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<document id="F295F75878E7D4C6125D7B9EFE653B34" ID-CLB-Dataset="68513" ID-DOI="10.5281/zenodo.6625539" ID-GBIF-Dataset="0a6d8930-783d-44bc-a461-7a9aaefaeff3" ID-ISBN="978-84-941892-3-4" ID-Zenodo-Dep="6625539" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1654714721633" checkinUser="carolina" docAuthor="Don E. Wilson, Thomas E. Lacher, Jr &amp; Russell A. Mittermeier" docDate="2016" docId="03822308B744FFFDFA15FBB9FB68F407" docLanguage="en" docName="hbmw_6_Leporidae_0062.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I, Barcelona: Lynx Edicions" docTitle="Pronolagus rupestris" docType="treatment" docVersion="10" lastPageNumber="111" masterDocId="FFBB5B70B747FFF9FFABFFBBFFCBF65E" masterDocTitle="Leporidae" masterLastPageNumber="148" masterPageNumber="107" pageNumber="110" updateTime="1699339205497" updateUser="ExternalLinkService">
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<mods:title id="6D36BCDADB0A009F49B69B5BBBF6A4FE">Leporidae</mods:title>
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<mods:namePart id="59CD850128B05BB52315468FC2F2957A">Don E. Wilson</mods:namePart>
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<mods:namePart id="37F032F594FB4DFBF8574C2EA1569A0D">Thomas E. Lacher, Jr</mods:namePart>
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<mods:namePart id="2EF4762BDB3DA31B199B538C173DAE36">Russell A. Mittermeier</mods:namePart>
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<mods:title id="3BFF7D1212984C65D119693D1542015A">Handbook of the Mammals of the World Volume 6 Lagomorphs and Rodents I</mods:title>
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<paragraph id="8B94921EB744FFFAFA15FBB9FA15F26E" blockId="3.[1468,2439,1026,1115]" box="[1470,1502,1026,1072]" pageId="3" pageNumber="110">
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<figureCitation id="13108E9BB744FFFAFA15FBB9FA15F26E" box="[1470,1502,1026,1072]" captionStart="Plate 5: Leporidae" captionStartId="22.[106,136,3348,3373]" captionTargetBox="[12,2744,14,3642]" captionTargetPageId="21" captionText="33. Indian Hare (Lepus nigricollis), 34. Burmese Hare (Lepus peguensis), 36. Scrub Hare (Lepus saxatilis), 37. African Savanna Hare (Lepus victoriae), 38. Yarkand Hare (Lepus yarkandensis), 39. Ethiopian Highland Hare (Lepus starcki), 40. Cape Hare (Lepus capensis), 41. Abyssinian Hare (Lepus habessinicus), 42. Tolai Hare (Lepus tolai), 43. Desert Hare (Lepus tibetanus), 44. Yunnan Hare (Lepus comus), 45. Woolly Hare (Lepus oiostolus), 46. Snowshoe Hare (Lepus americanus), 47. Black-tailed Jackrabbit (Lepus californicus), 48. Black Jackrabbit (Lepus insularis)" figureDoi="http://doi.org/10.5281/zenodo.6625695" httpUri="https://zenodo.org/record/6625695/files/figure.png" pageId="3" pageNumber="110">4.</figureCitation>
</heading>
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<subSubSection id="C331C195B744FFFAFA44FBB9F818F26E" box="[1519,2003,1026,1072]" pageId="3" pageNumber="110" type="vernacular_names">
<paragraph id="8B94921EB744FFFAFA44FBB9F818F26E" blockId="3.[1468,2439,1026,1115]" box="[1519,2003,1026,1072]" pageId="3" pageNumber="110">
<heading id="D0DC2572B744FFFAFA44FBB9F818F26E" box="[1519,2003,1026,1072]" pageId="3" pageNumber="110">
<vernacularName id="0528E230B744FFFAFA44FBB9F818F26E" box="[1519,2003,1026,1072]" pageId="3" pageNumber="110">Smiths Red Rock Hare</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAF7B0FBB9F64DF26E" box="[2075,2438,1026,1072]" pageId="3" pageNumber="110" type="nomenclature">
<paragraph id="8B94921EB744FFFAF7B0FBB9F64DF26E" blockId="3.[1468,2439,1026,1115]" box="[2075,2438,1026,1072]" pageId="3" pageNumber="110">
<heading id="D0DC2572B744FFFAF7B0FBB9F64DF26E" box="[2075,2438,1026,1072]" pageId="3" pageNumber="110">
<taxonomicName id="4C2BE99DB744FFFAF7B0FBB9F64DF26E" ID-CoL="4MT9Z" baseAuthorityName="A. Smith" baseAuthorityYear="1834" box="[2075,2438,1026,1072]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="rupestris">
<emphasis id="B95F4E0CB744FFFAF7B0FBB9F64DF26E" box="[2075,2438,1026,1072]" italics="true" pageId="3" pageNumber="110">Pronolagus rupestris</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAFA16FBF9F723F209" box="[1469,2280,1090,1111]" pageId="3" pageNumber="110" type="vernacular_names">
<paragraph id="8B94921EB744FFFAFA16FBF9F723F209" blockId="3.[1468,2439,1026,1115]" box="[1469,2280,1090,1111]" pageId="3" pageNumber="110">
<heading id="D0DC2572B744FFFAFA16FBF9F723F209" box="[1469,2280,1090,1111]" pageId="3" pageNumber="110">
<emphasis id="B95F4E0CB744FFFAFA16FBF9F9C3F209" bold="true" box="[1469,1544,1090,1111]" pageId="3" pageNumber="110">French:</emphasis>
<vernacularName id="0528E230B744FFFAF9B9FBF9F96DF209" box="[1554,1702,1090,1111]" pageId="3" pageNumber="110">Lapin de Smith</vernacularName>
/
<emphasis id="B95F4E0CB744FFFAF917FBF9F8DDF209" bold="true" box="[1724,1814,1090,1111]" pageId="3" pageNumber="110">German:</emphasis>
<vernacularName id="0528E230B744FFFAF88AFBF9F86EF209" box="[1825,1957,1090,1111]" pageId="3" pageNumber="110">Rotkaninchen</vernacularName>
/
<emphasis id="B95F4E0CB744FFFAF810FBF9F7DDF209" bold="true" box="[1979,2070,1090,1111]" pageId="3" pageNumber="110">Spanish:</emphasis>
<vernacularName id="0528E230B744FFFAF78BFBF9F723F209" box="[2080,2280,1090,1111]" pageId="3" pageNumber="110">Liebre roja de Smith</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAF7B5FB3DF5A5F2F9" box="[2078,2670,1158,1191]" pageId="3" pageNumber="110" type="reference_group">
<paragraph id="8B94921EB744FFFAF7B5FB3DF5A5F2F9" blockId="3.[2078,2674,1158,1585]" box="[2078,2670,1158,1191]" pageId="3" pageNumber="110">
<emphasis id="B95F4E0CB744FFFAF7B5FB3DF772F2F9" bold="true" box="[2078,2233,1158,1191]" pageId="3" pageNumber="110">Taxonomy.</emphasis>
<taxonomicName id="4C2BE99DB744FFFAF763FB3DF5A2F2F9" ID-CoL="6PPQ5" authorityName="A. Smith" authorityYear="1834" box="[2248,2665,1158,1191]" class="Mammalia" family="Leporidae" genus="Lepus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="rupestris">Lepus rupestris A. Smith, 1834</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAF7B4FB0DF63CF31A" pageId="3" pageNumber="110" type="materials_examined">
<paragraph id="8B94921EB744FFFAF7B4FB0DF63CF31A" blockId="3.[2078,2674,1158,1585]" pageId="3" pageNumber="110">
<materialsCitation id="3B439843B744FFFAF7B4FB0DF63CF31A" ID-GBIF-Occurrence="3866276301" pageId="3" pageNumber="110">“South Africa, rocky situations.” Refined by R. S. Hoffman and A. T. Smith in 2005 to “probably Van Rhynsdorp District, Western Cape Province, South Africa.”</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAF7B5FAF0F997F134" pageId="3" pageNumber="110" type="discussion">
<paragraph id="8B94921EB744FFFAF7B5FAF0F997F134" blockId="3.[2078,2674,1158,1585]" lastBlockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">
Formerly,
<taxonomicName id="4C2BE99DB744FFFAF715FAF0F691F332" baseAuthorityName="A. Smith" baseAuthorityYear="1834" box="[2238,2394,1355,1388]" class="Mammalia" family="Leporidae" genus="Lepus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="rupestris">P. rupestris</taxonomicName>
was included in
<taxonomicName id="4C2BE99DB744FFFAF7B5FACCF731F3CA" baseAuthorityName="I. Geoffroy Saint-Hilaire" baseAuthorityYear="1832" box="[2078,2298,1399,1428]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="crassicaudatus">P. crassicaudatus</taxonomicName>
.
<taxonomicName id="4C2BE99DB744FFFAF6A1FACCF5F2F3CA" authorityName="Hewitt" authorityYear="1927" box="[2314,2617,1399,1428]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="saundersiae">Pronolagus saundersiae</taxonomicName>
was until recently included as a subspecies of
<taxonomicName id="4C2BE99DB744FFFAF7B5FA7AF760F3BC" baseAuthorityName="A. Smith" baseAuthorityYear="1834" box="[2078,2219,1473,1506]" class="Mammalia" family="Leporidae" genus="Lepus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="rupestris">P. rupestris</taxonomicName>
but now has species status. Systematic position of the two widely disjunct populations needs to be clarified because probably no gene flow occurs between them. Distribution of
<taxonomicName id="4C2BE99DB744FFFAF6BCF98DF669F009" baseAuthorityName="A. Smith" baseAuthorityYear="1834" box="[2327,2466,1590,1623]" class="Mammalia" family="Leporidae" genus="Lepus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="rupestris">P. rupestris</taxonomicName>
does not overlap those of
<taxonomicName id="4C2BE99DB744FFFAF9CDF9DDF8CBF021" authorityName="Jameson" authorityYear="1907" box="[1638,1792,1638,1663]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="randensis">P. randensis</taxonomicName>
,
<taxonomicName id="4C2BE99DB744FFFAF8A4F9DDF80FF021" authorityName="Hewitt" authorityYear="1927" box="[1807,1988,1638,1663]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="saundersiae">P. saundersiae</taxonomicName>
, and
<taxonomicName id="4C2BE99DB744FFFAF7BAF9DDF727F021" baseAuthorityName="I. Geoffroy Saint-Hilaire" baseAuthorityYear="1832" box="[2065,2284,1638,1663]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="species" species="crassicaudatus">P. crassicaudatus</taxonomicName>
. As taxonomists are still trying to clarify the species differentiation in
<taxonomicName id="4C2BE99DB744FFFAF7B8F93EF76AF0F8" authorityName="Lyon" authorityYear="1904" box="[2067,2209,1669,1702]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="genus">Pronolagus</taxonomicName>
, the subspecific taxonomyis not elaborated yet. The original descriptions of the subspecies are often not very helpful as they are mostly based on few exterior characteristics and small numbers of individuals. It has been shown that the variability is clinal in more careful investigations. Hence, the distinction in subspecies might be arbitrary and unreasonable. Four subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAFA11F8CAF70EFE21" pageId="3" pageNumber="110" type="distribution">
<caption id="DF54C296B744FFFAFA11F8CAF70EFE21" ID-DOI="http://doi.org/10.5281/zenodo.6625551" ID-Zenodo-Dep="6625551" httpUri="https://zenodo.org/record/6625551/files/figure.png" inLine="true" pageId="3" pageNumber="110" targetBox="[1463,2053,1163,1576]" targetPageId="3">
<paragraph id="8B94921EB744FFFAFA11F8CAF881F1CC" blockId="3.[1461,2671,1590,3474]" box="[1466,1866,1905,1938]" pageId="3" pageNumber="110">
<emphasis id="B95F4E0CB744FFFAFA11F8CAF881F1CC" bold="true" box="[1466,1866,1905,1938]" pageId="3" pageNumber="110">Subspecies and Distribution.</emphasis>
</paragraph>
<paragraph id="8B94921EB744FFFAFA11F822F5EBFE57" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">
<taxonomicName id="4C2BE99DB744FFFAFA11F822F5EBFE57" authority="A. Smith, 1834" authorityName="A. Smith" authorityYear="1834" baseAuthorityName="A. Smith" baseAuthorityYear="1834" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="subSpecies" species="rupestris" subSpecies="rupestris">P. r. rupestrisA. Smith, 1834 — N &amp; NWSouthAfrica (NorthWestandNorthernCapeprovinces); recently, aspecimenhasbeensampledinSNamibia (Keetmanshoop), sothedistributioncouldbecontiguousfromSouthAfricaacrosstheOrangeRiver.</taxonomicName>
</paragraph>
<paragraph id="8B94921EB744FFFAFA11F7A8F6F1FE6E" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">
<taxonomicName id="4C2BE99DB744FFFAFA11F7A8F6F1FE6E" authority="Thomas, 1902" authorityName="Thomas" authorityYear="1902" baseAuthorityName="Thomas" baseAuthorityYear="1902" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="subSpecies" species="rupestris" subSpecies="curryi">P. r. curryiThomas, 1902 — NSouthAfrica (FreeStateP. r.).</taxonomicName>
</paragraph>
<paragraph id="8B94921EB744FFFAFA11F78CF758FE06" blockId="3.[1461,2671,1590,3474]" box="[1466,2195,2103,2136]" pageId="3" pageNumber="110">
<taxonomicName id="4C2BE99DB744FFFAFA11F78CF758FE06" authority="Thomas, 1902" authorityName="Thomas" authorityYear="1902" box="[1466,2195,2103,2136]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="subSpecies" species="rupestris" subSpecies="nytkae">P. r. nytkaeThomas, 1902 — EZambia, andNMalawi.</taxonomicName>
</paragraph>
<paragraph id="8B94921EB744FFFAFA11F7DDF70EFE21" blockId="3.[1461,2671,1590,3474]" box="[1466,2245,2150,2175]" pageId="3" pageNumber="110">
<taxonomicName id="4C2BE99DB744FFFAFA11F7DDF8E6FE21" authority="Kershaw, 1924" authorityName="Kershaw" authorityYear="1924" box="[1466,1837,2150,2175]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="110" phylum="Chordata" rank="subSpecies" species="rupestris" subSpecies="vallicola">P. r. vallicola Kershaw, 1924</taxonomicName>
— SW Kenya and C Tanzania.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C331C195B744FFFAFA12F735F83EFFE4" pageId="3" pageNumber="110" type="description">
<paragraph id="8B94921EB744FFFAFA12F735F83EFFE4" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">Descriptive notes. Head-body 380-540 mm, tail 50-115 mm, ear 80-110 mm, hindfoot 85-100 mm; weight 1.4-2.1 kg. Smiths Red Rock Hare is medium-sized and has thick, dense, woolly fur with reddish undertone. Dorsal pelage is grizzled brown anteriorly, rufous posteriorly, and bright rufous on rump. Flanks are paler, and ventral fur is rufous to whitish rufous. Head and ears are grayish brown, and cheeks are grayish white. Throat patch is brownish, and nuchal patch is rufous. Forelimbs are bright rufous, and hindlimbs are pale rufous. Tail is medium-sized and either black or black and dark red. Fur color varies geographically.</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAFA13F67AF604FC06" pageId="3" pageNumber="110" type="biology_ecology">
<paragraph id="8B94921EB744FFFAFA13F67AF604FC06" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">
Habitat. Solid rocks (“kopjes”) and rockyhillsides with boulders with grassy or scrubby vegetation. Rock crevices and boulders are essential habitat features, and gaps between them naturally fragment populations. Smiths Red Rock Hare is generally found at lower elevations in South Africa than Hewitts Red Rock Hare (
<taxonomicName id="4C2BE99DB744FFFAF69BF580F609FC06" baseAuthorityName="A. Smith" baseAuthorityYear="1834" box="[2352,2498,2619,2648]" class="Mammalia" family="Leporidae" genus="Pronolagus" kingdom="Animalia" order="Lagomorpha" pageId="3" pageNumber="111" phylum="Chordata" rank="species" species="rupestris">P. rupestris</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAFA13F5E5F6E5FD1A" pageId="3" pageNumber="110" type="food_feeding">
<paragraph id="8B94921EB744FFFAFA13F5E5F6E5FD1A" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">
<emphasis id="B95F4E0CB744FFFAFA13F5E5F90BFC21" bold="true" box="[1464,1728,2654,2687]" pageId="3" pageNumber="110">Food and Feeding.</emphasis>
Smiths Red Rock Hare is herbivorous. It forages near rocks, and its diet consists mainly of grasses, preferably fresh sprouts. One study offecal pellets from the Ngong Hills, Kenya, showed that
<taxonomicName id="4C2BE99DB744FFFAF788F516F77AFC90" box="[2083,2225,2733,2766]" class="Liliopsida" family="Poaceae" genus="Ischaemum" kingdom="Plantae" order="Poales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="genus">Ischaemum</taxonomicName>
afrum (
<taxonomicName id="4C2BE99DB744FFFAF689F516F65CFC90" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[2338,2455,2733,2766]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="3" pageNumber="110" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) was the main grass in the diet. A small amount of other grass species and a very small amount of dicotyledons also were recorded in the diet. Smiths Red Rock Hares avoid areas of dense vegetative cover and moribund vegetation when feeding.
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFAFA1CF4F0F667FB42" pageId="3" pageNumber="110" type="biology_ecology">
<paragraph id="8B94921EB744FFFAFA1CF4F0F94CFA71" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">Breeding. More than one male Smiths Red Rock Hare might accompany a female during the breeding season. Females give birth to young in nests lined with their fur and grass in small cavities in the ground. Young are likely altricial, with sparse covering of hair and eyes closed. Reproductive season is from spring to summer (September-February) in South Africa. Gestation lasts 35-45 days, and litter sizes are 1-2 young.</paragraph>
<paragraph id="8B94921EB744FFFAFA1EF38CF804FA20" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">Activity patterns. There is no specific information available for this species, but Smiths Red Rock Hare is no doubt nocturnal.</paragraph>
<paragraph id="8B94921EB744FFFAFA1DF33EF667FB42" blockId="3.[1461,2671,1590,3474]" pageId="3" pageNumber="110">
<emphasis id="B95F4E0CB744FFFAFA1DF33EF7BCFAF8" bold="true" box="[1462,2167,3205,3238]" pageId="3" pageNumber="110">Movements, Home range and Social organization.</emphasis>
Smiths Red Rock Hare has a wide variety of vocalizations. It emits a “tu...tu” sound when alarmed, butit grunts when it is disturbed. Young make “churring” sounds when trapped under rocks. Characteristic disc-like feces are deposited in latrines that might have a social function.
</paragraph>
</subSubSection>
<subSubSection id="C331C195B744FFFDFA1DF298FE08F7BD" lastPageId="4" lastPageNumber="111" pageId="3" pageNumber="110" type="conservation">
<paragraph id="8B94921EB744FFFDFA1DF298FE08F7BD" blockId="3.[1461,2671,1590,3474]" lastBlockId="4.[105,1312,292,604]" lastPageId="4" lastPageNumber="111" pageId="3" pageNumber="110">
<emphasis id="B95F4E0CB744FFFAFA1DF298F8D2FB1A" bold="true" box="[1462,1817,3363,3396]" pageId="3" pageNumber="110">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Smiths Red Rock Hare is widespread. In South Africa, overall population is greater than 10,000 adults. A population decline of more than 10% is predicted to occur by 2022 in South Africa. No data are available on the status of the East African population. Unregulated hunting poses a conservation threat. Habitat loss from construction of human dwellings is a secondary threat. Twenty percent of the habitat of Smiths Red Rock Hare has been lost since the 1900s, and future loss is predicted to continue at the same rate until 2022.
</paragraph>
</subSubSection>
<subSubSection id="C331C195B743FFFDFFC0FE4AFB68F407" pageId="4" pageNumber="111" type="reference_group">
<paragraph id="8B94921EB743FFFDFFC0FE4AFB68F407" blockId="4.[105,1312,292,604]" pageId="4" pageNumber="111">Bibliography. Angermann (2016), Boitani et al. (1999), Duthie (1997), Duthie &amp; Robinson (1990), Happold (2013c¢), Hoffmann &amp; Smith (2005), Lissovsky (2016), Matthee (2015), Matthee &amp; Robinson (1996), Matthee, Collins &amp; Keith (2004b), Smith &amp; Boyer (2008a), Smithers (1983), Stewart (1971b), Thomas (1902c).</paragraph>
</subSubSection>
</treatment>
</document>