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<document id="207A82CAFBFBE11CFDD6294CE8E4B0C7" ID-DOI="10.11646/zootaxa.4802.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3904926" ID-ZooBank="46448DF5-3090-4A4C-9893-F6E68DB8636C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592922952004" checkinUser="plazi" docAuthor="Zhu, Wenhui, Yao, Zhiyuan, Zheng, Guo &amp; Li, Shuqiang" docDate="2020" docId="03D2364EFFD67C4CCCEBF9A1FEAFFB46" docLanguage="en" docName="zootaxa.4802.1.7.pdf" docOrigin="Zootaxa 4802 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana muruo Zhu &amp; Yao &amp; Zheng &amp; Li 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="120" masterDocId="FFEB4E36FFD27C45CC7CFFB4FFC5FFE5" masterDocTitle="The Belisana spiders (Araneae: Pholcidae) from Tibet, China" masterLastPageNumber="128" masterPageNumber="111" pageNumber="115" updateTime="1700276892605" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="DDCB698373DFB0C270DB8A50EAC44B5E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9A4423F7D77B3A4E54075B1AA48036D4">
<mods:title id="7D320E9DC20AAECCD36DAAAB814E6832">The Belisana spiders (Araneae: Pholcidae) from Tibet, China</mods:title>
</mods:titleInfo>
<mods:name id="CB3699257CF64740EB293883074F1A9D" type="personal">
<mods:role id="75ED639E611CF682766591D06CDE9657">
<mods:roleTerm id="0A8D7CB9BAA0CD6C25DDBAEA8DB9DE73">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FB3B72325919BF310C36E44AED63BE37">Zhu, Wenhui</mods:namePart>
<mods:affiliation id="2E31428B97479BF811DCAC5EE9951B58">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
</mods:name>
<mods:name id="54F725B92E9407F56D5479CF1CA32686" type="personal">
<mods:role id="CD3190699C12EDB130268C2E351FFFF5">
<mods:roleTerm id="B3DBCBF435BA6A3513C67B1D3524A9D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="62F7FB3B50187A44F533FE592111AF0A">Yao, Zhiyuan</mods:namePart>
<mods:affiliation id="C8EC6AE3FDB2BFE1D9250349A2A7B839">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; yaozy @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 1631 - 0949</mods:affiliation>
</mods:name>
<mods:name id="E83499CF02C88FC2E60746F10AC5666D" type="personal">
<mods:role id="55177CBB914C16787C13E56D693B2DC1">
<mods:roleTerm id="B6791E494223CD63774B925627719783">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2610E30A8A0DA7C9B22D77B4AFDB473D">Zheng, Guo</mods:namePart>
<mods:affiliation id="CB87672F99D2588F13510B5A31715A44">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhengguo @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 9735 - 9700</mods:affiliation>
</mods:name>
<mods:name id="355521FB3286D181CB6184D47BAA3DCF" type="personal">
<mods:role id="C157B9267806A528EC08004644AEE3C0">
<mods:roleTerm id="DE7334BE3504447C2BCAC4E2E8C05293">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0FCF124655362A1F7DF097CAD711E928">Li, Shuqiang</mods:namePart>
<mods:affiliation id="61708C119393E4D120F07E946D845350">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
</mods:name>
<mods:typeOfResource id="6D30481C7BA59B47E8F8D6C5FA0B185E">text</mods:typeOfResource>
<mods:relatedItem id="267907F3E65513AED78A31E1F5A6E121" type="host">
<mods:titleInfo id="6233E14C0BC9816B8453F80152370F02">
<mods:title id="77B9FD791D2B5BEF76A66D03E1A4022F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="66E2DBD51A7C49AD524256C17208790D">
<mods:date id="A66699C341B8BA81A5542CE337E4C763">2020</mods:date>
<mods:detail id="1D1FD0FA0607252638D36D4284923FE6" type="pubDate">
<mods:number id="D006331AF6AEFB7FFFB769E5946A0F26">2020-06-22</mods:number>
</mods:detail>
<mods:detail id="7DDE27D65EFD51CC2490A8177562F303" type="volume">
<mods:number id="B8E5592FB17AE51E4094B219A414AE1E">4802</mods:number>
</mods:detail>
<mods:detail id="0E17478701EFA5987A230F6F026BD305" type="issue">
<mods:number id="70BEB582EB3B29532450F1D2387686F2">1</mods:number>
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<mods:start id="B6970CE4C61457454A63CB9A41475A12">111</mods:start>
<mods:end id="E87A70AE2BDE2D6A88B5A7327F1CCEA0">128</mods:end>
</mods:extent>
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<mods:classification id="50DBF32AE3CD457C9A422096385F5682">journal article</mods:classification>
<mods:identifier id="A4F29B3B7EB53D9C57DC6BD0B9221CCF" type="DOI">10.11646/zootaxa.4802.1.7</mods:identifier>
<mods:identifier id="CE9701F86ACAADDCF0B6AA42CD48B4AB" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E54EC2AD8C67237A9925411AFE8994F7" type="Zenodo-Dep">3904926</mods:identifier>
<mods:identifier id="66FCA3B97656F45C1AADC696DD6DE42D" type="ZooBank">46448DF5-3090-4A4C-9893-F6E68DB8636C</mods:identifier>
</mods:mods>
<treatment id="03D2364EFFD67C4CCCEBF9A1FEAFFB46" LSID="urn:lsid:plazi:treatment:03D2364EFFD67C4CCCEBF9A1FEAFFB46" httpUri="http://treatment.plazi.org/id/03D2364EFFD67C4CCCEBF9A1FEAFFB46" lastPageId="9" lastPageNumber="120" pageId="4" pageNumber="115">
<subSubSection id="C361D4D3FFD67C41CCEBF9A1FDF2F9CA" box="[151,567,1557,1584]" pageId="4" pageNumber="115" type="nomenclature">
<paragraph id="8BC48758FFD67C41CCEBF9A1FDF2F9CA" blockId="4.[151,567,1557,1619]" box="[151,567,1557,1584]" pageId="4" pageNumber="115">
<heading id="D08C3034FFD67C41CCEBF9A1FDF2F9CA" bold="true" box="[151,567,1557,1584]" fontSize="11" level="1" pageId="4" pageNumber="115" reason="1">
<emphasis id="B90F5B4AFFD67C41CCEBF9A1FDF2F9CA" bold="true" box="[151,567,1557,1584]" pageId="4" pageNumber="115">
<taxonomicName id="4C7BFCDBFFD67C41CCEBF9A1FE16F9CA" ID-CoL="93VJG" authority="Yao &amp; Li" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[151,467,1557,1584]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="muruo" status="sp. nov.">
<emphasis id="B90F5B4AFFD67C41CCEBF9A1FE9FF9CA" bold="true" box="[151,346,1557,1584]" italics="true" pageId="4" pageNumber="115">Belisana muruo</emphasis>
Yao &amp; Li
</taxonomicName>
<taxonomicNameLabel id="A23CE631FFD67C41CDA6F9A1FDF2F9CA" box="[474,567,1557,1583]" pageId="4" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361D4D3FFD67C41CCEBF98DFE93F9B6" box="[151,342,1592,1619]" pageId="4" pageNumber="115" type="description">
<paragraph id="8BC48758FFD67C41CCEBF98DFE93F9B6" blockId="4.[151,567,1557,1619]" box="[151,342,1592,1619]" pageId="4" pageNumber="115">
<figureCitation id="13409BDDFFD67C41CCEBF98DFF3FF9B6" box="[151,250,1592,1619]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="7.[151,250,1890,1915]" captionStartId-1="8.[151,250,1895,1920]" captionTargetBox-0="[221,1366,181,1864]" captionTargetBox-1="[219,1367,181,1870]" captionTargetId-0="figure@7.[221,1366,181,1864]" captionTargetId-1="figure@8.[219,1367,181,1870]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." captionText-1="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi-0="http://doi.org/10.5281/zenodo.3904936" figureDoi-1="http://doi.org/10.5281/zenodo.3904938" httpUri-0="https://zenodo.org/record/3904936/files/figure.png" httpUri-1="https://zenodo.org/record/3904938/files/figure.png" pageId="4" pageNumber="115">Figs 5–6</figureCitation>
,
<figureCitation id="13409BDDFFD67C41CD7AF98DFEDBF9B6" box="[262,286,1593,1619]" captionStart="FIGURE 11" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[205,1387,190,1908]" captionTargetId="figure@15.[199,1387,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Drawings of female genitalia in ventral and dorsal views. A–B. B. gyirong Zhang, Zhu &amp; Song, 2006; C–D. B. muruo sp. nov.; E–F. B. xigaze sp. nov.; G–H. B. zham sp. nov. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3904948" httpUri="https://zenodo.org/record/3904948/files/figure.png" pageId="4" pageNumber="115">11</figureCitation>
C–D
</paragraph>
</subSubSection>
<subSubSection id="C361D4D3FFD67C41CCEBF935FEC4F8E2" pageId="4" pageNumber="115" type="materials_examined">
<paragraph id="8BC48758FFD67C41CCEBF935FEC4F8E2" blockId="4.[151,1437,1664,2016]" pageId="4" pageNumber="115">
<emphasis id="B90F5B4AFFD67C41CCEBF935FE81F97E" bold="true" box="[151,324,1664,1691]" pageId="4" pageNumber="115">Type material.</emphasis>
<materialsCitation id="3B138D05FFD67C41CD36F935FEC9F906" collectingDate="2014-09-06" collectionCode="IZCAS" collectorName="J. Liu" country="China" county="Zayu County" elevation="3427" location="Male" municipality="Zhuwagen Town" pageId="4" pageNumber="115" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="B90F5B4AFFD67C41CD36F935FE05F97E" bold="true" box="[330,448,1665,1691]" pageId="4" pageNumber="115">
<typeStatus id="54C039FAFFD67C41CD36F935FE7EF97E" box="[330,443,1665,1691]" pageId="4" pageNumber="115" type="holotype">Holotype</typeStatus>
:
</emphasis>
<location id="8EA4D183FFD67C41CDBBF935FDC4F97E" LSID="urn:lsid:plazi:treatment:03D2364EFFD67C4CCCEBF9A1FEAFFB46:8EA4D183FFD67C41CDBBF935FDC4F97E" box="[455,513,1665,1691]" country="China" county="Zayu County" municipality="Zhuwagen Town" name="Male" pageId="4" pageNumber="115" stateProvince="Xizang">Male</location>
(
<collectionCode id="ED6A1F9DFFD67C41CE72F935FDA4F97E" box="[526,609,1665,1691]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="115">IZCAS</collectionCode>
Ar39801),
<location id="8EA4D183FFD67C41CEA3F935FC40F97E" LSID="urn:lsid:plazi:treatment:03D2364EFFD67C4CCCEBF9A1FEAFFB46:8EA4D183FFD67C41CEA3F935FC40F97E" box="[735,901,1665,1691]" country="China" county="Zayu County" municipality="Zhuwagen Town" name="Unnamed cave" pageId="4" pageNumber="115" stateProvince="Xizang">Unnamed cave</location>
(28°47
<emphasis id="B90F5B4AFFD67C41CFA4F934FC18F97E" bold="true" box="[984,989,1664,1691]" pageId="4" pageNumber="115">ꞌ</emphasis>
17.99
<emphasis id="B90F5B4AFFD67C41C860F934FBE3F97E" bold="true" box="[1052,1062,1664,1691]" pageId="4" pageNumber="115">ꞌꞌ</emphasis>
N, 98°31
<emphasis id="B90F5B4AFFD67C41C8F7F934FB55F97E" bold="true" box="[1163,1168,1664,1691]" pageId="4" pageNumber="115">ꞌ</emphasis>
04.98
<emphasis id="B90F5B4AFFD67C41C8B3F934FB1CF97E" bold="true" box="[1231,1241,1664,1691]" pageId="4" pageNumber="115">ꞌꞌ</emphasis>
E,
<elevation id="0056606BFFD67C41C88BF935FF75F95A" metricMagnitude="3" metricUnit="m" metricValue="3.4269999999999996" pageId="4" pageNumber="115" unit="m" value="3427.0">
elevation
<quantity id="4C832ABDFFD67C41C918F935FF75F95A" metricMagnitude="3" metricUnit="m" metricValue="3.4269999999999996" pageId="4" pageNumber="115" unit="m" value="3427.0">3427 m</quantity>
</elevation>
),
<quantity id="4C832ABDFFD67C41CCB9F911FF3BF95A" box="[197,254,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="115" unit="km" value="8.0">8 km</quantity>
east of
<location id="8EA4D183FFD67C41CD2BF911FE3EF95A" LSID="urn:lsid:plazi:treatment:03D2364EFFD67C4CCCEBF9A1FEAFFB46:8EA4D183FFD67C41CD2BF911FE3EF95A" box="[343,507,1701,1727]" country="China" county="Zayu County" municipality="Zhuwagen Town" name="Da Zhengtong" pageId="4" pageNumber="115" stateProvince="Xizang">Da Zhengtong</location>
,
<location id="8EA4D183FFD67C41CE7BF911FD6BF95A" LSID="urn:lsid:plazi:treatment:03D2364EFFD67C4CCCEBF9A1FEAFFB46:8EA4D183FFD67C41CE7BF911FD6BF95A" box="[519,686,1701,1727]" country="China" county="Zayu County" municipality="Zhuwagen Town" name="Muruo Village" pageId="4" pageNumber="115" stateProvince="Xizang">Muruo Village</location>
,
<collectingMunicipality id="6BA01D22FFD67C41CEC5F911FCBDF95A" box="[697,888,1701,1727]" pageId="4" pageNumber="115">Zhuwagen Town</collectingMunicipality>
,
<collectingCounty id="62A5FFD4FFD67C41CFF8F911FBDDF95A" box="[900,1048,1701,1727]" pageId="4" pageNumber="115">Zayu County</collectingCounty>
,
<emphasis id="B90F5B4AFFD67C41C858F910FBA6F95A" bold="true" box="[1060,1123,1700,1727]" pageId="4" pageNumber="115">
<collectingRegion id="49BF49BAFFD67C41C858F910FBA6F95A" box="[1060,1123,1700,1727]" country="China" name="Xizang" pageId="4" pageNumber="115">Tibet</collectingRegion>
</emphasis>
,
<emphasis id="B90F5B4AFFD67C41C80DF910FB7FF95A" bold="true" box="[1137,1210,1700,1727]" pageId="4" pageNumber="115">
<collectingCountry id="F36CC7C8FFD67C41C80DF910FB7FF95A" box="[1137,1210,1700,1727]" name="China" pageId="4" pageNumber="115">China</collectingCountry>
</emphasis>
,
<date id="FFC5A198FFD67C41C8B5F911FA5DF95A" box="[1225,1432,1701,1727]" pageId="4" pageNumber="115" value="2014-09-06">
<collectingDate id="EF815870FFD67C41C8B5F911FA5DF95A" box="[1225,1432,1701,1727]" pageId="4" pageNumber="115" value="2014-09-06">6 September 2014</collectingDate>
</date>
,
<collectorName id="268EE28EFFD67C41CCEBF97DFF1CF906" box="[151,217,1737,1763]" pageId="4" pageNumber="115">J. Liu</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B138D05FFD67C41CD69F97DFC93F906" box="[277,854,1737,1763]" collectionCode="IZCAS" country="China" pageId="4" pageNumber="115" specimenCount="2" specimenCount-male="2" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="B90F5B4AFFD67C41CD69F97DFE53F906" bold="true" box="[277,406,1737,1763]" pageId="4" pageNumber="115">
<typeStatus id="54C039FAFFD67C41CD69F97DFE54F906" box="[277,401,1737,1763]" pageId="4" pageNumber="115" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D7D4CD1FFD67C41CDE3F97DFE3DF906" box="[415,504,1737,1763]" pageId="4" pageNumber="115" type="male">2 males</specimenCount>
(
<collectionCode id="ED6A1F9DFFD67C41CE75F97DFD99F906" box="[521,604,1737,1763]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="115">IZCAS</collectionCode>
Ar39802–39803) and
</materialsCitation>
<materialsCitation id="3B138D05FFD67C41CF23F97DFF38F8E2" collectingDate="2014-09-06" collectionCode="IZCAS" collectorName="J. Liu" country="China" county="Zayu County" elevation="3427" location="Male" municipality="Zhuwagen Town" pageId="4" pageNumber="115" specimenCount="4" specimenCount-female="4" stateProvince="Xizang" typeStatus="paratype">
<specimenCount id="9D7D4CD1FFD67C41CF23F97DFC0BF906" box="[863,974,1737,1763]" pageId="4" pageNumber="115" type="female">4 females</specimenCount>
(
<collectionCode id="ED6A1F9DFFD67C41CFA2F97DFBF4F906" box="[990,1073,1737,1763]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="115">IZCAS</collectionCode>
Ar39804–39807), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361D4D3FFD67C41CCBBF8A5FBB9F8CE" box="[199,1148,1809,1835]" pageId="4" pageNumber="115" type="etymology">
<paragraph id="8BC48758FFD67C41CCBBF8A5FBB9F8CE" blockId="4.[151,1437,1664,2016]" box="[199,1148,1809,1835]" pageId="4" pageNumber="115">
<emphasis id="B90F5B4AFFD67C41CCBBF8A5FE8BF8CE" bold="true" box="[199,334,1809,1835]" pageId="4" pageNumber="115">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="54C039FAFFD67C41CEC9F8A5FD20F8CE" box="[693,741,1809,1835]" pageId="4" pageNumber="115">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C361D4D3FFD67C41CCBBF880FC11F83A" pageId="4" pageNumber="115" type="diagnosis">
<paragraph id="8BC48758FFD67C41CCBBF880FC11F83A" blockId="4.[151,1437,1664,2016]" pageId="4" pageNumber="115">
<emphasis id="B90F5B4AFFD67C41CCBBF880FE87F8AA" bold="true" box="[199,322,1844,1871]" pageId="4" pageNumber="115">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C7BFCDBFFD67C41CE26F882FBDCF8AA" authority="Zhang, Zhu &amp; Song, 2006" authorityName="Zhang, Zhu &amp; Song" authorityYear="2006" box="[602,1049,1845,1871]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="mainling">
<emphasis id="B90F5B4AFFD67C41CE26F882FD1AF8AA" box="[602,735,1845,1871]" italics="true" pageId="4" pageNumber="115">B. mainling</emphasis>
<bibRefCitation id="EFEAFAA9FFD67C41CE95F881FBDCF8AA" author="Zhang, F. &amp; Zhu, M. &amp; Song, D." box="[745,1049,1845,1871]" pageId="4" pageNumber="115" pagination="194 - 205" refId="ref6960" refString="Zhang, F., Zhu, M. &amp; Song, D. (2006) A review of pholcid spiders from Tibet, China (Araneae, Pholcidae). The Journal of Arachnology, 34, 194 - 205. https: // doi. org / 10.1636 / h 04 - 22.1" type="journal article" year="2006">Zhang, Zhu &amp; Song, 2006</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFEAFAA9FFD67C41C825F881FAA0F8AA" author="Zhang, F. &amp; Zhu, M. &amp; Song, D." box="[1113,1381,1845,1871]" pageId="4" pageNumber="115" pagination="194 - 205" refId="ref6960" refString="Zhang, F., Zhu, M. &amp; Song, D. (2006) A review of pholcid spiders from Tibet, China (Araneae, Pholcidae). The Journal of Arachnology, 34, 194 - 205. https: // doi. org / 10.1636 / h 04 - 22.1" type="journal article" year="2006">
Zhang
<emphasis id="B90F5B4AFFD67C41C8D6F882FB1BF8AA" box="[1194,1246,1845,1871]" italics="true" pageId="4" pageNumber="115">et al</emphasis>
. 2006: 203
</bibRefCitation>
, figs 32–40) with similar bulbal apophyses (
<figureCitation id="13409BDDFFD67C41CE31F8EDFD61F891" box="[589,676,1881,1908]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="4" pageNumber="115">Fig. 6C</figureCitation>
) and vulval pore plates (
<figureCitation id="13409BDDFFD67C41CFC6F8EDFBD3F896" box="[954,1046,1881,1907]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="4" pageNumber="115">Figs 6B</figureCitation>
,
<figureCitation id="13409BDDFFD67C41C85EF8EDFB91F896" box="[1058,1108,1881,1907]" captionStart="FIGURE 11" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[205,1387,190,1908]" captionTargetId="figure@15.[199,1387,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Drawings of female genitalia in ventral and dorsal views. A–B. B. gyirong Zhang, Zhu &amp; Song, 2006; C–D. B. muruo sp. nov.; E–F. B. xigaze sp. nov.; G–H. B. zham sp. nov. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3904948" httpUri="https://zenodo.org/record/3904948/files/figure.png" pageId="4" pageNumber="115">11D</figureCitation>
), but can be distinguished by presence of large subdisto-prolateral sclerite (arrow in
<figureCitation id="13409BDDFFD67C41CF7EF8C9FC92F872" box="[770,855,1916,1943]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="4" pageNumber="115">Fig. 5C</figureCitation>
) and retrolateral membranous flap (arrow f in
<figureCitation id="13409BDDFFD67C41C90CF8C9FF7FF85E" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="4" pageNumber="115">Fig. 5D</figureCitation>
) on procursus. Unlike
<taxonomicName id="4C7BFCDBFFD67C41CDBCF816FD81F85E" authorityName="Zhang, Zhu &amp; Song" authorityYear="2006" box="[448,580,1953,1979]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="115" phylum="Arthropoda" rank="species" species="mainling">
<emphasis id="B90F5B4AFFD67C41CDBCF816FD81F85E" box="[448,580,1953,1979]" italics="true" pageId="4" pageNumber="115">B. mainling</emphasis>
</taxonomicName>
male cheliceral distal apophyses face each other mesially (arrow da in
<figureCitation id="13409BDDFFD67C41C90CF815FF78F805" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="4" pageNumber="115">Fig. 6D</figureCitation>
), and epigynal pockets are more separated (arrow ep in
<figureCitation id="13409BDDFFD67C41CF52F871FC4CF83A" box="[814,905,1989,2015]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="4" pageNumber="115">Figs 6B</figureCitation>
,
<figureCitation id="13409BDDFFD67C41CFE8F871FC02F83A" box="[916,967,1989,2015]" captionStart="FIGURE 11" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[205,1387,190,1908]" captionTargetId="figure@15.[199,1387,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Drawings of female genitalia in ventral and dorsal views. A–B. B. gyirong Zhang, Zhu &amp; Song, 2006; C–D. B. muruo sp. nov.; E–F. B. xigaze sp. nov.; G–H. B. zham sp. nov. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3904948" httpUri="https://zenodo.org/record/3904948/files/figure.png" pageId="4" pageNumber="115">11D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF04D7D0FFD77C40CCEBF82DFAF1F81F" ID-DOI="http://doi.org/10.5281/zenodo.3904932" ID-Zenodo-Dep="3904932" httpUri="https://zenodo.org/record/3904932/files/figure.png" pageId="5" pageNumber="116" startId="5.[151,250,1945,1970]" targetBox="[203,1385,181,1920]" targetPageId="5">
<paragraph id="8BC48758FFD77C40CCEBF82DFAF1F81F" blockId="5.[151,1436,1945,2042]" pageId="5" pageNumber="116">
<emphasis id="B90F5B4AFFD77C40CCEBF82DFED1F857" bold="true" box="[151,276,1945,1970]" pageId="5" pageNumber="116">FIGURE 3.</emphasis>
<taxonomicName id="4C7BFCDBFFD77C40CD60F82EFE05F857" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[284,448,1946,1970]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="116" phylum="Arthropoda" rank="species" species="medog" status="sp. nov.">
<emphasis id="B90F5B4AFFD77C40CD60F82EFE05F857" box="[284,448,1946,1970]" italics="true" pageId="5" pageNumber="116">Belisana medog</emphasis>
</taxonomicName>
<emphasis id="B90F5B4AFFD77C40CDBBF82EFDDDF857" bold="true" box="[455,536,1946,1970]" pageId="5" pageNumber="116">
<taxonomicNameLabel id="A23CE631FFD77C40CDBBF82EFDDDF857" box="[455,536,1946,1970]" pageId="5" pageNumber="116" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Flipped right pedipalp (A. Prolateral view, arrow points at dorsal protruding part of procursus; B. Retrolateral view); C–D. Distal part of procursus (C. Prolateral view, arrow points at bifid distoprolateral process; D. Retrolateral view). b = bulb, e = embolus, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="DF04D7D0FFD47C43CCEBF82DFCDFF81F" ID-DOI="http://doi.org/10.5281/zenodo.3904934" ID-Zenodo-Dep="3904934" httpUri="https://zenodo.org/record/3904934/files/figure.png" pageId="6" pageNumber="117" startId="6.[151,250,1945,1970]" targetBox="[203,1385,181,1920]" targetPageId="6">
<paragraph id="8BC48758FFD47C43CCEBF82DFCDFF81F" blockId="6.[151,1436,1945,2042]" pageId="6" pageNumber="117">
<emphasis id="B90F5B4AFFD47C43CCEBF82DFED3F857" bold="true" box="[151,278,1945,1970]" pageId="6" pageNumber="117">FIGURE 4.</emphasis>
<taxonomicName id="4C7BFCDBFFD47C43CD5CF82EFE03F857" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[288,454,1946,1970]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="117" phylum="Arthropoda" rank="species" species="medog" status="sp. nov.">
<emphasis id="B90F5B4AFFD47C43CD5CF82EFE03F857" box="[288,454,1946,1970]" italics="true" pageId="6" pageNumber="117">Belisana medog</emphasis>
</taxonomicName>
<emphasis id="B90F5B4AFFD47C43CDB3F82EFDE6F857" bold="true" box="[463,547,1946,1970]" pageId="6" pageNumber="117">
<taxonomicNameLabel id="A23CE631FFD47C43CDB3F82EFDE6F857" box="[463,547,1946,1970]" pageId="6" pageNumber="117" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A. Bulb, prolateral view; B. Chelicerae, frontal view; C–E. Habitus (C. Dorsal view; D. Lateral view; E. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, pa = proximo-lateral apophysis. Scale bars: 0.10 (A–B), 0.20 (C–E).
</paragraph>
</caption>
<caption id="DF04D7D0FFD57C42CCEBF8D6FD1FF802" ID-DOI="http://doi.org/10.5281/zenodo.3904936" ID-Zenodo-Dep="3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="7" pageNumber="118" startId="7.[151,250,1890,1915]" targetBox="[221,1366,181,1864]" targetPageId="7">
<paragraph id="8BC48758FFD57C42CCEBF8D6FD1FF802" blockId="7.[151,1437,1890,2023]" pageId="7" pageNumber="118">
<emphasis id="B90F5B4AFFD57C42CCEBF8D6FED7F89E" bold="true" box="[151,274,1890,1915]" pageId="7" pageNumber="118">FIGURE 5.</emphasis>
<taxonomicName id="4C7BFCDBFFD57C42CD64F8D6FE7CF89F" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[280,441,1890,1914]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="118" phylum="Arthropoda" rank="species" species="muruo" status="sp. nov.">
<emphasis id="B90F5B4AFFD57C42CD64F8D6FE7CF89F" box="[280,441,1890,1914]" italics="true" pageId="7" pageNumber="118">Belisana muruo</emphasis>
</taxonomicName>
<emphasis id="B90F5B4AFFD57C42CDC3F8D7FDCAF89E" bold="true" box="[447,527,1891,1915]" pageId="7" pageNumber="118">
<taxonomicNameLabel id="A23CE631FFD57C42CDC3F8D7FDCAF89E" box="[447,527,1891,1915]" pageId="7" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D).
</paragraph>
</caption>
<caption id="DF04D7D0FFDA7C4DCCEBF8D3FC07F809" ID-DOI="http://doi.org/10.5281/zenodo.3904938" ID-Zenodo-Dep="3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="8" pageNumber="119" startId="8.[151,250,1895,1920]" targetBox="[219,1367,181,1870]" targetPageId="8">
<paragraph id="8BC48758FFDA7C4DCCEBF8D3FC07F809" blockId="8.[151,1437,1895,2028]" pageId="8" pageNumber="119">
<emphasis id="B90F5B4AFFDA7C4DCCEBF8D3FED0F865" bold="true" box="[151,277,1895,1920]" pageId="8" pageNumber="119">FIGURE 6.</emphasis>
<taxonomicName id="4C7BFCDBFFDA7C4DCD61F8DCFE05F865" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[285,448,1896,1920]" class="Arachnida" family="Pholcidae" genus="Belisana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="119" phylum="Arthropoda" rank="species" species="muruo" status="sp. nov.">
<emphasis id="B90F5B4AFFDA7C4DCD61F8DCFE05F865" box="[285,448,1896,1920]" italics="true" pageId="8" pageNumber="119">Belisana muruo</emphasis>
</taxonomicName>
<emphasis id="B90F5B4AFFDA7C4DCDB5F8DCFDD9F865" bold="true" box="[457,540,1896,1920]" pageId="8" pageNumber="119">
<taxonomicNameLabel id="A23CE631FFDA7C4DCDB5F8DCFDD9F865" box="[457,540,1896,1920]" pageId="8" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H).
</paragraph>
</caption>
<subSubSection id="C361D4D3FFDB7C4CCCBBFF2CFBDFFC0A" pageId="9" pageNumber="120" type="description">
<paragraph id="8BC48758FFDB7C4CCCBBFF2CFE93FD4E" blockId="9.[151,1437,152,1187]" pageId="9" pageNumber="120">
<emphasis id="B90F5B4AFFDB7C4CCCBBFF2CFE5AFF56" bold="true" box="[199,415,152,179]" pageId="9" pageNumber="120">Description. Male</emphasis>
(
<emphasis id="B90F5B4AFFDB7C4CCDCEFF2DFDDAFF56" bold="true" box="[434,543,153,179]" pageId="9" pageNumber="120">
<typeStatus id="54C039FAFFDB7C4CCDCEFF2DFDDFFF56" box="[434,538,153,179]" pageId="9" pageNumber="120" type="holotype">holotype</typeStatus>
,
</emphasis>
IZCAS Ar39801): Total length 2.13 (2.19 with clypeus), carapace 0.94 long, 0.78 wide, opisthosoma 1.19 long, 1.25 wide. Leg I: 7.41 (1.94 + 0.31 + 2.06 + 2.44 + 0.66), leg II: 5.54 (1.44 + 0.19 + 1.41 + 2.34 + 0.16), leg III: 3.23 (0.78 + 0.13 + 0.88 + 1.25 + 0.19), leg IV: 4.18 (1.31 + 0.19 + 1.03 + 1.34 + 0.31); tibia I L/d: 36. Habitus as in
<figureCitation id="13409BDDFFDB7C4CCD94FEB1FDEAFEFA" box="[488,559,261,287]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Figs 6</figureCitation>
E–F. Carapace pale, with pair of large brown marks; ocular area with brown stripe medially; sternum pale, without marks. Legs pale, without darker rings. Opisthosoma grey, with black spots dorsally and laterally. Distance PME-PME 0.10, diameter PME 0.07, distance PME-ALE 0.04, AME absent. Ocular area barely elevated. Thoracic furrow absent. Clypeus unmodified. Sternum slightly longer than wide (0.63/0.53). Chelicerae (
<figureCitation id="13409BDDFFDB7C4CCD63FE21FEB3FE55" box="[287,374,405,432]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Fig. 6D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of short, pointed distal apophyses each with hardened and pointed tip (distance between tips: 0.14). Pedipalps as in
<figureCitation id="13409BDDFFDB7C4CCFA7FE0DFBDBFE36" box="[987,1054,440,467]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="9" pageNumber="120">Figs 5</figureCitation>
A–B; coxa unmodified; trochanter with short ventral apophysis provided with large retrolateral apophysis (arrow in
<figureCitation id="13409BDDFFDB7C4CC840FE69FB53FE12" box="[1084,1174,476,503]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="9" pageNumber="120">Fig. 5B</figureCitation>
); femur with indistinct proximo-dorsal apophysis (arrow in
<figureCitation id="13409BDDFFDB7C4CCE53FDB5FD4DFDFE" box="[559,648,512,539]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="9" pageNumber="120">Fig. 5A</figureCitation>
); procursus (
<figureCitation id="13409BDDFFDB7C4CCF6BFDB5FC9EFDFE" box="[791,859,512,539]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="9" pageNumber="120">Figs 5</figureCitation>
A–D) simple proximally and complex distally, with large subdisto-prolateral sclerite (arrow in
<figureCitation id="13409BDDFFDB7C4CCE09FD91FD0EFDDA" box="[629,715,548,575]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="9" pageNumber="120">Fig. 5C</figureCitation>
), long, curved distal spine and retrolateral membranous flap (arrow f in
<figureCitation id="13409BDDFFDB7C4CCC87FDFDFE91FD86" box="[251,340,584,611]" captionStart="FIGURE 5" captionStartId="7.[151,250,1890,1915]" captionTargetBox="[221,1366,181,1864]" captionTargetId="figure@7.[221,1366,181,1864]" captionTargetPageId="7" captionText="FIGURE 5. Belisana muruo sp. nov., holotype male. A–B. Pedipalp (Bulb removed) (A. Prolateral view, arrow points at proximo-dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolateral apophysis on pedipalpal trochanter); C–D. Distal part of procursus (C. Prolateral view, arrow points at large subdisto-prolateral sclerite; D. Retrolateral view). f = flap, pr = procursus. Scale bars: 0.10 (A–B), 0.05 (C–D)." figureDoi="http://doi.org/10.5281/zenodo.3904936" httpUri="https://zenodo.org/record/3904936/files/figure.png" pageId="9" pageNumber="120">Fig. 5D</figureCitation>
); bulb (
<figureCitation id="13409BDDFFDB7C4CCDD1FDFDFDC0FD81" box="[429,517,585,612]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Fig. 6C</figureCitation>
) with hooked apophysis and simple embolus. Retrolateral trichobothria of tibia I at 17% proximally; legs with short vertical setae on tibiae, without spines or curved setae; tarsus I with 21 distinct pseudosegments.
</paragraph>
<paragraph id="8BC48758FFDB7C4CCCBBFD01FBFBFCBA" blockId="9.[151,1437,152,1187]" pageId="9" pageNumber="120">
<emphasis id="B90F5B4AFFDB7C4CCCBBFD01FEDBFD2A" bold="true" box="[199,286,693,719]" pageId="9" pageNumber="120">Female</emphasis>
(IZCAS Ar39804): Similar to male, habitus as in
<figureCitation id="13409BDDFFDB7C4CCF1CFD01FC63FD2A" box="[864,934,693,719]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Figs 6</figureCitation>
G–H. Total length 2.06 (2.19 with clypeus), carapace 0.88 long, 0.63 wide, opisthosoma 1.18 long, 1.09 wide. Tibia I: 2.66; tibia I L/d: 27. Distance PME-PME 0.10, diameter PME 0.06, distance PME-ALE 0.04, AME absent. Sternum slightly longer than wide (0.63/0.56). External female genitalia (
<figureCitation id="13409BDDFFDB7C4CCDBDFC95FDDAFCDE" box="[449,543,801,827]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Figs 6A</figureCitation>
,
<figureCitation id="13409BDDFFDB7C4CCE56FC95FD9DFCDE" box="[554,600,801,827]" captionStart="FIGURE 11" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[205,1387,190,1908]" captionTargetId="figure@15.[199,1387,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Drawings of female genitalia in ventral and dorsal views. A–B. B. gyirong Zhang, Zhu &amp; Song, 2006; C–D. B. muruo sp. nov.; E–F. B. xigaze sp. nov.; G–H. B. zham sp. nov. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3904948" httpUri="https://zenodo.org/record/3904948/files/figure.png" pageId="9" pageNumber="120">11C</figureCitation>
) simple and flat, with pair of indistinct pockets 0.18 apart (ep in
<figureCitation id="13409BDDFFDB7C4CC941FC95FA52FCDE" box="[1341,1431,801,827]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Figs 6B</figureCitation>
,
<figureCitation id="13409BDDFFDB7C4CCCEBFCF1FF0FFCBA" box="[151,202,837,863]" captionStart="FIGURE 11" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[205,1387,190,1908]" captionTargetId="figure@15.[199,1387,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Drawings of female genitalia in ventral and dorsal views. A–B. B. gyirong Zhang, Zhu &amp; Song, 2006; C–D. B. muruo sp. nov.; E–F. B. xigaze sp. nov.; G–H. B. zham sp. nov. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3904948" httpUri="https://zenodo.org/record/3904948/files/figure.png" pageId="9" pageNumber="120">11D</figureCitation>
). Vulva (
<figureCitation id="13409BDDFFDB7C4CCD53FCF1FE4FFCBA" box="[303,394,837,863]" captionStart="FIGURE 6" captionStartId="8.[151,250,1895,1920]" captionTargetBox="[219,1367,181,1870]" captionTargetId="figure@8.[219,1367,181,1870]" captionTargetPageId="8" captionText="FIGURE 6. Belisana muruo sp. nov., holotype male (C–F) and paratype female (A–B, G–H). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; E–H. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (A–D), 0.40 (E–H)." figureDoi="http://doi.org/10.5281/zenodo.3904938" httpUri="https://zenodo.org/record/3904938/files/figure.png" pageId="9" pageNumber="120">Figs 6B</figureCitation>
,
<figureCitation id="13409BDDFFDB7C4CCDEAFCF1FE03FCBA" box="[406,454,837,863]" captionStart="FIGURE 11" captionStartId="15.[151,250,1945,1970]" captionTargetBox="[205,1387,190,1908]" captionTargetId="figure@15.[199,1387,181,1919]" captionTargetPageId="15" captionText="FIGURE 11. Drawings of female genitalia in ventral and dorsal views. A–B. B. gyirong Zhang, Zhu &amp; Song, 2006; C–D. B. muruo sp. nov.; E–F. B. xigaze sp. nov.; G–H. B. zham sp. nov. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3904948" httpUri="https://zenodo.org/record/3904948/files/figure.png" pageId="9" pageNumber="120">11D</figureCitation>
) with wavy anterior arch and pair of elliptic pore plates.
</paragraph>
<paragraph id="8BC48758FFDB7C4CCCBBFCDCFE24FC2E" blockId="9.[151,1437,152,1187]" pageId="9" pageNumber="120">
<emphasis id="B90F5B4AFFDB7C4CCCBBFCDCFE84FC66" bold="true" box="[199,321,872,899]" pageId="9" pageNumber="120">Variation:</emphasis>
<materialsCitation id="3B138D05FFDB7C4CCD30FCDDFE18FC2E" collectionCode="IZCAS" collectorName="Tibia I" pageId="9" pageNumber="120" specimenCount="4" typeStatus="paratype">
<collectorName id="268EE28EFFDB7C4CCD30FCDDFE5CFC66" box="[332,409,873,899]" pageId="9" pageNumber="120">Tibia I</collectorName>
in one male
<typeStatus id="54C039FAFFDB7C4CCE45FCDDFD5CFC66" box="[569,665,873,899]" pageId="9" pageNumber="120" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED6A1F9DFFDB7C4CCED0FCDDFD3AFC66" box="[684,767,873,899]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="9" pageNumber="120">IZCAS</collectionCode>
Ar39802; leg I missing in another male
<typeStatus id="54C039FAFFDB7C4CC89CFCDDFA85FC66" box="[1248,1344,873,899]" pageId="9" pageNumber="120" type="paratype">paratype</typeStatus>
<collectionCode id="ED6A1F9DFFDB7C4CC937FCDDFA59FC66" box="[1355,1436,873,899]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="9" pageNumber="120">IZCAS</collectionCode>
Ar39803): 2.56.
<collectorName id="268EE28EFFDB7C4CCD33FC39FE5DFC42" box="[335,408,909,935]" pageId="9" pageNumber="120">Tibia I</collectorName>
in two female
<typeStatus id="54C039FAFFDB7C4CCE40FC39FD62FC42" box="[572,679,909,935]" pageId="9" pageNumber="120" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED6A1F9DFFDB7C4CCEC9FC39FCCDFC42" box="[693,776,909,935]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="9" pageNumber="120">IZCAS</collectionCode>
Ar39805–39806; leg I missing in the other female
<typeStatus id="54C039FAFFDB7C4CC940FC39FA59FC42" box="[1340,1436,909,935]" pageId="9" pageNumber="120" type="paratype">paratype</typeStatus>
<collectionCode id="ED6A1F9DFFDB7C4CCCEBFC05FF2DFC2E" box="[151,232,945,971]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="9" pageNumber="120">IZCAS</collectionCode>
Ar39807): 1.92, 2.56
</materialsCitation>
.
</paragraph>
<paragraph id="8BC48758FFDB7C4CCCBBFC61FBDFFC0A" blockId="9.[151,1437,152,1187]" box="[199,1050,980,1007]" pageId="9" pageNumber="120">
<emphasis id="B90F5B4AFFDB7C4CCCBBFC61FE40FC0A" bold="true" box="[199,389,980,1007]" pageId="9" pageNumber="120">Natural history.</emphasis>
The species was found in the aphotic zone inside the cave.
</paragraph>
</subSubSection>
<subSubSection id="C361D4D3FFDB7C4CCCBBFC4CFCF0FBF6" box="[199,821,1016,1043]" pageId="9" pageNumber="120" type="distribution">
<paragraph id="8BC48758FFDB7C4CCCBBFC4CFCF0FBF6" blockId="9.[151,1437,152,1187]" box="[199,821,1016,1043]" pageId="9" pageNumber="120">
<emphasis id="B90F5B4AFFDB7C4CCCBBFC4CFEA4FBF6" bold="true" box="[199,353,1016,1043]" pageId="9" pageNumber="120">Distribution.</emphasis>
<collectingCountry id="F36CC7C8FFDB7C4CCD1BFC4DFE6FFBF6" box="[359,426,1017,1043]" name="China" pageId="9" pageNumber="120">China</collectingCountry>
(
<collectingRegion id="49BF49BAFFDB7C4CCDC5FC4DFE3DFBF6" box="[441,504,1017,1043]" country="China" name="Xizang" pageId="9" pageNumber="120">Tibet</collectingRegion>
,
<typeStatus id="54C039FAFFDB7C4CCE7FFC4DFDF6FBF6" box="[515,563,1017,1043]" pageId="9" pageNumber="120">type</typeStatus>
locality only;
<figureCitation id="13409BDDFFDB7C4CCEAAFC4DFCEDFBF6" box="[726,808,1017,1043]" captionStart="FIGURE 12" captionStartId="16.[151,250,1215,1240]" captionTargetBox="[151,1435,181,1189]" captionTargetId="figure@16.[151,1435,181,1189]" captionTargetPageId="16" captionText="FIGURE 12. Distribution records of five species of the genus Belisana. 1. B. gyirong Zhang, Zhu &amp; Song, 2006; 2. B. medog sp. nov.; 3. B. muruo sp. nov.; 4. B. xigaze sp. nov.; 5. B. zham sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3904950" httpUri="https://zenodo.org/record/3904950/files/figure.png" pageId="9" pageNumber="120">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361D4D3FFDB7C4CCCBBFBA9FEAFFB46" pageId="9" pageNumber="120" type="discussion">
<paragraph id="8BC48758FFDB7C4CCCBBFBA9FEAFFB46" blockId="9.[151,1437,152,1187]" pageId="9" pageNumber="120">
<emphasis id="B90F5B4AFFDB7C4CCCBBFBA9FEFEFBD2" bold="true" box="[199,315,1053,1079]" pageId="9" pageNumber="120">Remarks.</emphasis>
This new species was found within cavities of rock walls inside the cave, while the other species in this paper were found on the underside of rocks in epigean environments. In contrast with these other longlegged species, this new species has remarkably short legs. This discrepancy probably reflects adaptation to different microhabitats.
</paragraph>
</subSubSection>
</treatment>
</document>
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