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<document id="2BD17B88718EE8DC2AA80A2FFD543647" ID-DOI="10.26107/RBZ-2019-0041" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576444" ID-ZooBank="AAA866A1-FBAF-486D-AB84-3AB989A269C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614778122966" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Grego, Jozef, Vermeulen, Jaap J., Reischütz, Alexander, Hunyadi, András &amp; Jochum, Adrienne" docDate="2019" docId="03BF447DFF91FFBBFC91FF48FE42FA92" docLanguage="en" docName="RafflesBZool.67.517-535.pdf" docOrigin="Raffles Bulletin of Zoology 67" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Tonkinospira suturata Pall-Gergely &amp; Grego 2019, new species" docType="treatment" docVersion="4" lastPageNumber="533" masterDocId="FF863C05FF9DFFABFFEEFA75FFE6FFC4" masterDocTitle="New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)" masterLastPageNumber="535" masterPageNumber="517" pageNumber="529" updateTime="1698878161501" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="CD8DB5F0611E47B159470C8DC914A2E7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0C1391FF1A144B4ECD3155990B04C4E9">
<mods:title id="E66AB90A08F6F9FE84FC29F1BA94AC8E">New Tonkinospira Jochum, Slapnik &amp; Páll-Gergely, 2014 species from Laos and Vietnam (Gastropoda: Pulmonata: Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name id="4AE24842C76066CF9063EAB1B7CCA416" type="personal">
<mods:role id="170CD49F851EA5798E5790B7C905CFEA">
<mods:roleTerm id="0E147773858ACC2A6F054A6EA45EF0B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7D863BE1915BCA5C6898967C8C932DBB">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="7FDB4BC2662DEC0278835FEA863AA184" type="personal">
<mods:role id="AB3DC8B73E5EB8935E9FD2BE0F9E8026">
<mods:roleTerm id="8264546785267DD9BA63DA1292B81188">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2C88E92DABF83E81D6007B1B7DEF672E">Grego, Jozef</mods:namePart>
</mods:name>
<mods:name id="D0B2CAC0DF8EECD1B6328AB80DB0A8DA" type="personal">
<mods:role id="700004850C32D20739058D7156060FBC">
<mods:roleTerm id="41A0219B3A21C33C68E3EC0508B32C30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="628488EACC2E4CD6ACA908E8C982DE1F">Vermeulen, Jaap J.</mods:namePart>
</mods:name>
<mods:name id="22282D9793E44DE628A02D569FD27C91" type="personal">
<mods:role id="54EF12CFFFE194F87786B6DC398D0517">
<mods:roleTerm id="DBD177B5E22AF02CDE1568764103CFF8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00875EEA6DA0CAAC3EB444ABFC6F3264">Reischütz, Alexander</mods:namePart>
</mods:name>
<mods:name id="5C558B0512EDE1CE5E2BE040CB12AC0A" type="personal">
<mods:role id="BF1FF7D8D681D4777DA1AF6AD927B9D8">
<mods:roleTerm id="F37D4E7C92794F8DE93269A391282BC5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BE5D061588E6289DDD9427BE61CE7E8">Hunyadi, András</mods:namePart>
</mods:name>
<mods:name id="9C2DBBEDD1651786148D600F8FE9593B" type="personal">
<mods:role id="A1F23B6307D1C25453C4AA07B7E664CD">
<mods:roleTerm id="E5AB852187EF9A5BA3C0C63B6123A44B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CD7FF2611B2E1B792554DA5F0D26D119">Jochum, Adrienne</mods:namePart>
</mods:name>
<mods:typeOfResource id="F20F781BB39A5A111604745EBB11BF63">text</mods:typeOfResource>
<mods:relatedItem id="C4DB1BB24A13EEED3FB1CD40647404E1" type="host">
<mods:titleInfo id="3EF2B37A8C2F5FC2ED3F74BF88356B40">
<mods:title id="9E0D815B3802F16680CC55F529609A5C">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="2D4CFC04ED09A108F37CEFAFE4C1FABB">
<mods:date id="B62C69880AE59A08D9EE8D67658CAD0F">2019</mods:date>
<mods:detail id="7D81C44DEFB7DE2E03D147E78E97978F" type="pubDate">
<mods:number id="9F6DC1A7893AFE61703C970C22CEF67C">2019-11-08</mods:number>
</mods:detail>
<mods:detail id="34B1B78BB4F5B22C1BB9484894E2912B" type="volume">
<mods:number id="EADF8D2E5D1AD49C2029BFB7EB76B152">67</mods:number>
</mods:detail>
<mods:extent id="A028D88898F7CD5BB9E6840054363E86" unit="page">
<mods:start id="22E6C9005907B4FE08E03750159AEC21">517</mods:start>
<mods:end id="6C1585BE500593F9CF824A160C503BF7">535</mods:end>
</mods:extent>
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<mods:classification id="8B5D3474DB99F3A904A5E670C67F3498">journal article</mods:classification>
<mods:identifier id="0264F38CA8729968C8E7CD539C670E0E" type="DOI">10.26107/RBZ-2019-0041</mods:identifier>
<mods:identifier id="7C77B109AEF8AB9E331F20E2DB12C43F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="A6DCED9DEEDDB13872CA6194330E7276" type="Zenodo-Dep">4576444</mods:identifier>
<mods:identifier id="6D4D29B20130D408FAA53601227597B7" type="ZooBank">AAA866A1-FBAF-486D-AB84-3AB989A269C1</mods:identifier>
</mods:mods>
<treatment id="03BF447DFF91FFBBFC91FF48FE42FA92" ID-DOI="http://doi.org/10.5281/zenodo.4576557" ID-Zenodo-Dep="4576557" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFBBFC91FF48FE42FA92" httpUri="http://treatment.plazi.org/id/03BF447DFF91FFBBFC91FF48FE42FA92" lastPageId="16" lastPageNumber="533" pageId="12" pageNumber="529">
<subSubSection id="C30CA6E0FF91FFA7FC91FF48FB2EFAB1" pageId="12" pageNumber="529" type="nomenclature">
<paragraph id="8BA9F56BFF91FFA7FC91FF48FB2EFAB1" blockId="12.[895,1416,1340,1429]" pageId="12" pageNumber="529">
<heading id="D0E14207FF91FFA7FC91FF48FB2EFAB1" bold="true" centered="true" fontSize="10" level="2" pageId="12" pageNumber="529" reason="2">
<emphasis id="B9622979FF91FFA7FC91FF48FB2EFAB1" bold="true" pageId="12" pageNumber="529">
<taxonomicName id="4C168EE8FF91FFA7FC91FF48FA65FA91" ID-CoL="7TCV2" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2019" box="[895,1411,1340,1365]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="529" phylum="Mollusca" rank="species" species="suturata" status="sp. nov.">
<emphasis id="B9622979FF91FFA7FC91FF48FB9EFA91" bold="true" box="[895,1144,1341,1365]" italics="true" pageId="12" pageNumber="529">Tonkinospira suturata</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="A2519402FF91FFA7FBACFF28FB2EFAB1" box="[1090,1224,1373,1397]" pageId="12" pageNumber="529" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FBBDFF08FB55FA51" box="[1107,1203,1405,1429]" pageId="12" pageNumber="529" type="description">
<paragraph id="8BA9F56BFF91FFA7FBBDFF08FB55FA51" blockId="12.[895,1416,1340,1429]" box="[1107,1203,1405,1429]" pageId="12" pageNumber="529">
(
<figureCitation id="132DE9EEFF91FFA7FBB5FF08FB4CFA51" box="[1115,1194,1405,1429]" captionStart="Fig" captionStartId="14.[136,171,2091,2113]" captionTargetBox="[154,1454,189,2067]" captionTargetId="figure-7@14.[154,1454,189,2067]" captionTargetPageId="14" captionText="Fig. 11. Tonkinospira suturata Páll-Gergely &amp; Grego, new species, HNHM 104405 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4576480" httpUri="https://zenodo.org/record/4576480/files/figure.png" pageId="12" pageNumber="529">Fig. 11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FCADFFC8FACBF9B1" pageId="12" pageNumber="529" type="materials_examined">
<paragraph id="8BA9F56BFF91FFA7FCADFFC8FACBF9B1" blockId="12.[834,1475,1469,1654]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADFFC8FC08FA11" bold="true" box="[835,1006,1469,1493]" pageId="12" pageNumber="529">Type material.</emphasis>
<materialsCitation id="3B7EFF36FF91FFA7FC19FFC8FACBF9B1" collectingDate="2017-02-11" collectionCode="JG" collectorName="J. Grego." country="Laos" elevation="160" latitude="17.563583" location="Earthquake Dome" longLatPrecision="1" longitude="104.871635" municipality="Tham Nam Don Cave" pageId="12" pageNumber="529" specimenCode="HNHM 104405, JG2" specimenCount="2" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="54AD4BC9FF91FFA7FC19FFC8FBBCFA11" box="[1015,1114,1469,1493]" pageId="12" pageNumber="529" type="holotype">Holotype</typeStatus>
(H:
<quantity id="4CEE588EFF91FFA7FB7EFFC8FB12FA11" box="[1168,1268,1469,1493]" metricMagnitude="-3" metricUnit="m" metricValue="1.41" pageId="12" pageNumber="529" unit="mm" value="1.41">1.41 mm</quantity>
, D:
<quantity id="4CEE588EFF91FFA7FAC8FFC8FA6CFA11" box="[1318,1418,1469,1493]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="12" pageNumber="529" unit="mm" value="1.84">1.84 mm</quantity>
,
<figureCitation id="132DE9EEFF91FFA7FA76FFC8FCB8FA31" captionStart="Fig" captionStartId="14.[136,171,2091,2113]" captionTargetBox="[154,1454,189,2067]" captionTargetId="figure-7@14.[154,1454,189,2067]" captionTargetPageId="14" captionText="Fig. 11. Tonkinospira suturata Páll-Gergely &amp; Grego, new species, HNHM 104405 (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4576480" httpUri="https://zenodo.org/record/4576480/files/figure.png" pageId="12" pageNumber="529">Fig. 11</figureCitation>
) (
<specimenCode id="DBB05D10FF91FFA7FC98FFA8FBC2FA31" box="[886,1060,1501,1525]" collectionCode="HNHM" country="Hungary" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="12" pageNumber="529">HNHM 104405</specimenCode>
),
<collectingCountry id="F301B5FBFF91FFA7FBD9FFABFB8BFA31" box="[1079,1133,1502,1525]" name="Laos" pageId="12" pageNumber="529">Laos</collectingCountry>
,
<collectingRegion id="49D23B89FF91FFA7FB94FFA8FA9FFA31" box="[1146,1401,1501,1525]" country="Laos" name="Khammouan" pageId="12" pageNumber="529">Khammouane Province</collectingRegion>
,
<collectingMunicipality id="6BCD6F11FF91FFA7FA6AFFA8FC01F9D1" pageId="12" pageNumber="529">Tham Nam Dôn Cave</collectingMunicipality>
,
<location id="8EC9A3B0FF91FFA7FC1EFF88FB56F9D1" LSID="urn:lsid:plazi:treatment:03BF447DFF91FFBBFC91FF48FE42FA92:8EC9A3B0FF91FFA7FC1EFF88FB56F9D1" box="[1008,1200,1533,1557]" country="Laos" latitude="17.563583" longLatPrecision="1" longitude="104.871635" municipality="Tham Nam Don Cave" name="Earthquake Dome" pageId="12" pageNumber="529" stateProvince="Khammouan">Earthquake Dome</location>
, sand sediments at bank of cave river,
<elevation id="003B1258FF91FFA7FC59FC68FBD7F9F1" box="[951,1073,1565,1589]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="529" unit="m" value="160.0">
<quantity id="4CEE588EFF91FFA7FC59FC68FC1FF9F1" box="[951,1017,1565,1589]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="529" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
,
<geoCoordinate id="EE2293ACFF91FFA7FBD2FC69FB21F9F1" box="[1084,1223,1564,1589]" degrees="17" direction="north" minutes="33.815" orientation="latitude" pageId="12" pageNumber="529" precision="1" value="17.563583">17°33.815′N</geoCoordinate>
,
<geoCoordinate id="EE2293ACFF91FFA7FB3EFC69FA84F9F1" box="[1232,1378,1564,1589]" degrees="104" direction="east" minutes="52.298" orientation="longitude" pageId="12" pageNumber="529" precision="1" value="104.871635">104°52.298′E</geoCoordinate>
(locality code:
<specimenCode id="DBB05D10FF91FFA7FC6BFC48FC52F991" box="[901,948,1597,1621]" collectionCode="JG" pageId="12" pageNumber="529">JG2</specimenCode>
B), coll.
<collectorName id="26E390BDFF91FFA7FBE1FC4BFB92F991" box="[1039,1140,1597,1621]" pageId="12" pageNumber="529">J. Grego.</collectorName>
,
<date id="FFA8D3ABFF91FFA7FB91FC48FAA4F991" box="[1151,1346,1597,1621]" pageId="12" pageNumber="529" value="2017-02-11">
<collectingDate id="EFEC2A43FF91FFA7FB91FC48FAA4F991" box="[1151,1346,1597,1621]" pageId="12" pageNumber="529" value="2017-02-11">11 February 2017</collectingDate>
</date>
; 2 damaged
<typeStatus id="54AD4BC9FF91FFA7FCAAFC2BFC4CF9B2" box="[836,938,1630,1654]" pageId="12" pageNumber="529" type="paratype">paratypes</typeStatus>
(coll.
<collectionCode id="ED076DAEFF91FFA7FC1DFC28FBF2F9B1" box="[1011,1044,1629,1653]" pageId="12" pageNumber="529">JG</collectionCode>
), same data as preceding.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFA7FCADFCE8FB09F8D1" pageId="12" pageNumber="529" type="diagnosis">
<paragraph id="8BA9F56BFF91FFA7FCADFCE8FB09F8D1" blockId="12.[834,1475,1693,1813]" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADFCE8FC5EF971" bold="true" box="[835,952,1693,1717]" pageId="12" pageNumber="529">Diagnosis.</emphasis>
A large
<taxonomicName id="4C168EE8FF91FFA7FBF3FCE8FB4DF971" authorityName="Jochum, Slapnik &amp; Pall-Gergely" authorityYear="2014" box="[1053,1195,1693,1717]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="529" phylum="Mollusca" rank="genus">
<emphasis id="B9622979FF91FFA7FBF3FCE8FB4DF971" box="[1053,1195,1693,1717]" italics="true" pageId="12" pageNumber="529">Tonkinospira</emphasis>
</taxonomicName>
species with a depressed conical shell, rounded body whorl, fine radial growth lines and without spiral striation, a wide umbilicus, and a peristome that is not adnate to penultimate whorl.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF91FFBBFCADFD48FEB1FB92" lastPageId="16" lastPageNumber="533" pageId="12" pageNumber="529" type="description">
<paragraph id="8BA9F56BFF91FFBBFCADFD48FEB1FB92" blockId="12.[834,1474,1853,2101]" lastBlockId="16.[136,775,766,1110]" lastPageId="16" lastPageNumber="533" pageId="12" pageNumber="529">
<emphasis id="B9622979FF91FFA7FCADFD48FC29F891" bold="true" box="[835,975,1853,1877]" pageId="12" pageNumber="529">Description.</emphasis>
Shell large for the genus; depressed conical with rounded body whorl; shell wider than tall; 4.25 whorls, regularly increasing, separated by a very deep suture; whorls shouldered; protoconch consists of 1.5 whorls, with extremely fine radial growth lines and without spiral striation (on the first whorl of protoconch some spiral striae visible, but they are not regular, probably due to erosion of the shell surface); teleoconch overall weakly sculptured, with irregular, fine radial growth lines, and relatively dense, regular, obscure spiral striation, ca.
<quantity id="4CEE588EFF8DFFBBFE8FF96BFE7DFCF2" box="[353,411,798,822]" metricMagnitude="-1" metricUnit="m" metricValue="5.588" pageId="16" pageNumber="533" unit="in" value="22.0">22 in</quantity>
number in the middle shell line from apertural view; aperture almost in alignment with penultimate whorl and very slightly oblique to shell axis in lateral view; aperture rounded-ovate, slightly free from the preceding whorl along the parietal margin; peristome very slightly expanded, not reflected; aperture not adnate to penultimate whorl, protruding into a truncated tuba (best seen from umbilical perspective); two weak, very low suggestions of parietal barriers are discernible; umbilicus wider than narrow.
</paragraph>
</subSubSection>
<caption id="DF69A5E3FF90FFA6FF9FF25EFB80F785" ID-DOI="http://doi.org/10.5281/zenodo.4576478" ID-Zenodo-Dep="4576478" box="[113,1126,2091,2113]" httpUri="https://zenodo.org/record/4576478/files/figure.png" pageId="13" pageNumber="530" startId="13.[113,148,2091,2113]" targetBox="[131,1430,189,2067]" targetPageId="13">
<paragraph id="8BA9F56BFF90FFA6FF9FF25EFB80F785" blockId="13.[113,1126,2091,2113]" box="[113,1126,2091,2113]" pageId="13" pageNumber="530">
Fig. 10.
<taxonomicName id="4C168EE8FF90FFA6FF28F25EFD2CF785" authority="Pall-Gergely &amp; A. Reischutz" authorityName="Pall-Gergely &amp; A. Reischutz" authorityYear="2019" box="[198,714,2091,2113]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="530" phylum="Mollusca" rank="species" species="raxajacki" status="sp. nov.">
<emphasis id="B9622979FF90FFA6FF28F25EFE4FF784" box="[198,425,2091,2112]" italics="true" pageId="13" pageNumber="530">Tonkinospira raxajacki</emphasis>
Páll-Gergely &amp; A. Reischütz
</taxonomicName>
<taxonomicNameLabel id="A2519402FF90FFA6FD3FF25EFCAFF785" box="[721,841,2091,2113]" pageId="13" pageNumber="530" rank="species">new species</taxonomicNameLabel>
, NHMW 113205 (holotype).
</paragraph>
</caption>
<caption id="DF69A5E3FF93FFA5FF66F25EFBD7F785" ID-DOI="http://doi.org/10.5281/zenodo.4576480" ID-Zenodo-Dep="4576480" box="[136,1073,2091,2113]" httpUri="https://zenodo.org/record/4576480/files/figure.png" pageId="14" pageNumber="531" startId="14.[136,171,2091,2113]" targetBox="[154,1454,189,2067]" targetPageId="14">
<paragraph id="8BA9F56BFF93FFA5FF66F25EFBD7F785" blockId="14.[136,1073,2091,2113]" box="[136,1073,2091,2113]" pageId="14" pageNumber="531">
Fig. 11.
<taxonomicName id="4C168EE8FF93FFA5FF30F25EFD71F784" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2019" box="[222,663,2091,2113]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="531" phylum="Mollusca" rank="species" species="suturata" status="sp. nov.">
<emphasis id="B9622979FF93FFA5FF30F25EFE50F784" box="[222,438,2091,2112]" italics="true" pageId="14" pageNumber="531">Tonkinospira suturata</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="A2519402FF93FFA5FD4CF25EFCFCF785" box="[674,794,2091,2113]" pageId="14" pageNumber="531" rank="species">new species</taxonomicNameLabel>
, HNHM 104405 (holotype).
</paragraph>
</caption>
<caption id="DF69A5E3FF92FFA4FF9FF25EFB68F785" ID-DOI="http://doi.org/10.5281/zenodo.4576484" ID-Zenodo-Dep="4576484" box="[113,1166,2091,2113]" httpUri="https://zenodo.org/record/4576484/files/figure.png" pageId="15" pageNumber="532" startId="15.[113,148,2091,2113]" targetBox="[131,1430,189,2067]" targetPageId="15">
<paragraph id="8BA9F56BFF92FFA4FF9FF25EFB68F785" blockId="15.[113,1166,2091,2113]" box="[113,1166,2091,2113]" pageId="15" pageNumber="532">
Fig. 12.
<taxonomicName id="4C168EE8FF92FFA4FF28F25EFD2CF785" authority="Pall-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2019" box="[198,714,2091,2113]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="15" pageNumber="532" phylum="Mollusca" rank="species" species="triangulata" status="sp. nov.">
<emphasis id="B9622979FF92FFA4FF28F25EFE5CF784" box="[198,442,2091,2112]" italics="true" pageId="15" pageNumber="532">Tonkinospira triangulata</emphasis>
Páll-Gergely &amp; Vermeulen
</taxonomicName>
,
<taxonomicNameLabel id="A2519402FF92FFA4FD38F25EFCABF785" box="[726,845,2091,2113]" pageId="15" pageNumber="532" rank="species">new species</taxonomicNameLabel>
, NHMUK 20190687 (holotype).
</paragraph>
</caption>
<caption id="DF69A5E3FF8DFFBBFF69F8C4FB76FD03" ID-DOI="http://doi.org/10.5281/zenodo.4576486" ID-Zenodo-Dep="4576486" box="[135,1168,689,711]" httpUri="https://zenodo.org/record/4576486/files/figure.png" pageId="16" pageNumber="533" startId="16.[135,170,689,711]" targetBox="[135,1474,189,674]" targetPageId="16">
<paragraph id="8BA9F56BFF8DFFBBFF69F8C4FB76FD03" blockId="16.[135,1168,689,711]" box="[135,1168,689,711]" pageId="16" pageNumber="533">
Fig. 13. Variability within
<taxonomicName id="4C168EE8FF8DFFBBFE7FF8C4FC73FD03" authority="Pall-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2019" box="[401,917,689,711]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="triangulata" status="sp. nov.">
<emphasis id="B9622979FF8DFFBBFE7FF8C4FD63FD02" box="[401,645,689,710]" italics="true" pageId="16" pageNumber="533">Tonkinospira triangulata</emphasis>
Páll-Gergely &amp; Vermeulen
</taxonomicName>
,
<taxonomicNameLabel id="A2519402FF8DFFBBFC4FF8C7FBF3FD03" box="[929,1045,689,711]" pageId="16" pageNumber="533" rank="species">new species</taxonomicNameLabel>
(JJV 6265).
</paragraph>
</caption>
<subSubSection id="C30CA6E0FF8DFFBBFF66FE0BFDADFB32" pageId="16" pageNumber="533" type="diagnosis">
<paragraph id="8BA9F56BFF8DFFBBFF66FE0BFDADFB32" blockId="16.[136,775,1150,1270]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFF66FE0BFE60FB52" bold="true" box="[136,390,1150,1174]" pageId="16" pageNumber="533">Differential diagnosis.</emphasis>
The most similar species in terms of shell size and the weak sculpture is
<taxonomicName id="4C168EE8FF8DFFBBFDD4FEEAFD53FB72" box="[570,693,1183,1206]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="tomasini">
<emphasis id="B9622979FF8DFFBBFDD4FEEAFD53FB72" box="[570,693,1183,1206]" italics="true" pageId="16" pageNumber="533">T. tomasini</emphasis>
</taxonomicName>
, which has a much more elevated spire, and adnate peristome and a more oblique aperture to the shell axis.
</paragraph>
</subSubSection>
<subSubSection id="C30CA6E0FF8DFFBBFF66FF6BFE42FA92" pageId="16" pageNumber="533" type="etymology">
<paragraph id="8BA9F56BFF8DFFBBFF66FF6BFE42FA92" blockId="16.[136,774,1310,1366]" pageId="16" pageNumber="533">
<emphasis id="B9622979FF8DFFBBFF66FF6BFEECFAF2" bold="true" box="[136,266,1310,1334]" pageId="16" pageNumber="533">Etymology.</emphasis>
The specific epithet
<taxonomicName id="4C168EE8FF8DFFBBFE05FF6AFDA2FAF2" authorityName="Pall-Gergely &amp; Grego" authorityYear="2019" box="[491,580,1311,1334]" class="Gastropoda" family="Gastrocoptidae" genus="Tonkinospira" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="533" phylum="Mollusca" rank="species" species="suturata">
<emphasis id="B9622979FF8DFFBBFE05FF6AFDA2FAF2" box="[491,580,1311,1334]" italics="true" pageId="16" pageNumber="533">suturata</emphasis>
</taxonomicName>
refers to the deep suture of the new species.
</paragraph>
</subSubSection>
</treatment>
</document>
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