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<document id="052E568B0262C7BD2197EFEB205685A5" ID-DOI="10.5281/zenodo.275889" ID-GBIF-Dataset="8fc1956b-d634-4644-9399-56b167f3d5b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="275889" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460188038450" checkinUser="plazi" docAuthor="Csősz, Sándor &amp; Schulz, Andreas" docDate="2010" docId="03A9DF047936FFE7FF70FE0AFE77D00A" docLanguage="en" docName="zt02401p029.pdf" docOrigin="Zootaxa 2401" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tetramorium densopilosum Radchenko &amp; Arakelian 1990" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FF90A77C7938FFF7FFE7FFB0FFCCD358" masterDocTitle="A taxonomic review of the Palaearctic Tetramorium ferox species-complex (Hymenoptera, Formicidae)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="15" updateTime="1698236371737" updateUser="plazi">
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<mods:title id="D768B9066DED5C04AC137152575641D0">A taxonomic review of the Palaearctic Tetramorium ferox species-complex (Hymenoptera, Formicidae)</mods:title>
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<mods:namePart id="3B88503508D44DE8FD2D57811A818C66">Csősz, Sándor</mods:namePart>
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<mods:namePart id="B584F58F1FB17B2082A80056B6A9870E">Schulz, Andreas</mods:namePart>
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<mods:date id="ED0D2781C421203A851293B6A35D5CB3">2010</mods:date>
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<treatment id="03A9DF047936FFE7FF70FE0AFE77D00A" ID-DOI="http://doi.org/10.5281/zenodo.6210298" ID-GBIF-Taxon="127655293" ID-Zenodo-Dep="6210298" LSID="urn:lsid:plazi:treatment:03A9DF047936FFE7FF70FE0AFE77D00A" httpUri="http://treatment.plazi.org/id/03A9DF047936FFE7FF70FE0AFE77D00A" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="C31A3D997936FFF9FF70FE0AFCA9D28C" box="[151,869,442,468]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BBF6E127936FFF9FF70FE0AFCA9D28C" blockId="14.[151,869,442,468]" box="[151,869,442,468]" pageId="14" pageNumber="15">
<heading id="D0F7D97E7936FFF9FF70FE0AFCA9D28C" bold="true" box="[151,869,442,468]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
<taxonomicName id="4C0015917936FFF9FF70FE0AFCA9D28C" ID-CoL="55PL5" authority="Radchenko &amp; Arakelian, 1990" authorityName="Radchenko &amp; Arakelian" authorityYear="1990" box="[151,869,442,468]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007936FFF9FF70FE0AFCA9D28C" bold="true" box="[151,869,442,468]" pageId="14" pageNumber="15">
<emphasis id="B974B2007936FFF9FF70FE0AFE2ED28C" bold="true" box="[151,482,442,468]" italics="true" pageId="14" pageNumber="15">Tetramorium densopilosum</emphasis>
<bibRefCitation id="EF9113E37936FFF9FE0DFE0AFCA9D28C" author="Radchenko" box="[490,869,442,468]" pageId="14" pageNumber="29" refString="Radchenko, A. G. &amp; Arakelyan, G. R. (1990) Ants of the Tetramorium ferox species-group (Hymenoptera, Formicidae) from Crimea and the Caucasus. [in Russian] Biologuichesky Zhurnal Armenii, 5, 371 - 378." type="journal article" year="1990">Radchenko &amp; Arakelian, 1990</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31A3D997936FFF9FF70FDB2FE36D101" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BBF6E127936FFF9FF70FDB2FE36D101" blockId="14.[151,1436,514,601]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB10493C7936FFF9FF70FDB2FE36D101" infered="true" pageId="14" pageNumber="15">
<taxonomicName id="4C0015917936FFF9FF70FDB2FC89D141" authority="Radchenko &amp; Arakelian, 1990: 372" authorityName="Radchenko &amp; Arakelian" authorityPageNumber="372" authorityYear="1990" box="[151,837,514,537]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="densopilosus">
<emphasis id="B974B2007936FFF9FF70FDB2FE78D141" box="[151,436,514,537]" italics="true" pageId="14" pageNumber="15">Tetramorium densopilosus</emphasis>
<treatmentCitation id="0AA148037936FFF9FE5AFDB2FC89D141" author="Radchenko" box="[445,837,514,537]" page="372" pageId="14" pageNumber="15" year="1990">
<bibRefCitation id="EF9113E37936FFF9FE5AFDB2FCC1D141" author="Radchenko" box="[445,781,514,537]" pageId="14" pageNumber="29" refString="Radchenko, A. G. &amp; Arakelyan, G. R. (1990) Ants of the Tetramorium ferox species-group (Hymenoptera, Formicidae) from Crimea and the Caucasus. [in Russian] Biologuichesky Zhurnal Armenii, 5, 371 - 378." type="journal article" year="1990">Radchenko &amp; Arakelian, 1990</bibRefCitation>
: 372
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="54BBD0B07936FFF9FCB5FDB2FC79D141" box="[850,949,514,537]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
queen and
<typeStatus id="54BBD0B07936FFF9FBD1FDB2FB5ED141" box="[1078,1170,514,537]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
worker, male (6w,
<quantity id="4CF8C3F77936FFF9FA8DFDB2FA5CD141" box="[1386,1424,514,537]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="14" pageNumber="15" unit="m" value="4.0">4m</quantity>
):
<collectingCountry id="F3172E827936FFF9FF20FD92FEEBD161" box="[199,295,546,569]" name="Armenia" pageId="14" pageNumber="15">Armenia</collectingCountry>
, Nature Reserve,
<date id="FFBE48D27936FFF9FE01FD92FD93D161" box="[486,607,546,569]" pageId="14" pageNumber="15" value="1986-06-14">14.VI.1986</date>
(A. Radchenko) [label: Arm SSR Khosrovskij, z-k No 237.86, A.Radchenko
<date id="FFBE48D27936FFF9FF20FDF2FED6D101" box="[199,282,578,601]" pageId="14" pageNumber="15" value="86-06-14">14.6.86</date>
] (examined) [SIZK].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31A3D997936FFF9FF70FD37FDF2D5BA" pageId="14" pageNumber="15" type="description">
<paragraph id="8BBF6E127936FFF9FF70FD37FB9ED0DA" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">
<emphasis id="B974B2007936FFF9FF70FD37FE79D1F9" bold="true" box="[151,437,647,673]" pageId="14" pageNumber="15">Description of workers</emphasis>
(figs 25–27). Medium to large size, CS 833 [763, 878]. Whole body and appendages light brown to brown. Head slightly longer than broad, CL/CW 1.04 [1.02, 1.05], with straight sides, straight posterior head margin and rounded corners. Eyes small, EYE 0.17 [0.17, 0.18]. Frons moderately wide, FR/ CS 0.40 [0.38, 0.41], frontal lobes usually as wide as the frons, rarely slightly broader, FL/FR 1.00 [1.00, 1.02]. Scape long, SL/CS 0.78 [0.76, 0.80], with a feeble dorsal carina basally. Promesonotal dorsum slightly convex, metanotal groove moderately deep. Propodeal denticles moderately long. Petiolar node very high, NOH/NOL 1.23 [1.06, 1.44] trapezoid seen in profile, steeply rounded backward.
</paragraph>
<paragraph id="8BBF6E127936FFF9FF23FC3DFBA4D764" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">Head longitudinally rugulose and microreticulate, posterior surface ruguloso-reticulate, ground surface microreticulate, appears dull. Mesosoma dorsum and mesopleura ruguloso-reticulate, ground surface coarsely microreticulate. Dorsum of petiolar node ruguloso-reticulate, ground surface microreticulate, median part smooth. Dorsum of postpetiole rugulose and microreticulate. First gastral tergite smooth, appears shiny. Ventral surface of head with a row of short setae only (without long C-shape hairs).</paragraph>
<paragraph id="8BBF6E127936FFF9FF23FBF7FBA2D644" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">
<emphasis id="B974B2007936FFF9FF23FBF7FE0BD739" bold="true" box="[196,455,1095,1121]" pageId="14" pageNumber="15">Description of gynes</emphasis>
(figs 22–24, 28). Medium to large size, CS 980 [970, 990]. Head, mesosoma and appendages light brown, gaster dark brown. Head broader than long, CL/CW 0.87 [0.83, 0.90] with sides and posterior margin straight. Head trapezoid in full face view, narrower at genae. Frons moderately wide, FR/CS 0.40 [0.38, 0.41]. Scape long, SL/CS 0.76 [0.74, 0.78], without a dorsal carina basally, its surface is smooth and shiny. Head slightly narrower than scutum, MW/CS 1.06 [1.05, 1.07]. Propodeal teeth long and acute. Petiole and postpetiole wide, CS/PEW 1.71 [1.68, 1.75], CS/PPW 1.28 [1.24, 1.33].
</paragraph>
<paragraph id="8BBF6E127936FFF9FF23FA98FBA4D6A7" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">Head dorsum, posterior surface and sides ruguloso-reticulate, ground surface microreticulate. Frons longitudinally rugulose and microreticulate. Scutum and scutellum longitudinally rugulose. Sides of mesosoma, rugoso-reticulate and microreticulate, ventral part of katepisternum always rugulose, or microreticulate. Dorsum of petiolar node and postpetiole coarsely reticulate and microreticulate, medially shiny. First gastral tergite smooth, appears shiny, 1st gastral tergite very densely covered by pubescent hairs. Ventral surface of head with a row of short setae only (without long C-shape hairs).</paragraph>
<paragraph id="8BBF6E127936FFF9FF23F9BAFDF2D5BA" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">
<emphasis id="B974B2007936FFF9FF23F9BAFE1ED57C" bold="true" box="[196,466,1546,1572]" pageId="14" pageNumber="15">Description of males.</emphasis>
Head sides feebly convex, posterior margin and corners rounded. Scutum much broader than head. Propodeal denticles short but acute, propodeum angulate in profile. Dorsum of petiolar with a transverse crest, dorsolaterally sharp and angulate. Whole body and appendages brownish black. Head, scutum, scutellum, anepi- and katepisternum as well as petiole and postpetiole finely rugulo-reticulate, between main sculpturation microreticulate. First gastral tergite smooth, appears shiny, 1st gastral tergite very densely covered by pubescent hairs.
</paragraph>
</subSubSection>
<subSubSection id="C31A3D997936FFF9FF23F95DFF39D49A" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BBF6E127936FFF9FF23F95DFDC5D475" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">
<emphasis id="B974B2007936FFF9FF23F95DFE8BD45F" bold="true" box="[196,327,1773,1799]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
The very dense pubescence on the first gastral tergite (Fig. 28) makes the
<taxonomicName id="4C0015917936FFF9FB3FF95EFA50D45F" box="[1240,1436,1774,1799]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007936FFF9FB3FF95EFA50D45F" box="[1240,1436,1774,1799]" italics="true" pageId="14" pageNumber="15">T. densopilosum</emphasis>
</taxonomicName>
gynes unique within this group.
</paragraph>
<paragraph id="8BBF6E127936FFF9FF23F889FF39D49A" blockId="14.[151,1437,647,1986]" pageId="14" pageNumber="15">
<taxonomicName id="4C0015917936FFF9FF23F889FDC7D40A" box="[196,523,1849,1874]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007936FFF9FF23F889FDC7D40A" box="[196,523,1849,1874]" italics="true" pageId="14" pageNumber="15">Tetramorium densopilosum</emphasis>
</taxonomicName>
workers cannot be confused with those of
<taxonomicName id="4C0015917936FFF9FBF5F889FB75D40A" box="[1042,1209,1849,1874]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="diomedeum">
<emphasis id="B974B2007936FFF9FBF5F889FB75D40A" box="[1042,1209,1849,1874]" italics="true" pageId="14" pageNumber="15">T. diomedeum</emphasis>
</taxonomicName>
or
<taxonomicName id="4C0015917936FFF9FB04F889FA92D40A" box="[1251,1374,1849,1874]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="aegeum">
<emphasis id="B974B2007936FFF9FB04F889FA92D40A" box="[1251,1374,1849,1874]" italics="true" pageId="14" pageNumber="15">T. aegeum</emphasis>
</taxonomicName>
. The general appearance of the head and mesosoma of
<taxonomicName id="4C0015917936FFF9FCC3F8EEFC34D42F" box="[804,1016,1886,1911]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007936FFF9FCC3F8EEFC34D42F" box="[804,1016,1886,1911]" italics="true" pageId="14" pageNumber="15">T. densopilosum</emphasis>
</taxonomicName>
workers is always rugulose and microreticulate, but head and mesosoma of the two latter species are always shiny and usually at least partly smooth.
</paragraph>
</subSubSection>
<subSubSection id="C31A3D997937FFE7FF70F8A7FA81D1CF" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BBF6E127937FFF8FF70F8A7FE63D4CA" blockId="15.[151,1436,1815,1938]" pageId="15" pageNumber="16">
<emphasis id="B974B2007937FFF8FF70F8A7FE92D477" bold="true" box="[151,350,1815,1839]" pageId="15" pageNumber="16">FIGURES 22–28.</emphasis>
<taxonomicName id="4C0015917937FFF8FE83F8A8FC16D477" authority="Radchenko &amp; Arakelian, 1990" authorityName="Radchenko &amp; Arakelian" authorityYear="1990" box="[356,986,1816,1839]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007937FFF8FE83F8A8FD44D477" box="[356,648,1816,1839]" italics="true" pageId="15" pageNumber="16">Tetramorium densopilosum</emphasis>
<bibRefCitation id="EF9113E37937FFF8FD68F8A8FC16D477" author="Radchenko" box="[655,986,1816,1839]" pageId="15" pageNumber="29" refString="Radchenko, A. G. &amp; Arakelyan, G. R. (1990) Ants of the Tetramorium ferox species-group (Hymenoptera, Formicidae) from Crimea and the Caucasus. [in Russian] Biologuichesky Zhurnal Armenii, 5, 371 - 378." type="journal article" year="1990">Radchenko &amp; Arakelian, 1990</bibRefCitation>
</taxonomicName>
. Gyne mesosoma, petiole and postpetiole, dorsal view (Fig. 22), lateral view (Fig. 23); gyne head, dorsal view (Fig. 24); worker head, dorsal view (Fig. 25); worker mesosoma, petiole and postpetiole, dorsal view (Fig. 26), lateral view (Fig. 27); dense pubescence pilosity on 1st gastral tergite of a gyne (Fig. 28).
</paragraph>
<paragraph id="8BBF6E127928FFE7FF23FF28FC83D234" blockId="16.[151,1437,152,850]" pageId="16" pageNumber="17">
<taxonomicName id="4C0015917928FFE7FF23FF28FE2FD3E9" box="[196,483,152,177]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="feroxoide">
<emphasis id="B974B2007928FFE7FF23FF28FE2FD3E9" box="[196,483,152,177]" italics="true" pageId="16" pageNumber="17">Tetramorium feroxoide</emphasis>
</taxonomicName>
workers usually bear feeble, often parallel rugulae on head and mesosoma. Discriminant function D(3a) = -0.1025 FL +0.0712 MW –0.0942 SPSP +8.1089 for
<taxonomicName id="4C0015917928FFE7FB6DFF0DFA87D38E" box="[1162,1355,189,214]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FB6DFF0DFA87D38E" box="[1162,1355,189,214]" italics="true" pageId="16" pageNumber="17">T. densopilosum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0015917928FFE7FA61FF0DFEC5D3A3" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="feroxoide">
<emphasis id="B974B2007928FFE7FA61FF0DFEC5D3A3" italics="true" pageId="16" pageNumber="17">T. feroxoide</emphasis>
</taxonomicName>
workers resulted in 100% success, and each case (56 individuals) was correctly identified. The mean D(3a) scores for
<taxonomicName id="4C0015917928FFE7FE46FEB8FD92D279" box="[417,606,264,289]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FE46FEB8FD92D279" box="[417,606,264,289]" italics="true" pageId="16" pageNumber="17">T. densopilosum</emphasis>
</taxonomicName>
workers (n=8) was -2.191 ±0.89 [-3.512, -0.918], for nest means (n=2) -2.186 ±0.01 [-2.193, -2.180]. The mean scores for
<taxonomicName id="4C0015917928FFE7FCEDFE9DFC54D21E" box="[778,920,301,326]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="feroxoide">
<emphasis id="B974B2007928FFE7FCEDFE9DFC54D21E" box="[778,920,301,326]" italics="true" pageId="16" pageNumber="17">T. feroxoide</emphasis>
</taxonomicName>
workers (n=48) was 2.191 ±0.99 [+0.012, +3.911], for nest mean (n=2) -2.186 ±0.01 [-2.193, -2.180].
</paragraph>
<paragraph id="8BBF6E127928FFE7FF23FEC8FBC4D114" blockId="16.[151,1437,152,850]" pageId="16" pageNumber="17">
Separation of
<taxonomicName id="4C0015917928FFE7FE8DFEC8FDE4D2C9" box="[362,552,376,401]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FE8DFEC8FDE4D2C9" box="[362,552,376,401]" italics="true" pageId="16" pageNumber="17">T. densopilosum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0015917928FFE7FD84FEC8FD75D2C9" box="[611,697,376,401]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B974B2007928FFE7FD84FEC8FD75D2C9" box="[611,697,376,401]" italics="true" pageId="16" pageNumber="17">T. ferox</emphasis>
</taxonomicName>
workers is very difficult. With 8 observed workers we are not able to show variation in
<taxonomicName id="4C0015917928FFE7FE2BFE2DFD5DD2EE" box="[460,657,413,438]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FE2BFE2DFD5DD2EE" box="[460,657,413,438]" italics="true" pageId="16" pageNumber="17">T. densopilosum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C0015917928FFE7FD46FE2DFC26D2EE" box="[673,1002,413,438]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FD46FE2DFC26D2EE" box="[673,1002,413,438]" italics="true" pageId="16" pageNumber="17">Tetramorium densopilosum</emphasis>
</taxonomicName>
workers have a slightly trapezoidal head and the petiolar node is slightly angulate anterordorsally (Fig. 26) in contrast to that of
<taxonomicName id="4C0015917928FFE7FB34FE72FAE5D283" box="[1235,1321,450,475]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B974B2007928FFE7FB34FE72FAE5D283" box="[1235,1321,450,475]" italics="true" pageId="16" pageNumber="17">T. ferox</emphasis>
</taxonomicName>
, which is rounded. The frontal carinae are more developed in
<taxonomicName id="4C0015917928FFE7FCEBFE58FC02D159" box="[780,974,488,513]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FCEBFE58FC02D159" box="[780,974,488,513]" italics="true" pageId="16" pageNumber="17">T. densopilosum</emphasis>
</taxonomicName>
, and also the erect hairs of mesosoma seem to be longer end denser than those of
<taxonomicName id="4C0015917928FFE7FD4AFDBDFCCBD17E" box="[685,775,525,550]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B974B2007928FFE7FD4AFDBDFCCBD17E" box="[685,775,525,550]" italics="true" pageId="16" pageNumber="17">T. ferox</emphasis>
</taxonomicName>
. A discriminant D(4b) function proves the separation between
<taxonomicName id="4C0015917928FFE7FEE7FD82FE99D113" box="[256,341,562,587]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B974B2007928FFE7FEE7FD82FE99D113" box="[256,341,562,587]" italics="true" pageId="16" pageNumber="17">T. ferox</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0015917928FFE7FE76FD82FD82D113" box="[401,590,562,587]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="densopilosum">
<emphasis id="B974B2007928FFE7FE76FD82FD82D113" box="[401,590,562,587]" italics="true" pageId="16" pageNumber="17">T. densopilosum</emphasis>
</taxonomicName>
: see differential diagnosis of
<taxonomicName id="4C0015917928FFE7FC4EFD82FC33D113" box="[937,1023,562,587]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ferox">
<emphasis id="B974B2007928FFE7FC4EFD82FC33D113" box="[937,1023,562,587]" italics="true" pageId="16" pageNumber="17">T. ferox</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBF6E127928FFE7FF23FDE8FA81D1CF" blockId="16.[151,1437,152,850]" pageId="16" pageNumber="17">
For the results of cumulative discriminant analysis for worker caste see
<tableCitation id="C6825BA97928FFE7FBC1FDE8FB47D12A" box="[1062,1163,600,626]" captionStart="TABLE 3" captionStartId="9.[151,239,153,177]" captionTargetBox="[155,1274,243,903]" captionTargetPageId="9" captionText="TABLE 3. Unstandardized row coefficients of Roots 1 – 4 achieved by discriminant D (11) analysis run on workers of five species of Tetramorium ferox - complex (T. aegeum, densopilosum, diomedum, ferox and feroxoide)." httpUri="http://table.plazi.org/id/DF7F3E9A7931FFFEFF70FF29FB7AD389" pageId="16" pageNumber="17" tableUuid="DF7F3E9A7931FFFEFF70FF29FB7AD389">Tables 3</tableCitation>
&amp;
<tableCitation id="C6825BA97928FFE7FB55FDE8FB0DD12A" box="[1202,1217,600,626]" captionStart="TABLE 4" captionStartId="9.[151,239,964,988]" captionTargetBox="[155,1374,1053,1445]" captionTargetPageId="9" captionText="TABLE 4. Discriminant D (11) scores for worker individuals of five species of Tetramorium ferox - complex (T. aegeum, densopilosum, diomedum, ferox and feroxoide) by Roots 1 - 4. Upper row: mean ± SD, lower row [Min, Max]." httpUri="http://table.plazi.org/id/DF7F3E9A7931FFFEFF70FC74FAE8D0A4" pageId="16" pageNumber="17" tableUuid="DF7F3E9A7931FFFEFF70FC74FAE8D0A4">4</tableCitation>
and for gynes see
<tableCitation id="C6825BA97928FFE7FF70FDCDFF35D1CF" box="[151,249,637,663]" captionStart="TABLE 5" captionStartId="10.[151,239,636,660]" captionTargetBox="[155,1288,726,1431]" captionTargetPageId="10" captionText="TABLE 5. Unstandardized row coefficients of Roots 1 – 4 achieved by discriminant D (12 b) analysis run on gynes of five species of Tetramorium ferox - complex (T. aegeum, densopilosum, diomedum, ferox and feroxoide)." httpUri="http://table.plazi.org/id/DF7F3E9A7932FFFDFF70FDCCFB7AD1EC" pageId="16" pageNumber="17" tableUuid="DF7F3E9A7932FFFDFF70FDCCFB7AD1EC">Tables 5</tableCitation>
&amp;
<tableCitation id="C6825BA97928FFE7FEF9FDCDFEE3D1CF" box="[286,303,637,663]" captionStart="TABLE 6" captionStartId="10.[151,240,1498,1522]" captionTargetBox="[155,1373,1587,1979]" captionTargetPageId="10" captionText="TABLE 6. Discriminant D (12 b) scores for gyne individuals of five species of Tetramorium ferox - complex (T. aegeum, densopilosum, diomedum, ferox and feroxoide) by Roots 1 - 4. Upper row: mean ± SD, lower row [Min, Max]." httpUri="http://table.plazi.org/id/DF7F3E9A7932FFFDFF70FA6AFAE8D549" pageId="16" pageNumber="17" tableUuid="DF7F3E9A7932FFFDFF70FA6AFAE8D549">6</tableCitation>
. Morphometric characters and indices for workers and gynes are provided in
<tableCitation id="C6825BA97928FFE7FB5CFDCDFADED1CF" box="[1211,1298,637,663]" captionStart="TABLE 7" captionStartId="24.[151,242,151,175]" captionTargetBox="[151,1438,270,1782]" captionTargetPageId="24" captionText="TABLE 7. Morphometric comparison (Μm) of Tetramorium workers. Abbreviations of morphometric characters in" httpUri="http://table.plazi.org/id/DF7F3E9A7920FFEFFF70FF27FA50D3F7" pageId="16" pageNumber="17" tableUuid="DF7F3E9A7920FFEFFF70FF27FA50D3F7">Table 7</tableCitation>
&amp;
<tableCitation id="C6825BA97928FFE7FAD0FDCDFA84D1CF" box="[1335,1352,637,663]" captionStart="TABLE 8" captionStartId="25.[151,243,407,431]" captionText="TABLE 8. Morphometric comparison (Μm) of Tetramorium gynes. Abbreviations of morphometric characters in" pageId="16" pageNumber="17">8</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A3D997928FFE7FF21FD12FBB0D1E4" box="[198,1148,674,700]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BBF6E127928FFE7FF21FD12FBB0D1E4" blockId="16.[151,1437,152,850]" box="[198,1148,674,700]" pageId="16" pageNumber="17">
<emphasis id="B974B2007928FFE7FF21FD12FEABD1E4" bold="true" box="[198,359,674,700]" pageId="16" pageNumber="17">Distribution.</emphasis>
This species is known from
<collectingCountry id="F3172E827928FFE7FD50FD12FCEDD1E4" box="[695,801,674,700]" name="Armenia" pageId="16" pageNumber="17">Armenia</collectingCountry>
,
<collectingCountry id="F3172E827928FFE7FCCAFD12FC47D1E4" box="[813,907,674,700]" name="Georgia" pageId="16" pageNumber="17">Georgia</collectingCountry>
and Eastern
<collectingCountry id="F3172E827928FFE7FBC5FD12FBB4D1E4" box="[1058,1144,674,700]" name="Turkey" pageId="16" pageNumber="17">Turkey</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C31A3D997928FFE7FF21FD77FE77D00A" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BBF6E127928FFE7FF21FD77FAD6D05F" blockId="16.[151,1437,152,850]" pageId="16" pageNumber="17">
<emphasis id="B974B2007928FFE7FF21FD77FE72D1B9" bold="true" box="[198,446,711,737]" pageId="16" pageNumber="17">Material examined.</emphasis>
GEORGIA—USSR,
<collectingCountry id="F3172E827928FFE7FD35FD78FCFFD1BA" box="[722,819,712,738]" name="Georgia" pageId="16" pageNumber="17">Georgia</collectingCountry>
, Dranda, Kodori stream,
<date id="FFBE48D27928FFE7FB8EFD78FB3CD1BA" box="[1129,1264,712,738]" pageId="16" pageNumber="17" value="1975-05-20">20.05.1975</date>
. leg. Zombori (1w /
<collectionCode id="ED11F6D77928FFE7FF3CFD5DFEFBD05F" LSID="urn:lsid:biocol.org:col:34991" box="[219,311,749,775]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="16" pageNumber="17">HNHM</collectionCode>
, 1w /
<collectionCode id="ED11F6D77928FFE7FE99FD5DFE00D05F" box="[382,460,749,775]" pageId="16" pageNumber="17">PCAS</collectionCode>
);
<collectingCountry id="F3172E827928FFE7FE07FD5DFD8DD05F" box="[480,577,749,775]" name="Georgia" pageId="16" pageNumber="17">Georgia</collectingCountry>
,
<collectionCode id="ED11F6D77928FFE7FDAAFD5DFD55D05F" box="[589,665,749,775]" pageId="16" pageNumber="17">USSR</collectionCode>
, Novy Afon,
<date id="FFBE48D27928FFE7FCDFFD5DFC73D05F" box="[824,959,749,775]" pageId="16" pageNumber="17" value="1975-05-24">24.05.1975</date>
, leg. Zombori (1w /
<collectionCode id="ED11F6D77928FFE7FB48FD5DFAC2D05F" LSID="urn:lsid:biocol.org:col:34991" box="[1199,1294,749,775]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="16" pageNumber="17">HNHM</collectionCode>
);
</paragraph>
<paragraph id="8BBF6E127928FFE7FF23FCA2FE77D00A" blockId="16.[151,1437,152,850]" pageId="16" pageNumber="17">
TURKEY—Agri, Cilli Gecidi
<quantity id="4CF8C3F77928FFE7FDB2FCA2FD52D074" box="[597,670,786,812]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="16" pageNumber="17" unit="km" value="30.0">30km</quantity>
N Dogubayazit 1600mH Strassenrand Steppe leg. Schulz,
<date id="FFBE48D27928FFE7FF70FC88FED7D00A" box="[151,283,824,850]" pageId="16" pageNumber="17" value="1989-06-30">30.06.1989</date>
(2q /
<collectionCode id="ED11F6D77928FFE7FE87FC88FE63D00A" box="[352,431,824,850]" pageId="16" pageNumber="17">PCAS</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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