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<document id="7E2239EF7FA18EEFE5A2450F508223A0" ID-DOI="10.5252/z2012n2a13" ID-ISSN="1638-9387" ID-Zenodo-Dep="5165517" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628214690286" checkinUser="felipe" docAuthor="Macpherson, Enrique" docDate="2012" docId="03848F29FF8CFFF15D664FBA3CC69411" docLanguage="en" docName="zoosystema.34.2.409-427.pdf" docOrigin="Zoosystema 34 (2)" docSource="http://dx.doi.org/10.5252/z2012n2a13" docStyle="DocumentStyle:EB3C2C73672D2E883259A050BD27978B.6:Zoosystema.2011-2012.journal_article" docStyleId="EB3C2C73672D2E883259A050BD27978B" docStyleName="Zoosystema.2011-2012.journal_article" docStyleVersion="6" docTitle="Galathea echinata Macpherson 2012, n. sp." docType="treatment" docVersion="3" lastPageNumber="416" masterDocId="FFBDF751FF8FFFF65E4D4C0A387C903B" masterDocTitle="New deep-sea squat lobsters of the genus Galathea Fabricius, 1793 (Decapoda, Galatheidae) from Vanuatu and New Caledonia" masterLastPageNumber="427" masterPageNumber="409" pageNumber="412" updateTime="1699035975347" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AE546AE137DA6B21841B0AEAF3F9A107">New deep-sea squat lobsters of the genus Galathea Fabricius, 1793 (Decapoda, Galatheidae) from Vanuatu and New Caledonia</mods:title>
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<mods:name id="E805C476041DAFD3EF63975684CCF5D0" type="personal">
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<mods:namePart id="2DAC8884A8A04B994EF6568FAEFA5984">Macpherson, Enrique</mods:namePart>
<mods:affiliation id="6C688DDF31EFEEDB160962AAF22DA9BA">Centre dEstudis Avançats de Blanes (CEAB-CSIC), c. acc. Cala San Francesc 14, E- 17300 Blanes (Spain) macpherson @ ceab. csic. es</mods:affiliation>
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<mods:title id="70FA0BF2F774F1E0A9E969BF2AAFC491">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="F5CE1673B5053AF0D3DE328381E2FA7C">
<mods:date id="5D284A6C0885FDB997525523C5F687C9">2012</mods:date>
<mods:detail id="C8C112AAA3AB2E4C46509C9737217FB4" type="pubDate">
<mods:number id="C378086DF713C511D249257ED01F158A">2012-06-30</mods:number>
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<mods:number id="EDAD866DD3A440C58BE5F7DA150ACF78">34</mods:number>
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<mods:classification id="1C832559E5C11E4B914E26655D940466">journal article</mods:classification>
<mods:identifier id="5395D0C29E70A10D6B9CC9E26C8AFB10" type="DOI">10.5252/z2012n2a13</mods:identifier>
<mods:identifier id="BF372104E546AD940CECF3A7F656AA08" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="D05CA492B32A10103614D9AE4BAEAA59" type="Zenodo-Dep">5165517</mods:identifier>
</mods:mods>
<treatment id="03848F29FF8CFFF15D664FBA3CC69411" LSID="urn:lsid:plazi:treatment:03848F29FF8CFFF15D664FBA3CC69411" httpUri="http://treatment.plazi.org/id/03848F29FF8CFFF15D664FBA3CC69411" lastPageId="7" lastPageNumber="416" pageId="3" pageNumber="412">
<subSubSection id="C3376DB4FF8CFFF55D664FBA3C4F93F0" box="[811,1075,944,971]" pageId="3" pageNumber="412" type="nomenclature">
<paragraph id="8B923E3FFF8CFFF55D664FBA3C4F93F0" blockId="3.[811,1075,944,1002]" box="[811,1075,944,971]" pageId="3" pageNumber="412">
<heading id="D0DA8953FF8CFFF55D664FBA3C4F93F0" box="[811,1075,944,971]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="412" reason="2">
<taxonomicName id="4C2D45BCFF8CFFF55D664FBA3B8E93F0" ID-CoL="c6efb651-3a57-4e47-8fdd-5c2197fda1dc" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[811,1010,944,971]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="echinata" status="sp. nov.">
<emphasis id="B959E22DFF8CFFF55D664FBA3B8E93F0" bold="true" box="[811,1010,944,971]" italics="true" pageId="3" pageNumber="412">Galathea echinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8CFFF55DB54FBB3C4F93F0" box="[1016,1075,945,971]" pageId="3" pageNumber="412" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3376DB4FF8CFFF55DCA4FDA3BAA93D1" box="[903,982,976,1002]" pageId="3" pageNumber="412" type="description">
<paragraph id="8B923E3FFF8CFFF55DCA4FDA3BAA93D1" blockId="3.[811,1075,944,1002]" box="[903,982,976,1002]" pageId="3" pageNumber="412">
(
<figureCitation id="131622BAFF8CFFF55DC24FDA3BB393D1" box="[911,975,976,1002]" captionStart="FIG" captionStartId="6.[113,124,1612,1629]" captionTargetBox="[124,1205,220,1557]" captionTargetId="figure-121@6.[113,1209,214,1562]" captionTargetPageId="6" captionText="FIG. 2. — Galathea echinata n. sp., holotype ♂, 5.1 mm (MNHN-IU-2009-591): A, carapace and abdomen, dorsal view; B, anterior part of sternal plastron; C, cephalic region, showing antennular and antennal peduncles, ventral view; D, right Mxp3, lateral view; E, right P1, dorsal view; F, distal part of P1 fingers, ventral view; G, right P2, lateral view; H, right P3, lateral view; I, right P4, lateral view. Scale bars: A, B, D, F, 2 mm; C, 1.5 mm; E, G, H, I, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5165521" httpUri="https://zenodo.org/record/5165521/files/figure.png" pageId="3" pageNumber="412">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3376DB4FF8CFFF55CE848003CCD9481" pageId="3" pageNumber="412" type="materials_examined">
<paragraph id="8B923E3FFF8CFFF55CE848003CE5946D" blockId="3.[677,1210,1033,1110]" pageId="3" pageNumber="412">
<materialsCitation id="3B453462FF8CFFF55CE848003CE5946D" collectingDate="2003-10-21" country="New Caledonia" elevation="371" latitude="-23.465334" location="New Caledonia" longLatPrecision="12" longitude="167.84833" pageId="3" pageNumber="412" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="5496809DFF8CFFF55CE848003B6F941B" box="[677,787,1034,1057]" pageId="3" pageNumber="412" type="holotype">HOLOTYPE</typeStatus>
. —
<emphasis id="B959E22DFF8CFFF55D0848033B949418" bold="true" box="[837,1000,1033,1059]" pageId="3" pageNumber="412">
<collectingCountry id="F33A7EAFFF8CFFF55D0848033B949418" box="[837,1000,1033,1059]" name="New Caledonia" pageId="3" pageNumber="412">New Caledonia</collectingCountry>
</emphasis>
. NORFOLK 2, stn DW2024,
<geoCoordinate id="EE1958F8FF8CFFF55D51482E3BE99407" box="[796,917,1060,1084]" degrees="23" direction="south" minutes="27.92" orientation="latitude" pageId="3" pageNumber="412" precision="9" value="-23.465334">23°27.92S</geoCoordinate>
,
<geoCoordinate id="EE1958F8FF8CFFF55DEE48293C519400" box="[931,1069,1059,1083]" degrees="167" direction="east" minutes="50.90" orientation="longitude" pageId="3" pageNumber="412" precision="9" value="167.84833">167°50.90E</geoCoordinate>
,
<quantity id="4CD593DAFF8CFFF55A76482E3CCB9400" box="[1083,1207,1060,1084]" metricMagnitude="2" metricUnit="m" metricValue="3.7049999999999996" metricValueMax="3.71" metricValueMin="3.7" pageId="3" pageNumber="412" unit="m" value="370.5" valueMax="371.0" valueMin="370.0">
<elevation id="0000D90CFF8CFFF55A76482E3CCB9400" box="[1083,1207,1060,1084]" metricMagnitude="2" metricUnit="m" metricValue="3.7049999999999996" metricValueMax="3.71" metricValueMin="3.7" pageId="3" pageNumber="412" unit="m" value="370.5" valueMax="371.0" valueMin="370.0">370-371 m</elevation>
</quantity>
,
<date id="FF9318FFFF8CFFF55CE848343B73946D" box="[677,783,1086,1110]" pageId="3" pageNumber="412" value="2003-10-21">
<collectingDate id="EFD7E117FF8CFFF55CE848343B73946D" box="[677,783,1086,1110]" pageId="3" pageNumber="412" value="2003-10-21">21.X.2003</collectingDate>
</date>
,
<specimenCount id="9D2BF5B6FF8CFFF55D5748343B3D946E" box="[794,833,1086,1110]" pageId="3" pageNumber="412" type="male">1 ♂</specimenCount>
,
<quantity id="4CD593DAFF8CFFF55D0348343BE0946D" box="[846,924,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="3" pageNumber="412" unit="mm" value="5.1">5.1 mm</quantity>
(MNHN-IU-2009-591).
</materialsCitation>
</paragraph>
<paragraph id="8B923E3FFF8CFFF55CE848643CCD9481" blockId="3.[677,1210,1133,1211]" pageId="3" pageNumber="412">
<typeStatus id="5496809DFF8CFFF55CE848643B7194BF" box="[677,781,1134,1157]" pageId="3" pageNumber="412" type="paratype">PARATYPES</typeStatus>
. —
<emphasis id="B959E22DFF8CFFF55D7048643BA694BC" bold="true" box="[829,986,1134,1159]" pageId="3" pageNumber="412">
<collectingCountry id="F33A7EAFFF8CFFF55D7048643BA694BC" box="[829,986,1134,1159]" name="New Caledonia" pageId="3" pageNumber="412">New Caledonia</collectingCountry>
</emphasis>
.
<collectionCode id="ED3CA6FAFF8CFFF55DA448643C5994BD" box="[1001,1061,1134,1158]" pageId="3" pageNumber="412">SMIB</collectionCode>
5, stn DW88,
<geoCoordinate id="EE1958F8FF8CFFF55CEB48823B65949B" box="[678,793,1160,1184]" degrees="22" direction="south" minutes="18.60" orientation="latitude" pageId="3" pageNumber="412" precision="9" value="-22.31">22°18.60S</geoCoordinate>
,
<geoCoordinate id="EE1958F8FF8CFFF55D6848823BDB949B" box="[805,935,1160,1184]" degrees="168" direction="east" minutes="40.20" orientation="longitude" pageId="3" pageNumber="412" precision="9" value="168.67">168°40.20E</geoCoordinate>
,
<quantity id="4CD593DAFF8CFFF55DF948823B86949B" box="[948,1018,1160,1184]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="412" unit="m" value="350.0">350 m</quantity>
,
<date id="FF9318FFFF8CFFF55A4B48823C01949B" box="[1030,1149,1160,1184]" pageId="3" pageNumber="412" value="1989-09-13">13.IX.1989</date>
,
<specimenCount id="9D2BF5B6FF8CFFF55AC748823CCE94A4" box="[1162,1202,1160,1184]" pageId="3" pageNumber="412" type="male">1 ♂</specimenCount>
,
<quantity id="4CD593DAFF8CFFF55CE848A83A899481" box="[677,757,1186,1210]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="3" pageNumber="412" unit="mm" value="4.6">4.6 mm</quantity>
, 1 ovig.
<specimenCount id="9D2BF5B6FF8CFFF55D0A48AC3B249481" box="[839,856,1190,1210]" pageId="3" pageNumber="412" type="female"></specimenCount>
, 5.0 mm (MNHN-IU-2009-592).
</paragraph>
</subSubSection>
<subSubSection id="C3376DB4FF8CFFF55CE848D93CEE94D1" box="[677,1170,1234,1258]" pageId="3" pageNumber="412" type="distribution">
<paragraph id="8B923E3FFF8CFFF55CE848D93CEE94D1" blockId="3.[677,1170,1234,1258]" box="[677,1170,1234,1258]" pageId="3" pageNumber="412">
DISTRIBUTION. —
<collectingCountry id="F33A7EAFFF8CFFF55D1348D93B8494D1" box="[862,1016,1234,1258]" name="New Caledonia" pageId="3" pageNumber="412">New Caledonia</collectingCountry>
, in
<quantity id="4CD593DAFF8CFFF55A5048D83CF294D1" box="[1053,1166,1234,1258]" metricMagnitude="2" metricUnit="m" metricValue="3.6049999999999995" metricValueMax="3.71" metricValueMin="3.5" pageId="3" pageNumber="412" unit="m" value="360.5" valueMax="371.0" valueMin="350.0">
350-
<specimenCount id="9D2BF5B6FF8CFFF55A0648D83CF294D1" box="[1099,1166,1234,1258]" pageId="3" pageNumber="412" type="generic">371 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3376DB4FF8CFFF55CE849083B429575" pageId="3" pageNumber="412" type="etymology">
<paragraph id="8B923E3FFF8CFFF55CE849083B429575" blockId="3.[677,1210,1282,1359]" pageId="3" pageNumber="412">ETYMOLOGY. — From the Latin “echinatus”, spiny, in reference to the numerous spines on the dorsal surface of the carapace.</paragraph>
</subSubSection>
<subSubSection id="C3376DB4FF8CFFF15CE849673C7C916A" lastPageId="7" lastPageNumber="416" pageId="3" pageNumber="412" type="description">
<paragraph id="8B923E3FFF8CFFF55CE849673B4995BE" blockId="3.[675,1212,1389,1700]" box="[677,821,1389,1415]" pageId="3" pageNumber="412">DESCRIPTION</paragraph>
<paragraph id="8B923E3FFF8CFFF35CE8498639F395DE" blockId="3.[675,1212,1389,1700]" lastBlockId="5.[111,649,849,1700]" lastPageId="5" lastPageNumber="414" pageId="3" pageNumber="412">Carapace, exclusive of rostrum, 1.2 times as long as broad; dorsal surface of gastric region slightly convex, anterior cardiac region concave; anterior and posterior cervical grooves distinct; ridges on gastric and anterior branchial regions scale-like, not well defined; mid-transverse ridge interrupted, preceded by cervical groove, followed by scale-like ridges and interrupted transverse ridge; all ridges finely setose, with some scattered long and thick plumose setae.Two pairs of small epigastric spines; one median protogastric, and one parahepatic spine on each side; two submedian metagastric spines; one postcervical spine on each side and two cardiac spines. Lateral margins medially convex, with five spines: two spines in front of and three strong spines behind indistinct end of anterior cervical groove; first anterolateral, strong, distinctly posterior to level of lateral limit of orbit; second small, just anterior to lateral end of anterior cervical groove, accompanying another small spine ventral to between first and second; one spine on anterior branchial region, and two spines on posterior branchial margin, last slightly smaller than others, situated at level of cardiac spines. Small spine on lateral limit of orbit; infra-orbital margin with one spine. Rostrum 2.2-2.3 times as long as broad, length 0.6 that of, breadth 0.3 that of carapace, dorsal surface nearly horizontal in lateral view, with small setiferous ridges; lateral margin with four relatively small teeth.</paragraph>
<caption id="DF526EB7FF8BFFF25E3C4A693CC7969A" ID-DOI="http://doi.org/10.5281/zenodo.5165519" ID-Zenodo-Dep="5165519" httpUri="https://zenodo.org/record/5165519/files/figure.png" pageId="4" pageNumber="413" startId="4.[113,124,1635,1652]" targetBox="[144,1189,221,1591]" targetPageId="4">
<paragraph id="8B923E3FFF8BFFF25E3C4A693CC7969A" blockId="4.[113,1211,1635,1697]" pageId="4" pageNumber="413">
FIG. 1. —
<taxonomicName id="4C2D45BCFF8BFFF25EF04A693921964F" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[189,349,1635,1652]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="barbellata" status="sp. nov.">
<emphasis id="B959E22DFF8BFFF25EF04A693921964F" box="[189,349,1635,1652]" italics="true" pageId="4" pageNumber="413">Galathea barbellata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8BFFF25F2D4A6939ED964F" box="[352,401,1635,1652]" pageId="4" pageNumber="413" rank="species">n. sp.</taxonomicNameLabel>
, holotype ovig. ♀, 4.9 mm (MNHN-IU-2009-587):
<emphasis id="B959E22DFF8BFFF25D6F4A693B53964F" bold="true" box="[802,815,1635,1652]" pageId="4" pageNumber="413">A</emphasis>
, carapace and abdomen,dorsal view;
<emphasis id="B959E22DFF8BFFF25A264A693C04964F" bold="true" box="[1131,1144,1635,1652]" pageId="4" pageNumber="413">B</emphasis>
, sternal plastron;
<emphasis id="B959E22DFF8BFFF25EF24A7338B196B1" bold="true" box="[191,205,1657,1674]" pageId="4" pageNumber="413">C</emphasis>
, cephalic region, showing antennular and antennal peduncles, ventral view;
<emphasis id="B959E22DFF8BFFF25D0E4A733B2D96B1" bold="true" box="[835,849,1657,1674]" pageId="4" pageNumber="413">D</emphasis>
, right Mxp3, lateral view;
<emphasis id="B959E22DFF8BFFF25A684A733C4D96B1" bold="true" box="[1061,1073,1657,1674]" pageId="4" pageNumber="413">E</emphasis>
, right P1, dorsal view;
<emphasis id="B959E22DFF8BFFF25EEC4A9A38D0969A" bold="true" box="[161,172,1680,1697]" pageId="4" pageNumber="413">F</emphasis>
, right P2, lateral view;
<emphasis id="B959E22DFF8BFFF25F2B4A9A3908969A" bold="true" box="[358,372,1680,1697]" pageId="4" pageNumber="413">G</emphasis>
, right P3, lateral view;
<emphasis id="B959E22DFF8BFFF25C634A9A3A40969A" bold="true" box="[558,572,1680,1697]" pageId="4" pageNumber="413">H</emphasis>
, right P4, lateral view.Scale bars: A, B, F, G, H, 1 mm; C, D, 0.5 mm; E, 2 mm.
</paragraph>
</caption>
<caption id="DF526EB7FF8AFFF35ECD4CD13CD690D7" box="[128,1194,219,236]" pageId="5" pageNumber="414" startId="5.[128,139,219,236]">
<paragraph id="8B923E3FFF8AFFF35ECD4CD13CD690D7" blockId="5.[128,1194,219,236]" box="[128,1194,219,236]" pageId="5" pageNumber="414">
TABLE 1. — Main differences among
<taxonomicName id="4C2D45BCFF8AFFF35FE34CD13A3290D7" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[430,590,219,236]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="barbellata" status="sp. nov.">
<emphasis id="B959E22DFF8AFFF35FE34CD13A3290D7" box="[430,590,219,236]" italics="true" pageId="5" pageNumber="414">Galathea barbellata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8AFFF35C1E4CD13AFF90D7" box="[595,643,219,236]" pageId="5" pageNumber="414" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C2D45BCFF8AFFF35CC14CD13A9690D7" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[652,746,219,236]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="echinata" status="sp. nov.">
<emphasis id="B959E22DFF8AFFF35CC14CD13A9690D7" box="[652,746,219,236]" italics="true" pageId="5" pageNumber="414">G. echinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8AFFF35CBD4CD13B5C90D7" box="[752,800,219,236]" pageId="5" pageNumber="414" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C2D45BCFF8AFFF35D644CD13BF090D7" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[809,908,219,236]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="profunda" status="n. sp. and g.">
<emphasis id="B959E22DFF8AFFF35D644CD13BF090D7" box="[809,908,219,236]" italics="true" pageId="5" pageNumber="414">G. profunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8AFFF35DDC4CD13B8290D7" box="[913,1022,219,236]" pageId="5" pageNumber="414" rank="genus">
n. sp. and
<taxonomicName id="4C2D45BCFF8AFFF35DA44CD13B8290D7" authority="Baba, 1990" authorityName="Baba" authorityYear="1990" box="[1001,1022,219,236]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B959E22DFF8AFFF35DA44CD13B8290D7" box="[1001,1022,219,236]" italics="true" pageId="5" pageNumber="414">G.</emphasis>
</taxonomicName>
</taxonomicNameLabel>
<emphasis id="B959E22DFF8AFFF35A4F4CD13C3D90D7" box="[1026,1089,219,236]" italics="true" pageId="5" pageNumber="414">robusta</emphasis>
Baba, 1990.
</paragraph>
</caption>
<paragraph id="8B923E3FFF8AFFF35F164D1E3CE591E4" pageId="5" pageNumber="414">
<table id="F92DCC9FFF8A00095E3C4D1E3CC991E4" box="[113,1205,276,479]" gridcols="5" gridrows="3" pageId="5" pageNumber="414">
<tr id="351D3C7DFF8A00095E3C4D1E3CC99113" box="[113,1205,276,296]" gridrow="0" pageId="5" pageNumber="414" rowspan-0="1">
<th id="76CC5501FF8A00095F164D1E3A669113" box="[347,538,276,296]" gridcol="1" gridrow="0" pageId="5" pageNumber="414">
<emphasis id="B959E22DFF8AFFF35F164D1E3A669113" bold="true" box="[347,538,276,296]" pageId="5" pageNumber="414">
<taxonomicName id="4C2D45BCFF8AFFF35F164D1E39A19113" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[347,477,276,296]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="barbellata" status="sp. nov.">
<emphasis id="B959E22DFF8AFFF35F164D1E39A19113" bold="true" box="[347,477,276,296]" italics="true" pageId="5" pageNumber="414">G. barbellata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8AFFF35FAE4D1E3A669113" box="[483,538,276,296]" pageId="5" pageNumber="414" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CC5501FF8A00095C664D1E3B7C9113" box="[555,768,276,296]" gridcol="2" gridrow="0" pageId="5" pageNumber="414">
<emphasis id="B959E22DFF8AFFF35C704D1E3A939113" bold="true" box="[573,751,276,296]" pageId="5" pageNumber="414">
<taxonomicName id="4C2D45BCFF8AFFF35C704D1E3ACD9113" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[573,689,276,296]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="echinata" status="sp. nov.">
<emphasis id="B959E22DFF8AFFF35C704D1E3ACD9113" bold="true" box="[573,689,276,296]" italics="true" pageId="5" pageNumber="414">G. echinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8AFFF35CFA4D1E3A939113" box="[695,751,276,296]" pageId="5" pageNumber="414" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CC5501FF8A00095D594D1E3BB39113" box="[788,975,276,296]" gridcol="3" gridrow="0" pageId="5" pageNumber="414">
<emphasis id="B959E22DFF8AFFF35D5A4D1E3BB09113" bold="true" box="[791,972,276,296]" pageId="5" pageNumber="414">
<taxonomicName id="4C2D45BCFF8AFFF35D5A4D1E3BF39113" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[791,911,276,296]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="profunda" status="sp. nov.">
<emphasis id="B959E22DFF8AFFF35D5A4D1E3BF39113" bold="true" box="[791,911,276,296]" italics="true" pageId="5" pageNumber="414">G. profunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF8AFFF35DD84D1E3BB09113" box="[917,972,276,296]" pageId="5" pageNumber="414" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76CC5501FF8A00095DAF4D1E3CC99113" box="[994,1205,276,296]" gridcol="4" gridrow="0" pageId="5" pageNumber="414">
<taxonomicName id="4C2D45BCFF8AFFF35A5B4D1E3CFC9113" authorityName="Baba" authorityYear="1990" box="[1046,1152,276,296]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="414" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B959E22DFF8AFFF35A5B4D1E3CFC9113" bold="true" box="[1046,1152,276,296]" italics="true" pageId="5" pageNumber="414">G. robusta</emphasis>
</taxonomicName>
</th>
</tr>
<table id="F92DCC9FFF8A00095E3C4D3B3CC99272" box="[113,1205,305,585]" gridrows="6" pageId="5" pageNumber="414">
<tr id="351D3C7DFF8A00095E3C4D3B3CC99190" box="[113,1205,305,427]" gridrow="1" pageId="5" pageNumber="414">
<th id="76CC5501FF8A00095E3C4D3B395B9190" box="[113,295,305,427]" gridcol="0" gridrow="1" pageId="5" pageNumber="414">Carapace: spines on gastric region</th>
<td id="76CC5501FF8A00095F164D3B3A669190" box="[347,538,305,427]" gridcol="1" gridrow="1" pageId="5" pageNumber="414">9-11 epigastric, 2 submedian protogastric and 1-2 parahepatic on each side</td>
<td id="76CC5501FF8A00095C664D3B3B7C9190" box="[555,768,305,427]" gridcol="2" gridrow="1" pageId="5" pageNumber="414">4 epigastric, 1 median protogastric, 1 parahepatic on each side and 2 submedian metagastric</td>
<td id="76CC5501FF8A00095D594D3B3BB39190" box="[788,975,305,427]" gridcol="3" gridrow="1" pageId="5" pageNumber="414">1 parahepatic on each side, 1 strong median protogastric</td>
<td id="76CC5501FF8A00095DAF4D3B3CC99190" box="[994,1205,305,427]" gridcol="4" gridrow="1" pageId="5" pageNumber="414">7-8 epigastric, 2 lateral protogastric on each side and 2 median mesogastric</td>
</tr>
<table id="F92DCC9FFF8A00095E3C4DB83CD6933B" box="[113,1194,434,768]" gridrows="3" pageId="5" pageNumber="414">
<tr id="351D3C7DFF8A00095E3C4DB83CC991E4" box="[113,1205,434,479]" gridrow="2" pageId="5" pageNumber="414">
<th id="76CC5501FF8A00095E3C4DB8395B91E4" box="[113,295,434,479]" gridcol="0" gridrow="2" pageId="5" pageNumber="414">Carapace: ridges on gastric region</th>
<td id="76CC5501FF8A00095F164DB83A6691E4" box="[347,538,434,479]" gridcol="1" gridrow="2" pageId="5" pageNumber="414">some uninterrupted and well defined</td>
<td id="76CC5501FF8A00095C664DB83B7C91E4" box="[555,768,434,479]" gridcol="2" gridrow="2" pageId="5" pageNumber="414">scale-like and not well defined</td>
<td id="76CC5501FF8A00095D594DB83BB391E4" box="[788,975,434,479]" gridcol="3" gridrow="2" pageId="5" pageNumber="414">some uninterrupted and well defined</td>
<td id="76CC5501FF8A00095DAF4DB83CC991E4" box="[994,1205,434,479]" gridcol="4" gridrow="2" pageId="5" pageNumber="414">some uninterrupted and well defined</td>
</tr>
<paragraph id="8B923E3FFF8AFFF35E3C4DED39849272" pageId="5" pageNumber="414">Carapace: spines 5 3 4 5 on branchial margin Sternum: anterior 1-2 spines unarmed unarmed unarmed margin of sternite 4 on each side</paragraph>
<paragraph id="8B923E3FFF8AFFF35E3C4E5A3C03933B" pageId="5" pageNumber="414">Distal spines on antennular article 1 3 well developed, and 1 small 2 well developed, and 1 small 3 well developed 2 well developed, and 1 small Spines on flexor margin of 3 Mxp merus 3 of moderated size 1 strong at midlength 1 strong at midlength, and 1-3 small 3 of moderated size P2 merus 4 times longer than broad 3 times longer than broad 3 times longer than broad&gt; 4 times longer than broad</paragraph>
</table>
</table>
</table>
</paragraph>
<paragraph id="8B923E3FFF8AFFF35EC549E1393A961F" blockId="5.[111,649,849,1700]" pageId="5" pageNumber="414">Pterygostomian flap rugose with sparse setae, anteriorly rounded.</paragraph>
<paragraph id="8B923E3FFF8AFFF35EC54A203BAE9390" blockId="5.[111,649,849,1700]" lastBlockId="5.[674,1212,849,1699]" pageId="5" pageNumber="414">Sternal plastron 1.3 times as long as broad, lateral limits divergent posteriorly. Sternite 3 1.3 times as broad as long, anterior margin bilobated. Sternite 4 6.2 times longer and 4.7 times broader than preceding sternite, 0.5 as long as broad; surface with a few short transverse ridges bearing long setae. Following sternites smooth.</paragraph>
<paragraph id="8B923E3FFF8AFFF35CF64FBA3C229452" blockId="5.[674,1212,849,1699]" pageId="5" pageNumber="414">Abdominal somites 2-4 each with two uninterrupted transverse ridges on tergite; somite 5 and 6 each with two ridges, posterior one uninterruped on somite 5 and both medially interrupted on somite 6. Telson 0.8 as long as broad, incompletely subdivided. Two pairs of male gonopods.</paragraph>
<paragraph id="8B923E3FFF8AFFF35CF648643C209493" blockId="5.[674,1212,849,1699]" pageId="5" pageNumber="414">Ocular peduncles 1.8 times longer than broad; cornea not dilated, as broad as peduncle.</paragraph>
<paragraph id="8B923E3FFF8AFFF35CF648A43A87957C" blockId="5.[674,1212,849,1699]" pageId="5" pageNumber="414">Basal article of antennular peduncle with two well-developed distal spines, distodorsal larger than distolateral, distomesial obsolete. Ultimate article with a few short setae not in tuft on distodorsal margin.</paragraph>
<paragraph id="8B923E3FFF8AFFF35CF649463C3C95DE" blockId="5.[674,1212,849,1699]" pageId="5" pageNumber="414">Article 1 of antennal peduncle with depressed ventral distomesial process not reaching distal margin of article 2. Article 2 with distolateral spine clearly larger than distomesial, barely reaching midlength of article 3. Articles 3 and 4 unarmed.</paragraph>
<paragraph id="8B923E3FFF8AFFF35CF649E13B1D9698" blockId="5.[674,1212,849,1699]" pageId="5" pageNumber="414">Mxp3 ischium with well-developed distal spine on extensor margin; crista dentata with 21 or 22 denticles. Merus subequal in length to ischium, with one strong spine at midlength of flexor margin; extensor margin with very small spine. Carpus spineless.</paragraph>
<caption id="DF526EB7FF89FFF05E3C4A463AFD969A" ID-DOI="http://doi.org/10.5281/zenodo.5165521" ID-Zenodo-Dep="5165521" httpUri="https://zenodo.org/record/5165521/files/figure.png" pageId="6" pageNumber="415" startId="6.[113,124,1612,1629]" targetBox="[124,1205,220,1557]" targetPageId="6">
<paragraph id="8B923E3FFF89FFF05E3C4A463AFD969A" blockId="6.[113,1211,1612,1697]" pageId="6" pageNumber="415">
FIG. 2. —
<taxonomicName id="4C2D45BCFF89FFF05E884A4639279666" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[197,347,1612,1629]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="415" phylum="Arthropoda" rank="species" species="echinata" status="sp. nov.">
<emphasis id="B959E22DFF89FFF05E884A4639279666" box="[197,347,1612,1629]" italics="true" pageId="6" pageNumber="415">Galathea echinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF89FFF05F2C4A4639E89666" box="[353,404,1612,1629]" pageId="6" pageNumber="415" rank="species">n. sp.</taxonomicNameLabel>
, holotype ♂, 5.1 mm (MNHN-IU-2009-591):
<emphasis id="B959E22DFF89FFF05D474A463B6B9666" bold="true" box="[778,791,1612,1629]" pageId="6" pageNumber="415">A</emphasis>
, carapace and abdomen, dorsal view;
<emphasis id="B959E22DFF89FFF05A2C4A463C129666" bold="true" box="[1121,1134,1612,1629]" pageId="6" pageNumber="415">B</emphasis>
, anterior part of sternal plastron;
<emphasis id="B959E22DFF89FFF05F734A693930964F" bold="true" box="[318,332,1635,1652]" pageId="6" pageNumber="415">C</emphasis>
, cephalic region, showing antennular and antennal peduncles, ventral view;
<emphasis id="B959E22DFF89FFF05D984A693B9F964F" bold="true" box="[981,995,1635,1652]" pageId="6" pageNumber="415">D</emphasis>
, right Mxp3, lateral view;
<emphasis id="B959E22DFF89FFF05E3C4A73380196B1" bold="true" box="[113,125,1657,1674]" pageId="6" pageNumber="415">E</emphasis>
, right P1, dorsal view;
<emphasis id="B959E22DFF89FFF05F734A73393596B1" bold="true" box="[318,329,1657,1674]" pageId="6" pageNumber="415">F</emphasis>
, distal part of P1 fingers, ventral view;
<emphasis id="B959E22DFF89FFF05CDD4A733AE296B1" bold="true" box="[656,670,1657,1674]" pageId="6" pageNumber="415">G</emphasis>
, right P2, lateral view;
<emphasis id="B959E22DFF89FFF05D104A733B1796B1" bold="true" box="[861,875,1657,1674]" pageId="6" pageNumber="415">H</emphasis>
, right P3, lateral view;
<emphasis id="B959E22DFF89FFF05A664A733C4D96B1" bold="true" box="[1067,1073,1657,1674]" pageId="6" pageNumber="415">I</emphasis>
, right P4, lateral view. Scale bars: A, B, D, F, 2 mm; C, 1.5 mm; E, G, H, I, 1 mm.
</paragraph>
</caption>
<paragraph id="8B923E3FFF88FFF15EC54CD2392E9350" blockId="7.[111,649,216,1700]" pageId="7" pageNumber="416">
P1 4.9 (males), 4.0 (females) times postorbital carapace length, relatively slender, subcylindrical, with long stiff setae on striae of all articles, and some scattered long and thick plumose setae. Merus 1.6- 1.9 times length of carapace, 1.8-2.3 times as long as carpus, with several rows of spines, dorsomesial row with strong spines. Carpus 0.7-0.9 length of palm, 2.4 (males), 1.5 (females) times longer than broad, dorsal surface with row of small spines; mesial surface with well-developed spines; some small spines along lateral margin. Palm 2.9-3.8 (males), 4.5 (females) times longer than broad; small dorsal spines roughly in three rows: mesial, dorsal and lateral; lateral row not continued on to fixed finger. Fingers 0.7 (males), 0.6 (females) as long as palm, distally spooned, touching each other with intermeshing teeth when closed; opposable margins gaping (
<typeStatus id="5496809DFF88FFF15E8B4EF839569336" box="[198,298,754,781]" pageId="7" pageNumber="416" type="holotype">holotype</typeStatus>
); when gaping, fixed finger with eminence at distal third and movable finger with process at proximal third; mesial margin of movable finger unarmed.
</paragraph>
<paragraph id="8B923E3FFF88FFF15EC54F7B3BDF910A" blockId="7.[111,649,216,1700]" lastBlockId="7.[675,1209,216,337]" pageId="7" pageNumber="416">P2-4 relatively short, somewhat compressed, setose, sparsely with thick long plumose setae on all articles. Meri successively shorter posteriorly (P3 merus 0.8 length of P2 merus, P4 merus 0.9 length of P3 merus), equally broad on P2-4; P2 merus 0.6 carapace length, 3.2 times as long as broad, 1.2 times longer than P2 propodus; P3 merus 2.9 times as long as broad, 1.0 times length of P3 propodus; P4 merus 2.5 times as long as broad, 0.8 length of P4 propodus; dorsal margins with row of 5-6 proximally diminishing spines on P2-3, with terminal spine only on P4; dorsolateral surface unarmed on P2-4; ventrolateral margins with strong terminal spine; ventromesial margin with small terminal spine on P2 only. Carpi with four or five (on P2-3) and 1-3 (on P4) spines on extensor margin, distal-most larger; dorsolateral surface with row of 3-4 small granules paralleling extensor margin on P2-4; flexor distal margins with very small distal spine. Propodi subequal in length on P2-4, each 4.5-5.0 times as long as broad; extensor margin with one or two proximal spines on P2-3, unarmed on P4; flexor margin with 6-8 spines, terminal one paired with another smaller spine mesial to it. Dactyli subequal in length, 0.6 length of propodi, ending in incurved, strong, sharp spine; flexor margin with five or six proximally diminishing teeth, terminal one prominent; each tooth with seta-like movable spine.</paragraph>
<paragraph id="8B923E3FFF88FFF15CF64D3D3C7C916A" blockId="7.[675,1209,216,337]" box="[699,1024,311,337]" pageId="7" pageNumber="416">Epipods absent on pereopods.</paragraph>
</subSubSection>
<subSubSection id="C3376DB4FF88FFF15CE84D7D3CC69411" pageId="7" pageNumber="416" type="discussion">
<paragraph id="8B923E3FFF88FFF15CE84D7D3B7591B4" blockId="7.[674,1212,375,1066]" box="[677,777,375,401]" pageId="7" pageNumber="416">REMARKS</paragraph>
<paragraph id="8B923E3FFF88FFF15CE84D9C3C279337" blockId="7.[674,1212,375,1066]" pageId="7" pageNumber="416">
<taxonomicName id="4C2D45BCFF88FFF15CE84D9C3B19918B" authority="Macpherson, 2012" authorityName="Macpherson" authorityYear="2012" box="[677,869,406,432]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="echinata" status="sp. nov.">
<emphasis id="B959E22DFF88FFF15CE84D9C3B19918B" box="[677,869,406,432]" italics="true" pageId="7" pageNumber="416">Galathea echinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A5F56FF88FFF15D234D9C3BD7918B" box="[878,939,406,432]" pageId="7" pageNumber="416" rank="species">n. sp.</taxonomicNameLabel>
belongs to the group of species having an interrupted ridge between the anteriormost branchial marginal spines directly behind the anterior cervical groove, the rostrum with four lateral spines, the carapace lateral margin with a small spine between the anterolateral and anteriormost branchial marginal spines, gastric striae not scale-like, and the antennular basal article with two spines. The closest relative is
<taxonomicName id="4C2D45BCFF88FFF15A044E9E3B5892F6" authority="Baba, 1990" authorityName="Baba" authorityYear="1990" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B959E22DFF88FFF15A044E9E3CCB9296" box="[1097,1207,659,686]" italics="true" pageId="7" pageNumber="416">G. robusta</emphasis>
Baba, 1990
</taxonomicName>
from
<collectingCountry id="F33A7EAFFF88FFF15D264EB93B9092F6" box="[875,1004,691,717]" name="Madagascar" pageId="7" pageNumber="416">Madagascar</collectingCountry>
(see above for the comparative material used), from which it is easily distinguished by the following (
<tableCitation id="C6AF0B84FF88FFF15DB44EF83C369337" box="[1017,1098,754,780]" captionStart="TABLE" captionStartId="5.[128,139,219,236]" captionText="TABLE 1. — Main differences among Galathea barbellata n. sp., G. echinata n. sp., G. profunda n. sp. and G. robusta Baba, 1990." pageId="7" pageNumber="416">Table 1</tableCitation>
):
</paragraph>
<paragraph id="8B923E3FFF88FFF15CE84F193BC193F0" blockId="7.[674,1212,375,1066]" pageId="7" pageNumber="416">
the branchial margin of the carapace has three strong spines instead of five spines of moderate size; the ridges on gastric and anterior branchial regions are scale-like and not well defined in the new species, whereas some of these ridges are uninterrupted and well defined in
<taxonomicName id="4C2D45BCFF88FFF15D044FBB3BC993F1" authorityName="Baba" authorityYear="1990" box="[841,949,944,971]" class="Malacostraca" family="Galatheidae" genus="Galathea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="416" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="B959E22DFF88FFF15D044FBB3BC993F1" box="[841,949,944,971]" italics="true" pageId="7" pageNumber="416">G. robusta</emphasis>
</taxonomicName>
;
</paragraph>
<paragraph id="8B923E3FFF88FFF15CE84FDB3CC69411" blockId="7.[674,1212,375,1066]" pageId="7" pageNumber="416"> the flexor margin of the Mxp3 merus bears a strong spine at midlength, instead of three spines of moderated size (proximal and median subequal).</paragraph>
</subSubSection>
</treatment>
</document>