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<document id="3F5FBAF66E37CDA4B791E0C401EA1EEB" ID-CLB-Dataset="2489" ID-DOI="10.21248/contrib.entomol.66.2.201-255" ID-GBIF-Dataset="991c89c8-c376-46a5-9aa8-6dfbbf623c23" ID-ISSN="0005-805X" ID-Zenodo-Dep="5212606" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629211447587" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="0391026EFFB2FFB7FF5570EFFD92FC47" docLanguage="en" docName="ContribEntomol.66.2.201-255.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Anaulacaspis sinuata Assing 2016, spec. nov." docType="treatment" docVersion="15" lastPageNumber="229" masterDocId="FFA87A16FFA9FFABFFE8713CFFB4FFA6" masterDocTitle="Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="255" masterPageNumber="201" pageNumber="228" updateTime="1699087475910" updateUser="ExternalLinkService" zenodo-license-document="Abstyles">
<mods:mods id="01FB0BEC2F9B848F0F521756A3462C82" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7419471DC764F9A66003738CC5D29A5E">
<mods:title id="64560FE133AFA584E5F28F03C73EB8EB">Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="6223B419FD612DA5D886CD264CED3E6C" type="personal">
<mods:role id="8F94D128399AA4520F932B76643BA97C">
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</mods:role>
<mods:namePart id="94E38B90813463F99960820AD3B8A4F0">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="A71E3E2567FE60954864BBA4DA50B9D6">text</mods:typeOfResource>
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<mods:title id="EF1E4FBF33B46E3E04F676F2984DA7C3">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="184FB8AF44D71C8AF59D9FDC69F08702">
<mods:date id="05CD43270936492EF159EE7DD2E4EFE5">2016</mods:date>
<mods:detail id="EDCF3C56C8C29CC0F4BC3714EFF3EA8A" type="pubDate">
<mods:number id="87B5A015D2C1E2B0562629DD54984D71">2016-12-20</mods:number>
</mods:detail>
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<mods:number id="F84C4DC90C7FB14A83EBF4C2003E4779">66</mods:number>
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<mods:number id="693EC936519BDB8012553A345AFC8CCA">2</mods:number>
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<mods:start id="9AF16A0FF2792C05A0CC50959D172AFF">201</mods:start>
<mods:end id="9D70A9DC2B0C3376862C4155302242AB">255</mods:end>
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<mods:location id="8B414C00C643C0BD34838806DF60E894">
<mods:url id="0A1992BB1CA8D77D0247CB650F04EE13">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
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<mods:classification id="676F2D00B4C3AD7678FC5C2FB22D8AFB">journal article</mods:classification>
<mods:identifier id="4DBC06E6F88DE99ECA00935FF5975FFB" type="CLB-Dataset">2489</mods:identifier>
<mods:identifier id="1642D1D2B59B0B6C67FBEA6CAAE711BB" type="DOI">10.21248/contrib.entomol.66.2.201-255</mods:identifier>
<mods:identifier id="0C7C7583AD12C27220AE90F1B14273E6" type="GBIF-Dataset">991c89c8-c376-46a5-9aa8-6dfbbf623c23</mods:identifier>
<mods:identifier id="CF5EC1A141C82E921DF78D61B5342F90" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="7C0C6571041EF962684BC8AA271F0CAC" type="Zenodo-Dep">5212606</mods:identifier>
</mods:mods>
<treatment id="0391026EFFB2FFB7FF5570EFFD92FC47" ID-DOI="http://doi.org/10.5281/zenodo.5884797" ID-GBIF-Taxon="192454157" ID-Zenodo-Dep="5884797" LSID="urn:lsid:plazi:treatment:0391026EFFB2FFB7FF5570EFFD92FC47" httpUri="http://treatment.plazi.org/id/0391026EFFB2FFB7FF5570EFFD92FC47" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228">
<subSubSection id="C322E0F3FFB2FFB0FF5570EFFDBCFE48" box="[189,520,467,494]" pageId="27" pageNumber="228" type="nomenclature">
<paragraph id="8B87B378FFB2FFB0FF5570EFFDBCFE48" blockId="27.[189,520,467,527]" box="[189,520,467,494]" pageId="27" pageNumber="228">
<heading id="D0CF0414FFB2FFB0FF5570EFFDBCFE48" box="[189,520,467,494]" centered="true" fontSize="11" level="2" pageId="27" pageNumber="228" reason="3">
<taxonomicName id="4C38C8FBFFB2FFB0FF5570EFFE2CFE48" ID-CoL="DJKW" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[189,408,467,494]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="sinuata" status="sp. nov.">
<emphasis id="B94C6F6AFFB2FFB0FF5570EFFE2CFE48" box="[189,408,467,494]" italics="true" pageId="27" pageNumber="228">Anaulacaspis sinuata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FD211FFB2FFB0FE7770EFFDBCFE48" box="[415,520,467,494]" pageId="27" pageNumber="228" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FFB2FFB0FF5570C5FEDCFDA9" box="[189,360,505,527]" pageId="27" pageNumber="228" type="description">
<paragraph id="8B87B378FFB2FFB0FF5570C5FEDCFDA9" blockId="27.[189,520,467,527]" box="[189,360,505,527]" pageId="27" pageNumber="228">
(
<figureCitation id="1303AFFDFFB2FFB0FF2970C5FE97FDA9" box="[193,291,505,527]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="27" pageNumber="228">Figs 52–56</figureCitation>
,
<figureCitation id="1303AFFDFFB2FFB0FEC270C5FED6FDA9" box="[298,354,505,527]" captionStart="Map 2" captionStartId="27.[151,196,2023,2044]" captionTargetBox="[151,1398,1241,2014]" captionTargetId="figure-455@27.[151,1398,1241,2014]" captionTargetPageId="27" captionText="Map 2: Distributions of Anaulacaspis nigra (black circles), A. pseudonigra (white circle), A. reticulata (white triangle), A. libanotica (black triangles), A. caucasica (white diamonds), A. truncata (black diamonds), and A. sinuata (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="27" pageNumber="228">Map 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FFB2FFB0FF7F7315FE9CFCC7" pageId="27" pageNumber="228" type="materials_examined">
<paragraph id="8B87B378FFB2FFB0FF7F7315FE9CFCC7" blockId="27.[151,756,553,865]" pageId="27" pageNumber="228">
<emphasis id="B94C6F6AFFB2FFB0FF7F7315FE9BFDE7" bold="true" box="[151,303,553,577]" pageId="27" pageNumber="228">Type material</emphasis>
:
<materialsCitation id="3B50B925FFB2FFB0FED67315FF6AFD07" ID-GBIF-Occurrence="3449483351" collectionCode="cAss" collectorName="A. Senglet" country="Iran" determinerName="V. Assing" latitude="35.533333" location="E de Marivan" longLatPrecision="1193" longitude="46.333332" pageId="27" pageNumber="228" specimenCount="1" stateProvince="KORDESTAN" typeStatus="holotype">
<typeStatus id="54830DDAFFB2FFB0FED67315FE15FDE7" box="[318,417,553,577]" pageId="27" pageNumber="228" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3E78F1FFB2FFB0FE407316FE09FDE7" box="[424,445,554,577]" pageId="27" pageNumber="228">♂</specimenCount>
: “
<collectingCountry id="F32FF3E8FFB2FFB0FE3C7315FDA1FDE7" box="[468,533,553,577]" name="Iran" pageId="27" pageNumber="228">IRAN</collectingCountry>
<collectingRegion id="49FC7D9AFFB2FFB0FDF57316FD0CFDE3" box="[541,696,554,581]" country="Iran" name="Kordestan" pageId="27" pageNumber="228">KORDESTÂN</collectingRegion>
,
<location id="8EE7E5A3FFB2FFB0FD2C7316FF47FDC7" LSID="urn:lsid:plazi:treatment:0391026EFFB2FFB7FF5570EFFD92FC47:8EE7E5A3FFB2FFB0FD2C7316FF47FDC7" country="Iran" latitude="35.533333" longLatPrecision="1193" longitude="46.333332" name="E de Marivan" pageId="27" pageNumber="228" stateProvince="KORDESTAN">E de Marivân</location>
,
<geoCoordinate id="EE0CD5BFFFB2FFB0FEEC7375FEEFFDC7" box="[260,347,585,609]" degrees="35" direction="north" minutes="32" orientation="latitude" pageId="27" pageNumber="228" precision="925" value="35.533333">35°32'N</geoCoordinate>
,
<geoCoordinate id="EE0CD5BFFFB2FFB0FE837375FE0AFDC7" box="[363,446,585,609]" degrees="46" direction="east" minutes="20" orientation="longitude" pageId="27" pageNumber="228" precision="925" value="46.333332">46°20'E</geoCoordinate>
,
<collectorName id="26CDD6AEFFB2FFB0FE267375FD8FFDC7" box="[462,571,585,609]" pageId="27" pageNumber="228">A. Senglet</collectorName>
<date id="FF8695B8FFB2FFB0FDAF7375FD27FDC7" box="[583,659,585,609]" pageId="27" pageNumber="228" value="75-09-16">16.9.75</date>
/ Holotypus ♂
<taxonomicName id="4C38C8FBFFB2FFB0FF1C7355FE66FD27" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[244,466,617,641]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="sinuata" status="sp. nov.">
<emphasis id="B94C6F6AFFB2FFB0FF1C7355FE66FD27" box="[244,466,617,641]" italics="true" pageId="27" pageNumber="228">Anaulacaspis sinuata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27FD211FFB2FFB0FE317356FDA5FD27" box="[473,529,618,641]" pageId="27" pageNumber="228" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E5B5FBCFFB2FFB0FDA57356FD1BFD27" box="[589,687,617,641]" pageId="27" pageNumber="228">V. Assing</determinerName>
2016” (
<collectionCode id="ED292BBDFFB2FFB0FF7773B5FF62FD07" box="[159,214,649,673]" pageId="27" pageNumber="228">cAss</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B50B925FFB2FFB0FF0F73B6FD77FD07" ID-GBIF-Occurrence="3449483588" box="[231,707,649,674]" collectingDate="2008-06-09" collectingDateMax="2011-10-15" collectingDateMin="2008-06-09" collectionCode="MNB" collectorName="Frisch &amp; Serri" country="Iran" determinerName="V. Assing" elevation="1890" latitude="35.533333" location="Sanandaj - Divandarreh" longLatPrecision="1193" longitude="35.761944" pageId="27" pageNumber="228" specimenCount="1" stateProvince="Kordestan Province" typeStatus="paratype">
<typeStatus id="54830DDAFFB2FFB0FF0F73B6FEE5FD04" box="[231,337,650,674]" pageId="27" pageNumber="228" type="paratype">Paratypes</typeStatus>
: 1 ♀: same data as holotype (cAss);
</materialsCitation>
<materialsCitation id="3B50B925FFB2FFB0FD2173B5FD44FCA7" ID-GBIF-Occurrence="3455129321" collectingDate="2008-06-09" collectionCode="MNB" collectorName="Frisch &amp; Serri" country="Iran" elevation="1890" location="Sanandaj - Divandarreh" longitude="35.761944" municipality="Sanandaj - Divandarreh" pageId="27" pageNumber="228" specimenCount="3" stateProvince="Kordestan Province">
<specimenCount id="9D3E78F1FFB2FFB0FD2173B5FD5AFD07" box="[713,750,649,673]" pageId="27" pageNumber="228">1 ♂</specimenCount>
,
<specimenCount id="9D3E78F1FFB2FFB0FF7F7395FF64FD67" box="[151,208,681,705]" pageId="27" pageNumber="228">2 ♀♀</specimenCount>
: “
<collectingCountry id="F32FF3E8FFB2FFB0FF0D7395FE9CFD67" box="[229,296,681,705]" name="Iran" pageId="27" pageNumber="228">IRAN</collectingCountry>
,
<collectingRegion id="49FC7D9AFFB2FFB0FED97395FDB0FD67" box="[305,516,681,705]" country="Iran" name="Kordestan" pageId="27" pageNumber="228">Kordestan Province</collectingRegion>
,
<collectingMunicipality id="6BE32902FFB2FFB0FDE67395FF0EFD47" country="Iran" latitude="35.533333" longLatPrecision="1193" longitude="35.761944" name="Sanandaj - Divandarreh" pageId="27" pageNumber="228" stateProvince="Kordestan Province">Sanandaj – Divandarreh</collectingMunicipality>
road,
<locationDeviation id="9A35AA5AFFB2FFB0FEEC73F5FE44FD47" box="[260,496,713,737]" location="Divandarreh" pageId="27" pageNumber="228">
<quantity id="4CC01E9DFFB2FFB0FEEC73F5FEF2FD47" box="[260,326,713,737]" metricMagnitude="4" metricUnit="m" metricValue="2.1" pageId="27" pageNumber="228" unit="km" value="21.0">21 km</quantity>
S Divandarreh
</locationDeviation>
,
<quantity id="4CC01E9DFFB2FFB0FE1573F5FDFAFD47" box="[509,590,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="27" pageNumber="228" unit="m" value="1890.0">
<elevation id="0015544BFFB2FFB0FE1573F5FDFAFD47" box="[509,590,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.89" pageId="27" pageNumber="228" unit="m" value="1890.0">1890 m</elevation>
</quantity>
, N
<geoCoordinate id="EE0CD5BFFFB2FFB0FD9173F5FD40FD47" box="[633,756,713,737]" degrees="35" direction="east" minutes="45" orientation="longitude" pageId="27" pageNumber="228" precision="15" seconds="43" value="35.761944">35°45'43&quot; E</geoCoordinate>
047°04'43&quot;,
<date id="FF8695B8FFB2FFB0FEFD73D5FE3EFCA7" box="[277,394,745,769]" pageId="27" pageNumber="228" value="2008-06-09">
<collectingDate id="EFC26C50FFB2FFB0FEFD73D5FE3EFCA7" box="[277,394,745,769]" pageId="27" pageNumber="228" value="2008-06-09">06.09.2008</collectingDate>
</date>
, lg.
<collectorName id="26CDD6AEFFB2FFB0FE5F73D5FE4DFCA7" box="[439,505,745,769]" pageId="27" pageNumber="228">Frisch</collectorName>
&amp;
<collectorName id="26CDD6AEFFB2FFB0FDF473D5FDE4FCA7" box="[540,592,745,769]" pageId="27" pageNumber="228">Serri</collectorName>
” (
<collectionCode id="ED292BBDFFB2FFB0FD8173D5FD1CFCA7" box="[617,680,745,769]" pageId="27" pageNumber="228">MNB</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="3B50B925FFB2FFB0FF7F7235FE90FCC7" ID-GBIF-Occurrence="3449483673" collectingDate="2011-10-15" collectingDateMax="2011-10-15" collectingDateMin="2008-06-09" collectionCode="MNB" collectorName="Frisch" country="Iran" determinerName="V. Assing" elevation="1740" latitude="35.533333" location="Lorestan Province" longLatPrecision="1193" longitude="35.761944" pageId="27" pageNumber="228" specimenCount="1" stateProvince="Lorestan Province" typeStatus="paratype">
2 ♂♂: “
<collectingCountry id="F32FF3E8FFB2FFB0FF027235FE99FC87" box="[234,301,777,801]" name="Iran" pageId="27" pageNumber="228">IRAN</collectingCountry>
,
<collectingRegion id="49FC7D9AFFB2FFB0FED47236FDB6FC87" box="[316,514,777,801]" country="Iran" name="Lorestan" pageId="27" pageNumber="228">Lorestan Province</collectingRegion>
,
<locationDeviation id="9A35AA5AFFB2FFB0FDF97235FD45FC87" box="[529,753,777,801]" location="Borujerd" pageId="27" pageNumber="228">
<quantity id="4CC01E9DFFB2FFB0FDF97235FDE7FC87" box="[529,595,777,801]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="27" pageNumber="228" unit="km" value="20.0">20 km</quantity>
SW Borujerd
</locationDeviation>
,
<quantity id="4CC01E9DFFB2FFB0FF7F7215FF5CFCE7" box="[151,232,809,833]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="27" pageNumber="228" unit="m" value="1740.0">
<elevation id="0015544BFFB2FFB0FF7F7215FF5CFCE7" box="[151,232,809,833]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="27" pageNumber="228" unit="m" value="1740.0">1740 m</elevation>
</quantity>
,
<geoCoordinate id="EE0CD5BFFFB2FFB0FF197216FED9FCE7" box="[241,365,809,833]" degrees="33" direction="north" minutes="46" orientation="latitude" pageId="27" pageNumber="228" precision="15" seconds="23" value="33.773056">N 33°46'23&quot;</geoCoordinate>
<geoCoordinate id="EE0CD5BFFFB2FFB0FE9A7216FE4CFCE7" box="[370,504,809,833]" degrees="048" direction="east" minutes="39" orientation="longitude" pageId="27" pageNumber="228" precision="15" seconds="06" value="48.65167">E 048°39'06&quot;</geoCoordinate>
,
<date id="FF8695B8FFB2FFB0FDEA7215FDC2FCE7" box="[514,630,809,833]" pageId="27" pageNumber="228" value="2011-10-15">
<collectingDate id="EFC26C50FFB2FFB0FDEA7215FDC2FCE7" box="[514,630,809,833]" pageId="27" pageNumber="228" value="2011-10-15">15.10.2011</collectingDate>
</date>
, leg.
<collectorName id="26CDD6AEFFB2FFB0FD417215FD5FFCE7" box="[681,747,809,833]" pageId="27" pageNumber="228">Frisch</collectorName>
” (
<collectionCode id="ED292BBDFFB2FFB0FF777275FF6AFCC7" box="[159,222,841,865]" collectionName="MNB" pageId="27" pageNumber="228">MNB</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FFB2FFB0FF7F72B5FF6DFC47" pageId="27" pageNumber="228" type="etymology">
<paragraph id="8B87B378FFB2FFB0FF7F72B5FF6DFC47" blockId="27.[151,756,905,993]" pageId="27" pageNumber="228">
<emphasis id="B94C6F6AFFB2FFB0FF7F72B5FEBAFC07" bold="true" box="[151,270,905,929]" pageId="27" pageNumber="228">Etymology</emphasis>
: The specific epithet (Latin, adjective) alludes to the sinuate ventral process of the aedeagus (lateral view).
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FFB2FFB7FF7F7535FDE2FCE7" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228" type="description">
<paragraph id="8B87B378FFB2FFB0FF7F7535FE27FB07" blockId="27.[151,757,1033,1186]" pageId="27" pageNumber="228">
<emphasis id="B94C6F6AFFB2FFB0FF7F7535FEADFB87" bold="true" box="[151,281,1033,1057]" pageId="27" pageNumber="228">Description</emphasis>
: Body length 2.0–
<quantity id="4CC01E9DFFB2FFB0FE057535FDF4FB87" box="[493,576,1033,1057]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="27" pageNumber="228" unit="mm" value="2.3">2.3 mm</quantity>
; length of forebody 1.0–
<quantity id="4CC01E9DFFB2FFB0FEE17515FEEFFBE7" box="[265,347,1065,1089]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="27" pageNumber="228" unit="mm" value="1.1">1.1 mm</quantity>
. Coloration: body black, sometimes with the postero-sutural portion of the elytra indistinctly and diffusely paler; legs blackish-brown; antennae black. Integument very glossy.
</paragraph>
<paragraph id="8B87B378FFB2FFB0FCF271D5FC1BFDA7" blockId="27.[794,1399,233,1185]" pageId="27" pageNumber="228">
Head weakly transverse, 1.05–1.10 times as broad as long, without appreciable sexual dimorphism; punctation extremely fine and moderately dense; interstices without microsculpture. Eyes slightly longer than postocular region in dorsal view. Antennae approximately
<quantity id="4CC01E9DFFB2FFB0FACD7055FAC3FE27" box="[1317,1399,361,385]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="27" pageNumber="228" unit="mm" value="0.8">0.8 mm</quantity>
long; antennomere IV approximately as long as broad; antennomeres V–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long.
</paragraph>
<paragraph id="8B87B378FFB2FFB0FCF27336FBDEFD27" blockId="27.[794,1399,233,1185]" pageId="27" pageNumber="228">Pronotum 1.10–1.15 times as broad as long and approximately as broad as head; sexual dimorphism indistinct; punctation moderately dense and fine, usually slightly more distinct than that of head.</paragraph>
<paragraph id="8B87B378FFB2FFB0FCF273B5FAE3FD67" blockId="27.[794,1399,233,1185]" pageId="27" pageNumber="228">Elytra slightly longer than pronotum; punctation moderately dense and very fine. Hind wings fully developed.</paragraph>
<paragraph id="8B87B378FFB2FFB0FCF273F5FC46FC27" blockId="27.[794,1399,233,1185]" pageId="27" pageNumber="228">
Abdomen slightly narrower than elytra; punctation fine and distinct, denser in anterior than in posterior portions of tergites; tergite VIII (
<figureCitation id="1303AFFDFFB2FFB0FB097235FA9AFC87" box="[1249,1326,777,801]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="27" pageNumber="228">Fig. 52</figureCitation>
) with postero-lateral cluster of long thin setae on either side, posterior margin truncate or indistinctly convex, with thin marginal setae.
</paragraph>
<paragraph id="8B87B378FFB2FFB0FCF272B6FC10FBE7" blockId="27.[794,1399,233,1185]" pageId="27" pageNumber="228">
♂: pronotum usually with very fine and shallow median sulcus; sternite VIII (
<figureCitation id="1303AFFDFFB2FFB0FBB07295FB11FC67" box="[1112,1189,937,961]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="27" pageNumber="228">Fig. 53</figureCitation>
) strongly, convexly produced posteriorly; median lobe of aedeagus (
<figureCitation id="1303AFFDFFB2FFB0FCC972D5FC2EFBA7" box="[801,922,1001,1025]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="27" pageNumber="228">Figs 54–55</figureCitation>
) small,
<quantity id="4CC01E9DFFB2FFB0FC1972D5FB39FBA7" box="[1009,1165,1001,1025]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="27" pageNumber="228" unit="mm" value="0.22" valueMax="0.23" valueMin="0.21">0.21–0.23 mm</quantity>
long; ventral process sinuate in lateral view, broad and apically not incised in ventral view.
</paragraph>
<paragraph id="8B87B378FFB2FFB0FCF27576FC61FB07" blockId="27.[794,1399,233,1185]" pageId="27" pageNumber="228">
♀: pronotum with or without very indistinct median sulcus; sternite VIII (
<figureCitation id="1303AFFDFFB2FFB0FBF97555FBEFFB27" box="[1041,1115,1129,1153]" captionStart="Figs 49–69" captionStartId="13.[151,191,1999,2021]" captionTargetBox="[158,1391,246,1961]" captionTargetId="figure-41@13.[151,1398,232,1971]" captionTargetPageId="13" captionText="Figs 49–69: Anaulacaspsis libanotica (49–51), A. sinuata (52–56), A. nigerrima (57–61), and A. persica (62–69): male tergite VIII (49, 52, 57, 62–63); male sternite VIII (50, 53, 58, 64); female sternite VIII (51, 56); median lobe of aedeagus in lateral and in ventral view (54–55, 59–61, 65–69). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5212616" httpUri="https://zenodo.org/record/5212616/files/figure.png" pageId="27" pageNumber="228">Fig. 56</figureCitation>
) with distinctly convex posterior margin.
</paragraph>
<caption id="DF47E3F0FFB2FFB0FF7F76DBFDBDF791" ID-DOI="http://doi.org/10.5281/zenodo.5212628" ID-Zenodo-Dep="5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="27" pageNumber="228" startId="27.[151,196,2023,2044]" targetBox="[151,1398,1241,2014]" targetPageId="27">
<paragraph id="8B87B378FFB2FFB0FF7F76DBFDBDF791" blockId="27.[151,1399,2022,2103]" pageId="27" pageNumber="228">
<emphasis id="B94C6F6AFFB2FFB0FF7F76DBFF61F85D" bold="true" box="[151,213,2022,2044]" pageId="27" pageNumber="228">Map 2</emphasis>
: Distributions of
<taxonomicName id="4C38C8FBFFB2FFB0FE9676DAFD85F85A" box="[382,561,2022,2044]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B94C6F6AFFB2FFB0FE9676DAFD85F85A" box="[382,561,2022,2044]" italics="true" pageId="27" pageNumber="228">Anaulacaspis nigra</emphasis>
</taxonomicName>
(black circles),
<taxonomicName id="4C38C8FBFFB2FFB0FD2F76DBFCE0F85D" authorityName="Assing" authorityYear="2016" box="[711,852,2022,2044]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="pseudonigra">
<emphasis id="B94C6F6AFFB2FFB0FD2F76DBFCE0F85D" box="[711,852,2022,2044]" italics="true" pageId="27" pageNumber="228">A. pseudonigra</emphasis>
</taxonomicName>
(white circle),
<taxonomicName id="4C38C8FBFFB2FFB0FC0C76DBFBECF85D" authorityName="Assing" authorityYear="2016" box="[996,1112,2022,2044]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="reticulata">
<emphasis id="B94C6F6AFFB2FFB0FC0C76DBFBECF85D" box="[996,1112,2022,2044]" italics="true" pageId="27" pageNumber="228">A. reticulata</emphasis>
</taxonomicName>
(white triangle),
<taxonomicName id="4C38C8FBFFB2FFB0FB1776DBFAC3F85D" authorityName="Assing" authorityYear="2016" box="[1279,1399,2022,2044]" class="Insecta" family="Staphylinidae" genus="Melagria" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="libanotica">
<emphasis id="B94C6F6AFFB2FFB0FB1776DBFAC3F85D" box="[1279,1399,2022,2044]" italics="true" pageId="27" pageNumber="228">A. libanotica</emphasis>
</taxonomicName>
(black triangles),
<taxonomicName id="4C38C8FBFFB2FFB0FEA87938FE00F7BF" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[320,436,2052,2073]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="caucasica">
<emphasis id="B94C6F6AFFB2FFB0FEA87938FE00F7BF" box="[320,436,2052,2073]" italics="true" pageId="27" pageNumber="228">A. caucasica</emphasis>
</taxonomicName>
(white diamonds),
<taxonomicName id="4C38C8FBFFB2FFB0FD9A7938FD69F7BF" box="[626,733,2052,2073]" class="Insecta" family="Staphylinidae" genus="Anaulacaspsis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="truncata">
<emphasis id="B94C6F6AFFB2FFB0FD9A7938FD69F7BF" box="[626,733,2052,2073]" italics="true" pageId="27" pageNumber="228">A. truncata</emphasis>
</taxonomicName>
(black diamonds), and
<taxonomicName id="4C38C8FBFFB2FFB0FC2C7938FB90F7BF" authorityName="Assing" authorityYear="2016" box="[964,1060,2052,2073]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="sinuata">
<emphasis id="B94C6F6AFFB2FFB0FC2C7938FB90F7BF" box="[964,1060,2052,2073]" italics="true" pageId="27" pageNumber="228">A. sinuata</emphasis>
</taxonomicName>
(stars) in the West Palaearctic and Middle Asia, based on revised records.
</paragraph>
</caption>
<paragraph id="8B87B378FFB5FFB7FF5571D5FDE2FCE7" blockId="28.[189,794,233,833]" pageId="28" pageNumber="229">
<emphasis id="B94C6F6AFFB5FFB7FF5571D5FE2CFEA7" bold="true" box="[189,408,233,257]" pageId="28" pageNumber="229">Comparative notes</emphasis>
:
<taxonomicName id="4C38C8FBFFB5FFB7FE4771D5FD23FEA7" authorityName="Assing" authorityYear="2016" box="[431,663,233,257]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="sinuata">
<emphasis id="B94C6F6AFFB5FFB7FE4771D5FD23FEA7" box="[431,663,233,257]" italics="true" pageId="28" pageNumber="229">Anaulacaspis sinuata</emphasis>
</taxonomicName>
is reliably distinguished from other dark-coloured species of the
<taxonomicName id="4C38C8FBFFB5FFB7FF557015FEA5FEE7" box="[189,273,297,321]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B94C6F6AFFB5FFB7FF557015FEA5FEE7" box="[189,273,297,321]" italics="true" pageId="28" pageNumber="229">A. nigra</emphasis>
</taxonomicName>
group only by the shape of the median lobe of the aedeagus. In external appearance (blackish-brown legs; glossy and finely punctured forebody, it is similar to
<taxonomicName id="4C38C8FBFFB5FFB7FF5570B5FE93FE07" baseAuthorityName="Fagel" baseAuthorityYear="1969" box="[189,295,393,417]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigrina">
<emphasis id="B94C6F6AFFB5FFB7FF5570B5FE93FE07" box="[189,295,393,417]" italics="true" pageId="28" pageNumber="229">A. nigrina</emphasis>
</taxonomicName>
, from which it additionally differs by less dense punctation of the head and pronotum, usually uniformly black elytra, a truncate posterior margin of the abdominal tergite VIII, and a posteriorly more strongly produced male sternite VIII. In coloration, glossy appearance, fine punctation, the absence of a distinct sexual dimorphism of the head and pronotum, and the lateral aspect of the ventral process of the aedeagus,
<taxonomicName id="4C38C8FBFFB5FFB7FDF87355FDCFFD27" authorityName="Assing" authorityYear="2016" box="[528,635,617,641]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="sinuata">
<emphasis id="B94C6F6AFFB5FFB7FDF87355FDCFFD27" box="[528,635,617,641]" italics="true" pageId="28" pageNumber="229">A. sinuata</emphasis>
</taxonomicName>
also resembles
<taxonomicName id="4C38C8FBFFB5FFB7FF5573B5FEF6FD07" baseAuthorityName="BERNHAUER" baseAuthorityYear="1908" box="[189,322,649,673]" class="Insecta" family="Staphylinidae" genus="Falagria" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="229" phylum="Arthropoda" rank="species" species="nigerrima">
<emphasis id="B94C6F6AFFB5FFB7FF5573B5FEF6FD07" box="[189,322,649,673]" italics="true" pageId="28" pageNumber="229">A. nigerrima</emphasis>
</taxonomicName>
, from which it is distinguished by the shape of tergite VIII, and the distinctly smaller median lobe of the aedeagus with a more strongly curved (lateral view), apically less acute (lateral view), smoothly converging lateral margins (ventral view), and apically not distinctly pointed ventral process (ventral view).
</paragraph>
</subSubSection>
<subSubSection id="C322E0F3FFB5FFB7FF557255FD92FC47" pageId="28" pageNumber="229" type="distribution">
<paragraph id="8B87B378FFB5FFB7FF557255FD92FC47" blockId="28.[189,794,873,994]" pageId="28" pageNumber="229">
<emphasis id="B94C6F6AFFB5FFB7FF557255FEF2FC27" bold="true" box="[189,326,873,897]" pageId="28" pageNumber="229">Distribution</emphasis>
: The currently known distribution is confined to few localities in
<collectingRegion id="49FC7D9AFFB5FFB7FDE572B5FDCFFC07" box="[525,635,905,929]" country="Iran" name="Kordestan" pageId="28" pageNumber="229">Kordestan</collectingRegion>
and
<collectingRegion id="49FC7D9AFFB5FFB7FD5572B6FCAEFC07" box="[701,794,906,929]" country="Iran" name="Lorestan" pageId="28" pageNumber="229">Lorestan</collectingRegion>
provinces,
<collectingCountry id="F32FF3E8FFB5FFB7FEDB7296FED4FC67" box="[307,352,938,961]" name="Iran" pageId="28" pageNumber="229">Iran</collectingCountry>
(
<figureCitation id="1303AFFDFFB5FFB7FE997295FE0CFC67" box="[369,440,937,961]" captionStart="Map 2" captionStartId="27.[151,196,2023,2044]" captionTargetBox="[151,1398,1241,2014]" captionTargetId="figure-455@27.[151,1398,1241,2014]" captionTargetPageId="27" captionText="Map 2: Distributions of Anaulacaspis nigra (black circles), A. pseudonigra (white circle), A. reticulata (white triangle), A. libanotica (black triangles), A. caucasica (white diamonds), A. truncata (black diamonds), and A. sinuata (stars) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212628" httpUri="https://zenodo.org/record/5212628/files/figure.png" pageId="28" pageNumber="229">Map 2</figureCitation>
). The altitudes indicated on the labels range from
<quantity id="4CC01E9DFFB5FFB7FE9372F5FD95FC47" box="[379,545,969,993]" metricMagnitude="3" metricUnit="m" metricValue="1.815" metricValueMax="1.89" metricValueMin="1.7399999999999998" pageId="28" pageNumber="229" unit="m" value="1815.0" valueMax="1890.0" valueMin="1740.0">1740 to 1890 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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