" ID-ZooBank="9EF23FE9-DDD8-46D4-A275-09A35243BF30" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.tables_approvedBy="diego" IM.treatments_approvedBy="diego" approvalRequired="3" approvalRequired_for_textStreams="3" checkinTime="1680677550347" checkinUser="plazi" docAuthor="Richards, Stephen J., Donnellan, Stephen C. & Oliver, Paul M." docDate="2023" docId="03D60841A03FFFF037CFF94CC0FBFE99" docLanguage="en" docName="zootaxa.5263.2.1.pdf" docOrigin="Zootaxa 5263 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Litoria lisae Richards & Donnellan & Oliver 2023, sp. nov." docType="treatment" docVersion="3" lastPageNumber="175" masterDocId="FFEF7039A02CFFE93758FFA7C117FF84" masterDocTitle="Five new species of the pelodryadid genus Litoria Tschudi from the southern versant of Papua New Guinea’s Central Cordillera, with observations on the diversification of reproductive strategies in Melanesian treefrogs" masterLastPageNumber="190" masterPageNumber="151" pageNumber="170" updateTime="1681154595770" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Five new species of the pelodryadid genus Litoria Tschudi from the southern versant of Papua New Guinea’s Central Cordillera, with observations on the diversification of reproductive strategies in Melanesian treefrogs</mods:title>
|
||
</mods:titleInfo>
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<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Richards, Stephen J.</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-0251-3884</mods:nameIdentifier>
|
||
<mods:affiliation>Herpetology Department, South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia.</mods:affiliation>
|
||
<mods:nameIdentifier type="email">steve.richards@samuseum.sa.gov.au</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Donnellan, Stephen C.</mods:namePart>
|
||
<mods:affiliation>Herpetology Department, South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Oliver, Paul M.</mods:namePart>
|
||
<mods:affiliation>Centre for Planetary Health and Food Security, Griffith University, 170 Kessels Rd, Brisbane, Queensland 4121, and Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, Queensland, 4101 Australia.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-04-05</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5263</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>151</mods:start>
|
||
<mods:end>190</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
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||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.2.1</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5263.2.1</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7800918</mods:identifier>
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<mods:identifier type="ZooBank">9EF23FE9-DDD8-46D4-A275-09A35243BF30</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:03D60841A03FFFF037CFF94CC0FBFE99" httpUri="http://treatment.plazi.org/id/03D60841A03FFFF037CFF94CC0FBFE99" lastPageId="25" lastPageNumber="175" pageId="19" pageNumber="170">
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<subSubSection pageId="19" pageNumber="170" type="nomenclature">
|
||
<paragraph blockId="19.[151,1211,1770,1905]" box="[151,405,1770,1797]" pageId="19" pageNumber="170">
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||
<heading bold="true" box="[151,405,1770,1797]" fontSize="11" level="1" pageId="19" pageNumber="170" reason="1">
|
||
<emphasis bold="true" box="[151,405,1770,1797]" pageId="19" pageNumber="170">
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[151,297,1770,1797]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="19" pageNumber="170" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis bold="true" box="[151,297,1770,1797]" italics="true" pageId="19" pageNumber="170">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[312,405,1771,1797]" pageId="19" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1211,1770,1905]" box="[151,322,1807,1833]" pageId="19" pageNumber="170">Lisa’s Treefrog</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="170" type="description">
|
||
<paragraph blockId="19.[151,1211,1770,1905]" box="[151,275,1843,1869]" pageId="19" pageNumber="170">
|
||
<figureCitation box="[151,275,1843,1869]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStart-2="FIGURE 13" captionStartId-0="20.[151,250,1619,1643]" captionStartId-1="22.[151,250,1829,1853]" captionStartId-2="23.[151,250,1280,1304]" captionTargetBox-0="[176,1412,453,1594]" captionTargetBox-1="[160,1427,345,1806]" captionTargetBox-2="[174,1415,890,1252]" captionTargetId-0="figure-139@20.[176,1412,453,1595]" captionTargetId-1="figure-118@22.[160,1427,345,1806]" captionTargetId-2="figure-357@23.[168,1420,885,1257]" captionTargetPageId-0="20" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="FIGURE 11. Litoria lisae sp. nov. in life, A) holotype (SAMA R71689), B–C), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards." captionText-1="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." captionText-2="FIGURE 13. Wave form (amplitude over time; top) and spectrogram (frequency over time; bottom) of two types of advertisement call produced by Litoria lisae sp. nov. (SAMA R71689) recorded at an air temperature of 20.5°C. Note distinctly clumped nature of pulses in chirp call (left) compared to buzz call (right)." figureDoi-0="http://doi.org/10.5281/zenodo.7800946" figureDoi-1="http://doi.org/10.5281/zenodo.7800948" figureDoi-2="http://doi.org/10.5281/zenodo.7800952" httpUri-0="https://zenodo.org/record/7800946/files/figure.png" httpUri-1="https://zenodo.org/record/7800948/files/figure.png" httpUri-2="https://zenodo.org/record/7800952/files/figure.png" pageId="19" pageNumber="170">Figs 11–13</figureCitation>
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1211,1770,1905]" box="[151,1211,1878,1905]" pageId="19" pageNumber="170">
|
||
https://zoobank.org/
|
||
<uri box="[379,1211,1878,1905]" pageId="19" pageNumber="170">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid box="[649,1211,1878,1905]" pageId="19" pageNumber="170">C64BD388-84E0-4288-A6CD-F70DFEABDC35</uuid>
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||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="20" lastPageNumber="171" pageId="19" pageNumber="170" type="materials_examined">
|
||
<paragraph blockId="19.[151,1436,1950,2013]" pageId="19" pageNumber="170">
|
||
<materialsCitation collectingDate="2017-05-16" collectionCode="SAMA, FN" collectorName="S. J. Richards" country="Papua New Guinea" elevation="1030" latitude="-6.4619" location="Arakubi Quarry" longLatPrecision="7" longitude="143.256" pageId="19" pageNumber="170" specimenCode="R71689, SJR15385" specimenCount="1" stateProvince="Southern Highlands" typeStatus="holotype">
|
||
<emphasis bold="true" box="[151,267,1951,1977]" pageId="19" pageNumber="170">
|
||
<typeStatus box="[151,262,1951,1977]" pageId="19" pageNumber="170">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode box="[277,356,1951,1977]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="19" pageNumber="170" type="Museum">SAMA</collectionCode>
|
||
<specimenCode box="[367,456,1951,1977]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19" pageNumber="170">R71689</specimenCode>
|
||
(
|
||
<collectionCode box="[474,511,1951,1977]" pageId="19" pageNumber="170">FN</collectionCode>
|
||
<specimenCode box="[522,641,1950,1977]" collectionCode="SJR" pageId="19" pageNumber="170">SJR15385</specimenCode>
|
||
),
|
||
<location LSID="urn:lsid:plazi:treatment:03D60841A03FFFF037CFF94CC0FBFE99:8EA0EF8CA03FFFFA35CEF838C25BF83D" box="[662,844,1951,1977]" country="Papua New Guinea" latitude="-6.4619" longLatPrecision="7" longitude="143.256" name="Arakubi Quarry" pageId="19" pageNumber="170" stateProvince="Southern Highlands">Arakubi Quarry</location>
|
||
,
|
||
<collectingRegion box="[859,1198,1951,1977]" country="Papua New Guinea" name="Southern Highlands" pageId="19" pageNumber="170">Southern Highlands Province</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1213,1436,1951,1977]" name="Papua New Guinea" pageId="19" pageNumber="170">Papua New Guinea</collectingCountry>
|
||
(
|
||
<geoCoordinate box="[159,267,1987,2013]" degrees="6.4619" direction="south" orientation="latitude" pageId="19" pageNumber="170" precision="5" value="-6.4619">6.4619°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[278,415,1986,2013]" degrees="143.2560" direction="east" orientation="longitude" pageId="19" pageNumber="170" precision="5" value="143.256">143.2560°E</geoCoordinate>
|
||
;
|
||
<elevation box="[426,569,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="19" pageNumber="170" unit="m" value="1030.0">
|
||
<quantity box="[426,511,1987,2013]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="19" pageNumber="170" unit="m" value="1030.0">1030 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) by
|
||
<collectorName box="[622,769,1987,2013]" pageId="19" pageNumber="170">S.J. Richards</collectorName>
|
||
on
|
||
<date box="[811,964,1987,2013]" pageId="19" pageNumber="170" value="2017-05-16">
|
||
<collectingDate box="[811,964,1987,2013]" pageId="19" pageNumber="170" value="2017-05-16">16 May 2017</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph blockId="20.[151,1436,151,429]" pageId="20" pageNumber="171">
|
||
<materialsCitation collectingDate="2015-07-01" collectionCode="SAMA" pageId="20" pageNumber="171" specimenCode="R71688, SJR14931" specimenCount="2" typeStatus="paratype">
|
||
<emphasis bold="true" box="[199,326,151,177]" pageId="20" pageNumber="171">
|
||
<typeStatus box="[199,321,151,177]" pageId="20" pageNumber="171">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(n = 8, all adult males)
|
||
<collectionCode box="[607,686,151,177]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="20" pageNumber="171" type="Museum">SAMA</collectionCode>
|
||
<specimenCode box="[696,784,151,177]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R71688</specimenCode>
|
||
(
|
||
<specimenCode box="[801,922,151,177]" collectionCode="SJR" pageId="20" pageNumber="171">SJR14931</specimenCode>
|
||
), same details as holotype but collected on
|
||
<date pageId="20" pageNumber="171" value="2015-07-01">
|
||
<collectingDate pageId="20" pageNumber="171" value="2015-07-01">1 July 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2015-05-17" collectionCode="FN" collectorName="C. Dahl & S. J. Richards" country="Papua New Guinea" elevation="1400" latitude="-6.4402" location="Kilometer Point" longLatPrecision="7" longitude="143.224" municipality="Iagifu Ridge" pageId="20" pageNumber="171" specimenCode="R71690, SJR15388" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
|
||
<specimenCode box="[276,365,187,213]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R71690</specimenCode>
|
||
(
|
||
<collectionCode box="[380,417,187,213]" pageId="20" pageNumber="171">FN</collectionCode>
|
||
<specimenCode box="[425,544,186,213]" collectionCode="SJR" pageId="20" pageNumber="171">SJR15388</specimenCode>
|
||
),
|
||
<location LSID="urn:lsid:plazi:treatment:03D60841A03FFFF037CFF94CC0FBFE99:8EA0EF8CA038FFFD356CFF1CC3F0FF51" box="[564,743,187,213]" country="Papua New Guinea" latitude="-6.4402" longLatPrecision="7" longitude="143.224" municipality="Iagifu Ridge" name="Kilometer Point" pageId="20" pageNumber="171" stateProvince="Southern Highlands">Kilometer Point</location>
|
||
107,
|
||
<collectingMunicipality box="[807,952,187,213]" pageId="20" pageNumber="171">Iagifu Ridge</collectingMunicipality>
|
||
,
|
||
<collectingRegion box="[963,1296,187,213]" country="Papua New Guinea" name="Southern Highlands" pageId="20" pageNumber="171">Southern Highlands Province</collectingRegion>
|
||
,
|
||
<collectingCountry name="Papua New Guinea" pageId="20" pageNumber="171">Papua New Guinea</collectingCountry>
|
||
(
|
||
<geoCoordinate box="[250,358,223,249]" degrees="6.4402" direction="south" orientation="latitude" pageId="20" pageNumber="171" precision="5" value="-6.4402">6.4402°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[373,510,223,249]" degrees="143.2240" direction="east" orientation="longitude" pageId="20" pageNumber="171" precision="5" value="143.224">143.2240°E</geoCoordinate>
|
||
;
|
||
<elevation box="[524,680,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="171" unit="m" value="1400.0">
|
||
<quantity box="[524,619,223,249]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="171" unit="m" value="1400.0">1,400 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) by
|
||
<collectorName box="[739,829,223,249]" pageId="20" pageNumber="171">C. Dahl</collectorName>
|
||
and
|
||
<collectorName box="[891,1041,223,249]" pageId="20" pageNumber="171">S.J. Richards</collectorName>
|
||
on
|
||
<date box="[1090,1248,222,249]" pageId="20" pageNumber="171" value="2015-05-17">
|
||
<collectingDate box="[1090,1248,222,249]" pageId="20" pageNumber="171" value="2015-05-17">17 May 2015</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2001-10-23" collectionCode="FN" collectorName="S. J. Richards" country="Papua New Guinea" elevation="1360" latitude="-6.4446" location="Iagifu Ridge" longLatPrecision="7" longitude="143.2193" pageId="20" pageNumber="171" specimenCode="R60719-60720, SJR2381-2382, R71686, SJR2127" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
|
||
<specimenCode box="[1264,1436,223,249]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R60719–60720</specimenCode>
|
||
(
|
||
<collectionCode box="[159,196,259,285]" pageId="20" pageNumber="171">FN</collectionCode>
|
||
<specimenCode box="[206,379,259,285]" collectionCode="SJR" pageId="20" pageNumber="171">SJR2381–2382</specimenCode>
|
||
) and
|
||
<specimenCode box="[447,535,259,285]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R71686</specimenCode>
|
||
(
|
||
<collectionCode box="[553,590,259,285]" pageId="20" pageNumber="171">FN</collectionCode>
|
||
<specimenCode box="[600,705,259,285]" collectionCode="SJR" pageId="20" pageNumber="171">SJR2127</specimenCode>
|
||
), summit of
|
||
<location LSID="urn:lsid:plazi:treatment:03D60841A03FFFF037CFF94CC0FBFE99:8EA0EF8CA038FFFD340FFEA4C2FDFE99" box="[855,1002,259,285]" country="Papua New Guinea" latitude="-6.4446" longLatPrecision="7" longitude="143.2193" name="Iagifu Ridge" pageId="20" pageNumber="171" stateProvince="Southern Highlands">Iagifu Ridge</location>
|
||
,
|
||
<collectingRegion box="[1015,1353,259,285]" country="Papua New Guinea" name="Southern Highlands" pageId="20" pageNumber="171">Southern Highlands Province</collectingRegion>
|
||
,
|
||
<collectingCountry name="Papua New Guinea" pageId="20" pageNumber="171">Papua New Guinea</collectingCountry>
|
||
(
|
||
<geoCoordinate box="[309,417,295,321]" degrees="6.4446" direction="south" orientation="latitude" pageId="20" pageNumber="171" precision="5" value="-6.4446">6.4446°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[429,566,295,321]" degrees="143.2193" direction="east" orientation="longitude" pageId="20" pageNumber="171" precision="5" value="143.2193">143.2193°E</geoCoordinate>
|
||
;
|
||
<elevation box="[578,723,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="20" pageNumber="171" unit="m" value="1360.0">
|
||
<quantity box="[578,664,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.3599999999999999" pageId="20" pageNumber="171" unit="m" value="1360.0">1360 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) by
|
||
<collectorName box="[778,926,295,321]" pageId="20" pageNumber="171">S.J. Richards</collectorName>
|
||
on
|
||
<date box="[970,1159,295,321]" pageId="20" pageNumber="171" value="2001-10-23">
|
||
<collectingDate box="[970,1159,295,321]" pageId="20" pageNumber="171" value="2001-10-23">23 October 2001</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCode box="[1167,1344,295,321]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R60719–60720</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
and
|
||
<date pageId="20" pageNumber="171" value="2002-05-27">27 May 2002</date>
|
||
<materialsCitation collectingDate="2001-10-29" collectionCode="FN, PNGNM" collectorName="S. Richards" country="Papua New Guinea" elevation="830" latitude="-6.8145" location="Gobe Ridge" longLatPrecision="7" longitude="143.7743" pageId="20" pageNumber="171" specimenCode="R60716, SJR2443, R71687, SJR2445, SJR2444" specimenCount="1" stateProvince="Southern Highlands" typeStatus="paratype">
|
||
respectively;
|
||
<specimenCode box="[425,513,331,357]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R60716</specimenCode>
|
||
(
|
||
<collectionCode box="[529,566,331,357]" pageId="20" pageNumber="171">FN</collectionCode>
|
||
<specimenCode box="[574,679,331,357]" collectionCode="SJR" pageId="20" pageNumber="171">SJR2443</specimenCode>
|
||
),
|
||
<specimenCode box="[699,787,331,357]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="20" pageNumber="171">R71687</specimenCode>
|
||
(
|
||
<collectionCode box="[803,840,331,357]" pageId="20" pageNumber="171">FN</collectionCode>
|
||
<specimenCode box="[848,951,330,357]" collectionCode="SJR" pageId="20" pageNumber="171">SJR2445</specimenCode>
|
||
) and
|
||
<collectionCode box="[1014,1115,331,357]" pageId="20" pageNumber="171">PNGNM</collectionCode>
|
||
(
|
||
<collectionCode box="[1131,1168,331,357]" pageId="20" pageNumber="171">FN</collectionCode>
|
||
<specimenCode box="[1176,1281,331,357]" collectionCode="SJR" pageId="20" pageNumber="171">SJR2444</specimenCode>
|
||
),
|
||
<location LSID="urn:lsid:plazi:treatment:03D60841A03FFFF037CFF94CC0FBFE99:8EA0EF8CA038FFFD324DFEECC48BFEE1" box="[1301,1436,331,357]" country="Papua New Guinea" latitude="-6.8145" longLatPrecision="7" longitude="143.7743" name="Gobe Ridge" pageId="20" pageNumber="171" stateProvince="Southern Highlands">Gobe Ridge</location>
|
||
summit,
|
||
<collectingRegion box="[249,581,367,393]" country="Papua New Guinea" name="Southern Highlands" pageId="20" pageNumber="171">Southern Highlands Province</collectingRegion>
|
||
,
|
||
<collectingCountry box="[593,809,367,393]" name="Papua New Guinea" pageId="20" pageNumber="171">Papua New Guinea</collectingCountry>
|
||
(
|
||
<geoCoordinate box="[824,931,366,393]" degrees="6.8145" direction="south" orientation="latitude" pageId="20" pageNumber="171" precision="5" value="-6.8145">6.8145°S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[943,1080,367,393]" degrees="143.7743" direction="east" orientation="longitude" pageId="20" pageNumber="171" precision="5" value="143.7743">143.7743°E</geoCoordinate>
|
||
;
|
||
<elevation box="[1091,1219,367,393]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="20" pageNumber="171" unit="m" value="830.0">
|
||
<quantity box="[1091,1162,367,393]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="20" pageNumber="171" unit="m" value="830.0">830 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
) by
|
||
<collectorName box="[1272,1402,367,393]" pageId="20" pageNumber="171">S. Richards</collectorName>
|
||
on
|
||
<date box="[151,342,403,429]" pageId="20" pageNumber="171" value="2001-10-29">
|
||
<collectingDate box="[151,342,403,429]" pageId="20" pageNumber="171" value="2001-10-29">29 October 2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7800946" ID-Zenodo-Dep="7800946" httpUri="https://zenodo.org/record/7800946/files/figure.png" pageId="20" pageNumber="171" startId="20.[151,250,1619,1643]" targetBox="[176,1412,453,1594]" targetPageId="20">
|
||
<paragraph blockId="20.[151,1437,1619,1679]" pageId="20" pageNumber="171">
|
||
<emphasis bold="true" box="[151,281,1619,1643]" pageId="20" pageNumber="171">FIGURE 11</emphasis>
|
||
.
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[295,421,1619,1643]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[295,421,1619,1643]" italics="true" pageId="20" pageNumber="171">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[428,510,1619,1643]" pageId="20" pageNumber="171">
|
||
<taxonomicNameLabel box="[428,510,1619,1643]" pageId="20" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life,
|
||
<emphasis bold="true" box="[593,611,1619,1643]" pageId="20" pageNumber="171">A</emphasis>
|
||
) holotype (SAMA R71689),
|
||
<emphasis bold="true" box="[913,962,1619,1643]" pageId="20" pageNumber="171">B–C</emphasis>
|
||
), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="20" pageNumber="171" type="diagnosis">
|
||
<paragraph blockId="20.[151,1437,1725,2040]" pageId="20" pageNumber="171">
|
||
<emphasis bold="true" box="[199,320,1725,1751]" pageId="20" pageNumber="171">Diagnosis.</emphasis>
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[332,470,1725,1751]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[332,470,1725,1751]" italics="true" pageId="20" pageNumber="171">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[479,570,1725,1751]" pageId="20" pageNumber="171">
|
||
<taxonomicNameLabel box="[479,570,1725,1751]" pageId="20" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is distinguished from all other
|
||
<taxonomicName authorityName="sensu Tyler & Davies" authorityYear="1978" box="[930,1007,1726,1751]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="20" pageNumber="171" phylum="Chordata" rank="genus">
|
||
<emphasis box="[930,1007,1726,1751]" italics="true" pageId="20" pageNumber="171">Litoria</emphasis>
|
||
</taxonomicName>
|
||
by the following unique combination of characters: size small (SVL of males 27.0–
|
||
<quantity box="[677,782,1761,1787]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="20" pageNumber="171" unit="mm" value="29.7">29.7 mm</quantity>
|
||
); dorsum in life green with darker green spots; vomerine teeth absent; finger webbing extensive; prominent dermal fold along outer edge of foot; pigmentation on nictitating membrane restricted to narrow band at dorsal margin; advertisement call a single note containing either evenly spaced or discretely clumped pulses; and is genetically diagnosable from
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[888,1082,1869,1895]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="haematogaster" status="sp. nov.">
|
||
<emphasis box="[888,1082,1869,1895]" italics="true" pageId="20" pageNumber="171">L. haematogaster</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1089,1179,1869,1895]" pageId="20" pageNumber="171">
|
||
<taxonomicNameLabel box="[1089,1179,1869,1895]" pageId="20" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at 65 sites and from
|
||
<taxonomicName authorityName="Johnston & Richards" authorityYear="1994" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="20" pageNumber="171" phylum="Chordata" rank="species" species="majikthise">
|
||
<emphasis italics="true" pageId="20" pageNumber="171">L. majikthise</emphasis>
|
||
</taxonomicName>
|
||
at 64 sites in the 787 base pair alignment of mitochondrial
|
||
<emphasis box="[924,976,1905,1931]" italics="true" pageId="20" pageNumber="171">ND4</emphasis>
|
||
gene and flanking tRNA (
|
||
<tableCitation box="[1268,1351,1905,1931]" captionStart="TABLE 1" captionStartId="7.[151,239,467,493]" captionTargetBox="[151,2022,539,1721]" captionText="TABLE 1. Diagnostic sites from the ND4 alignment for species diagnoses. A) Litoria naispela sp. nov. versus L. singadanae; B) L. nigropunctata, L. daraiensis sp. nov., L. gracilis sp. nov., C) L. lisae sp. nov., L. haematogaster sp. nov., L. majikthise. Sites are numbered according to the alignment used in the phylogenetic analysis. A) 1 1 1 1 1 1 1 2 2 2 2 2 2 2 3 3 3 3 3 3 3 3 4 4 4 4 4 4 4 4 4 4 5 5 5 5 5 5 5 5 5 6 6 6 6 6 6 6 0 0 1 2 3 3 4 0 0 2 5 8 8 8 1 2 3 4 5 6 7 9 0 1 2 2 3 5 5 8 8 9 0 1 1 4 4 5 6 8 9 0 1 3 4 5 6 7 1 2 6 1 1 6 5 5 8 3 0 0 1 3 6 6 1 3 2 7 0 5 9 0 2 4 1 0 4 2 7 9 9 2 8 1 7 3 5 8 2 7 6 7 5 5 7 1 L. n A T G T T A G C T G A G A T C G C A T A A G G T C G A C T T G A C A G C T T T C A T T C T C T C L. s G C A C C G T T C A G A A C T A T T C T G A A C T A A T C G A G T G A T C C C T G C C T C T C T B) 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 9 9 9 0 0 0 0 0 1 3 3 4 4 5 5 6 7 8 9 0 0 1 1 2 3 5 5 5 6 6 6 6 6 7 1 4 5 1 2 6 8 9 6 3 9 3 5 4 5 0 5 5 9 5 8 1 8 3 6 3 6 7 0 1 2 5 8 2 L. n C C C A T G C C G A A T C C C T T T A T T T T A A A C A A C C G C G L. g T C C G C C C T A C/T T C T T C T C T C/T C T C/T C T T A C/T G A T T G A A L. d T T T G C T C T A T A C T T T C C C T A C T T T T G T G A C T A A A" pageId="20" pageNumber="171">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="24" lastPageNumber="175" pageId="20" pageNumber="171" type="description">
|
||
<paragraph blockId="20.[151,1437,1725,2040]" lastBlockId="21.[151,1436,151,429]" lastPageId="21" lastPageNumber="172" pageId="20" pageNumber="171">
|
||
<emphasis bold="true" box="[199,489,1941,1967]" pageId="20" pageNumber="171">
|
||
Description of
|
||
<typeStatus box="[378,484,1941,1967]" pageId="20" pageNumber="171">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male (
|
||
<figureCitation box="[638,740,1941,1967]" captionStart="FIGURE 11" captionStartId="20.[151,250,1619,1643]" captionTargetBox="[176,1412,453,1594]" captionTargetId="figure-139@20.[176,1412,453,1595]" captionTargetPageId="20" captionText="FIGURE 11. Litoria lisae sp. nov. in life, A) holotype (SAMA R71689), B–C), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800946" httpUri="https://zenodo.org/record/7800946/files/figure.png" pageId="20" pageNumber="171">Fig. 11A</figureCitation>
|
||
) with vocal slits and pale brown nuptial pads. Morphometric data are summarised in
|
||
<tableCitation box="[413,498,1977,2003]" captionStart="TABLE 4" captionStartId="21.[151,239,476,500]" captionTargetPageId="21" captionText="TABLE 4. Measurements and ratios of the type series of Litoria lisae sp. nov. SAMA R71689 is the holotype." pageId="20" pageNumber="171">Table 4</tableCitation>
|
||
. Body very slender. Limbs long (TL/SVL 0.56). Head narrower than long (HW/SVL 0.28; HL/SV 0.33, HW/HL 0.87). Vomerine teeth absent. Tongue moderately large, oval, with broad posterior notch. Vocal slits short, located laterally in floor of mouth, extending from slightly posterior of angle of jaws to slightly anterior of angle of jaws. Snout narrowly rounded in dorsal view (
|
||
<figureCitation box="[905,1010,187,213]" captionStart="FIGURE 12" captionStartId="22.[151,250,1829,1853]" captionTargetBox="[160,1427,345,1806]" captionTargetId="figure-118@22.[160,1427,345,1806]" captionTargetPageId="22" captionText="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="21" pageNumber="172">Fig. 12A</figureCitation>
|
||
), bluntly rounded (nearly truncate) in lateral view (
|
||
<figureCitation box="[297,398,223,249]" captionStart="FIGURE 12" captionStartId="22.[151,250,1829,1853]" captionTargetBox="[160,1427,345,1806]" captionTargetId="figure-118@22.[160,1427,345,1806]" captionTargetPageId="22" captionText="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="21" pageNumber="172">Fig. 12B</figureCitation>
|
||
); canthus rostralis broadly rounded, nearly straight; loreal region steep, slightly concave; lips slightly flared; nostrils much closer to tip of snout than eyes, oriented anterolaterally; internarial distance equal to distance from external naris to eye (EN/IN 1.0); eyes moderately small (EYE/SVL 0.11) but prominent, protruding in dorsal and ventral views; pupil horizontal, pigmentation on nictitating membrane restricted to narrow band along dorsal edge. Tympanum small (TYM/SVL 0.05), slightly less than one half diameter of eye (TYM/EYE = 0.45), distinct except dorsal edge of annulus obscured by thick, strongly curved supratympanic ridge.
|
||
</paragraph>
|
||
<caption box="[151,1331,476,501]" pageId="21" pageNumber="172" startId="21.[151,239,476,500]" targetBox="[164,1423,514,1437]" targetIsTable="true" targetPageId="21">
|
||
<paragraph blockId="21.[151,1331,476,501]" box="[151,1331,476,501]" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[151,266,476,500]" pageId="21" pageNumber="172">TABLE 4.</emphasis>
|
||
Measurements and ratios of the type series of
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[763,894,476,500]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="21" pageNumber="172" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[763,894,476,500]" italics="true" pageId="21" pageNumber="172">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[901,986,477,501]" pageId="21" pageNumber="172">
|
||
<taxonomicNameLabel box="[901,986,477,501]" pageId="21" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
SAMA R71689 is the holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="21" pageNumber="172">
|
||
<table box="[164,1423,514,1437]" gridcols="12" gridrows="27" pageId="21" pageNumber="172">
|
||
<tr box="[164,1423,514,537]" gridrow="0" pageId="21" pageNumber="172" rowspan-10="1" rowspan-11="1">
|
||
<th box="[164,270,514,537]" gridcol="0" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[176,258,514,536]" pageId="21" pageNumber="172">SAMA #</emphasis>
|
||
</th>
|
||
<th box="[298,371,514,537]" gridcol="1" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[298,371,515,536]" pageId="21" pageNumber="172">R71689</emphasis>
|
||
</th>
|
||
<th box="[401,474,514,537]" gridcol="2" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[401,474,515,536]" pageId="21" pageNumber="172">R71690</emphasis>
|
||
</th>
|
||
<th box="[503,576,514,537]" gridcol="3" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[503,576,515,536]" pageId="21" pageNumber="172">R71688</emphasis>
|
||
</th>
|
||
<th box="[606,679,514,537]" gridcol="4" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[606,679,515,536]" pageId="21" pageNumber="172">R60719</emphasis>
|
||
</th>
|
||
<th box="[708,781,514,537]" gridcol="5" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[708,781,515,536]" pageId="21" pageNumber="172">R60720</emphasis>
|
||
</th>
|
||
<th box="[811,884,514,537]" gridcol="6" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[811,884,515,536]" pageId="21" pageNumber="172">R60718</emphasis>
|
||
</th>
|
||
<th box="[911,997,514,537]" gridcol="7" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[911,997,515,537]" pageId="21" pageNumber="172">PNGNM</emphasis>
|
||
</th>
|
||
<th box="[1024,1097,514,537]" gridcol="8" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[1024,1097,515,536]" pageId="21" pageNumber="172">R71687</emphasis>
|
||
</th>
|
||
<th box="[1127,1200,514,537]" gridcol="9" gridrow="0" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[1127,1200,515,536]" pageId="21" pageNumber="172">R71686</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr box="[164,1423,549,571]" gridrow="1" pageId="21" pageNumber="172">
|
||
<th box="[164,270,549,571]" gridcol="0" gridrow="1" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[200,234,549,571]" pageId="21" pageNumber="172">Sex</emphasis>
|
||
</th>
|
||
<td box="[298,371,549,571]" gridcol="1" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[401,474,549,571]" gridcol="2" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[503,576,549,571]" gridcol="3" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[606,679,549,571]" gridcol="4" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[708,781,549,571]" gridcol="5" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[811,884,549,571]" gridcol="6" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[911,997,549,571]" gridcol="7" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[1024,1097,549,571]" gridcol="8" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[1127,1200,549,571]" gridcol="9" gridrow="1" pageId="21" pageNumber="172">M</td>
|
||
<td box="[1238,1293,549,571]" gridcol="10" gridrow="1" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[1238,1293,549,570]" pageId="21" pageNumber="172">Mean</emphasis>
|
||
</td>
|
||
<td box="[1327,1423,549,571]" gridcol="11" gridrow="1" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[1327,1389,549,570]" pageId="21" pageNumber="172">Range</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr box="[164,1423,584,606]" gridrow="2" pageId="21" pageNumber="172">
|
||
<th box="[164,270,584,606]" gridcol="0" gridrow="2" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[196,240,584,606]" pageId="21" pageNumber="172">SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,584,606]" gridcol="1" gridrow="2" pageId="21" pageNumber="172">27.3</td>
|
||
<td box="[401,474,584,606]" gridcol="2" gridrow="2" pageId="21" pageNumber="172">27.4</td>
|
||
<td box="[503,576,584,606]" gridcol="3" gridrow="2" pageId="21" pageNumber="172">27.0</td>
|
||
<td box="[606,679,584,606]" gridcol="4" gridrow="2" pageId="21" pageNumber="172">28.5</td>
|
||
<td box="[708,781,584,606]" gridcol="5" gridrow="2" pageId="21" pageNumber="172">29.7</td>
|
||
<td box="[811,884,584,606]" gridcol="6" gridrow="2" pageId="21" pageNumber="172">29.1</td>
|
||
<td box="[911,997,584,606]" gridcol="7" gridrow="2" pageId="21" pageNumber="172">27.2</td>
|
||
<td box="[1024,1097,584,606]" gridcol="8" gridrow="2" pageId="21" pageNumber="172">29.3</td>
|
||
<td box="[1127,1200,584,606]" gridcol="9" gridrow="2" pageId="21" pageNumber="172">28.9</td>
|
||
<td box="[1238,1293,584,606]" gridcol="10" gridrow="2" pageId="21" pageNumber="172">28.3</td>
|
||
<td box="[1327,1423,584,606]" gridcol="11" gridrow="2" pageId="21" pageNumber="172">27.0–29.7</td>
|
||
</tr>
|
||
<tr box="[164,1423,618,641]" gridrow="3" pageId="21" pageNumber="172">
|
||
<th box="[164,270,618,641]" gridcol="0" gridrow="3" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[203,233,619,640]" pageId="21" pageNumber="172">TL</emphasis>
|
||
</th>
|
||
<td box="[298,371,618,641]" gridcol="1" gridrow="3" pageId="21" pageNumber="172">15.4</td>
|
||
<td box="[401,474,618,641]" gridcol="2" gridrow="3" pageId="21" pageNumber="172">15.8</td>
|
||
<td box="[503,576,618,641]" gridcol="3" gridrow="3" pageId="21" pageNumber="172">15.8</td>
|
||
<td box="[606,679,618,641]" gridcol="4" gridrow="3" pageId="21" pageNumber="172">16.6</td>
|
||
<td box="[708,781,618,641]" gridcol="5" gridrow="3" pageId="21" pageNumber="172">16.5</td>
|
||
<td box="[811,884,618,641]" gridcol="6" gridrow="3" pageId="21" pageNumber="172">17.2</td>
|
||
<td box="[911,997,618,641]" gridcol="7" gridrow="3" pageId="21" pageNumber="172">15.8</td>
|
||
<td box="[1024,1097,618,641]" gridcol="8" gridrow="3" pageId="21" pageNumber="172">16.4</td>
|
||
<td box="[1127,1200,618,641]" gridcol="9" gridrow="3" pageId="21" pageNumber="172">16.3</td>
|
||
<td box="[1238,1293,618,641]" gridcol="10" gridrow="3" pageId="21" pageNumber="172">16.2</td>
|
||
<td box="[1327,1423,618,641]" gridcol="11" gridrow="3" pageId="21" pageNumber="172">15.4–17.2</td>
|
||
</tr>
|
||
<tr box="[164,1423,653,675]" gridrow="4" pageId="21" pageNumber="172">
|
||
<th box="[164,270,653,675]" gridcol="0" gridrow="4" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[201,234,653,674]" pageId="21" pageNumber="172">HL</emphasis>
|
||
</th>
|
||
<td box="[298,371,653,675]" gridcol="1" gridrow="4" pageId="21" pageNumber="172">8.9</td>
|
||
<td box="[401,474,653,675]" gridcol="2" gridrow="4" pageId="21" pageNumber="172">9.2</td>
|
||
<td box="[503,576,653,675]" gridcol="3" gridrow="4" pageId="21" pageNumber="172">9.2</td>
|
||
<td box="[606,679,653,675]" gridcol="4" gridrow="4" pageId="21" pageNumber="172">9.5</td>
|
||
<td box="[708,781,653,675]" gridcol="5" gridrow="4" pageId="21" pageNumber="172">8.7</td>
|
||
<td box="[811,884,653,675]" gridcol="6" gridrow="4" pageId="21" pageNumber="172">10.5</td>
|
||
<td box="[911,997,653,675]" gridcol="7" gridrow="4" pageId="21" pageNumber="172">9.1</td>
|
||
<td box="[1024,1097,653,675]" gridcol="8" gridrow="4" pageId="21" pageNumber="172">9.6</td>
|
||
<td box="[1127,1200,653,675]" gridcol="9" gridrow="4" pageId="21" pageNumber="172">9.8</td>
|
||
<td box="[1238,1293,653,675]" gridcol="10" gridrow="4" pageId="21" pageNumber="172">9.4</td>
|
||
<td box="[1327,1423,653,675]" gridcol="11" gridrow="4" pageId="21" pageNumber="172">8.7–10.5</td>
|
||
</tr>
|
||
<tr box="[164,1423,688,710]" gridrow="5" pageId="21" pageNumber="172">
|
||
<th box="[164,270,688,710]" gridcol="0" gridrow="5" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[197,237,688,709]" pageId="21" pageNumber="172">HW</emphasis>
|
||
</th>
|
||
<td box="[298,371,688,710]" gridcol="1" gridrow="5" pageId="21" pageNumber="172">7.7</td>
|
||
<td box="[401,474,688,710]" gridcol="2" gridrow="5" pageId="21" pageNumber="172">8.5</td>
|
||
<td box="[503,576,688,710]" gridcol="3" gridrow="5" pageId="21" pageNumber="172">8.4</td>
|
||
<td box="[606,679,688,710]" gridcol="4" gridrow="5" pageId="21" pageNumber="172">8.4</td>
|
||
<td box="[708,781,688,710]" gridcol="5" gridrow="5" pageId="21" pageNumber="172">8.7</td>
|
||
<td box="[811,884,688,710]" gridcol="6" gridrow="5" pageId="21" pageNumber="172">9.2</td>
|
||
<td box="[911,997,688,710]" gridcol="7" gridrow="5" pageId="21" pageNumber="172">8.7</td>
|
||
<td box="[1024,1097,688,710]" gridcol="8" gridrow="5" pageId="21" pageNumber="172">8.3</td>
|
||
<td box="[1127,1200,688,710]" gridcol="9" gridrow="5" pageId="21" pageNumber="172">8.4</td>
|
||
<td box="[1238,1293,688,710]" gridcol="10" gridrow="5" pageId="21" pageNumber="172">8.5</td>
|
||
<td box="[1327,1423,688,710]" gridcol="11" gridrow="5" pageId="21" pageNumber="172">7.7–9.2</td>
|
||
</tr>
|
||
<tr box="[164,1423,722,744]" gridrow="6" pageId="21" pageNumber="172">
|
||
<th box="[164,270,722,744]" gridcol="0" gridrow="6" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[194,241,723,744]" pageId="21" pageNumber="172">EYE</emphasis>
|
||
</th>
|
||
<td box="[298,371,722,744]" gridcol="1" gridrow="6" pageId="21" pageNumber="172">3.1</td>
|
||
<td box="[401,474,722,744]" gridcol="2" gridrow="6" pageId="21" pageNumber="172">3.5</td>
|
||
<td box="[503,576,722,744]" gridcol="3" gridrow="6" pageId="21" pageNumber="172">3.2</td>
|
||
<td box="[606,679,722,744]" gridcol="4" gridrow="6" pageId="21" pageNumber="172">3.5</td>
|
||
<td box="[708,781,722,744]" gridcol="5" gridrow="6" pageId="21" pageNumber="172">3.1</td>
|
||
<td box="[811,884,722,744]" gridcol="6" gridrow="6" pageId="21" pageNumber="172">3.3</td>
|
||
<td box="[911,997,722,744]" gridcol="7" gridrow="6" pageId="21" pageNumber="172">3.3</td>
|
||
<td box="[1024,1097,722,744]" gridcol="8" gridrow="6" pageId="21" pageNumber="172">3.5</td>
|
||
<td box="[1127,1200,722,744]" gridcol="9" gridrow="6" pageId="21" pageNumber="172">3.5</td>
|
||
<td box="[1238,1293,722,744]" gridcol="10" gridrow="6" pageId="21" pageNumber="172">3.3</td>
|
||
<td box="[1327,1423,722,744]" gridcol="11" gridrow="6" pageId="21" pageNumber="172">3.1–3.5</td>
|
||
</tr>
|
||
<tr box="[164,1423,757,779]" gridrow="7" pageId="21" pageNumber="172">
|
||
<th box="[164,270,757,779]" gridcol="0" gridrow="7" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[191,244,757,778]" pageId="21" pageNumber="172">TYM</emphasis>
|
||
</th>
|
||
<td box="[298,371,757,779]" gridcol="1" gridrow="7" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[401,474,757,779]" gridcol="2" gridrow="7" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[503,576,757,779]" gridcol="3" gridrow="7" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[606,679,757,779]" gridcol="4" gridrow="7" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[708,781,757,779]" gridcol="5" gridrow="7" pageId="21" pageNumber="172">1.2</td>
|
||
<td box="[811,884,757,779]" gridcol="6" gridrow="7" pageId="21" pageNumber="172">1.5</td>
|
||
<td box="[911,997,757,779]" gridcol="7" gridrow="7" pageId="21" pageNumber="172">1.5</td>
|
||
<td box="[1024,1097,757,779]" gridcol="8" gridrow="7" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[1127,1200,757,779]" gridcol="9" gridrow="7" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[1238,1293,757,779]" gridcol="10" gridrow="7" pageId="21" pageNumber="172">1.5</td>
|
||
<td box="[1327,1423,757,779]" gridcol="11" gridrow="7" pageId="21" pageNumber="172">1.2–1.6</td>
|
||
</tr>
|
||
<tr box="[164,1423,792,813]" gridrow="8" pageId="21" pageNumber="172">
|
||
<th box="[164,270,792,813]" gridcol="0" gridrow="8" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[201,232,792,813]" pageId="21" pageNumber="172">EN</emphasis>
|
||
</th>
|
||
<td box="[298,371,792,813]" gridcol="1" gridrow="8" pageId="21" pageNumber="172">2.4</td>
|
||
<td box="[401,474,792,813]" gridcol="2" gridrow="8" pageId="21" pageNumber="172">2.9</td>
|
||
<td box="[503,576,792,813]" gridcol="3" gridrow="8" pageId="21" pageNumber="172">2.8</td>
|
||
<td box="[606,679,792,813]" gridcol="4" gridrow="8" pageId="21" pageNumber="172">2.9</td>
|
||
<td box="[708,781,792,813]" gridcol="5" gridrow="8" pageId="21" pageNumber="172">2.7</td>
|
||
<td box="[811,884,792,813]" gridcol="6" gridrow="8" pageId="21" pageNumber="172">3.0</td>
|
||
<td box="[911,997,792,813]" gridcol="7" gridrow="8" pageId="21" pageNumber="172">2.8</td>
|
||
<td box="[1024,1097,792,813]" gridcol="8" gridrow="8" pageId="21" pageNumber="172">2.9</td>
|
||
<td box="[1127,1200,792,813]" gridcol="9" gridrow="8" pageId="21" pageNumber="172">3.0</td>
|
||
<td box="[1238,1293,792,813]" gridcol="10" gridrow="8" pageId="21" pageNumber="172">2.8</td>
|
||
<td box="[1327,1423,792,813]" gridcol="11" gridrow="8" pageId="21" pageNumber="172">2.4–3.0</td>
|
||
</tr>
|
||
<tr box="[164,1423,826,848]" gridrow="9" pageId="21" pageNumber="172">
|
||
<th box="[164,270,826,848]" gridcol="0" gridrow="9" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[205,230,826,847]" pageId="21" pageNumber="172">IN</emphasis>
|
||
</th>
|
||
<td box="[298,371,826,848]" gridcol="1" gridrow="9" pageId="21" pageNumber="172">2.4</td>
|
||
<td box="[401,474,826,848]" gridcol="2" gridrow="9" pageId="21" pageNumber="172">3.0</td>
|
||
<td box="[503,576,826,848]" gridcol="3" gridrow="9" pageId="21" pageNumber="172">2.5</td>
|
||
<td box="[606,679,826,848]" gridcol="4" gridrow="9" pageId="21" pageNumber="172">3.0</td>
|
||
<td box="[708,781,826,848]" gridcol="5" gridrow="9" pageId="21" pageNumber="172">2.9</td>
|
||
<td box="[811,884,826,848]" gridcol="6" gridrow="9" pageId="21" pageNumber="172">3.0</td>
|
||
<td box="[911,997,826,848]" gridcol="7" gridrow="9" pageId="21" pageNumber="172">2.8</td>
|
||
<td box="[1024,1097,826,848]" gridcol="8" gridrow="9" pageId="21" pageNumber="172">2.9</td>
|
||
<td box="[1127,1200,826,848]" gridcol="9" gridrow="9" pageId="21" pageNumber="172">3.0</td>
|
||
<td box="[1238,1293,826,848]" gridcol="10" gridrow="9" pageId="21" pageNumber="172">2.8</td>
|
||
<td box="[1327,1423,826,848]" gridcol="11" gridrow="9" pageId="21" pageNumber="172">2.4–3.0</td>
|
||
</tr>
|
||
<tr box="[164,1423,861,882]" gridrow="10" pageId="21" pageNumber="172">
|
||
<th box="[164,270,861,882]" gridcol="0" gridrow="10" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[196,239,861,882]" pageId="21" pageNumber="172">4TD</emphasis>
|
||
</th>
|
||
<td box="[298,371,861,882]" gridcol="1" gridrow="10" pageId="21" pageNumber="172">1.2</td>
|
||
<td box="[401,474,861,882]" gridcol="2" gridrow="10" pageId="21" pageNumber="172">1.2</td>
|
||
<td box="[503,576,861,882]" gridcol="3" gridrow="10" pageId="21" pageNumber="172">1.1</td>
|
||
<td box="[606,679,861,882]" gridcol="4" gridrow="10" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[708,781,861,882]" gridcol="5" gridrow="10" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[811,884,861,882]" gridcol="6" gridrow="10" pageId="21" pageNumber="172">1.3</td>
|
||
<td box="[911,997,861,882]" gridcol="7" gridrow="10" pageId="21" pageNumber="172">1.3</td>
|
||
<td box="[1024,1097,861,882]" gridcol="8" gridrow="10" pageId="21" pageNumber="172">1.3</td>
|
||
<td box="[1127,1200,861,882]" gridcol="9" gridrow="10" pageId="21" pageNumber="172">1.3</td>
|
||
<td box="[1238,1293,861,882]" gridcol="10" gridrow="10" pageId="21" pageNumber="172">1.3</td>
|
||
<td box="[1327,1423,861,882]" gridcol="11" gridrow="10" pageId="21" pageNumber="172">1.1–1.4</td>
|
||
</tr>
|
||
<tr box="[164,1423,895,917]" gridrow="11" pageId="21" pageNumber="172">
|
||
<th box="[164,270,895,917]" gridcol="0" gridrow="11" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[198,238,895,916]" pageId="21" pageNumber="172">4TP</emphasis>
|
||
</th>
|
||
<td box="[298,371,895,917]" gridcol="1" gridrow="11" pageId="21" pageNumber="172">0.9</td>
|
||
<td box="[401,474,895,917]" gridcol="2" gridrow="11" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[503,576,895,917]" gridcol="3" gridrow="11" pageId="21" pageNumber="172">0.7</td>
|
||
<td box="[606,679,895,917]" gridcol="4" gridrow="11" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[708,781,895,917]" gridcol="5" gridrow="11" pageId="21" pageNumber="172">0.9</td>
|
||
<td box="[811,884,895,917]" gridcol="6" gridrow="11" pageId="21" pageNumber="172">0.9</td>
|
||
<td box="[911,997,895,917]" gridcol="7" gridrow="11" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[1024,1097,895,917]" gridcol="8" gridrow="11" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[1127,1200,895,917]" gridcol="9" gridrow="11" pageId="21" pageNumber="172">1.0</td>
|
||
<td box="[1238,1293,895,917]" gridcol="10" gridrow="11" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[1327,1423,895,917]" gridcol="11" gridrow="11" pageId="21" pageNumber="172">0.7–1.0</td>
|
||
</tr>
|
||
<tr box="[164,1423,930,952]" gridrow="12" pageId="21" pageNumber="172">
|
||
<th box="[164,270,930,952]" gridcol="0" gridrow="12" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[196,238,930,951]" pageId="21" pageNumber="172">3FD</emphasis>
|
||
</th>
|
||
<td box="[298,371,930,952]" gridcol="1" gridrow="12" pageId="21" pageNumber="172">1.5</td>
|
||
<td box="[401,474,930,952]" gridcol="2" gridrow="12" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[503,576,930,952]" gridcol="3" gridrow="12" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[606,679,930,952]" gridcol="4" gridrow="12" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[708,781,930,952]" gridcol="5" gridrow="12" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[811,884,930,952]" gridcol="6" gridrow="12" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[911,997,930,952]" gridcol="7" gridrow="12" pageId="21" pageNumber="172">1.6</td>
|
||
<td box="[1024,1097,930,952]" gridcol="8" gridrow="12" pageId="21" pageNumber="172">1.4</td>
|
||
<td box="[1127,1200,930,952]" gridcol="9" gridrow="12" pageId="21" pageNumber="172">1.7</td>
|
||
<td box="[1238,1293,930,952]" gridcol="10" gridrow="12" pageId="21" pageNumber="172">1.5</td>
|
||
<td box="[1327,1423,930,952]" gridcol="11" gridrow="12" pageId="21" pageNumber="172">1.4–1.7</td>
|
||
</tr>
|
||
<tr box="[164,1423,965,986]" gridrow="13" pageId="21" pageNumber="172">
|
||
<th box="[164,270,965,986]" gridcol="0" gridrow="13" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[198,237,965,986]" pageId="21" pageNumber="172">3FP</emphasis>
|
||
</th>
|
||
<td box="[298,371,965,986]" gridcol="1" gridrow="13" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[401,474,965,986]" gridcol="2" gridrow="13" pageId="21" pageNumber="172">1.0</td>
|
||
<td box="[503,576,965,986]" gridcol="3" gridrow="13" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[606,679,965,986]" gridcol="4" gridrow="13" pageId="21" pageNumber="172">0.8</td>
|
||
<td box="[708,781,965,986]" gridcol="5" gridrow="13" pageId="21" pageNumber="172">1.0</td>
|
||
<td box="[811,884,965,986]" gridcol="6" gridrow="13" pageId="21" pageNumber="172">1.0</td>
|
||
<td box="[911,997,965,986]" gridcol="7" gridrow="13" pageId="21" pageNumber="172">1.0</td>
|
||
<td box="[1024,1097,965,986]" gridcol="8" gridrow="13" pageId="21" pageNumber="172">1.1</td>
|
||
<td box="[1127,1200,965,986]" gridcol="9" gridrow="13" pageId="21" pageNumber="172">1.0</td>
|
||
<td box="[1238,1293,965,986]" gridcol="10" gridrow="13" pageId="21" pageNumber="172">0.9</td>
|
||
<td box="[1327,1423,965,986]" gridcol="11" gridrow="13" pageId="21" pageNumber="172">0.8–1.1</td>
|
||
</tr>
|
||
<tr box="[164,1423,999,1021]" gridrow="14" pageId="21" pageNumber="172">
|
||
<th box="[164,270,999,1021]" gridcol="0" gridrow="14" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[178,259,999,1021]" pageId="21" pageNumber="172">TL/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,999,1021]" gridcol="1" gridrow="14" pageId="21" pageNumber="172">0.56</td>
|
||
<td box="[401,474,999,1021]" gridcol="2" gridrow="14" pageId="21" pageNumber="172">0.58</td>
|
||
<td box="[503,576,999,1021]" gridcol="3" gridrow="14" pageId="21" pageNumber="172">0.59</td>
|
||
<td box="[606,679,999,1021]" gridcol="4" gridrow="14" pageId="21" pageNumber="172">0.58</td>
|
||
<td box="[708,781,999,1021]" gridcol="5" gridrow="14" pageId="21" pageNumber="172">0.56</td>
|
||
<td box="[811,884,999,1021]" gridcol="6" gridrow="14" pageId="21" pageNumber="172">0.59</td>
|
||
<td box="[911,997,999,1021]" gridcol="7" gridrow="14" pageId="21" pageNumber="172">0.58</td>
|
||
<td box="[1024,1097,999,1021]" gridcol="8" gridrow="14" pageId="21" pageNumber="172">0.56</td>
|
||
<td box="[1127,1200,999,1021]" gridcol="9" gridrow="14" pageId="21" pageNumber="172">0.56</td>
|
||
<td box="[1238,1293,999,1021]" gridcol="10" gridrow="14" pageId="21" pageNumber="172">0.57</td>
|
||
<td box="[1327,1423,999,1021]" gridcol="11" gridrow="14" pageId="21" pageNumber="172">0.56–0.59</td>
|
||
</tr>
|
||
<tr box="[164,1423,1034,1056]" gridrow="15" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1034,1056]" gridcol="0" gridrow="15" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[173,263,1034,1056]" pageId="21" pageNumber="172">HW/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1034,1056]" gridcol="1" gridrow="15" pageId="21" pageNumber="172">0.28</td>
|
||
<td box="[401,474,1034,1056]" gridcol="2" gridrow="15" pageId="21" pageNumber="172">0.31</td>
|
||
<td box="[503,576,1034,1056]" gridcol="3" gridrow="15" pageId="21" pageNumber="172">0.31</td>
|
||
<td box="[606,679,1034,1056]" gridcol="4" gridrow="15" pageId="21" pageNumber="172">0.29</td>
|
||
<td box="[708,781,1034,1056]" gridcol="5" gridrow="15" pageId="21" pageNumber="172">0.29</td>
|
||
<td box="[811,884,1034,1056]" gridcol="6" gridrow="15" pageId="21" pageNumber="172">0.32</td>
|
||
<td box="[911,997,1034,1056]" gridcol="7" gridrow="15" pageId="21" pageNumber="172">0.32</td>
|
||
<td box="[1024,1097,1034,1056]" gridcol="8" gridrow="15" pageId="21" pageNumber="172">0.28</td>
|
||
<td box="[1127,1200,1034,1056]" gridcol="9" gridrow="15" pageId="21" pageNumber="172">0.29</td>
|
||
<td box="[1238,1293,1034,1056]" gridcol="10" gridrow="15" pageId="21" pageNumber="172">0.30</td>
|
||
<td box="[1327,1423,1034,1056]" gridcol="11" gridrow="15" pageId="21" pageNumber="172">0.28–0.32</td>
|
||
</tr>
|
||
<tr box="[164,1423,1068,1090]" gridrow="16" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1068,1090]" gridcol="0" gridrow="16" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[176,259,1068,1090]" pageId="21" pageNumber="172">HL/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1068,1090]" gridcol="1" gridrow="16" pageId="21" pageNumber="172">0.33</td>
|
||
<td box="[401,474,1068,1090]" gridcol="2" gridrow="16" pageId="21" pageNumber="172">0.34</td>
|
||
<td box="[503,576,1068,1090]" gridcol="3" gridrow="16" pageId="21" pageNumber="172">0.34</td>
|
||
<td box="[606,679,1068,1090]" gridcol="4" gridrow="16" pageId="21" pageNumber="172">0.33</td>
|
||
<td box="[708,781,1068,1090]" gridcol="5" gridrow="16" pageId="21" pageNumber="172">0.29</td>
|
||
<td box="[811,884,1068,1090]" gridcol="6" gridrow="16" pageId="21" pageNumber="172">0.36</td>
|
||
<td box="[911,997,1068,1090]" gridcol="7" gridrow="16" pageId="21" pageNumber="172">0.33</td>
|
||
<td box="[1024,1097,1068,1090]" gridcol="8" gridrow="16" pageId="21" pageNumber="172">0.33</td>
|
||
<td box="[1127,1200,1068,1090]" gridcol="9" gridrow="16" pageId="21" pageNumber="172">0.34</td>
|
||
<td box="[1238,1293,1068,1090]" gridcol="10" gridrow="16" pageId="21" pageNumber="172">0.33</td>
|
||
<td box="[1327,1423,1068,1090]" gridcol="11" gridrow="16" pageId="21" pageNumber="172">0.29–0.36</td>
|
||
</tr>
|
||
<tr box="[164,1423,1103,1125]" gridrow="17" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1103,1125]" gridcol="0" gridrow="17" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[178,257,1103,1125]" pageId="21" pageNumber="172">HW/HL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1103,1125]" gridcol="1" gridrow="17" pageId="21" pageNumber="172">0.87</td>
|
||
<td box="[401,474,1103,1125]" gridcol="2" gridrow="17" pageId="21" pageNumber="172">0.92</td>
|
||
<td box="[503,576,1103,1125]" gridcol="3" gridrow="17" pageId="21" pageNumber="172">0.91</td>
|
||
<td box="[606,679,1103,1125]" gridcol="4" gridrow="17" pageId="21" pageNumber="172">0.88</td>
|
||
<td box="[708,781,1103,1125]" gridcol="5" gridrow="17" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[811,884,1103,1125]" gridcol="6" gridrow="17" pageId="21" pageNumber="172">0.88</td>
|
||
<td box="[911,997,1103,1125]" gridcol="7" gridrow="17" pageId="21" pageNumber="172">0.96</td>
|
||
<td box="[1024,1097,1103,1125]" gridcol="8" gridrow="17" pageId="21" pageNumber="172">0.86</td>
|
||
<td box="[1127,1200,1103,1125]" gridcol="9" gridrow="17" pageId="21" pageNumber="172">0.86</td>
|
||
<td box="[1238,1293,1103,1125]" gridcol="10" gridrow="17" pageId="21" pageNumber="172">0.90</td>
|
||
<td box="[1327,1423,1103,1125]" gridcol="11" gridrow="17" pageId="21" pageNumber="172">0.86–1.00</td>
|
||
</tr>
|
||
<tr box="[164,1423,1138,1160]" gridrow="18" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1138,1160]" gridcol="0" gridrow="18" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[169,266,1138,1160]" pageId="21" pageNumber="172">EYE/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1138,1160]" gridcol="1" gridrow="18" pageId="21" pageNumber="172">0.11</td>
|
||
<td box="[401,474,1138,1160]" gridcol="2" gridrow="18" pageId="21" pageNumber="172">0.13</td>
|
||
<td box="[503,576,1138,1160]" gridcol="3" gridrow="18" pageId="21" pageNumber="172">0.12</td>
|
||
<td box="[606,679,1138,1160]" gridcol="4" gridrow="18" pageId="21" pageNumber="172">0.12</td>
|
||
<td box="[708,781,1138,1160]" gridcol="5" gridrow="18" pageId="21" pageNumber="172">0.10</td>
|
||
<td box="[811,884,1138,1160]" gridcol="6" gridrow="18" pageId="21" pageNumber="172">0.11</td>
|
||
<td box="[911,997,1138,1160]" gridcol="7" gridrow="18" pageId="21" pageNumber="172">0.12</td>
|
||
<td box="[1024,1097,1138,1160]" gridcol="8" gridrow="18" pageId="21" pageNumber="172">0.12</td>
|
||
<td box="[1127,1200,1138,1160]" gridcol="9" gridrow="18" pageId="21" pageNumber="172">0.12</td>
|
||
<td box="[1238,1293,1138,1160]" gridcol="10" gridrow="18" pageId="21" pageNumber="172">0.12</td>
|
||
<td box="[1327,1423,1138,1160]" gridcol="11" gridrow="18" pageId="21" pageNumber="172">0.10–0.13</td>
|
||
</tr>
|
||
<tr box="[164,1423,1172,1194]" gridrow="19" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1172,1194]" gridcol="0" gridrow="19" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[166,269,1172,1194]" pageId="21" pageNumber="172">TYM/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1172,1194]" gridcol="1" gridrow="19" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[401,474,1172,1194]" gridcol="2" gridrow="19" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[503,576,1172,1194]" gridcol="3" gridrow="19" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[606,679,1172,1194]" gridcol="4" gridrow="19" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[708,781,1172,1194]" gridcol="5" gridrow="19" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[811,884,1172,1194]" gridcol="6" gridrow="19" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[911,997,1172,1194]" gridcol="7" gridrow="19" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[1024,1097,1172,1194]" gridcol="8" gridrow="19" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[1127,1200,1172,1194]" gridcol="9" gridrow="19" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[1238,1293,1172,1194]" gridcol="10" gridrow="19" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[1327,1423,1172,1194]" gridcol="11" gridrow="19" pageId="21" pageNumber="172">0.04–0.06</td>
|
||
</tr>
|
||
<tr box="[164,1423,1207,1229]" gridrow="20" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1207,1229]" gridcol="0" gridrow="20" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[164,270,1207,1229]" pageId="21" pageNumber="172">TYM/EYE</emphasis>
|
||
</th>
|
||
<td box="[298,371,1207,1229]" gridcol="1" gridrow="20" pageId="21" pageNumber="172">0.45</td>
|
||
<td box="[401,474,1207,1229]" gridcol="2" gridrow="20" pageId="21" pageNumber="172">0.46</td>
|
||
<td box="[503,576,1207,1229]" gridcol="3" gridrow="20" pageId="21" pageNumber="172">0.50</td>
|
||
<td box="[606,679,1207,1229]" gridcol="4" gridrow="20" pageId="21" pageNumber="172">0.40</td>
|
||
<td box="[708,781,1207,1229]" gridcol="5" gridrow="20" pageId="21" pageNumber="172">0.39</td>
|
||
<td box="[811,884,1207,1229]" gridcol="6" gridrow="20" pageId="21" pageNumber="172">0.45</td>
|
||
<td box="[911,997,1207,1229]" gridcol="7" gridrow="20" pageId="21" pageNumber="172">0.45</td>
|
||
<td box="[1024,1097,1207,1229]" gridcol="8" gridrow="20" pageId="21" pageNumber="172">0.40</td>
|
||
<td box="[1127,1200,1207,1229]" gridcol="9" gridrow="20" pageId="21" pageNumber="172">0.46</td>
|
||
<td box="[1238,1293,1207,1229]" gridcol="10" gridrow="20" pageId="21" pageNumber="172">0.44</td>
|
||
<td box="[1327,1423,1207,1229]" gridcol="11" gridrow="20" pageId="21" pageNumber="172">0.39–0.50</td>
|
||
</tr>
|
||
<tr box="[164,1423,1241,1263]" gridrow="21" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1241,1263]" gridcol="0" gridrow="21" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[186,249,1241,1263]" pageId="21" pageNumber="172">EN/IN</emphasis>
|
||
</th>
|
||
<td box="[298,371,1241,1263]" gridcol="1" gridrow="21" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[401,474,1241,1263]" gridcol="2" gridrow="21" pageId="21" pageNumber="172">0.97</td>
|
||
<td box="[503,576,1241,1263]" gridcol="3" gridrow="21" pageId="21" pageNumber="172">1.12</td>
|
||
<td box="[606,679,1241,1263]" gridcol="4" gridrow="21" pageId="21" pageNumber="172">0.97</td>
|
||
<td box="[708,781,1241,1263]" gridcol="5" gridrow="21" pageId="21" pageNumber="172">0.93</td>
|
||
<td box="[811,884,1241,1263]" gridcol="6" gridrow="21" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[911,997,1241,1263]" gridcol="7" gridrow="21" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[1024,1097,1241,1263]" gridcol="8" gridrow="21" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[1127,1200,1241,1263]" gridcol="9" gridrow="21" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[1238,1293,1241,1263]" gridcol="10" gridrow="21" pageId="21" pageNumber="172">1.00</td>
|
||
<td box="[1327,1423,1241,1263]" gridcol="11" gridrow="21" pageId="21" pageNumber="172">0.93–1.12</td>
|
||
</tr>
|
||
<tr box="[164,1423,1276,1298]" gridrow="22" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1276,1298]" gridcol="0" gridrow="22" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[171,265,1276,1298]" pageId="21" pageNumber="172">4TD/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1276,1298]" gridcol="1" gridrow="22" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[401,474,1276,1298]" gridcol="2" gridrow="22" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[503,576,1276,1298]" gridcol="3" gridrow="22" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[606,679,1276,1298]" gridcol="4" gridrow="22" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[708,781,1276,1298]" gridcol="5" gridrow="22" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[811,884,1276,1298]" gridcol="6" gridrow="22" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[911,997,1276,1298]" gridcol="7" gridrow="22" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[1024,1097,1276,1298]" gridcol="8" gridrow="22" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[1127,1200,1276,1298]" gridcol="9" gridrow="22" pageId="21" pageNumber="172">0.04</td>
|
||
<td box="[1238,1293,1276,1298]" gridcol="10" gridrow="22" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[1327,1423,1276,1298]" gridcol="11" gridrow="22" pageId="21" pageNumber="172">0.04–0.05</td>
|
||
</tr>
|
||
<tr box="[164,1423,1311,1333]" gridrow="23" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1311,1333]" gridcol="0" gridrow="23" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[173,263,1311,1333]" pageId="21" pageNumber="172">4TD/4TP</emphasis>
|
||
</th>
|
||
<td box="[298,371,1311,1333]" gridcol="1" gridrow="23" pageId="21" pageNumber="172">1.33</td>
|
||
<td box="[401,474,1311,1333]" gridcol="2" gridrow="23" pageId="21" pageNumber="172">1.50</td>
|
||
<td box="[503,576,1311,1333]" gridcol="3" gridrow="23" pageId="21" pageNumber="172">1.57</td>
|
||
<td box="[606,679,1311,1333]" gridcol="4" gridrow="23" pageId="21" pageNumber="172">1.75</td>
|
||
<td box="[708,781,1311,1333]" gridcol="5" gridrow="23" pageId="21" pageNumber="172">1.56</td>
|
||
<td box="[811,884,1311,1333]" gridcol="6" gridrow="23" pageId="21" pageNumber="172">1.44</td>
|
||
<td box="[911,997,1311,1333]" gridcol="7" gridrow="23" pageId="21" pageNumber="172">1.63</td>
|
||
<td box="[1024,1097,1311,1333]" gridcol="8" gridrow="23" pageId="21" pageNumber="172">1.63</td>
|
||
<td box="[1127,1200,1311,1333]" gridcol="9" gridrow="23" pageId="21" pageNumber="172">1.30</td>
|
||
<td box="[1238,1293,1311,1333]" gridcol="10" gridrow="23" pageId="21" pageNumber="172">1.52</td>
|
||
<td box="[1327,1423,1311,1333]" gridcol="11" gridrow="23" pageId="21" pageNumber="172">1.30–1.75</td>
|
||
</tr>
|
||
<tr box="[164,1423,1345,1367]" gridrow="24" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1345,1367]" gridcol="0" gridrow="24" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[172,264,1345,1367]" pageId="21" pageNumber="172">3FD/SVL</emphasis>
|
||
</th>
|
||
<td box="[298,371,1345,1367]" gridcol="1" gridrow="24" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[401,474,1345,1367]" gridcol="2" gridrow="24" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[503,576,1345,1367]" gridcol="3" gridrow="24" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[606,679,1345,1367]" gridcol="4" gridrow="24" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[708,781,1345,1367]" gridcol="5" gridrow="24" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[811,884,1345,1367]" gridcol="6" gridrow="24" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[911,997,1345,1367]" gridcol="7" gridrow="24" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[1024,1097,1345,1367]" gridcol="8" gridrow="24" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[1127,1200,1345,1367]" gridcol="9" gridrow="24" pageId="21" pageNumber="172">0.06</td>
|
||
<td box="[1238,1293,1345,1367]" gridcol="10" gridrow="24" pageId="21" pageNumber="172">0.05</td>
|
||
<td box="[1327,1423,1345,1367]" gridcol="11" gridrow="24" pageId="21" pageNumber="172">0.05–0.06</td>
|
||
</tr>
|
||
<tr box="[164,1423,1380,1402]" gridrow="25" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1380,1402]" gridcol="0" gridrow="25" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[174,261,1380,1402]" pageId="21" pageNumber="172">3FD/3FP</emphasis>
|
||
</th>
|
||
<td box="[298,371,1380,1402]" gridcol="1" gridrow="25" pageId="21" pageNumber="172">1.88</td>
|
||
<td box="[401,474,1380,1402]" gridcol="2" gridrow="25" pageId="21" pageNumber="172">1.60</td>
|
||
<td box="[503,576,1380,1402]" gridcol="3" gridrow="25" pageId="21" pageNumber="172">1.75</td>
|
||
<td box="[606,679,1380,1402]" gridcol="4" gridrow="25" pageId="21" pageNumber="172">2.00</td>
|
||
<td box="[708,781,1380,1402]" gridcol="5" gridrow="25" pageId="21" pageNumber="172">1.68</td>
|
||
<td box="[811,884,1380,1402]" gridcol="6" gridrow="25" pageId="21" pageNumber="172">1.40</td>
|
||
<td box="[911,997,1380,1402]" gridcol="7" gridrow="25" pageId="21" pageNumber="172">1.60</td>
|
||
<td box="[1024,1097,1380,1402]" gridcol="8" gridrow="25" pageId="21" pageNumber="172">1.27</td>
|
||
<td box="[1127,1200,1380,1402]" gridcol="9" gridrow="25" pageId="21" pageNumber="172">1.70</td>
|
||
<td box="[1238,1293,1380,1402]" gridcol="10" gridrow="25" pageId="21" pageNumber="172">1.65</td>
|
||
<td box="[1327,1423,1380,1402]" gridcol="11" gridrow="25" pageId="21" pageNumber="172">1.27–2.00</td>
|
||
</tr>
|
||
<tr box="[164,1423,1414,1437]" gridrow="26" pageId="21" pageNumber="172">
|
||
<th box="[164,270,1414,1437]" gridcol="0" gridrow="26" pageId="21" pageNumber="172">
|
||
<emphasis bold="true" box="[172,262,1414,1436]" pageId="21" pageNumber="172">3FD/4TD</emphasis>
|
||
</th>
|
||
<td box="[298,371,1414,1437]" gridcol="1" gridrow="26" pageId="21" pageNumber="172">1.25</td>
|
||
<td box="[401,474,1414,1437]" gridcol="2" gridrow="26" pageId="21" pageNumber="172">1.33</td>
|
||
<td box="[503,576,1414,1437]" gridcol="3" gridrow="26" pageId="21" pageNumber="172">1.27</td>
|
||
<td box="[606,679,1414,1437]" gridcol="4" gridrow="26" pageId="21" pageNumber="172">1.14</td>
|
||
<td box="[708,781,1414,1437]" gridcol="5" gridrow="26" pageId="21" pageNumber="172">1.14</td>
|
||
<td box="[811,884,1414,1437]" gridcol="6" gridrow="26" pageId="21" pageNumber="172">1.08</td>
|
||
<td box="[911,997,1414,1437]" gridcol="7" gridrow="26" pageId="21" pageNumber="172">1.23</td>
|
||
<td box="[1024,1097,1414,1437]" gridcol="8" gridrow="26" pageId="21" pageNumber="172">1.08</td>
|
||
<td box="[1127,1200,1414,1437]" gridcol="9" gridrow="26" pageId="21" pageNumber="172">1.31</td>
|
||
<td box="[1238,1293,1414,1437]" gridcol="10" gridrow="26" pageId="21" pageNumber="172">1.20</td>
|
||
<td box="[1327,1423,1414,1437]" gridcol="11" gridrow="26" pageId="21" pageNumber="172">1.08–1.33</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1437,1482,2013]" pageId="21" pageNumber="172">
|
||
Skin of dorsum and dorsal surfaces of limbs very finely granular (
|
||
<figureCitation box="[920,1022,1482,1509]" captionStart="FIGURE 11" captionStartId="20.[151,250,1619,1643]" captionTargetBox="[176,1412,453,1594]" captionTargetId="figure-139@20.[176,1412,453,1595]" captionTargetPageId="20" captionText="FIGURE 11. Litoria lisae sp. nov. in life, A) holotype (SAMA R71689), B–C), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800946" httpUri="https://zenodo.org/record/7800946/files/figure.png" pageId="21" pageNumber="172">Fig. 11A</figureCitation>
|
||
). Ventrally throat, chest and abdomen coarsely granular; ventral surfaces of limbs with scattered low tubercles, most prominent on posteroventral margins of thighs. Broken white dermal ridge below vent not extending onto adjacent regions of upper thighs; clump of ~15 prominent white tubercles below dermal ridge; heels with five (right) and seven (left) low, white tubercles; outer edge of foot with series of low, pale tubercles extending from heel nearly to distal subarticular tubercle of Toe 5, those on right tarsus partially fused to form short (
|
||
<quantity box="[701,788,1662,1689]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="172" unit="mm" value="3.5">3.5 mm</quantity>
|
||
) ridge; outer edge of hand with row of small pale tubercles extending from mid-point along forearm to distal subarticular tubercle on Finger 4.
|
||
</paragraph>
|
||
<paragraph blockId="21.[151,1437,1482,2013]" lastBlockId="22.[151,1437,150,321]" lastPageId="22" lastPageNumber="173" pageId="21" pageNumber="172">
|
||
Fingers moderately short with expanded terminal discs (3FD/SVL 0.05; 3FD/3FP 1.88) with distinct marginal grooves; subarticular tubercles strongly bilobed; relative lengths of fingers: 3>4>2>1 (
|
||
<figureCitation box="[1109,1211,1770,1797]" captionStart="FIGURE 12" captionStartId="22.[151,250,1829,1853]" captionTargetBox="[160,1427,345,1806]" captionTargetId="figure-118@22.[160,1427,345,1806]" captionTargetPageId="22" captionText="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="21" pageNumber="172">Fig. 12C</figureCitation>
|
||
). Webbing on inside of Finger 4 reaching to just above distal subarticular tubercle, on outside of Finger 3 reaching to top of distal subarticular tubercle and continuing to disc as thick flange (
|
||
<figureCitation box="[851,956,1842,1869]" captionStart="FIGURE 12" captionStartId="22.[151,250,1829,1853]" captionTargetBox="[160,1427,345,1806]" captionTargetId="figure-118@22.[160,1427,345,1806]" captionTargetPageId="22" captionText="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="21" pageNumber="172">Fig. 12C</figureCitation>
|
||
), on inside of Finger 3 reaching to small proximal subarticular tubercle, on outside of Finger 2 reaching to halfway between distal subarticular tubercle and disc, and between fingers 1 and 2 limited to basal fringe. Hand with well-defined narrow inner, and less well defined but detectable outer, metacarpal tubercles. Nuptial pad on Finger 1 pale brown, elongate (
|
||
<quantity box="[1179,1267,1950,1977]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="21" pageNumber="172" unit="mm" value="1.5">1.5 mm</quantity>
|
||
long), broader distally than proximally. Toes with expanded terminal discs with terminal grooves, smaller than those on fingers (4TD/SVL 0.04, 3FD/4TD 1.25) (
|
||
<figureCitation box="[544,651,151,177]" captionStart="FIGURE 12" captionStartId="22.[151,250,1829,1853]" captionTargetBox="[160,1427,345,1806]" captionTargetId="figure-118@22.[160,1427,345,1806]" captionTargetPageId="22" captionText="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="22" pageNumber="173">Fig. 12D</figureCitation>
|
||
); relative lengths: 4>5=3>2>1. Webbing on inside of Toe 5 reaches nearly to disc, on both sides of Toe 4 to base of distal subarticular tubercle, on outside of Toe 3 nearly to base of disc, on inside of Toe 3 to halfway between proximal and distal subarticular tubercles, on outside of Toe 2 to slightly more than halfway to distal subarticular tubercle, and between toes 1 and 2 reduced to a basal fringe. Foot with welldefined oval inner metatarsal tubercle, poorly defined outer tubercle (
|
||
<figureCitation box="[929,1034,295,321]" captionStart="FIGURE 12" captionStartId="22.[151,250,1829,1853]" captionTargetBox="[160,1427,345,1806]" captionTargetId="figure-118@22.[160,1427,345,1806]" captionTargetPageId="22" captionText="FIGURE 12. Litoria lisae sp. nov. holotype (SAMA R71689), A) dorsal and B) lateral view of head (scale bars = 5 mm), C) palmar view of hand, D) plantar view of foot (scale bars = 1 mm)." figureDoi="http://doi.org/10.5281/zenodo.7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="22" pageNumber="173">Fig. 12D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7800948" ID-Zenodo-Dep="7800948" httpUri="https://zenodo.org/record/7800948/files/figure.png" pageId="22" pageNumber="173" startId="22.[151,250,1829,1853]" targetBox="[160,1427,345,1806]" targetPageId="22">
|
||
<paragraph blockId="22.[151,1436,1829,1890]" pageId="22" pageNumber="173">
|
||
<emphasis bold="true" box="[151,282,1829,1853]" pageId="22" pageNumber="173">FIGURE 12</emphasis>
|
||
.
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[296,421,1830,1854]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="22" pageNumber="173" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[296,421,1830,1854]" italics="true" pageId="22" pageNumber="173">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[428,509,1830,1854]" pageId="22" pageNumber="173">
|
||
<taxonomicNameLabel box="[428,509,1830,1854]" pageId="22" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype (SAMA R71689),
|
||
<emphasis bold="true" box="[802,820,1829,1853]" pageId="22" pageNumber="173">A</emphasis>
|
||
) dorsal and
|
||
<emphasis bold="true" box="[949,975,1829,1853]" pageId="22" pageNumber="173">B)</emphasis>
|
||
lateral view of head (scale bars = 5 mm),
|
||
<emphasis bold="true" box="[1409,1427,1829,1853]" pageId="22" pageNumber="173">C</emphasis>
|
||
) palmar view of hand,
|
||
<emphasis bold="true" box="[373,391,1866,1890]" pageId="22" pageNumber="173">D</emphasis>
|
||
) plantar view of foot (scale bars = 1 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="22.[151,1436,1936,2034]" lastBlockId="23.[151,1437,151,861]" lastPageId="23" lastPageNumber="174" pageId="22" pageNumber="173">
|
||
<emphasis box="[199,353,1936,1962]" italics="true" pageId="22" pageNumber="173">Colour in life</emphasis>
|
||
(
|
||
<figureCitation box="[370,475,1936,1962]" captionStart="FIGURE 11" captionStartId="20.[151,250,1619,1643]" captionTargetBox="[176,1412,453,1594]" captionTargetId="figure-139@20.[176,1412,453,1595]" captionTargetPageId="20" captionText="FIGURE 11. Litoria lisae sp. nov. in life, A) holotype (SAMA R71689), B–C), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800946" httpUri="https://zenodo.org/record/7800946/files/figure.png" pageId="22" pageNumber="173">Fig. 11A</figureCitation>
|
||
). Dorsal surfaces lime green becoming yellow green laterally and on dorsal surfaces of limbs, green on forelimbs terminating sharply at wrists; dorsal and lateral surfaces with scattered darker green spots; white to yellowish-white bar extends from beneath eye posteriorly to base of forearm; white mid-lateral stripe extends from approximately midway between front and hind limbs to groin; ventrally white anteriorly, except broad pale-brown band around edges of throat, extending onto proximal portions of forelimbs; abdomen yellow posterolaterally and in hidden surfaces of limbs and axillae, yellow in axillae extending approximately halfway along posterior surfaces of forearms; enlarged tubercles on outer edges of limbs and around vent white. Dorsal surfaces of hands and feet stippled with brown spots and blotches, patches of yellowish green dorsally on fingers 3 and 4. Iris ivory with paler ivory rim around pupil and dense dark brown reticulations.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,151,861]" pageId="23" pageNumber="174">
|
||
<emphasis box="[199,455,367,393]" italics="true" pageId="23" pageNumber="174">Colour in preservative.</emphasis>
|
||
In preservative dorsal and lateral surfaces mottled darker and lighter blue, brown pigment on throat prominent, much darker than in life; patch of brown pigment midlaterally that was nearly undetectable in life now evident; yellow of posteroventral surfaces and limbs now creamy, pale bar under eye white, more prominent than in life; throat and abdomen creamy with white tubercles.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,151,861]" pageId="23" pageNumber="174">
|
||
<emphasis bold="true" box="[199,319,510,536]" pageId="23" pageNumber="174">Variation.</emphasis>
|
||
All of the
|
||
<typeStatus box="[447,554,511,537]" pageId="23" pageNumber="174">paratypes</typeStatus>
|
||
are adult males. Morphometric variation is limited (
|
||
<tableCitation box="[1154,1241,511,537]" captionStart="TABLE 4" captionStartId="21.[151,239,476,500]" captionTargetPageId="21" captionText="TABLE 4. Measurements and ratios of the type series of Litoria lisae sp. nov. SAMA R71689 is the holotype." pageId="23" pageNumber="174">Table 4</tableCitation>
|
||
) but snout shape is variable –the
|
||
<typeStatus box="[330,428,547,573]" pageId="23" pageNumber="174">holotype</typeStatus>
|
||
has the most narrowly rounded snout of the type series – in most specimens the tip of the snout is bluntly rounded or in
|
||
<specimenCount box="[499,667,583,609]" pageId="23" pageNumber="174" type="generic">two specimens</specimenCount>
|
||
(SAMA R60716, R71690) near-truncate. All specimens exhibit the distinct white bar beneath the eyes, and in life are yellow on the hidden surfaces of the legs and have extensive areas of yellow on the posterior surfaces of the abdomen (
|
||
<figureCitation box="[743,845,655,681]" captionStart="FIGURE 11" captionStartId="20.[151,250,1619,1643]" captionTargetBox="[176,1412,453,1594]" captionTargetId="figure-139@20.[176,1412,453,1595]" captionTargetPageId="20" captionText="FIGURE 11. Litoria lisae sp. nov. in life, A) holotype (SAMA R71689), B–C), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800946" httpUri="https://zenodo.org/record/7800946/files/figure.png" pageId="23" pageNumber="174">Fig. 11C</figureCitation>
|
||
). However, dorsal and lateral colour in life is highly variable with some specimens being much darker green than the
|
||
<typeStatus box="[898,999,691,717]" pageId="23" pageNumber="174">holotype</typeStatus>
|
||
, or with more prominent darker green spots and large brown patches dorsally or laterally (e.g., SAMA R71690,
|
||
<figureCitation box="[960,1059,727,753]" captionStart="FIGURE 11" captionStartId="20.[151,250,1619,1643]" captionTargetBox="[176,1412,453,1594]" captionTargetId="figure-139@20.[176,1412,453,1595]" captionTargetPageId="20" captionText="FIGURE 11. Litoria lisae sp. nov. in life, A) holotype (SAMA R71689), B–C), paratype SAMA R71690 in dorsolateral and ventral views. Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800946" httpUri="https://zenodo.org/record/7800946/files/figure.png" pageId="23" pageNumber="174">Fig. 11B</figureCitation>
|
||
). In preservative types range from pale to dark blue with scattered darker flecks and blotches dorsally; intensity of dark-brown pigmentation on throat is also variable, ranging from pale brown clumps of pigmentation in SAMA R71686, to intense dark brown band of pigmentation laterally on the throat in the
|
||
<typeStatus box="[623,721,835,861]" pageId="23" pageNumber="174">holotype</typeStatus>
|
||
and SAMA R71690.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7800952" ID-Zenodo-Dep="7800952" httpUri="https://zenodo.org/record/7800952/files/figure.png" pageId="23" pageNumber="174" startId="23.[151,250,1280,1304]" targetBox="[174,1415,890,1252]" targetPageId="23">
|
||
<paragraph blockId="23.[151,1436,1280,1376]" pageId="23" pageNumber="174">
|
||
<emphasis bold="true" box="[151,279,1280,1304]" pageId="23" pageNumber="174">FIGURE 13</emphasis>
|
||
. Wave form (amplitude over time; top) and spectrogram (frequency over time; bottom) of two types of advertisement call produced by
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[334,461,1316,1340]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[334,461,1316,1340]" italics="true" pageId="23" pageNumber="174">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[470,553,1316,1340]" pageId="23" pageNumber="174">
|
||
<taxonomicNameLabel box="[470,553,1316,1340]" pageId="23" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(SAMA R71689) recorded at an air temperature of 20.5°C. Note distinctly clumped nature of pulses in chirp call (left) compared to buzz call (right).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="23.[151,1437,1422,2025]" pageId="23" pageNumber="174">
|
||
<emphasis bold="true" box="[199,426,1422,1449]" pageId="23" pageNumber="174">Advertisement call.</emphasis>
|
||
We recorded 17 calls produced by the
|
||
<typeStatus box="[854,950,1422,1448]" pageId="23" pageNumber="174">holotype</typeStatus>
|
||
at an air temperature of 22.0°C and 13 calls produced by
|
||
<collectionCode box="[292,370,1459,1485]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="23" pageNumber="174" type="Museum">SAMA</collectionCode>
|
||
<specimenCode box="[375,462,1458,1484]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="23" pageNumber="174">R71688</specimenCode>
|
||
at an air temperature of 20.5°
|
||
<collectorName box="[784,869,1458,1485]" pageId="23" pageNumber="174">C. Calls</collectorName>
|
||
of both animals were similar so data were combined for analysis but, where relevant, differences are noted.
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[752,887,1494,1520]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="23" pageNumber="174" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[752,887,1494,1520]" italics="true" pageId="23" pageNumber="174">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[894,982,1495,1521]" pageId="23" pageNumber="174">
|
||
<taxonomicNameLabel box="[894,982,1495,1521]" pageId="23" pageNumber="174" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
produces two distinct call types (
|
||
<figureCitation box="[1348,1428,1494,1521]" captionStart="FIGURE 13" captionStartId="23.[151,250,1280,1304]" captionTargetBox="[174,1415,890,1252]" captionTargetId="figure-357@23.[168,1420,885,1257]" captionTargetPageId="23" captionText="FIGURE 13. Wave form (amplitude over time; top) and spectrogram (frequency over time; bottom) of two types of advertisement call produced by Litoria lisae sp. nov. (SAMA R71689) recorded at an air temperature of 20.5°C. Note distinctly clumped nature of pulses in chirp call (left) compared to buzz call (right)." figureDoi="http://doi.org/10.5281/zenodo.7800952" httpUri="https://zenodo.org/record/7800952/files/figure.png" pageId="23" pageNumber="174">Fig. 13</figureCitation>
|
||
) at highly variable intervals (inter call interval 0.42–
|
||
<geoCoordinate box="[727,809,1531,1557]" degrees="31.22" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-31.22">31.22 s</geoCoordinate>
|
||
; mean =
|
||
<geoCoordinate box="[915,984,1530,1557]" degrees="6.55" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-6.55">6.55 s</geoCoordinate>
|
||
, SD = 7.22, n = 27), and there appeared to be no pattern to the sequence in which the two call types were produced.
|
||
<collectorName box="[994,1049,1566,1592]" pageId="23" pageNumber="174">Each</collectorName>
|
||
call type is a single, short note but the rate and pattern of pulse production within notes differ markedly.
|
||
<collectorName box="[913,956,1602,1628]" pageId="23" pageNumber="174">The</collectorName>
|
||
first call type is a short ‘buzz’ comprising a rapidly repeated series of evenly spaced pulses, with approximately
|
||
<geoCoordinate box="[906,968,1638,1664]" degrees="0.006" direction="north" orientation="latitude" pageId="23" pageNumber="174" precision="55" value="0.006">0.006</geoCoordinate>
|
||
<geoCoordinate box="[967,1042,1638,1664]" degrees="0.009" direction="west" orientation="longitude" pageId="23" pageNumber="174" precision="55" value="-0.009">–0.009</geoCoordinate>
|
||
s between pulses, while the second call type is a more slowly repeated series of pulses which are ‘clumped’ into discrete groups of 2–3 (
|
||
<figureCitation box="[1289,1370,1674,1701]" captionStart="FIGURE 13" captionStartId="23.[151,250,1280,1304]" captionTargetBox="[174,1415,890,1252]" captionTargetId="figure-357@23.[168,1420,885,1257]" captionTargetPageId="23" captionText="FIGURE 13. Wave form (amplitude over time; top) and spectrogram (frequency over time; bottom) of two types of advertisement call produced by Litoria lisae sp. nov. (SAMA R71689) recorded at an air temperature of 20.5°C. Note distinctly clumped nature of pulses in chirp call (left) compared to buzz call (right)." figureDoi="http://doi.org/10.5281/zenodo.7800952" httpUri="https://zenodo.org/record/7800952/files/figure.png" pageId="23" pageNumber="174">Fig. 13</figureCitation>
|
||
) with approximately
|
||
<geoCoordinate box="[319,401,1711,1737]" degrees="0.004" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="55" value="-0.004">0.004 s</geoCoordinate>
|
||
between pulses within groups and
|
||
<geoCoordinate box="[802,870,1711,1737]" degrees="0.03" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-0.03">0.03 s</geoCoordinate>
|
||
between discrete pulse groups.
|
||
<collectorName box="[1231,1335,1710,1736]" pageId="23" pageNumber="174">Although</collectorName>
|
||
the total length of both call types overlaps extensively the first call type contains more pulses that are produced at a faster rate.
|
||
<collectorName box="[151,187,1783,1809]" pageId="23" pageNumber="174">We</collectorName>
|
||
here refer to call type 1 as buzz calls, and call type 2 as chirp calls; they are each described in more detail below.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1422,2025]" pageId="23" pageNumber="174">
|
||
Eleven buzz calls from both animals combined were 0.12–
|
||
<geoCoordinate box="[855,921,1819,1845]" degrees="2.34" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-2.34">2.34 s</geoCoordinate>
|
||
long (mean 0.18, SD 0.03) and contained 10– 22 pulses (mean 16.36, SD 3.69) produced at a rate of 72.3–103.9 pulses/s (mean 87.62, SD 10.53). Although the structure of buzz calls produced by the
|
||
<typeStatus box="[594,692,1890,1916]" pageId="23" pageNumber="174">holotype</typeStatus>
|
||
and SAMA R71688 was extremely similar, the three buzz calls of SAMA R71688 were generally shorter (0.13–
|
||
<geoCoordinate box="[662,729,1927,1953]" degrees="0.17" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-0.17">0.17 s</geoCoordinate>
|
||
vs. 0.16–
|
||
<geoCoordinate box="[838,905,1926,1953]" degrees="0.24" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-0.24">0.24 s</geoCoordinate>
|
||
) and contained fewer pulses (10–13 vs. 16–22) produced at a slower rate (72.29–74.38 vs. 83.76–103.89 pulses/s) than eight buzz calls produced by the
|
||
<typeStatus box="[1325,1427,1962,1988]" pageId="23" pageNumber="174">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="23.[151,1437,1422,2025]" lastBlockId="24.[151,1437,150,2013]" lastPageId="24" lastPageNumber="175" pageId="23" pageNumber="174">
|
||
Nineteen chirp calls from both animals combined were 0.11–
|
||
<geoCoordinate box="[860,925,1999,2025]" degrees="0.17" direction="south" orientation="latitude" pageId="23" pageNumber="174" precision="555" value="-0.17">0.17 s</geoCoordinate>
|
||
long (mean 0.15, SD 0.02) and contained 6–12 pulses (mean 8.68, SD 1.82) produced at a rate of 37.26–69.56 pulses/s (mean 52.58, SD 9.71). Again, the structure of chirp calls produced by the
|
||
<typeStatus box="[483,579,187,213]" pageId="24" pageNumber="175">holotype</typeStatus>
|
||
and SAMA R71688 was extremely similar and there was extensive overlap in call length between the 10 chirp calls produced by SAMA R71688 (
|
||
<geoCoordinate box="[895,957,223,249]" degrees="0.107" direction="north" orientation="latitude" pageId="24" pageNumber="175" precision="55" value="0.107">0.107</geoCoordinate>
|
||
<geoCoordinate box="[957,1035,223,249]" degrees="0.166" direction="west" orientation="longitude" pageId="24" pageNumber="175" precision="55" value="-0.166">–0.166</geoCoordinate>
|
||
s) and the
|
||
<typeStatus box="[1154,1250,223,249]" pageId="24" pageNumber="175">holotype</typeStatus>
|
||
(
|
||
<geoCoordinate box="[1265,1327,223,249]" degrees="0.118" direction="north" orientation="latitude" pageId="24" pageNumber="175" precision="55" value="0.118">0.118</geoCoordinate>
|
||
<geoCoordinate box="[1326,1402,223,249]" degrees="0.176" direction="west" orientation="longitude" pageId="24" pageNumber="175" precision="55" value="-0.176">–0.176</geoCoordinate>
|
||
s). However, chirp calls of SAMA R71688 contained fewer pulses (6–8 vs. 9–12) produced at a slower rate (37.26–55.04 vs. 52.63–69.56 pulses/s) than the
|
||
<typeStatus box="[531,631,295,321]" pageId="24" pageNumber="175">holotype</typeStatus>
|
||
. Whether these call types serve different acoustic functions is not known.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,150,2013]" pageId="24" pageNumber="175">
|
||
<emphasis bold="true" box="[199,365,330,356]" pageId="24" pageNumber="175">Comparisons.</emphasis>
|
||
In its moderate size (male SVL 27.0–
|
||
<quantity box="[800,904,331,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="24" pageNumber="175" unit="mm" value="29.7">29.7 mm</quantity>
|
||
), slender body, green and brown dorsal colour, extensively webbed fingers, and males lacking a rostral spike,
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[876,1016,367,393]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[876,1016,367,393]" italics="true" pageId="24" pageNumber="175">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1026,1118,367,393]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[1026,1118,367,393]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
most closely resembles the following 11 species:
|
||
<taxonomicName authorityName="Richards & Donnellan" authorityYear="2020" box="[393,486,403,429]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="aplini">
|
||
<emphasis box="[393,486,403,429]" italics="true" pageId="24" pageNumber="175">L. aplini</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[499,645,403,429]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="daraiensis" status="sp. nov.">
|
||
<emphasis box="[499,645,403,429]" italics="true" pageId="24" pageNumber="175">L. daraiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[650,738,403,429]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[650,738,403,429]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[751,865,403,429]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis box="[751,865,403,429]" italics="true" pageId="24" pageNumber="175">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[878,1072,403,429]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="haematogaster" status="sp. nov.">
|
||
<emphasis box="[878,1072,403,429]" italics="true" pageId="24" pageNumber="175">L. haematogaster</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1078,1166,403,429]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[1078,1166,403,429]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1962" box="[1179,1244,403,428]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="iris">
|
||
<emphasis box="[1179,1244,403,428]" italics="true" pageId="24" pageNumber="175">L. iris</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Johnston & Richards" authorityYear="1994" box="[1257,1401,403,429]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="majikthise">
|
||
<emphasis box="[1257,1401,403,429]" italics="true" pageId="24" pageNumber="175">L. majikthise</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName baseAuthorityName="Meyer" baseAuthorityYear="1875" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="nigropunctata">
|
||
<emphasis box="[1414,1430,403,428]" italics="true" pageId="24" pageNumber="175">L</emphasis>
|
||
.
|
||
<emphasis box="[151,309,439,464]" italics="true" pageId="24" pageNumber="175">nigropunctata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Dennis & Cunningham" authorityYear="2006" box="[322,453,439,465]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="richardsi">
|
||
<emphasis box="[322,453,439,465]" italics="true" pageId="24" pageNumber="175">L. richardsi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Richards" authorityYear="2005" box="[466,623,439,465]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="singadanae">
|
||
<emphasis box="[466,623,439,465]" italics="true" pageId="24" pageNumber="175">L. singadanae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Gunther" authorityYear="2004" box="[637,780,439,464]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="umarensis">
|
||
<emphasis box="[637,780,439,464]" italics="true" pageId="24" pageNumber="175">L. umarensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Gunther" authorityYear="2004" box="[833,923,439,464]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="verae">
|
||
<emphasis box="[833,923,439,464]" italics="true" pageId="24" pageNumber="175">L. verae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[937,1074,439,465]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[937,1074,439,465]" italics="true" pageId="24" pageNumber="175">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1080,1169,439,465]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[1080,1169,439,465]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is also morphologically similar to the new species described below and is compared with it in that species’ account.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,150,2013]" pageId="24" pageNumber="175">
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[199,337,511,537]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[199,337,511,537]" italics="true" pageId="24" pageNumber="175">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[346,436,511,537]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[346,436,511,537]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from
|
||
<taxonomicName authorityName="Richards & Donnellan" authorityYear="2020" box="[746,842,511,537]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="aplini">
|
||
<emphasis box="[746,842,511,537]" italics="true" pageId="24" pageNumber="175">L. aplini</emphasis>
|
||
</taxonomicName>
|
||
by its smaller body size (adult males 27.0–
|
||
<quantity box="[1337,1436,511,537]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="24" pageNumber="175" unit="mm" value="29.7">29.7 mm</quantity>
|
||
vs. males
|
||
<quantity box="[267,430,546,573]" metricMagnitude="-2" metricUnit="m" metricValue="3.35" metricValueMax="3.51" metricValueMin="3.19" pageId="24" pageNumber="175" unit="mm" value="33.5" valueMax="35.1" valueMin="31.9">31.9–35.1 mm</quantity>
|
||
SVL), less prominent dermal fold along outer edges of limbs, hidden surfaces of limbs predominantly yellow (vs. predominantly blue with dark-brown mottling), and advertisement call a single buzz or chirp containing either evenly spaced or discretely clumped pulses (vs. advertisement call a short buzz normally followed by 1–7 clicks); from
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[494,640,655,681]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="daraiensis" status="sp. nov.">
|
||
<emphasis box="[494,640,655,681]" italics="true" pageId="24" pageNumber="175">L. daraiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[647,737,655,681]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[647,737,655,681]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by its larger size (male SVL> 27.0 mm vs.
|
||
<quantity box="[1237,1341,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="2.3899999999999997" pageId="24" pageNumber="175" unit="mm" value="23.9">23.9 mm</quantity>
|
||
), having (vs. lacking) a distinct white bar below the eye, and distinct advertisement call (a single buzz or chirp containing either evenly spaced or discretely clumped pulses vs. 10–11 distinctly pulsed chattering notes); from
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[1281,1396,727,753]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="gracilis" status="sp. nov.">
|
||
<emphasis box="[1281,1396,727,753]" italics="true" pageId="24" pageNumber="175">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by having a predominantly green dorsum with darker green and brown spots (vs. predominantly brown with green spots), having (vs. lacking) a distinct white bar beneath the eye, and advertisement call a single buzz or chirp containing either evenly spaced or discretely clumped pulses (vs. call containing 2–5 distinctly pulsed notes); from
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="haematogaster" status="sp. nov.">
|
||
<emphasis italics="true" pageId="24" pageNumber="175">L. haematogaster</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[322,410,871,897]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[322,410,871,897]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by lacking (vs. having) large conical tubercles along outer edge of tarsi and arms, and hidden surfaces of limbs yellow (vs. red); from
|
||
<taxonomicName baseAuthorityName="Tyler" baseAuthorityYear="1962" box="[587,652,907,932]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="iris">
|
||
<emphasis box="[587,652,907,932]" italics="true" pageId="24" pageNumber="175">L. iris</emphasis>
|
||
</taxonomicName>
|
||
by having hidden surfaces of limbs predominantly yellow (vs. posterior of thighs blue, red, or yellow, frequently blotched with white or purple), violet spots in axilla and groin absent (vs. present) and advertisement call a single buzz or chirp containing either evenly spaced or discretely clumped pulses (vs. advertisement call a series of up to 10 notes of variable length, with long notes preceding or following short notes); from
|
||
<taxonomicName authorityName="Johnston & Richards" authorityYear="1994" box="[300,445,1051,1077]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="majikthise">
|
||
<emphasis box="[300,445,1051,1077]" italics="true" pageId="24" pageNumber="175">L. majikthise</emphasis>
|
||
</taxonomicName>
|
||
by its smaller size (
|
||
<taxonomicName authorityName="Johnston & Richards" authorityYear="1994" box="[681,827,1051,1077]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="majikthise">
|
||
<emphasis box="[681,696,1051,1076]" italics="true" pageId="24" pageNumber="175">L</emphasis>
|
||
.
|
||
<emphasis box="[713,827,1051,1077]" italics="true" pageId="24" pageNumber="175">majikthise</emphasis>
|
||
</taxonomicName>
|
||
males
|
||
<quantity box="[910,1032,1050,1077]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.5" metricValueMin="3.0" pageId="24" pageNumber="175" unit="mm" value="32.5" valueMax="35.0" valueMin="30.0">30–35 mm</quantity>
|
||
SVL), having posterior surfaces of thighs yellow (vs. uniform red), and lacking violet patches on posteroventral surfaces of abdomen (vs. present); from
|
||
<taxonomicName baseAuthorityName="Meyer" baseAuthorityYear="1875" box="[214,402,1123,1149]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="nigropunctata">
|
||
<emphasis box="[214,229,1123,1148]" italics="true" pageId="24" pageNumber="175">L</emphasis>
|
||
.
|
||
<emphasis box="[246,402,1123,1148]" italics="true" pageId="24" pageNumber="175">nigropunctata</emphasis>
|
||
</taxonomicName>
|
||
by having dermal folds along outer margins of tarsi distinctly crenulated (vs. dermal folds forming low dermal ridge), prominent white bar below eye present (vs. absent) and advertisement call a single buzz or chirp containing either evenly spaced or discretely clumped pulses (vs. advertisement call comprising an irregular succession of clicks and buzzes;
|
||
<bibRefCitation author="Menzies, J. I." box="[526,688,1231,1257]" pageId="24" pageNumber="175" pagination="291 - 300" refId="ref24254" refString="Menzies, J. I. (1972) Papuan tree frogs of the Litoria nigropunctata group. Herpetologica, 28, 291 - 300." type="journal article" year="1972">Menzies 1972</bibRefCitation>
|
||
); from
|
||
<taxonomicName authorityName="Dennis & Cunningham" authorityYear="2006" box="[773,905,1231,1257]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="richardsi">
|
||
<emphasis box="[773,905,1231,1257]" italics="true" pageId="24" pageNumber="175">L. richardsi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Richards" authorityYear="2005" box="[963,1122,1231,1257]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="singadanae">
|
||
<emphasis box="[963,1122,1231,1257]" italics="true" pageId="24" pageNumber="175">L. singadanae</emphasis>
|
||
</taxonomicName>
|
||
in having a smaller (TYM/ EYE 0.39–0.50 vs. 0.65–0.81), pigmented (vs. substantially transparent) tympanum, and further from
|
||
<taxonomicName authorityName="Dennis & Cunningham" authorityYear="2006" box="[1305,1437,1267,1293]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="richardsi">
|
||
<emphasis box="[1305,1437,1267,1293]" italics="true" pageId="24" pageNumber="175">L. richardsi</emphasis>
|
||
</taxonomicName>
|
||
in lacking (vs. having) irregular black lines on dorsum and extensive black markings ventrolaterally, and from
|
||
<taxonomicName authorityName="Richards" authorityYear="2005" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="singadanae">
|
||
<emphasis italics="true" pageId="24" pageNumber="175">L. singadanae</emphasis>
|
||
</taxonomicName>
|
||
in lacking (vs. having) extensive area of orange on posteroventral surfaces in life; from
|
||
<taxonomicName authorityName="Gunther" authorityYear="2004" box="[1292,1436,1339,1364]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="umarensis">
|
||
<emphasis box="[1292,1436,1339,1364]" italics="true" pageId="24" pageNumber="175">L. umarensis</emphasis>
|
||
</taxonomicName>
|
||
in having posterior surfaces of thighs yellow (vs. brown), and dorsum with green and brown spots (vs. normally uniform green); and from
|
||
<taxonomicName authorityName="Gunther" authorityYear="2004" box="[447,540,1411,1436]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="verae">
|
||
<emphasis box="[447,540,1411,1436]" italics="true" pageId="24" pageNumber="175">L. verae</emphasis>
|
||
</taxonomicName>
|
||
in its smaller size (
|
||
<taxonomicName authorityName="Gunther" authorityYear="2004" box="[763,856,1411,1436]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="verae">
|
||
<emphasis box="[763,856,1411,1436]" italics="true" pageId="24" pageNumber="175">L. verae</emphasis>
|
||
</taxonomicName>
|
||
males
|
||
<quantity box="[937,1058,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" pageId="24" pageNumber="175" unit="mm" value="34.0" valueMax="35.0" valueMin="33.0">33–35 mm</quantity>
|
||
SVL), lacking (vs. having) small brown spots aligned transversely on dorsum, and having yellow (vs. orange) on posteroventral surfaces in life.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="175" type="distribution">
|
||
<paragraph blockId="24.[151,1437,150,2013]" pageId="24" pageNumber="175">
|
||
<emphasis bold="true" box="[199,494,1482,1509]" pageId="24" pageNumber="175">Distribution and ecology.</emphasis>
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[501,637,1483,1509]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[501,637,1483,1509]" italics="true" pageId="24" pageNumber="175">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[644,732,1483,1509]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[644,732,1483,1509]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from lower montane forest on Gobe Ridge and Iagifu Ridge (including Arakubi) in the Kikori River basin of southern
|
||
<collectingCountry box="[859,1071,1519,1545]" name="Papua New Guinea" pageId="24" pageNumber="175">Papua New Guinea</collectingCountry>
|
||
(
|
||
<figureCitation box="[1086,1153,1519,1545]" captionStart="FIGURE 4" captionStartId="9.[151,250,1739,1763]" captionTargetBox="[151,1431,676,1716]" captionTargetId="figure-179@9.[151,1436,676,1716]" captionTargetPageId="9" captionText="FIGURE 4. Map showing the distributions of Litoria species newly described species in this paper, and Litoria singadanae from the Huon Peninsula, Papua New Guniea. Blue square = Litoria daraiensis sp. nov., yellow triangle = Litoria gracilis sp. nov., red circle = Litoria haematogaster sp. nov., pale blue circles = Litoria lisae sp. nov., pink diamond = Litoria naispela sp. nov. and purple diamond = Litoria singadanae. Arrows indicate the type localities for Litoria gracilis sp. nov. and Litoria lisae sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.7800926" httpUri="https://zenodo.org/record/7800926/files/figure.png" pageId="24" pageNumber="175">Fig. 4</figureCitation>
|
||
), where they appear to be restricted to limestone karst habitats (
|
||
<figureCitation box="[559,640,1555,1581]" captionStart="FIGURE 14" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[327,1260,309,1913]" captionTargetId="figure-88@25.[327,1260,309,1913]" captionTargetPageId="25" captionText="FIGURE 14. Lower montane forest on limestone substrate at Iagifu Ridge, Papua New Guinea. Males of Litoria lisae sp. nov. called from low foliage in the forest along the trail illustrated A), and from deep within crevices and sinkholes in the karst, such as the large crevice indicated in B). Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800954" httpUri="https://zenodo.org/record/7800954/files/figure.png" pageId="24" pageNumber="175">Fig. 14</figureCitation>
|
||
). Although several individuals were seen on foliage near a shallow forest pool, calling males were encountered only on low foliage in areas with no evidence of surface water suitable for egg deposition (
|
||
<figureCitation box="[279,378,1627,1653]" captionStart="FIGURE 14" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[327,1260,309,1913]" captionTargetId="figure-88@25.[327,1260,309,1913]" captionTargetPageId="25" captionText="FIGURE 14. Lower montane forest on limestone substrate at Iagifu Ridge, Papua New Guinea. Males of Litoria lisae sp. nov. called from low foliage in the forest along the trail illustrated A), and from deep within crevices and sinkholes in the karst, such as the large crevice indicated in B). Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800954" httpUri="https://zenodo.org/record/7800954/files/figure.png" pageId="24" pageNumber="175">Fig. 14A</figureCitation>
|
||
) and have been heard vocalising from hidden refuges deep within limestone sinkholes (
|
||
<figureCitation box="[1324,1424,1627,1653]" captionStart="FIGURE 14" captionStartId="25.[151,250,1936,1960]" captionTargetBox="[327,1260,309,1913]" captionTargetId="figure-88@25.[327,1260,309,1913]" captionTargetPageId="25" captionText="FIGURE 14. Lower montane forest on limestone substrate at Iagifu Ridge, Papua New Guinea. Males of Litoria lisae sp. nov. called from low foliage in the forest along the trail illustrated A), and from deep within crevices and sinkholes in the karst, such as the large crevice indicated in B). Photos by S. Richards." figureDoi="http://doi.org/10.5281/zenodo.7800954" httpUri="https://zenodo.org/record/7800954/files/figure.png" pageId="24" pageNumber="175">Fig. 14B</figureCitation>
|
||
). Females have not been detected, and neither eggs nor larvae have been encountered. It is possible that this species breeds in small, subterranean waterbodies within the karst terrain, but this remains to be confirmed. Frog communities at the known localities for
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[445,526,1735,1761]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[445,526,1735,1761]" italics="true" pageId="24" pageNumber="175">L. lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[532,620,1735,1761]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[532,620,1735,1761]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are otherwise dominated by direct-developing microhylid species.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,150,2013]" pageId="24" pageNumber="175">
|
||
<emphasis bold="true" box="[199,470,1770,1797]" pageId="24" pageNumber="175">IUCN Red List status.</emphasis>
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[479,619,1771,1797]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="24" pageNumber="175" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[479,619,1771,1797]" italics="true" pageId="24" pageNumber="175">Litoria lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[628,719,1771,1797]" pageId="24" pageNumber="175">
|
||
<taxonomicNameLabel box="[628,719,1771,1797]" pageId="24" pageNumber="175" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from two areas within the Kikori River basin where extensive forest cover remains. However, logging operations are occurring widely within this catchment so until the species’ distribution, habitat requirements and any potential threats are better documented we recommend that it be listed as Data Deficient by the IUCN.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="25" lastPageNumber="176" pageId="24" pageNumber="175" type="etymology">
|
||
<paragraph blockId="24.[151,1437,150,2013]" pageId="24" pageNumber="175">
|
||
<emphasis bold="true" box="[199,334,1915,1941]" pageId="24" pageNumber="175">Etymology.</emphasis>
|
||
The name lisae is an honorific for the senior author’s wife, Lisa Capon, in gratitude for her ongoing support of his research activities.
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,150,2013]" lastBlockId="25.[151,1436,151,285]" lastPageId="25" lastPageNumber="176" pageId="24" pageNumber="175">
|
||
<emphasis bold="true" box="[199,475,1986,2013]" pageId="24" pageNumber="175">Molecular divergences.</emphasis>
|
||
Based on analyses of a 787 base pair alignment from the mitochondrial
|
||
<emphasis box="[1277,1331,1987,2013]" italics="true" pageId="24" pageNumber="175">ND4</emphasis>
|
||
gene and flanking tRNA
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[322,404,151,177]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="25" pageNumber="176" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[322,404,151,177]" italics="true" pageId="25" pageNumber="176">L. lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[410,499,151,177]" pageId="25" pageNumber="176">
|
||
<taxonomicNameLabel box="[410,499,151,177]" pageId="25" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most closely related to the clade including
|
||
<taxonomicName authorityName="Johnston & Richards" authorityYear="1994" box="[1005,1148,151,177]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="25" pageNumber="176" phylum="Chordata" rank="species" species="majikthise">
|
||
<emphasis box="[1005,1148,151,177]" italics="true" pageId="25" pageNumber="176">L. majikthise</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[1201,1396,151,177]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="25" pageNumber="176" phylum="Chordata" rank="species" species="haematogaster" status="sp. nov.">
|
||
<emphasis box="[1201,1396,151,177]" italics="true" pageId="25" pageNumber="176">L. haematogaster</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="25" pageNumber="176">
|
||
<taxonomicNameLabel pageId="25" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, however it is deeply genetically divergent from both of these taxa (
|
||
<emphasis box="[953,984,187,213]" italics="true" pageId="25" pageNumber="176">dA</emphasis>
|
||
between the taxa of 0.13 and 0.14,
|
||
<tableCitation captionStart="TABLE 2" captionStartId="9.[151,239,151,175]" captionTargetPageId="9" captionText="TABLE 2. Average net genetic distances (dA) between new Litoria species described herein and near relatives, based on data from the mitochondrial ND4 gene and flanking tRNA." pageId="25" pageNumber="176">Table 2</tableCitation>
|
||
). The two sequenced samples of
|
||
<taxonomicName authority="Richards & Donnellan & Oliver, 2023" authorityName="Richards & Donnellan & Oliver" authorityYear="2023" box="[548,632,223,249]" class="Amphibia" family="Pelodryadidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="25" pageNumber="176" phylum="Chordata" rank="species" species="lisae" status="sp. nov.">
|
||
<emphasis box="[548,632,223,249]" italics="true" pageId="25" pageNumber="176">L. lisae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[640,731,223,249]" pageId="25" pageNumber="176">
|
||
<taxonomicNameLabel box="[640,731,223,249]" pageId="25" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from localities 70 kilometres apart also showed some genetic divergence (p-distance 0.024).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |