164 lines
21 KiB
XML
164 lines
21 KiB
XML
<document id="6D7F9D590448A6552E091AFCF6965415" ID-CLB-Dataset="2489" ID-DOI="10.21248/contrib.entomol.66.2.201-255" ID-GBIF-Dataset="991c89c8-c376-46a5-9aa8-6dfbbf623c23" ID-ISSN="0005-805X" ID-Zenodo-Dep="5212606" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629211447587" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2016" docId="0391026EFF8FFF8DFF0B71CFFB7BFC47" docLanguage="en" docName="ContribEntomol.66.2.201-255.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 66 (2)" docSource="https://www.contributions-to-entomology.org/article/view/1904" docStyle="DocumentStyle:05A48F5F6902F879680B6E64765AA512.2:ContribEntomol.2013-2016.journal_article" docStyleId="05A48F5F6902F879680B6E64765AA512" docStyleName="ContribEntomol.2013-2016.journal_article" docStyleVersion="2" docTitle="Anaulacaspis cristata Assing 2016, spec. nov." docType="treatment" docVersion="10" lastPageNumber="239" masterDocId="FFA87A16FFA9FFABFFE8713CFFB4FFA6" masterDocTitle="Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="255" masterPageNumber="201" pageNumber="239" updateTime="1699087475910" updateUser="ExternalLinkService" zenodo-license-document="Abstyles">
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<mods:titleInfo id="E0847EA61AD80825EB9DC65E301C07AE">
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<mods:title id="4D4F164399DEE136B5C6F60095AF97DC">Revision of the Anaulacaspis species of the Palaearctic region (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:name id="2EE09A69B7E90DA9049B9EFCB213B934" type="personal">
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<mods:namePart id="264946864BDE8B78424681073AB13CC0">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="BB9152462F73942ABDE2CA94832FEE79">text</mods:typeOfResource>
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<mods:title id="A8AE60224DB1E296E41576A105804AAB">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="DE9BC80FF33CA52E204F96020C1C50EA">
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<mods:date id="CDC110C6B36E1378F16A36496A3B61B5">2016</mods:date>
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<mods:number id="19F54BAA27F40D28CAB366B6DBF91534">2016-12-20</mods:number>
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<mods:number id="8B4D2FAB541A6FA9915C37BA8AED1CF5">66</mods:number>
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<mods:url id="85DCD027F5A2398C3B7DBAC27219319D">https://www.contributions-to-entomology.org/article/view/1904</mods:url>
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<mods:classification id="96BB8E3C98F2F5042BA63EB38A37DC0F">journal article</mods:classification>
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<mods:identifier id="94FB2B3A35B1A9BEAC290EEF60069F60" type="CLB-Dataset">2489</mods:identifier>
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<mods:identifier id="7A59A7EF639796D76F13D2BBB4D5918B" type="DOI">10.21248/contrib.entomol.66.2.201-255</mods:identifier>
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<treatment id="0391026EFF8FFF8DFF0B71CFFB7BFC47" ID-DOI="http://doi.org/10.5281/zenodo.5884821" ID-GBIF-Taxon="192454165" ID-Zenodo-Dep="5884821" LSID="urn:lsid:plazi:treatment:0391026EFF8FFF8DFF0B71CFFB7BFC47" httpUri="http://treatment.plazi.org/id/0391026EFF8FFF8DFF0B71CFFB7BFC47" lastPageNumber="239" pageId="38" pageNumber="239">
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<subSubSection id="C322E0F3FF8FFF8DFF0B71CFFE16FE89" pageId="38" pageNumber="239" type="nomenclature">
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<paragraph id="8B87B378FF8FFF8DFF0B71CFFD9AFEA8" blockId="38.[227,558,243,303]" box="[227,558,243,270]" pageId="38" pageNumber="239">
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<heading id="D0CF0414FF8FFF8DFF0B71CFFD9AFEA8" box="[227,558,243,270]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="239" reason="3">
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<taxonomicName id="4C38C8FBFF8FFF8DFF0B71CFFE0AFEA8" ID-CoL="DJJ6" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" box="[227,446,243,270]" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="cristata" status="sp. nov.">
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<emphasis id="B94C6F6AFF8FFF8DFF0B71CFFE0AFEA8" box="[227,446,243,270]" italics="true" pageId="38" pageNumber="239">Anaulacaspis cristata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27FD211FF8FFF8DFE2D71CFFD9AFEA8" box="[453,558,243,270]" pageId="38" pageNumber="239" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8B87B378FF8FFF8DFF0B7025FE16FE89" blockId="38.[227,558,243,303]" box="[227,418,281,303]" pageId="38" pageNumber="239">
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(
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<figureCitation id="1303AFFDFF8FFF8DFF0F7025FEE9FE89" box="[231,349,281,303]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="38" pageNumber="239">Figs 132–135</figureCitation>
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,
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<figureCitation id="1303AFFDFF8FFF8DFE8D7025FE29FE89" box="[357,413,281,303]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="38" pageNumber="239">Map 5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C322E0F3FF8FFF8DFF557075FDEBFE67" pageId="38" pageNumber="239" type="materials_examined">
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<paragraph id="8B87B378FF8FFF8DFF557075FDEBFE67" blockId="38.[189,794,329,449]" pageId="38" pageNumber="239">
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<emphasis id="B94C6F6AFF8FFF8DFF557075FEE9FEC7" bold="true" box="[189,349,329,353]" pageId="38" pageNumber="239">Type material</emphasis>
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:
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<materialsCitation id="3B50B925FF8FFF8DFE9B7075FDEBFE67" ID-GBIF-Occurrence="3449483336" collectingDate="2007-07-05" collectionCode="MNB" collectorName="H. Muhle" country="PAK" county="Thiee" determinerName="V. Assing" elevation="1760" latitude="36.0229" location="Northern Areas" longLatPrecision="6" longitude="74.1681" municipality="Ghizar valley" pageId="38" pageNumber="239" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54830DDAFF8FFF8DFE9B7075FE62FEC7" box="[371,470,329,353]" pageId="38" pageNumber="239" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D3E78F1FF8FFF8DFE0E7076FDB5FEC7" box="[486,513,330,353]" pageId="38" pageNumber="239">♂:</specimenCount>
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“
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<collectingCountry id="F32FF3E8FF8FFF8DFDF27075FDFBFEC7" box="[538,591,329,353]" pageId="38" pageNumber="239">PAK</collectingCountry>
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,
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<location id="8EE7E5A3FF8FFF8DFD8A7075FCA2FEC7" LSID="urn:lsid:plazi:treatment:0391026EFF8FFF8DFF0B71CFFB7BFC47:8EE7E5A3FF8FFF8DFD8A7075FCA2FEC7" box="[610,790,329,353]" country="PAK" county="Thiee" latitude="36.0229" longLatPrecision="6" longitude="74.1681" municipality="Ghizar valley" name="Northern Areas" pageId="38" pageNumber="239">Northern Areas</location>
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,
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<collectingMunicipality id="6BE32902FF8FFF8DFF557055FEE0FE27" box="[189,340,361,385]" pageId="38" pageNumber="239">Ghizar valley</collectingMunicipality>
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,
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<collectingCounty id="62E6CBF4FF8FFF8DFE8F7055FE12FE27" box="[359,422,361,385]" pageId="38" pageNumber="239">Thiee</collectingCounty>
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,
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<geoCoordinate id="EE0CD5BFFF8FFF8DFE527055FD98FE27" box="[442,556,361,385]" degrees="36.0229" direction="north" orientation="latitude" pageId="38" pageNumber="239" precision="5" value="36.0229">36.0229°N</geoCoordinate>
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,
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<geoCoordinate id="EE0CD5BFFF8FFF8DFDA87055FD19FE27" box="[576,685,361,385]" degrees="74.1681" direction="east" orientation="longitude" pageId="38" pageNumber="239" precision="5" value="74.1681">74.1681°E</geoCoordinate>
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,
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<date id="FF8695B8FF8FFF8DFD297055FCADFE27" box="[705,793,361,385]" pageId="38" pageNumber="239" value="2007-07-05">
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<collectingDate id="EFC26C50FF8FFF8DFD297055FCADFE27" box="[705,793,361,385]" pageId="38" pageNumber="239" value="2007-07-05">5.7.2007</collectingDate>
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</date>
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<quantity id="4CC01E9DFF8FFF8DFF5570B5FEBAFE07" box="[189,270,393,417]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="38" pageNumber="239" unit="m" value="1760.0">
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<elevation id="0015544BFF8FFF8DFF5570B5FEBAFE07" box="[189,270,393,417]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="38" pageNumber="239" unit="m" value="1760.0">1760 m</elevation>
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</quantity>
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, leg.
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<collectorName id="26CDD6AEFF8FFF8DFEAF70B6FE1AFE07" box="[327,430,393,417]" pageId="38" pageNumber="239">H. Mühle</collectorName>
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/ Holotypus ♂
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<taxonomicName id="4C38C8FBFF8FFF8DFDB270B5FF52FE67" authority="Assing, 2016" authorityName="Assing" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="cristata" status="sp. nov.">
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<emphasis id="B94C6F6AFF8FFF8DFDB270B5FF52FE67" italics="true" pageId="38" pageNumber="239">Anaulacaspis cristata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27FD211FF8FFF8DFF047096FE91FE67" box="[236,293,426,449]" pageId="38" pageNumber="239" rank="species">sp. n.</taxonomicNameLabel>
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, det.
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<determinerName id="7E5B5FBCFF8FFF8DFEB67096FE74FE67" box="[350,448,425,449]" pageId="38" pageNumber="239">V. Assing</determinerName>
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2016” (
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<collectionCode id="ED292BBDFF8FFF8DFDFA7095FDE7FE67" box="[530,595,425,449]" collectionName="MNB" pageId="38" pageNumber="239">MNB</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C322E0F3FF8FFF8DFF5570D5FE49FDE7" pageId="38" pageNumber="239" type="etymology">
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<paragraph id="8B87B378FF8FFF8DFF5570D5FE49FDE7" blockId="38.[189,794,489,577]" pageId="38" pageNumber="239">
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<emphasis id="B94C6F6AFF8FFF8DFF5570D5FE80FDA7" bold="true" box="[189,308,489,513]" pageId="38" pageNumber="239">Etymology</emphasis>
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: The specific epithet is an adjective derived from the Latin noun crista. It alludes to the enormous crista apicalis of the aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C322E0F3FF8FFF8DFF557355FC63FC27" pageId="38" pageNumber="239" type="description">
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<paragraph id="8B87B378FF8FFF8DFF557355FE3BFCE7" blockId="38.[189,795,617,1281]" pageId="38" pageNumber="239">
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<emphasis id="B94C6F6AFF8FFF8DFF557355FE8BFD27" bold="true" box="[189,319,617,641]" pageId="38" pageNumber="239">Description</emphasis>
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: Body length
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<quantity id="4CC01E9DFF8FFF8DFE0D7355FD8DFD27" box="[485,569,617,641]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="38" pageNumber="239" unit="mm" value="2.3">2.3 mm</quantity>
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; length of forebody
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<quantity id="4CC01E9DFF8FFF8DFF5573B5FEA5FD07" box="[189,273,649,673]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="38" pageNumber="239" unit="mm" value="1.2">1.2 mm</quantity>
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. Coloration: head pale reddish-brown; pronotum reddish; elytra yellowish, with a rather pale and diffusely delimited medio-lateral infuscate spot; abdomen reddish with blackish segments VI and VII; legs yellowish; antennae brown with the basal three antennomeres pale-reddish.
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</paragraph>
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<paragraph id="8B87B378FF8FFF8DFF557275FDC6FBA7" blockId="38.[189,795,617,1281]" pageId="38" pageNumber="239">Head approximately 1.1 times as broad as long, probably with weak sexual dimorphism. Eyes indistinctly longer than postocular region in dorsal view. Antennae: antennomere IV weakly oblong; antennomeres V–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long.</paragraph>
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<paragraph id="8B87B378FF8FFF8DFF557536FE26FBC7" blockId="38.[189,795,617,1281]" pageId="38" pageNumber="239">Pronotum approximately 1.1 times as broad as long and approximately as broad as head, evidently with distinct sexual dimorphism.</paragraph>
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<paragraph id="8B87B378FF8FFF8DFF557555FDDBFB07" blockId="38.[189,795,617,1281]" pageId="38" pageNumber="239">Elytra approximately as long as pronotum; punctation dense and very fine. Hind wings present.</paragraph>
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<paragraph id="8B87B378FF8FFF8DFF557595FB95FE87" blockId="38.[189,795,617,1281]" lastBlockId="38.[831,1437,233,641]" pageId="38" pageNumber="239">
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Abdomen narrower than elytra; punctation distinct; tergite VIII (
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<figureCitation id="1303AFFDFF8FFF8DFEAB75F5FE2EFB47" box="[323,410,1225,1249]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="38" pageNumber="239">Fig. 132</figureCitation>
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) rather weakly transverse, posterior margin truncate, postero-laterally without a cluster of long thin setae on either side, marginal setae unmodified and moderately long.
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</paragraph>
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<paragraph id="8B87B378FF8FFF8DFCD77016FC06FDC7" blockId="38.[831,1437,233,641]" pageId="38" pageNumber="239">
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♂: head with distinct punctation and with weakly pronounced median impression posteriorly; pronotum with extensive and distinct median impression in posterior two-thirds and with dense and distinct punctation; posterior margin of sternite VIII (
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<figureCitation id="1303AFFDFF8FFF8DFB5E7095FABAFE67" box="[1206,1294,425,449]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="38" pageNumber="239">Fig. 133</figureCitation>
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) indistinctly, obtusely pointed in the middle; median lobe of aedeagus (
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<figureCitation id="1303AFFDFF8FFF8DFCAF70D5FC62FDA7" box="[839,982,489,513]" captionStart="Figs 113–135" captionStartId="16.[189,229,1970,1992]" captionTargetBox="[194,1432,241,1954]" captionTargetId="figure-134@16.[190,1436,232,1969]" captionText="Figs 113–135: Anaulacaspsis seclusa (113–120; 115–116: holotype of A. klapperichi; 113–114, 117–120: Iran), A. gratilla (121–131; 121–124: lectotype of A. lutzi; 128–131: nanistic male from Azerbaijan), and A. cristata (132–135): male tergite VIII (113, 121, 128, 132); male sternite VIII (114, 122, 129, 133); median lobe of aedeagus in lateral and in ventral view (115–118, 123–126, 130–131, 134–135); female tergite VIII (119); female sternite VIII (120, 127). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5884771" httpUri="https://zenodo.org/record/5884771/files/figure.png" pageId="38" pageNumber="239">Figs 134–135</figureCitation>
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)
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<quantity id="4CC01E9DFF8FFF8DFC0370D5FBFEFDA7" box="[1003,1098,489,513]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="38" pageNumber="239" unit="mm" value="0.35">0.35 mm</quantity>
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long; ventral process narrow, short (in relation to capsule), and weakly curved in lateral view, of triangular shape in ventral view; crista apicalis enormous.
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</paragraph>
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<paragraph id="8B87B378FF8FFF8DFCD77356FC7AFD27" blockId="38.[831,1437,233,641]" box="[831,974,617,641]" pageId="38" pageNumber="239">♀: unknown.</paragraph>
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<paragraph id="8B87B378FF8FFF8DFCD77395FC63FC27" blockId="38.[831,1436,681,897]" pageId="38" pageNumber="239">
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<emphasis id="B94C6F6AFF8FFF8DFCD77395FBA6FD67" bold="true" box="[831,1042,681,705]" pageId="38" pageNumber="239">Comparative notes</emphasis>
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: As can be inferred from the shapes and chaetotaxy of tergite and sternite VIII, as well as from the general morphology of the aedeagus,
|
||
<taxonomicName id="4C38C8FBFF8FFF8DFAB873D5FCDDFC87" authorityName="Assing" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Anaulacaspis" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="cristata">
|
||
<emphasis id="B94C6F6AFF8FFF8DFAB873D5FCDDFC87" italics="true" pageId="38" pageNumber="239">A. cristata</emphasis>
|
||
</taxonomicName>
|
||
belongs to the
|
||
<taxonomicName id="4C38C8FBFF8FFF8DFBE57235FBCBFC87" box="[1037,1151,777,801]" class="Insecta" family="Staphylinidae" genus="Falagria" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="239" phylum="Arthropoda" rank="species" species="naevula">
|
||
<emphasis id="B94C6F6AFF8FFF8DFBE57235FBCBFC87" box="[1037,1151,777,801]" italics="true" pageId="38" pageNumber="239">A. naevula</emphasis>
|
||
</taxonomicName>
|
||
group. Among the species of this group, it is characterized particularly by the morphology of the aedeagus, above all the enormous crista apicalis.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C322E0F3FF8FFF8DFCD77295FB7BFC47" pageId="38" pageNumber="239" type="distribution">
|
||
<paragraph id="8B87B378FF8FFF8DFCD77295FB7BFC47" blockId="38.[831,1436,937,993]" pageId="38" pageNumber="239">
|
||
<emphasis id="B94C6F6AFF8FFF8DFCD77295FC7CFC67" bold="true" box="[831,968,937,961]" pageId="38" pageNumber="239">Distribution</emphasis>
|
||
: The
|
||
<typeStatus id="54830DDAFF8FFF8DFBEE7296FB80FC64" box="[1030,1076,938,962]" pageId="38" pageNumber="239">type</typeStatus>
|
||
locality is situated in North
|
||
<collectingCountry id="F32FF3E8FF8FFF8DFA8C7295FCDFFC47" name="Pakistan" pageId="38" pageNumber="239">Pakistan</collectingCountry>
|
||
(
|
||
<figureCitation id="1303AFFDFF8FFF8DFC9172F5FC0BFC47" box="[889,959,969,993]" captionStart="Map 5" captionStartId="38.[189,234,2025,2046]" captionTargetBox="[189,1436,1328,2016]" captionTargetId="figure-506@38.[189,1436,1328,2016]" captionTargetPageId="38" captionText="Map 5: Distributions of Anaulacaspis flavomarginata (white diamonds), A. pamphylica (black star), A. inexpectata (white circles), and A. persica (black circles) in the West Palaearctic and Middle Asia, based on revised records." figureDoi="http://doi.org/10.5281/zenodo.5212639" httpUri="https://zenodo.org/record/5212639/files/figure.png" pageId="38" pageNumber="239">Map 5</figureCitation>
|
||
) at an altitude of
|
||
<quantity id="4CC01E9DFF8FFF8DFB9172F5FB7EFC47" box="[1145,1226,969,993]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="38" pageNumber="239" unit="m" value="1760.0">1760 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |