285 lines
38 KiB
XML
285 lines
38 KiB
XML
<document id="A63882AE517AE07D48602F35F1DF8A57" ID="10.11646/zootaxa.4168.2.2" ID-DOI="10.11646/zootaxa.4168.2.2" ID-GBIF-Dataset="8caff7dc-a5bc-4194-a7ec-739e149d63d3" ID-ISSN="1175-5326" ID-Zenodo-Dep="262140" ID-ZooBank="BEC1B33E-ACB6-4A35-813A-413D08610617" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473921602333" checkinUser="plazi" docAuthor="Doughty, Paul, Ellis, Ryan J. & Oliver, Paul M." docDate="2016" docId="03AABA6D1809AA6CFF5BFC14FE966746" docLanguage="en" docName="zootaxa.4168.2.2.pdf" docOrigin="Zootaxa 4168 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Crenadactylus ocellatus Gray 1845" docType="treatment" docVersion="7" lastPageNumber="250" masterDocId="FF93C2151801AA67FFCCFFE5FFCD6263" masterDocTitle="Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia" masterLastPageNumber="278" masterPageNumber="239" pageNumber="247" updateTime="1698385905195" updateUser="plazi">
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<mods:mods id="467756DD1D36FAC8D9F91CA7D349AAE8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BF2EB3ED1397677B2CFB754A89B89AA5">
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<mods:title id="1868EF1B8CBFBE0112C9229F4FFB0F87">Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia</mods:title>
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<mods:name id="00CC9DC6A1D09FC56FFE58D63C4AA28C" type="personal">
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<mods:role id="5D3BB34E48E251D151B3246FD1FE1017">
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<mods:roleTerm id="2B65FE32E75811FC1A184D59E27F17FB">Author</mods:roleTerm>
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<mods:namePart id="107582EC9D167866730B43115E965804">Doughty, Paul</mods:namePart>
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<mods:roleTerm id="6FAE85620CB8386B32AB349EB17FBDF4">Author</mods:roleTerm>
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<mods:namePart id="4A4CA46E6E8010AB6BD143A49ABB96DA">Ellis, Ryan J.</mods:namePart>
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</mods:name>
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<mods:name id="68777135BE2642882D26752B80406421" type="personal">
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<mods:roleTerm id="1F60EB818E95204699053162CF024FA9">Author</mods:roleTerm>
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<mods:namePart id="4549189BCF7C18C07F2F37EC92ED4B14">Oliver, Paul M.</mods:namePart>
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<mods:typeOfResource id="91FD36B4DC044AB00A7862D8F62847EC">text</mods:typeOfResource>
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<mods:title id="E6FB829C0FB0829532EE4131FD552524">Zootaxa</mods:title>
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<mods:part id="8C0698078E20BB6FA982CFD393D1CF39">
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<mods:date id="359BE085573D299001C21262F80F7743">2016</mods:date>
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<mods:detail id="03C880A0E2BCDD9A7D5B1C6C9C95A9EE" type="volume">
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<mods:number id="84626D750CB48E233367414687359304">4168</mods:number>
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<mods:detail id="BAA3A1007DC4E66E15DA1E2D503EE829" type="issue">
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<mods:number id="5E10F6E9398FB3E4297AE06B62B4E2BD">2</mods:number>
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<mods:classification id="05661D1C7F6F6D3C5E6F3FF92FCECFB2">journal article</mods:classification>
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<mods:identifier id="239C00E5E1340EFEC718BB350A58C5F2" type="DOI">10.11646/zootaxa.4168.2.2</mods:identifier>
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<mods:identifier id="EDD47A5380F9437510E3D7ABC9436D9C" type="GBIF-Dataset">8caff7dc-a5bc-4194-a7ec-739e149d63d3</mods:identifier>
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<mods:identifier id="B05D350BA1DE2101BB33251FDBC60F1B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03AABA6D1809AA6CFF5BFC14FE966746" ID-DOI="http://doi.org/10.5281/zenodo.6060770" ID-GBIF-Taxon="124551737" ID-Zenodo-Dep="6060770" LSID="urn:lsid:plazi:treatment:03AABA6D1809AA6CFF5BFC14FE966746" httpUri="http://treatment.plazi.org/id/03AABA6D1809AA6CFF5BFC14FE966746" lastPageId="11" lastPageNumber="250" pageId="8" pageNumber="247">
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<subSubSection id="C31958F01809AA6FFF5BFC14FE86662C" pageId="8" pageNumber="247" type="nomenclature">
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<paragraph id="8BBC0B7B1809AA6FFF5BFC14FD966668" blockId="8.[151,603,1009,1104]" box="[151,603,1009,1035]" pageId="8" pageNumber="247">
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<heading id="D0F4BC171809AA6FFF5BFC14FD966668" bold="true" box="[151,603,1009,1035]" fontSize="11" level="1" pageId="8" pageNumber="247" reason="1">
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<taxonomicName id="4C0370F81809AA6FFF5BFC14FD966668" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" box="[151,603,1009,1035]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="247" phylum="Chordata" rank="species" species="ocellatus">
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<emphasis id="B977D7691809AA6FFF5BFC14FD966668" bold="true" box="[151,603,1009,1035]" pageId="8" pageNumber="247">
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<emphasis id="B977D7691809AA6FFF5BFC14FE756668" bold="true" box="[151,440,1009,1035]" italics="true" pageId="8" pageNumber="247">Crenadactylus ocellatus</emphasis>
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(
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<bibRefCitation id="EF92768A1809AA6FFE0BFC14FD9E6668" author="Gray" box="[455,595,1009,1035]" pageId="8" pageNumber="247" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Trustees if the British Museum / Edward Newman, London, xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BBC0B7B1809AA6FFF5BFBF1FE86662C" blockId="8.[151,603,1009,1104]" pageId="8" pageNumber="247">
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South-western clawless geckos
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<figureCitation id="133817FE1809AA6FFF5BFBD2FF296633" box="[151,228,1079,1104]" captionStart="FIGURE 4" captionStartId="7.[151,250,1853,1875]" captionTargetBox="[166,1420,210,1826]" captionTargetId="figure@7.[150,1436,193,1843]" captionTargetPageId="7" captionText="FIGURE 4. Photographs of Crenadactylus ocellatus species-group members in life (left to right, top to bottom): A) C. ocellatus, Koolyanobbing, WA (photo—R. J. Ellis), B) C. horni, Trephina Gorge, NT (photo—G. M. Shea), C) occidentalis sp. nov., 70 km S Exmouth, WA (photo—G. Gaikhorst), D) C. tuberculatus sp. nov., Cape Range National Park, WA (photo—R. J. Ellis), E) C. pilbarensis sp. nov., Burrup Peninsula, WA (photo—B. Maryan), F) C. rostralis, Geikie Gorge National Park, WA (photo—P. M. Oliver), G) C. naso, Koolan Island, WA (photo—G. Gaikhorst), H) C. naso, Mitchell Plateau, WA (Photo— G. M. Shea)." httpUri="https://zenodo.org/record/262144/files/figure.png" pageId="8" pageNumber="247">Figs. 4</figureCitation>
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A, 5, 6, 7
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31958F01809AA6FFF5BFB65FDDD66B0" pageId="8" pageNumber="247" type="reference_group">
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<paragraph id="8BBC0B7B1809AA6FFF5BFB65FE3366F5" blockId="8.[151,849,1152,1235]" box="[151,510,1152,1174]" pageId="8" pageNumber="247">
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<taxonomicName id="4C0370F81809AA6FFF5BFB65FE3366F5" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" box="[151,510,1152,1174]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="247" phylum="Chordata" rank="species" species="ocellatus">
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<emphasis id="B977D7691809AA6FFF5BFB65FE4B66F5" box="[151,390,1152,1174]" italics="true" pageId="8" pageNumber="247">Diplodactylus ocellatus</emphasis>
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<bibRefCitation id="EF92768A1809AA6FFE41FB65FE3366F5" author="Gray" box="[397,510,1152,1174]" pageId="8" pageNumber="247" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Trustees if the British Museum / Edward Newman, London, xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
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</taxonomicName>
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||
</paragraph>
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<paragraph id="8BBC0B7B1809AA6FFF5BFB7BFDDD66B0" blockId="8.[151,849,1152,1235]" pageId="8" pageNumber="247">
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Synonymy:
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<taxonomicName id="4C0370F81809AA6FFEDEFB7BFD4C66D7" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" box="[274,641,1182,1204]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="247" phylum="Chordata" rank="species" species="bilineatus">
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<emphasis id="B977D7691809AA6FFEDEFB7BFDC766D7" box="[274,522,1182,1204]" italics="true" pageId="8" pageNumber="247">Diplodactylus bilineatus</emphasis>
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<bibRefCitation id="EF92768A1809AA6FFDDCFB7BFD4C66D7" author="Gray" box="[528,641,1182,1204]" pageId="8" pageNumber="247" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Trustees if the British Museum / Edward Newman, London, xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
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</taxonomicName>
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(fide
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<bibRefCitation id="EF92768A1809AA6FFD71FB7BFC8766D7" author="Gunther" box="[701,842,1182,1204]" pageId="8" pageNumber="247" refString="Gunther, A. (1867) Additions to the knowledge of Australian reptiles and fishes. Annals and Magazine of Natural History, 20, 45 - 68." type="journal article" year="1867">Günther 1867</bibRefCitation>
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) [‘South-west’ of
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<bibRefCitation id="EF92768A1809AA6FFE88FB58FDC466B0" box="[324,521,1213,1235]" pageId="8" pageNumber="247" refString="Oliver, P., Adams, M. & Doughty, P. (2010) Molecular evidence for ten species and Oligo-Miocene vicariance within a nominal Australian gecko species (Crenadactylus ocellatus, Diplodactylidae). BMC Evolutionary Biology, 10, 386. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 10 - 386" type="journal article">
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Oliver
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<emphasis id="B977D7691809AA6FFE47FB5BFE7366B0" box="[395,446,1213,1235]" italics="true" pageId="8" pageNumber="247">et al.</emphasis>
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(2010)
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</bibRefCitation>
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]
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31958F01809AA6FFF5BFB1AFB1167E6" pageId="8" pageNumber="247" type="materials_examined">
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||
<paragraph id="8BBC0B7B1809AA6FFF5BFB1AFDDE675E" blockId="8.[151,1436,1279,2025]" pageId="8" pageNumber="247">
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||
<materialsCitation id="3B6B01261809AA6FFF5BFB1AFDDE675E" ID-GBIF-Occurrence="1311491181" collectionCode="BMNH" location="Buchanan" pageId="8" pageNumber="247" specimenCode="BMNH 1947.3" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<emphasis id="B977D7691809AA6FFF5BFB1AFEC6677B" bold="true" box="[151,267,1279,1304]" pageId="8" pageNumber="247">
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<typeStatus id="54B8B5D91809AA6FFF5BFB1AFECB677B" box="[151,262,1279,1304]" pageId="8" pageNumber="247" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<specimenCode id="DBA5A3001809AA6FFED9FAE5FE0E677B" box="[277,451,1279,1304]" collectionCode="BMNH" pageId="8" pageNumber="247">
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||
<collectionCode id="ED1293BE1809AA6FFED9FAE5FEA46774" box="[277,361,1280,1303]" pageId="8" pageNumber="247">BMNH</collectionCode>
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||
1947.3
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</specimenCode>
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.6.68 from
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<collectingRegion id="49C7C5991809AA6FFD88FAE5FCDE677B" box="[580,787,1279,1304]" country="Australia" name="Western Australia" pageId="8" pageNumber="247">
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<collectingRegion id="49C7C5991809AA6FFD88FAE5FD526774" box="[580,671,1280,1303]" country="American Samoa" name="Western" pageId="8" pageNumber="247">Western</collectingRegion>
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Australia
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</collectingRegion>
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. Specimen obtained from the
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<location id="8EDC5DA01809AA6FFBBEFB1AFB2F677B" LSID="urn:lsid:plazi:treatment:03AABA6D1809AA6CFF5BFC14FE966746:8EDC5DA01809AA6FFBBEFB1AFB2F677B" box="[1138,1250,1279,1304]" name="Buchanan" pageId="8" pageNumber="247" stateProvince="Western Australia">Buchanan</location>
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collection from WA, collection date not specified.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BBC0B7B1809AA6FFF0BFAA2FB1167E6" blockId="8.[151,1436,1279,2025]" pageId="8" pageNumber="247">
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<materialsCitation id="3B6B01261809AA6FFF0BFAA2FB1167E6" ID-GBIF-Occurrence="1311491182" collectionCode="BMNH" location="Houtman's Abrolhos" pageId="8" pageNumber="247" specimenCode="BMNH 1947.3" specimenCount="2" stateProvince="Western Australia" typeStatus="syntype">
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<emphasis id="B977D7691809AA6FFF0BFAA2FE866703" bold="true" box="[199,331,1351,1376]" pageId="8" pageNumber="247">Synonomy.</emphasis>
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<taxonomicName id="4C0370F81809AA6FFE94FAA2FDBC6703" box="[344,625,1351,1376]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="247" phylum="Chordata" rank="species" species="bilineatus">
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||
<emphasis id="B977D7691809AA6FFE94FAA2FDBC6703" box="[344,625,1351,1376]" italics="true" pageId="8" pageNumber="247">Diplodactylus bilineatus</emphasis>
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</taxonomicName>
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.
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<typeStatus id="54B8B5D91809AA6FFD49FAADFD216703" box="[645,748,1352,1376]" pageId="8" pageNumber="247" type="syntype">Syntypes</typeStatus>
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(2),
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<bibRefCitation id="EF92768A1809AA6FFCE3FAADFC7D6703" author="Gray" box="[815,944,1351,1376]" pageId="8" pageNumber="247" refString="Gray, J. E. (1845) Catalogue of the Specimens of Lizards in the Collection of the British Museum. Trustees if the British Museum / Edward Newman, London, xxviii + 289 pp." type="book" year="1845">Gray 1845</bibRefCitation>
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||
,
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<specimenCode id="DBA5A3001809AA6FFC0DFAADFBBC6703" box="[961,1137,1351,1376]" collectionCode="BMNH" pageId="8" pageNumber="247">
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||
<collectionCode id="ED1293BE1809AA6FFC0DFAADFBD8673C" box="[961,1045,1352,1375]" pageId="8" pageNumber="247">BMNH</collectionCode>
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||
1947.3
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||
</specimenCode>
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||
.6.69 (
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||
<specimenCount id="9D05C0F21809AA6FFB72FAADFA9C6703" box="[1214,1361,1351,1376]" pageId="8" pageNumber="247" type="generic">2 specimens</specimenCount>
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||
) from
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<location id="8EDC5DA01809AA6FFF5BFA89FE4967E6" LSID="urn:lsid:plazi:treatment:03AABA6D1809AA6CFF5BFC14FE966746:8EDC5DA01809AA6FFF5BFA89FE4967E6" box="[151,388,1388,1413]" name="Houtman's Abrolhos" pageId="8" pageNumber="247" stateProvince="Western Australia">Houtman’s Abrolhos</location>
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||
,
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||
<collectingRegion id="49C7C5991809AA6FFE43FA88FD9667E6" box="[399,603,1388,1413]" country="Australia" name="Western Australia" pageId="8" pageNumber="247">
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||
<collectingRegion id="49C7C5991809AA6FFE43FA88FE2767E7" box="[399,490,1389,1412]" country="American Samoa" name="Western" pageId="8" pageNumber="247">Western</collectingRegion>
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||
Australia
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||
</collectingRegion>
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. Collected by ‘Mr. Gilbert’, collection date not specified.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31958F01809AA6FFF0BFA6AFDB864C7" pageId="8" pageNumber="247" type="diagnosis">
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||
<paragraph id="8BBC0B7B1809AA6FFF0BFA6AFDB864C7" blockId="8.[151,1436,1279,2025]" pageId="8" pageNumber="247">
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||
<emphasis id="B977D7691809AA6FFF0BFA6AFE8C67CB" bold="true" box="[199,321,1423,1448]" pageId="8" pageNumber="247">Diagnosis.</emphasis>
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||
A moderately large (to
|
||
<quantity id="4CFBA69E1809AA6FFD94FA6AFD7267C4" box="[600,703,1423,1448]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="8" pageNumber="247" unit="mm" value="35.5">35.5 mm</quantity>
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||
SVL) species of
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||
<taxonomicName id="4C0370F81809AA6FFC46FA6AFBE167CB" box="[906,1068,1423,1448]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="247" phylum="Chordata" rank="genus">
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||
<emphasis id="B977D7691809AA6FFC46FA6AFBE167CB" box="[906,1068,1423,1448]" italics="true" pageId="8" pageNumber="247">Crenadactylus</emphasis>
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||
</taxonomicName>
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||
with wide head (HW/HL 0.52– 0.70) and short trunk length (ILL/SVL 0.37–0.50). Rostral in full contact with nostril, internasal (if present) not extending beyond supranasal, 1 or 2 granular postmentals, dorsal scales homogeneous with smooth to weak keels, usually no pre-cloacal pores visible but some males with 2, when present pore-bearing scales in contact at midline, no enlarged tubercles on original tails. Ground colour tan and dark brown; dorsal pattern comprised of poorlydefined longitudinal pale and dark stripes, lateral zones pale with irregular dark stippling, pattern heavily overlain with intermixed pale and dark scales giving an irregular appearance, pale spots or ocelli comprised of 3–6 pale scales usually present in dorsolateral zone.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31958F01809AA6EFF0BF94AFDEB6462" lastPageId="9" lastPageNumber="248" pageId="8" pageNumber="247" type="description">
|
||
<paragraph id="8BBC0B7B1809AA6EFF0BF94AFD7C63B3" blockId="8.[151,1436,1279,2025]" lastBlockId="9.[151,1437,151,2034]" lastPageId="9" lastPageNumber="248" pageId="8" pageNumber="247">
|
||
<emphasis id="B977D7691809AA6FFF0BF94AFE9464AB" bold="true" box="[199,345,1711,1736]" pageId="8" pageNumber="247">Description.</emphasis>
|
||
A moderately built, small-bodied gecko (to
|
||
<quantity id="4CFBA69E1809AA6FFCABF94AFC0264A4" box="[871,975,1711,1736]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="8" pageNumber="247" unit="mm" value="35.5">35.5 mm</quantity>
|
||
SVL); body elongate, oblong in crosssection; head relatively wide (HeadW/HeadL 0.52–0.71), moderately long (HeadL/SVL 0.27–0.33), moderately depressed (HeadD/HeadL 0.27–0.35); in dorsal view, widest at ear openings; head not distinct from neck with neck only weakly constricted; loreal region convex; snout moderately short (SnL/HeadL 0.32–0.40), tapering to rounded tip; head depth ~1.5–2 times eye diameter; eye moderately large; rostral ~2.5 times wider than high, in broad contact with nostril, dorsal edge concave to accommodate supranasals, rostral groove extends from dorsal edge of rostral to ~50% of rostral height; 0 or 1 small internasals; supranasals horizontally elongate, 2–3 times wider than long; nostrils small, similar in size to postnasals, directed laterally, in contact with rostral, supranasal, 2–4 postnasals and first supralabial; supralabials 7–9, antero-dorso edge of first supralabial in contact with nostril; second supralabial rectangular, wider than high, length equal to first supralabial; supralabials 3–5 slightly smaller than first and second, gradually decreasing in size, all wider than high; scales on crown small and slightly rounded, ~one-third the size of dorsal scales, increasing in size anteriorly onto snout and supraocular regions; scales on snout flat and slightly enlarged, 1.5–2 times larger than scales on crown; supraocular scales enlarged; 1 or 2 soft spines on posterior eyelid; mental triangular to trapezoidal in shape, widest anteriorly, not extending past ventral edge of first infralabial; postmental not enlarged relative to other scales adjacent to infralabials; gulars small near infralabials, decreasing to granular, ~0.1–0.2 the size of scales on ventrum, juxtaposed and slightly rounded; gulars flattened from mental to eye, becoming more rounded from eye to neck; infralabials 7–9, first largest and squareshaped, gradually decreasing in size posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BBC0B7B1808AA6EFF0BFE39FD8060AF" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="248">Limbs short, distance between limbs close to half of SVL (ILL/SVL 0.37–0.50); scales on anterior surfaces of forelimbs elongate, broad and rounded, slightly imbricate, decreasing in size distally, ~2 times the size of scales elsewhere on forelimb; scales on dorsal surface of hind limbs rounded and slightly raised, decreasing in size and becoming more rounded distally; scales on anterior surface of thigh flat and circular, slightly larger in area than scales on hindlimbs elsewhere; fingers 5; toes 5; claws absent from all digits, digits short; fourth finger lamellae in 6 rows; fourth toe lamellae in 8 rows; apical plates much wider than digit; two pairs of dorsal apical plates; one pair of ventral apical plates, round to ovate.</paragraph>
|
||
<paragraph id="8BBC0B7B1808AA6EFF0BFD32FD81613E" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="248">Scales on dorsum and lateral surfaces small and homogeneous, smooth to weakly keeled, ~2.5 times wider than deep, juxtaposed; scales slightly raised, highest point at posterior edge of scale, widest at midpoint of scale; scales on ventrum homogeneous, flat, imbricate, ~equal to 1.5 times the size of dorsal scales in area, widest at midpoint, with rounded posterior edge.</paragraph>
|
||
<paragraph id="8BBC0B7B1808AA6EFF0BFC82FCE86673" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="248">
|
||
Tail moderately long, maximum width one-quarter to one-third along tail, gradually tapering to a fine point, round in cross-section; scales on dorsal surface of tail ~2–3 times size compared to dorsum, homogeneous, flattened, imbricate, square to oblong in shape, arranged in transverse rows; ventral tail scales flattened, ~2–3 times the size of dorsal tail scales; single enlarged cloacal spur per side, ~3–4 times the size of surrounding scales; no visible pre-cloacal pores (except on
|
||
<specimenCount id="9D05C0F21808AA6EFDE0FC1DFD4F6673" box="[556,642,1015,1040]" pageId="9" pageNumber="248" type="male">2 males</specimenCount>
|
||
with 2 pores).
|
||
</paragraph>
|
||
<paragraph id="8BBC0B7B1808AA6EFF0BFBF9FDEB6462" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="248">
|
||
<emphasis id="B977D7691808AA6EFF0BFBF9FE286657" box="[199,485,1052,1077]" italics="true" pageId="9" pageNumber="248">Colouration and pattern.</emphasis>
|
||
Dorsal background colouration tan and dark brown; light brown dorsolateral lines edged with dark brown emanate from back of eye to tail, continues forwards past eye as pale canthal stripe, terminating above nostril; poorly-defined silver-grey vertebral stripe discernible in some specimens, 2–3 scales wide, bordered by poorly defined dark brown paravertebral lines or dark edges of dorsolateral lines; dorsolateral zone with 7–12 widely spaced ocelli between eye and tail, variably defined ocelli cluster comprised of 3–6 pale scales, often lacking or loosely bordered by darker scales; lateral zone with poorly-defined lateral line that extends from postero-ventral edge of eye above limbs to tail, continuing forward as dark loreal stripes to nostril; below lateral stripes on flanks a mixture of darkly pigmented scales with grey and pale scales; crown and snout a mixture of dark brown, brown and pale grey scales; supralabials pale with dark stippling, infralabials moderately pigmented; ventral edge of orbit dark brown; limbs have same intermixture of dark and pale scales with numerous scattered small pale ocelli, weak on front limbs, more prominent on rear; top of digits have a network of dark lines; ventrum pale with fine to heavy dark brown flecking, not forming conspicuous lines; heavier flecking on rear limbs and cloacal region and tail; tail continues dorsal pattern including ocelli on dorso-lateral stripes, dark vertebral zone and light brown paravertebral zone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31958F01808AA6EFF0BF9E9FB2C6591" pageId="9" pageNumber="248" type="distribution">
|
||
<paragraph id="8BBC0B7B1808AA6EFF0BF9E9FB2C6591" blockId="9.[151,1437,151,2034]" pageId="9" pageNumber="248">
|
||
<emphasis id="B977D7691808AA6EFF0BF9E9FEAD6446" bold="true" box="[199,352,1548,1573]" pageId="9" pageNumber="248">Distribution.</emphasis>
|
||
This species occurs in the south-west of
|
||
<collectingCountry id="F3144BEB1808AA6EFC8CF9E9FC6A6446" box="[832,935,1548,1573]" name="Australia" pageId="9" pageNumber="248">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="133817FE1808AA6EFC75F9E9FBCC6446" box="[953,1025,1548,1573]" captionStart="FIGURE 2" captionStartId="2.[151,250,1920,1942]" captionTargetBox="[154,1425,895,1886]" captionTargetId="figure@2.[150,1436,895,1898]" captionTargetPageId="2" captionText="FIGURE 2. Distribution of the Crenadactylus ocellatus species-group including new species described in this paper. Darker shades denote areas of high topographic complexity, lighter shades lower complexity. Colours for taxa match those in Fig. 1. Questionable type location of Ebenavia horni holotype indicated by ‘? ’." httpUri="https://zenodo.org/record/262142/files/figure.png" pageId="9" pageNumber="248">Fig. 2</figureCitation>
|
||
). It is absent from the mesic southwestern corner, extending only as far south-west as Waroona (WAM R88483) then extending eastwards including records from Kojanup, Tambelup and Fitzgerald River National Park. The western-most extent of the distribution is from an old record in Israelite Bay at Cape Arid National Park (WAM R14176) and a more recent record from Balladonia at the edge of the Nullarbor Plain (WAM R157901). It extends into the wheatbelt and to the arid interzone, with records from near Mount Manning and Bowgada Nature Reserves, and an isolated record from Walganna Rock in the north-east (WAM R135103, genotyped as
|
||
<taxonomicName id="4C0370F81808AA6EFCBCF907FC3B6499" box="[880,1014,1761,1786]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="248" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B977D7691808AA6EFCBCF907FC3B6499" box="[880,1014,1761,1786]" italics="true" pageId="9" pageNumber="248">C. ocellatus</emphasis>
|
||
</taxonomicName>
|
||
and shown in
|
||
<figureCitation id="133817FE1808AA6EFB56F904FB146499" box="[1178,1241,1761,1786]" captionStart="FIGURE 3" captionStartId="6.[151,250,1768,1790]" captionTargetBox="[309,1276,193,1747]" captionTargetId="figure@6.[309,1276,193,1748]" captionTargetPageId="6" captionText="FIGURE 3. Morphological variation within the Crenadactylus ocellatus species group (left to right, top to bottom): A) typical homogeneous dorsal scalation of Crenadactylus species (except C. tuberculatus), B) heterogeneous dorsal scalation of C. tuberculatus sp. nov. with enlarged tubercles, C) typical internasal configuration of Crenadactylus species (except C. horni) showing small internasal when present (upper left) or supranasals in contact (lower right), D) internasal configuration of C. horni showing enlarged internasal extending beyond posterior edge of supranasals, E) typical postmental configuration of Crenadactylus species (except C. pilbarensis) showing granular gular scales, F) postmental configuration of C. pilbarensis sp. nov. showing enlarged postmental, G) typical snout scalation in lateral view (exception C. rostralis, C. naso [part]) showing rostral-nostril contact, H) snout scalation in lateral view of C. rostralis (and C. naso [part]) showing excluded rostral contact with nostril. (Drawings: A, B – K. Aplin; C – F, R. J. Ellis; G, H – R. E. Johnstone.)" httpUri="https://zenodo.org/record/262143/files/figure.png" pageId="9" pageNumber="248">Fig. 3</figureCitation>
|
||
A of
|
||
<bibRefCitation id="EF92768A1808AA6EFAD9F904FF19657E" author="Oliver" pageId="9" pageNumber="248" refString="Oliver, P., Adams, M. & Doughty, P. (2010) Molecular evidence for ten species and Oligo-Miocene vicariance within a nominal Australian gecko species (Crenadactylus ocellatus, Diplodactylidae). BMC Evolutionary Biology, 10, 386. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 10 - 386" type="journal article" year="2010">
|
||
Oliver
|
||
<emphasis id="B977D7691808AA6EFAA8F907FA516499" box="[1380,1436,1761,1786]" italics="true" pageId="9" pageNumber="248">et al.</emphasis>
|
||
2010
|
||
</bibRefCitation>
|
||
). Occurs on most of the Houtman Abrolhos Islands (off Geraldton) on which surveys have been conducted.
|
||
<emphasis id="B977D7691808AA6EFEDDF8CDFEBE6522" bold="true" box="[273,371,1832,1857]" pageId="9" pageNumber="248">Habitat.</emphasis>
|
||
Found in open woodland habitats throughout its range including areas dominated by
|
||
<emphasis id="B977D7691808AA6EFF5BF8AEFEEB6507" box="[151,294,1867,1892]" italics="true" pageId="9" pageNumber="248">Xanthorrhea</emphasis>
|
||
grass trees,
|
||
<taxonomicName id="4C0370F81808AA6EFE78F8AEFDFD6507" box="[436,560,1867,1892]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B977D7691808AA6EFE78F8AEFDFD6507" box="[436,560,1867,1892]" italics="true" pageId="9" pageNumber="248">Eucalyptus</emphasis>
|
||
</taxonomicName>
|
||
(jarrah, mallee, marri, tuart, wandoo) and spinifex grass (
|
||
<taxonomicName id="4C0370F81808AA6EFB0BF8AEFAD76507" box="[1223,1306,1867,1892]" class="Liliopsida" family="Poaceae" genus="Triodia" kingdom="Animalia" order="Poales" pageId="9" pageNumber="248" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B977D7691808AA6EFB0BF8AEFAD76507" box="[1223,1306,1867,1892]" italics="true" pageId="9" pageNumber="248">Triodia</emphasis>
|
||
</taxonomicName>
|
||
) on a wide variety of soft (sandy and loamy) and hard stony substrates (e.g. laterite). Collectors’ notes record specimens have been found under ground cover such as fallen and rotten logs, woodpiles, leaf litter, granite boulders, limestone slabs, sheets of tin, railway sleepers and rubbish. One specimen (WAM R72276) was found one meter high under the bark of a gimlet tree, and other records mention ‘under bark’ indicating some climbing ability.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7C5BF3180BAA6DFF5BFBCDFB89665D" box="[151,1092,1064,1086]" httpUri="https://zenodo.org/record/262145/files/figure.png" pageId="10" pageNumber="249" targetBox="[178,1401,193,1055]" targetPageId="10">
|
||
<paragraph id="8BBC0B7B180BAA6DFF5BFBCDFB89665D" blockId="10.[151,1092,1064,1086]" box="[151,1092,1064,1086]" pageId="10" pageNumber="249">
|
||
<emphasis id="B977D769180BAA6DFF5BFBCDFEDF665E" bold="true" box="[151,274,1064,1086]" pageId="10" pageNumber="249">FIGURE 5.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C0370F8180BAA6DFE55FBCDFD44665D" box="[409,649,1064,1086]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="249" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B977D769180BAA6DFE55FBCDFD44665D" box="[409,649,1064,1086]" italics="true" pageId="10" pageNumber="249">Diplodactylus ocellatus</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1947.3.6.68) (scale bar = 10 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7C5BF3180BAA6DFF5BF83CFB8C658C" box="[151,1089,2009,2031]" httpUri="https://zenodo.org/record/262146/files/figure.png" pageId="10" pageNumber="249" targetBox="[168,1412,1136,1974]" targetPageId="10">
|
||
<paragraph id="8BBC0B7B180BAA6DFF5BF83CFB8C658C" blockId="10.[151,1089,2009,2031]" box="[151,1089,2009,2031]" pageId="10" pageNumber="249">
|
||
<emphasis id="B977D769180BAA6DFF5BF83CFEDF658D" bold="true" box="[151,274,2009,2031]" pageId="10" pageNumber="249">FIGURE 6.</emphasis>
|
||
Syntype of
|
||
<taxonomicName id="4C0370F8180BAA6DFE41F83CFD4B658C" box="[397,646,2009,2031]" class="Reptilia" family="Diplodactylidae" genus="Diplodactylus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="249" phylum="Chordata" rank="species" species="bilineatus">
|
||
<emphasis id="B977D769180BAA6DFE41F83CFD4B658C" box="[397,646,2009,2031]" italics="true" pageId="10" pageNumber="249">Diplodactylus bilineatus</emphasis>
|
||
</taxonomicName>
|
||
(BMNH 1947.3.6.69) (scale bar = 10 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF7C5BF3180AAA6CFF5BFC75FC8561A7" httpUri="https://zenodo.org/record/262147/files/figure.png" pageId="11" pageNumber="250" targetBox="[163,1420,196,887]" targetPageId="11">
|
||
<paragraph id="8BBC0B7B180AAA6CFF5BFC75FC8561A7" blockId="11.[151,1436,912,964]" pageId="11" pageNumber="250">
|
||
<emphasis id="B977D769180AAA6CFF5BFC75FED961C6" bold="true" box="[151,276,912,934]" pageId="11" pageNumber="250">FIGURE 7.</emphasis>
|
||
Variation in
|
||
<taxonomicName id="4C0370F8180AAA6CFE51FC75FD5E61C5" box="[413,659,912,934]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B977D769180AAA6CFE51FC75FD5E61C5" box="[413,659,912,934]" italics="true" pageId="11" pageNumber="250">Crenadactylus ocellatus</emphasis>
|
||
</taxonomicName>
|
||
(scale bar = 10 mm). From left to right: WAM R132810, WAM R152975, WAM R135182, WAM R127614, WAM R152974, WAM R151338.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31958F0180AAA6CFF0BFC09FA896666" box="[199,1348,1004,1029]" pageId="11" pageNumber="250" type="etymology">
|
||
<paragraph id="8BBC0B7B180AAA6CFF0BFC09FA896666" blockId="11.[151,1437,1004,1317]" box="[199,1348,1004,1029]" pageId="11" pageNumber="250">
|
||
<emphasis id="B977D769180AAA6CFF0BFC09FE836666" bold="true" box="[199,334,1004,1029]" pageId="11" pageNumber="250">Etymology.</emphasis>
|
||
The specific name
|
||
<taxonomicName id="4C0370F8180AAA6CFDE5FC09FD436666" box="[553,654,1004,1029]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B977D769180AAA6CFDE5FC09FD436666" box="[553,654,1004,1029]" italics="true" pageId="11" pageNumber="250">ocellatus</emphasis>
|
||
</taxonomicName>
|
||
refers to the pale scattered blotches or ‘ocelli’ on the dorsum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31958F0180AAA6CFF0BFBF5FE966746" pageId="11" pageNumber="250" type="discussion">
|
||
<paragraph id="8BBC0B7B180AAA6CFF0BFBF5FE966746" blockId="11.[151,1437,1004,1317]" pageId="11" pageNumber="250">
|
||
<emphasis id="B977D769180AAA6CFF0BFBF5FEF6664A" bold="true" box="[199,315,1040,1065]" pageId="11" pageNumber="250">Remarks.</emphasis>
|
||
Our experience with this species indicates they are reasonably common compared to other
|
||
<taxonomicName id="4C0370F8180AAA6CFF5BFBD1FEF4662E" box="[151,313,1076,1101]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="genus">
|
||
<emphasis id="B977D769180AAA6CFF5BFBD1FEF4662E" box="[151,313,1076,1101]" italics="true" pageId="11" pageNumber="250">Crenadactylus</emphasis>
|
||
</taxonomicName>
|
||
species. Within the genus,
|
||
<taxonomicName id="4C0370F8180AAA6CFDA2FBD0FD39662E" box="[622,756,1076,1101]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="species" species="ocellatus">
|
||
<emphasis id="B977D769180AAA6CFDA2FBD0FD39662E" box="[622,756,1076,1101]" italics="true" pageId="11" pageNumber="250">C. ocellatus</emphasis>
|
||
</taxonomicName>
|
||
is exceptional for its dorsal patterning, with its intermixture of dark and pale scales and less well-defined longitudinal stripes (
|
||
<figureCitation id="133817FE180AAA6CFC54FBBDFC2D6613" box="[920,992,1112,1137]" captionStart="FIGURE 7" captionStartId="11.[151,250,912,934]" captionTargetBox="[163,1420,196,887]" captionTargetId="figure@11.[161,1426,193,891]" captionTargetPageId="11" captionText="FIGURE 7. Variation in Crenadactylus ocellatus (scale bar = 10 mm). From left to right: WAM R 132810, WAM R 152975, WAM R 135182, WAM R 127614, WAM R 152974, WAM R 151338." httpUri="https://zenodo.org/record/262147/files/figure.png" pageId="11" pageNumber="250">Fig. 7</figureCitation>
|
||
). In contrast, most other
|
||
<emphasis id="B977D769180AAA6CFAC4FBBDFA516612" box="[1288,1436,1112,1137]" italics="true" pageId="11" pageNumber="250">Crendactylus</emphasis>
|
||
species have relatively ‘clean’ longitudinal lines with possibly the exception of
|
||
<taxonomicName id="4C0370F8180AAA6CFBDBFB98FB0966F6" box="[1047,1220,1148,1173]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="species" species="tuberculatus" status="sp. nov.">
|
||
<emphasis id="B977D769180AAA6CFBDBFB98FB0966F6" box="[1047,1220,1148,1173]" italics="true" pageId="11" pageNumber="250">C. tuberculatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2446A12180AAA6CFB07FB99FAE866F6" box="[1227,1317,1148,1173]" pageId="11" pageNumber="250" rank="species">
|
||
<emphasis id="B977D769180AAA6CFB07FB99FAE866F6" bold="true" box="[1227,1317,1148,1173]" pageId="11" pageNumber="250">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="133817FE180AAA6CFAF9FB99FAB466F6" box="[1333,1401,1148,1173]" captionStart="FIGURE 4" captionStartId="7.[151,250,1853,1875]" captionTargetBox="[166,1420,210,1826]" captionTargetId="figure@7.[150,1436,193,1843]" captionTargetPageId="7" captionText="FIGURE 4. Photographs of Crenadactylus ocellatus species-group members in life (left to right, top to bottom): A) C. ocellatus, Koolyanobbing, WA (photo—R. J. Ellis), B) C. horni, Trephina Gorge, NT (photo—G. M. Shea), C) occidentalis sp. nov., 70 km S Exmouth, WA (photo—G. Gaikhorst), D) C. tuberculatus sp. nov., Cape Range National Park, WA (photo—R. J. Ellis), E) C. pilbarensis sp. nov., Burrup Peninsula, WA (photo—B. Maryan), F) C. rostralis, Geikie Gorge National Park, WA (photo—P. M. Oliver), G) C. naso, Koolan Island, WA (photo—G. Gaikhorst), H) C. naso, Mitchell Plateau, WA (Photo— G. M. Shea)." httpUri="https://zenodo.org/record/262144/files/figure.png" pageId="11" pageNumber="250">Fig. 4</figureCitation>
|
||
D). It is likely this broken colour pattern may be beneficial for camouflage owing to this species’ ground-dwelling habits. In contrast, the striped patterns in other
|
||
<taxonomicName id="4C0370F8180AAA6CFD70FB21FC9266BE" box="[700,863,1220,1245]" class="Reptilia" family="Diplodactylidae" genus="Crenadactylus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="genus">
|
||
<emphasis id="B977D769180AAA6CFD70FB21FC9266BE" box="[700,863,1220,1245]" italics="true" pageId="11" pageNumber="250">Crenadactylus</emphasis>
|
||
</taxonomicName>
|
||
taxa are likely beneficial to species that climb in spinifex clumps and are likely convergent with the phasmid geckos in the genus
|
||
<taxonomicName id="4C0370F8180AAA6CFBFDFB0DFAD46762" authority="Fitzinger" authorityName="Fitzinger" box="[1073,1305,1256,1281]" class="Reptilia" family="Diplodactylidae" genus="Strophurus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="250" phylum="Chordata" rank="genus">
|
||
<emphasis id="B977D769180AAA6CFBFDFB0DFB666762" box="[1073,1195,1256,1281]" italics="true" pageId="11" pageNumber="250">Strophurus</emphasis>
|
||
Fitzinger
|
||
</taxonomicName>
|
||
which also occur in spinifex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |