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<mods:title id="328C8F62BD896ECF96DF37E78F944124">Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)</mods:title>
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<mods:namePart id="E7CC8D132EAD558A115A27A4FF33829B">Lauren A. Esposito</mods:namePart>
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<mods:namePart id="EB489F0593B6C5950327884B24507D53">Humberto Y. Yamaguti</mods:namePart>
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<mods:namePart id="1305935FDB6F952C50E601A5AA74BA03">Cláudio A. Souza</mods:namePart>
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<mods:namePart id="0FE1D6A7AF37F3DD8858982565D9C4CC">Ricardo Pinto-Da-Rocha</mods:namePart>
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<mods:namePart id="3D43CE9396FD8B62B931AA4B759041B0">Lorenzo Prendini</mods:namePart>
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<treatment id="8F65ED57FFBDB12C38F3C8DFB7796C7C" ID-DOI="http://doi.org/10.5281/zenodo.4610716" ID-GBIF-Taxon="132953584" ID-Zenodo-Dep="4610716" LSID="urn:lsid:plazi:treatment:8F65ED57FFBDB12C38F3C8DFB7796C7C" httpUri="http://treatment.plazi.org/id/8F65ED57FFBDB12C38F3C8DFB7796C7C" lastPageId="120" lastPageNumber="120" pageId="119" pageNumber="119">
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<paragraph id="07735C41FFBDB12338F3C8DFB16B6B58" blockId="119.[147,596,1267,1325]" pageId="119" pageNumber="119">
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<taxonomicName id="C0CC27C2FFBDB12338F3C8DFB2F86B7E" ID-CoL="5995K" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[147,420,1267,1291]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB12338F3C8DFB2F86B7E" bold="true" box="[147,420,1267,1291]" italics="true" pageId="119" pageNumber="119">Troglorhopalurus lacrau</emphasis>
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(
<bibRefCitation id="635D21B0FFBDB12339D3C8D8B2F76B58" author="Lourenco and Pinto-da-Rocha" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="119" pageNumber="119" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997</bibRefCitation>
),
</emphasis>
comb. nov.
</paragraph>
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<subSubSection id="4FD60FCAFFBDB1233811C96BB1766BF4" pageId="119" pageNumber="119" type="description">
<paragraph id="07735C41FFBDB1233811C96BB12A6B15" blockId="119.[113,630,1351,1409]" box="[113,630,1351,1376]" pageId="119" pageNumber="119">
<figureCitation id="9FF740C4FFBDB1233811C96BB3B66B15" box="[113,234,1351,1376]" captionStart="FIG" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="119" pageNumber="119">
<figureCitation id="9FF740C4FFBDB1233811C96BB38E6B15" box="[113,210,1351,1376]" captionStart="FIG. 1" captionStartId="16.[107,147,1517,1538]" captionTargetBox="[188,1135,224,1487]" captionTargetId="figure@16.[142,1179,224,1487]" captionTargetPageId="16" captionText="FIG. 1. Representative species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971, habitus in life. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂, Isla Mona, Puerto Rico. B. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂, Guanambí, Bahía, Brazil. C. Jaguajir agamemnon (C. L. Koch, 1839), comb. nov., ♀, Parque Zoobotânico de Teresina, Piauí, Brazil. D. Jaguajir rochae (Borelli, 1910), comb. nov., ♂, Jeremoabo, Bahía, Brazil. E. Physoctonus debilis, ♀, Guanambí, Bahía, Brazil. F. Rhopalurus laticauda Thorell, 1876., ♀, Alter do Chão, Santarem, Pará, Brazil. G. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀, Lapa do Bode, Itaeté, Bahía, Brazil. H. Troglorhopalurus translucidus Lourenço et al., 2004, ♀, Gruta Rio dos Pombos, Igatu, Andaraí, Bahía, Brazil." httpUri="https://zenodo.org/record/851384/files/figure.png" pageId="119" pageNumber="119">Figures 1</figureCitation>
G
</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233896C964B2456B15" box="[246,281,1352,1376]" captionStart="FIG" captionStartId="17.[111,153,1486,1507]" captionTargetBox="[155,1166,224,1455]" captionTargetId="figure@17.[111,1214,224,1455]" captionTargetPageId="17" captionText="FIG. 2. Representative habitats of species in the New World buthid scorpion family Rhopalurusinae Bücherl, 1971. A. Isla Mona, Puerto Rico, habitat of Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov. B. Sierra de Cubitas, Camagüey, Cuba, habitat of Heteroctenus junceus (Herbst, 1800), comb. nov. C. Guanambí, Bahía, Brazil, habitat of Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., and Jaguajir rochae (Borelli, 1910), comb. nov. D. Exu, Pernambuco, Brazil, habitat of J. rochae and Physoctonus debilis (C.L. Koch, 1840). E. Santarém, Pará, Brazil, habitat of Rhopalurus laticauda Thorell, 1876. F. Normandia, Roraima, Brazil, habitat of R. laticauda. G. Lapa do Bode, Itaeté, Bahía, Brazil, habitat of Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov. H. Gruta do Lapão, Lençóis, Bahía, Brazil, habitat of Troglorhopalurus translucidus Lourenço et al., 2004." httpUri="https://zenodo.org/record/851386/files/figure.png" pageId="119" pageNumber="119">2G</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233944C964B26F6B15" box="[292,307,1352,1376]" captionStart="FIG" captionStartId="24.[107,149,1416,1437]" captionTargetBox="[138,1207,234,1374]" captionTargetId="figure@24.[107,1211,224,1383]" captionTargetPageId="24" captionText="FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C.L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses)." httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="119" pageNumber="119">9</figureCitation>
,
<figureCitation id="9FF740C4FFBDB123395FC964B2376B15" box="[319,363,1352,1376]" captionStart="FIG" captionStartId="26.[109,151,1396,1417]" captionTargetBox="[109,1213,224,1368]" captionTargetId="figure@26.[109,1213,224,1368]" captionTargetPageId="26" captionText="FIG. 10. Rhopalurusinae Bücherl, 1971, sternites III (AC, E, F), V and VI (D), ventral aspect. A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). B. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (MZSP). C, D. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). E. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 1 mm (A, E, F), 2 mm (B, C, D)." httpUri="https://zenodo.org/record/851402/files/figure.png" pageId="119" pageNumber="119">10E</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233916C964B2C26B15" box="[374,414,1352,1376]" captionStart="FIG" captionStartId="28.[108,150,1167,1188]" captionTargetBox="[108,1213,224,1136]" captionTargetId="figure@28.[108,1213,224,1136]" captionTargetPageId="28" captionText="FIG. 11. Rhopalurusinae Bücherl, 1971, dextral pecten, ventral aspect (A, C, E, G) and pectinal teeth, dorsal aspect (B, D, F). A. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♀ (AMNH). C. Rhopalurus laticauda Thorell, 1876, ♂ (MZSP) D. Jaguajir rochae (Borelli, 1910), comb. nov., (AMNH). E. Rhopalurus laticauda Thorell, 1876, ♂ (AMCC [LP 2845]). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (AMCC [LP 3260]). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851404/files/figure.png" pageId="119" pageNumber="119">11F</figureCitation>
,
<figureCitation id="9FF740C4FFBDB12339C9C964B2846B15" box="[425,472,1352,1376]" captionStart="FIG" captionStartId="29.[107,149,984,1005]" captionTargetBox="[107,1211,224,956]" captionTargetId="figure@29.[107,1211,224,956]" captionTargetPageId="29" captionText="FIG. 12. Rhopalurusinae Bücherl, 1971, pectinal teeth, dorsal aspect, fine structure. A. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., (MZSP). B. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). C. Rhopalurus laticauda Thorell, 1876, (MZSP). D. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha 1997), comb. nov., ♀ (MZSP). E. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). F. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP)." httpUri="https://zenodo.org/record/851406/files/figure.png" pageId="119" pageNumber="119">12D</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233984C964B1506B15" box="[484,524,1352,1376]" captionStart="FIG" captionStartId="32.[107,149,1598,1619]" captionTargetBox="[220,1108,224,1568]" captionTargetId="figure@32.[118,1207,224,1568]" captionTargetPageId="32" captionText="FIG. 14. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus garridoi (Armas, 1974), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851410/files/figure.png" pageId="119" pageNumber="119">14F</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233A77C964B1626B15" box="[535,574,1352,1376]" captionStart="FIG" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="119" pageNumber="119">17F</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233A2AC964B12D6B15" box="[586,625,1352,1376]" captionStart="FIG" captionStartId="38.[107,149,1554,1575]" captionTargetBox="[108,1212,224,1525]" captionTargetId="figure@38.[108,1212,224,1526]" captionTargetPageId="38" captionText="FIG. 18. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus bonettii (Armas, 1999), comb. nov., ♂ (AMNH). C. Heteroctenus junceus (Herbst, 1800), comb. nov., ♂ (AMNH). D. Heteroctenus junceus (Herbst, 1800), ♀ (AMNH). E. Heteroctenus princeps (Karsch, 1879), comb. nov., ♂ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851418/files/figure.png" pageId="119" pageNumber="119">18F</figureCitation>
,
</paragraph>
<paragraph id="07735C41FFBDB12338DCC945B1766BF4" blockId="119.[113,630,1351,1409]" box="[188,554,1385,1409]" pageId="119" pageNumber="119">
<figureCitation id="9FF740C4FFBDB12338DCC945B3B76BF4" box="[188,235,1385,1409]" captionStart="FIG" captionStartId="44.[107,149,1352,1373]" captionTargetBox="[108,1212,224,1323]" captionTargetId="figure@44.[108,1212,224,1323]" captionTargetPageId="44" captionText="FIG. 21. Rhopalurusinae Bücherl, 1971, telson, lateral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851424/files/figure.png" pageId="119" pageNumber="119">21K</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233896C945B2796BF4" box="[246,293,1385,1409]" captionStart="FIG" captionStartId="45.[108,147,1513,1534]" captionTargetBox="[176,1145,224,1484]" captionTargetId="figure@45.[108,1212,224,1484]" captionTargetPageId="45" captionText="FIG. 22. Rhopalurusinae Bücherl, 1971, telson, ventral aspect. A. Heteroctenus abudi (Armas and Marcano Fondeur, 1987), comb. nov., ♂ (AMNH). B. Heteroctenus junceus (Herbst, 1800), ♂ (AMNH). C. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov., ♂ (MZSP). D. Jaguajir agamemnon (C.L. Koch, 1839), comb. nov., ♀ (AMNH). E. Jaguajir pintoi (Mello-Leitão, 1932), comb. nov., ♂ (MZSP). F. Jaguajir rochae (Borelli, 1910), comb. nov., ♀ (AMNH). G. Physoctonus debilis (C.L. Koch, 1840), ♀ (AMNH). H. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). I. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH). J. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). K. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). L. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851426/files/figure.png" pageId="119" pageNumber="119">22K</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233950C945B2036BF4" box="[304,351,1385,1409]" captionStart="FIG" captionStartId="114.[108,150,859,880]" captionTargetBox="[108,1212,224,828]" captionTargetId="figure@114.[108,1212,224,828]" captionTargetPageId="114" captionText="FIG. 61. Troglorhopalurus Lourenço et al., 2004, pedipalp chela, retrolateral aspect. A. T. lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). B. Troglorhopalurus translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851504/files/figure.png" pageId="119" pageNumber="119">61A</figureCitation>
,
<figureCitation id="9FF740C4FFBDB123390BC945B2886BF4" box="[363,468,1385,1409]" captionStart="FIG" captionStartId="115.[108,150,1567,1588]" captionTargetBox="[108,1212,224,1539]" captionTargetId="figure@115.[108,1212,224,1539]" captionTargetPageId="115" captionText="FIG. 62. Troglorhopalurus Lourenço et al., 2004, metasoma and telson, dorsal (A, C), ventral (B, D) and lateral (E, F) aspects. A, B, E. T. lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). C, D, F. T. translucidus Lourenço et al., 2004, ♀ (MZSP)." httpUri="https://zenodo.org/record/851506/files/figure.png" pageId="119" pageNumber="119">62A, B, E</figureCitation>
,
<figureCitation id="9FF740C4FFBDB1233980C945B1766BF4" box="[480,554,1385,1409]" captionStart="FIG" captionStartId="117.[108,150,1678,1699]" captionTargetBox="[108,1212,224,1650]" captionTargetId="figure@117.[108,1212,224,1650]" captionTargetPageId="117" captionText="FIG. 63. Troglorhopalurus Lourenço et al., 2004, habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. T. lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (AMNH). C, D. T. translucidus Lourenço et al., 2004, ♀ (MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/851508/files/figure.png" pageId="119" pageNumber="119">63A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFBDB123380BC9B3B0216F68" pageId="119" pageNumber="119" type="reference_group">
<paragraph id="07735C41FFBDB123380BC9B3B16B6BC2" blockId="119.[107,635,1439,1741]" box="[107,567,1439,1463]" pageId="119" pageNumber="119">
<taxonomicName id="C0CC27C2FFBDB123380BC9B3B2C66BC3" authority="Lourenco" authorityName="Lourenco" box="[107,410,1439,1463]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB123380BC9B3B2706BC2" box="[107,300,1439,1463]" italics="true" pageId="119" pageNumber="119">Rhopalurus lacrau</emphasis>
Lourenço
</taxonomicName>
and Pinto-da-
</paragraph>
<paragraph id="07735C41FFBDB12338FBC9EEB23268B8" blockId="119.[107,635,1439,1741]" pageId="119" pageNumber="119">
Rocha, 1997: 182183, figs. 13, 5, 7, 9, 11, 14;
<bibRefCitation id="635D21B0FFBDB12338A2C9C8B2D06B89" author="Kovarik" bookContentInfo="175 pp." box="[194,396,1508,1532]" firstAuthor="Kovarik" journalOrPublisher="Madagaskar: Jihlava" pageId="119" pageNumber="119" refId="ref66846" refString="Kovarik, F. 1998. Stiri [Scorpions]. Madagaskar: Jihlava. 175 pp. [in Czech]" title="Stiri [Scorpions]" type="book" year="1998">Kovařík, 1998: 118</bibRefCitation>
;
<bibRefCitation id="635D21B0FFBDB12339F8C9C9B398686A" author="Fet and Lowe" editor="V. Fet &amp; W. D. Sissom" firstAuthor="Fet" journalOrPublisher="New York: New York Entomological Society" pageId="119" pageNumber="119" pagination="54 - 286" refId="ref65304" refString="Fet, V., and G. Lowe. 2000. Family Buthidae C. L. Koch, 1837. In V. Fet, W. D. Sissom, G., Lowe, and M. E. Braunwalder, Catalog of the scorpions of the world (1758 - 1998): 54 - 286. New York: New York Entomological Society." title="Family Buthidae C. L. Koch, 1837" type="book chapter" volumeTitle="G., Lowe, and M. E" year="2000">Fet and Lowe, 2000: 220</bibRefCitation>
;
<bibRefCitation id="635D21B0FFBDB12338AECA2BB2CC686A" author="Lourenco" box="[206,400,1543,1567]" firstAuthor="Lourenco" journalOrPublisher="Euscorpius" pageId="119" pageNumber="119" pagination="1 - 14" part="73" refId="ref68250" refString="Lourenco, W. R. 2008. The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae). Euscorpius 73: 1 - 14." title="The geographic pattern of distribution of the genus Rhopalurus Thorell, 1876 in the Guayana-Amazon region (Scorpiones: Buthidae)" type="journal article" year="2008">Lourenço, 2008: 3</bibRefCitation>
;
<bibRefCitation id="635D21B0FFBDB12339FCCA2BB3986837" author="Prendini" etAl="et al." firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="119" pageNumber="119" pagination="206 - 224" part="37" refId="ref70522" refString="Prendini, L., L. A. Esposito, J. Huff, and E. S. Volschenk. 2009. Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola. Journal of Arachnology 37: 206 - 224." title="Redescription of Rhopalurus abudi (Scorpiones, Buthidae), with first description of the male and first record from mainland Hispaniola" type="journal article" year="2009">Prendini et al., 2009: 222</bibRefCitation>
, 223;
<bibRefCitation id="635D21B0FFBDB1233962CA06B1476834" author="Brazil and Porto" bookContentInfo="84 pp." box="[258,539,1577,1602]" firstAuthor="Brazil" journalOrPublisher="Salvador, BA, Brazil: EDUFBA" pageId="119" pageNumber="119" refId="ref64569" refString="Brazil, T. K., and Porto, T. J. 2010. Os escorpioes. Salvador, BA, Brazil: EDUFBA. 84 pp." title="Os escorpioes" type="book" year="2010">Brazil and Porto, 2010: 50</bibRefCitation>
, 62;
<bibRefCitation id="635D21B0FFBDB12338FBCA61B2E76811" author="Porto" box="[155,443,1612,1636]" etAl="et al." firstAuthor="Porto" journalOrPublisher="Check List" pageId="119" pageNumber="119" pagination="292 - 297" part="6" refId="ref70138" refString="Porto, T. J., T. K. Brazil, and R. M. Lira-da-Silva. 2010. Scorpions, state of Bahia, northeastern Brazil. Check List 6 (2): 292 - 297." title="Scorpions, state of Bahia, northeastern Brazil" type="journal article" year="2010">Porto et al., 2010: 293295</bibRefCitation>
,
<tableCitation id="4A4E69FAFFBDB12339A6CA60B1526811" box="[454,526,1612,1636]" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="119" pageNumber="119" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
;
<bibRefCitation id="635D21B0FFBDB1233A7BCA61B21C68F2" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Acta Biologica Paranaense" pageId="119" pageNumber="119" pagination="69 - 76" part="43" refId="ref68287" refString="Lourenco, W. R. 2014. The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species. Acta Biologica Paranaense 43: 69 - 76." title="The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species" type="journal article" year="2014">Lourenço, 2014: 69</bibRefCitation>
, 7375, fig. 12;
<bibRefCitation id="635D21B0FFBDB1233988CA43B20768DF" author="Gallao and Bichuette" firstAuthor="Gallao" journalOrPublisher="Zoologia" pageId="119" pageNumber="119" pagination="1 - 13" part="33" refId="ref65887" refString="Gallao, J. E., and M. E. Bichuette. 2016. On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae). Zoologia 33 (6): 1 - 13." title="On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae)" type="journal article" year="2016">Gallão and Bichuette, 2016: 2</bibRefCitation>
, 3, 911, figs. 1, 18;
<bibRefCitation id="635D21B0FFBDB1233A55CABEB23668B8" author="Ubinski" etAl="et al." firstAuthor="Ubinski" journalOrPublisher="Cytogenetic and Genome Research" pageId="119" pageNumber="119" pagination="83 - 155" part="148" refId="ref72292" refString="Ubinski, C. V., L. S. Carvalho, and M. C. Schneider. 2016. Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil. Cytogenetic and Genome Research 148: 83 - 155. [doi: 10.1159 / 000446523]" title="Chromosome evolution in Brazilian scorpions of the genus Rhopalurus and related genera (Scorpiones: Buthidae). Abstracts of the 21 st International Chromosome Conference, July 10 - 13, Foz do Iguacu, Brazil" type="journal article" year="2016">Ubinski et al., 2016: 122</bibRefCitation>
.
</paragraph>
<paragraph id="07735C41FFBDB1233ACCCCC8B0216F68" blockId="119.[684,1173,227,286]" pageId="119" pageNumber="119">
<taxonomicName id="C0CC27C2FFBDB1233ACCCCC8B7026E8E" authority="Lourenco, 2014: 71" authorityName="Lourenco" authorityPageNumber="71" authorityYear="2014" box="[684,1118,227,252]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFBDB1233ACCCCC8B03C6E89" box="[684,864,228,252]" italics="true" pageId="119" pageNumber="119">Rhopalurus brejo</emphasis>
<bibRefCitation id="635D21B0FFBDB1233B08CCC8B7026E8E" author="Lourenco" box="[872,1118,227,252]" firstAuthor="Lourenco" journalOrPublisher="Acta Biologica Paranaense" pageId="119" pageNumber="119" pagination="69 - 76" part="43" refId="ref68287" refString="Lourenco, W. R. 2014. The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species. Acta Biologica Paranaense 43: 69 - 76." title="The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species" type="journal article" year="2014">Lourenço, 2014: 7175</bibRefCitation>
</taxonomicName>
, figs. 112; syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFBDB1233AACCD17B7EA6F8C" pageId="119" pageNumber="119" type="materials_examined">
<paragraph id="07735C41FFBDB1233AACCD17B7EA6F8C" blockId="119.[684,1214,314,1741]" pageId="119" pageNumber="119">
TYPE MATERIAL:
<emphasis id="35B88053FFBDB1233BE6CD16B0B36F27" bold="true" box="[902,1007,314,338]" pageId="119" pageNumber="119">BRAZIL:</emphasis>
<taxonomicName id="C0CC27C2FFBDB1233B95CD16B7EB6F27" box="[1013,1207,314,338]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B95CD16B7EB6F27" box="[1013,1207,314,338]" italics="true" pageId="119" pageNumber="119">Rhopalurus lacrau</emphasis>
</taxonomicName>
: Holotype ♀ (MZSP 15175),
<emphasis id="35B88053FFBDB1233B98CD70B7646F01" box="[1016,1080,348,372]" italics="true" pageId="119" pageNumber="119">Bahía</emphasis>
: Município Itaeté: Lapa do Bode (inside the cave), 12°56S 41°04W, 7.ix.1993, E. Trajluio.
<taxonomicName id="C0CC27C2FFBDB1233C60CDB2B7EB6FC3" authorityName="Lourenco" authorityYear="2014" box="[1024,1207,414,438]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFBDB1233C60CDB2B7EB6FC3" box="[1024,1207,414,438]" italics="true" pageId="119" pageNumber="119">Rhopalurus brejo</emphasis>
</taxonomicName>
: Holotype ♀ (MNHN),
<emphasis id="35B88053FFBDB1233BD7CDECB0AB6FA2" box="[951,1015,448,471]" italics="true" pageId="119" pageNumber="119">Ceará</emphasis>
: Chapada (serra) do Araripe, Brejo Grande, S of Santana, ii.1964.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFBDB1233AACCE2FB08D681D" pageId="119" pageNumber="119" type="diagnosis">
<paragraph id="07735C41FFBDB1233AACCE2FB08D681D" blockId="119.[684,1214,314,1741]" pageId="119" pageNumber="119">
DIAGNOSIS:
<taxonomicName id="C0CC27C2FFBDB1233B3ACE2FB7306C6E" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[858,1132,515,539]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B3ACE2FB7306C6E" box="[858,1132,515,539]" italics="true" pageId="119" pageNumber="119">Troglorhopalurus lacrau</emphasis>
</taxonomicName>
differs from its sister species,
<taxonomicName id="C0CC27C2FFBDB1233BDBCE09B73F6C49" box="[955,1123,548,572]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233BDBCE09B73F6C49" box="[955,1123,548,572]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
, as follows. The soma and appendages are shorter and broader in
<taxonomicName id="C0CC27C2FFBDB1233B55CE44B0C16C0A" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[821,925,615,639]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B55CE44B0C16C0A" box="[821,925,615,639]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
than
<taxonomicName id="C0CC27C2FFBDB1233B89CE44B7C86C0A" box="[1001,1172,615,639]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233B89CE44B7C86C0A" box="[1001,1172,615,639]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
, in which the soma and appendages are elongate and slender: total body length is about 40 mm in
<taxonomicName id="C0CC27C2FFBDB1233AACCEE0B0736C96" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[716,815,715,739]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233AACCEE0B0736C96" box="[716,815,715,739]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
and 60 mm in
<taxonomicName id="C0CC27C2FFBDB1233B86CEE0B7D76C96" box="[998,1163,715,739]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233B86CEE0B7D76C96" box="[998,1163,715,739]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
; the pedipalp is 4.5× longer than the carapace in
<taxonomicName id="C0CC27C2FFBDB1233CC9CEC1B1AE6D53" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233CC9CEC1B1AE6D53" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
but 6.5× longer in
<taxonomicName id="C0CC27C2FFBDB1233B80CF23B7D46D53" box="[992,1160,782,806]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233B80CF23B7D46D53" box="[992,1160,782,806]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
; the pedipalp chela manus is incrassate and slightly shorter than the chela movable finger in
<taxonomicName id="C0CC27C2FFBDB1233CC8CF7DB1AD6DFF" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233CC8CF7DB1AD6DFF" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
but very slender, the manus almost half the length of the movable finger in
<taxonomicName id="C0CC27C2FFBDB1233C58CFB8B18F6DB8" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233C58CFB8B18F6DB8" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
; the pedipalp patella is 3.8× longer than wide in
<taxonomicName id="C0CC27C2FFBDB1233B68CFFBB0356D9B" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[776,873,982,1006]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B68CFFBB0356D9B" box="[776,873,982,1006]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, but 6× longer in
<taxonomicName id="C0CC27C2FFBDB1233C58CFFBB18E6A7A" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233C58CFFBB18E6A7A" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
; metasomal segment V is 2.7× longer than wide in
<taxonomicName id="C0CC27C2FFBDB1233B67C836B0356A44" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[775,873,1049,1073]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B67C836B0356A44" box="[775,873,1049,1073]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, but 4.5× longer in
<taxonomicName id="C0CC27C2FFBDB1233C2AC836B1B96A27" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233C2AC836B1B96A27" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
. The retromedian and secondary accessory carinae of the pedipalp chela manus are weakly granular in
<taxonomicName id="C0CC27C2FFBDB1233BEDC852B0AD6AE0" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[909,1009,1149,1173]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233BEDC852B0AD6AE0" box="[909,1009,1149,1173]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, and absent in
<taxonomicName id="C0CC27C2FFBDB1233CC9C852B06A6AC3" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233CC9C852B06A6AC3" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
. Eight subrows of denticles are present in the median denticle row of the chela fixed finger of
<taxonomicName id="C0CC27C2FFBDB1233B33C8CEB0E96A8C" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[851,949,1249,1273]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B33C8CEB0E96A8C" box="[851,949,1249,1273]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, whereas nine subrows are present in
<taxonomicName id="C0CC27C2FFBDB1233B3AC92FB75E6B6E" box="[858,1026,1283,1307]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233B3AC92FB75E6B6E" box="[858,1026,1283,1307]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
. Macrosetae on the fixed and movable fingers of the pedipalp chela are shorter than the chela width in
<taxonomicName id="C0CC27C2FFBDB1233CC8C96AB1AD6B0A" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233CC8C96AB1AD6B0A" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, but longer in
<taxonomicName id="C0CC27C2FFBDB1233BC0C944B7196B0A" box="[928,1093,1383,1407]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233BC0C944B7196B0A" box="[928,1093,1383,1407]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
. Macrosetae of the telson vesicle are shorter than the subaculear tubercle in
<taxonomicName id="C0CC27C2FFBDB1233BD3C986B74B6BB4" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[947,1047,1449,1473]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233BD3C986B74B6BB4" box="[947,1047,1449,1473]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, but longer in
<taxonomicName id="C0CC27C2FFBDB1233ACCC9E0B00D6B96" box="[684,849,1483,1507]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233ACCC9E0B00D6B96" box="[684,849,1483,1507]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
. The subaculear tubercle of the telson is blunt in
<taxonomicName id="C0CC27C2FFBDB1233B1EC9C1B0BF6871" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[894,995,1516,1540]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233B1EC9C1B0BF6871" box="[894,995,1516,1540]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, but has a sharply pointed tip in
<taxonomicName id="C0CC27C2FFBDB1233B00CA22B7576853" box="[864,1035,1550,1574]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233B00CA22B7576853" box="[864,1035,1550,1574]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
. The posterior third of the aculeus is curved in
<taxonomicName id="C0CC27C2FFBDB1233C46CA1CB7D56832" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[1062,1161,1583,1607]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233C46CA1CB7D56832" box="[1062,1161,1583,1607]" italics="true" pageId="119" pageNumber="119">T. lacrau</emphasis>
</taxonomicName>
, but straight in
<taxonomicName id="C0CC27C2FFBDB1233B48CA7DB097681D" box="[808,971,1616,1640]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="translucidus">
<emphasis id="35B88053FFBDB1233B48CA7DB097681D" box="[808,971,1616,1640]" italics="true" pageId="119" pageNumber="119">T. translucidus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFBDB12C3AACCA5EB3846C99" lastPageId="120" lastPageNumber="120" pageId="119" pageNumber="119" type="distribution">
<paragraph id="07735C41FFBDB12C3AACCA5EB3846C99" blockId="119.[684,1214,314,1741]" lastBlockId="120.[107,637,228,1741]" lastPageId="120" lastPageNumber="120" pageId="119" pageNumber="119">
DISTRIBUTION:
<taxonomicName id="C0CC27C2FFBDB1233BE7CA5EB7FC68FF" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[903,1184,1650,1674]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="119" pageNumber="119" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFBDB1233BE7CA5EB7FC68FF" box="[903,1184,1650,1674]" italics="true" pageId="119" pageNumber="119">Troglorhopalurus lacrau</emphasis>
</taxonomicName>
is endemic to Brazil, and known from only two populations, in the states of Bahía and Ceará (fig. 9A). All except two specimens were collected from two connected caves, Lapa do Bode Cave and Gruta Escondida in the Município Itaeté of Bahía. A single, dead individual was collected from Bob Cave, ca. 20 km northeast (
<bibRefCitation id="635D21B0FFB2B12C3886CDA5B1426FD4" author="Gallao and Bichuette" box="[230,542,393,417]" firstAuthor="Gallao" journalOrPublisher="Zoologia" pageId="120" pageNumber="120" pagination="1 - 13" part="33" refId="ref65887" refString="Gallao, J. E., and M. E. Bichuette. 2016. On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae). Zoologia 33 (6): 1 - 13." title="On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae)" type="journal article" year="2016">Gallão and Bichuette, 2016</bibRefCitation>
), also in Município Itaeté. The second population of
<taxonomicName id="C0CC27C2FFB2B12C3A07CD87B3ED6F91" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFB2B12C3A07CD87B3ED6F91" italics="true" pageId="120" pageNumber="120">T. lacrau</emphasis>
</taxonomicName>
is represented by the type locality and only known specimen of its junior synonym,
<taxonomicName id="C0CC27C2FFB2B12C380BCE23B39F6C53" authorityName="Lourenco" authorityYear="2014" box="[107,195,526,550]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFB2B12C380BCE23B39F6C53" box="[107,195,526,550]" italics="true" pageId="120" pageNumber="120">R. brejo</emphasis>
</taxonomicName>
, in the state of Ceará. The two localities are 700 km apart but the region between them, the Serra do Espinhaço, is poorly sampled. Assuming the record from Ceará is reliable, this species may be discovered in the Serra do Espinhaço when it is more thoroughly surveyed.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFB2B12C38EBCED9B2AB6A63" pageId="120" pageNumber="120" type="biology_ecology">
<paragraph id="07735C41FFB2B12C38EBCED9B2AB6A63" blockId="120.[107,637,228,1741]" pageId="120" pageNumber="120">
ECOLOGY: Most of the known specimens of
<taxonomicName id="C0CC27C2FFB2B12C380BCF3BB3886D5B" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[107,212,790,814]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFB2B12C380BCF3BB3886D5B" box="[107,212,790,814]" italics="true" pageId="120" pageNumber="120">T. lacrau</emphasis>
</taxonomicName>
were collected under stones inside limestone caves. Despite extensive searches, no specimens were found outside the type locality, Lapa do Bode Cave (
<bibRefCitation id="635D21B0FFB2B12C3913CF55B3FA6DC7" author="Gallao and Bichuette" firstAuthor="Gallao" journalOrPublisher="Zoologia" pageId="120" pageNumber="120" pagination="1 - 13" part="33" refId="ref65887" refString="Gallao, J. E., and M. E. Bichuette. 2016. On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae). Zoologia 33 (6): 1 - 13." title="On the enigmatic troglobitic scorpion Troglorhopalurus translucidus: distribution, description of adult females, life history and comments on Rhopalurus lacrau (Scorpiones: Buthidae)" type="journal article" year="2016">Gallão and Bichuette, 2016</bibRefCitation>
), where this species has a well-established population (fig. 2G). The habitat, distribution and habitus are consistent with the troglophile ecomorphotype (
<bibRefCitation id="635D21B0FFB2B12C394DCFD2B2806A63" author="Prendini" box="[301,476,1022,1046]" firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="120" pageNumber="120" pagination="173 - 188" part="29" refId="ref70328" refString="Prendini, L. 2001 a. Further additions to the scorpion fauna of Trinidad and Tobago. Journal of Arachnology 29 (2): 173 - 188." title="Further additions to the scorpion fauna of Trinidad and Tobago" type="journal article" year="2001">Prendini, 2001</bibRefCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFB2B12C38EBC833B28668B9" pageId="120" pageNumber="120" type="discussion">
<paragraph id="07735C41FFB2B12C38EBC833B28668B9" blockId="120.[107,637,228,1741]" pageId="120" pageNumber="120">
REMARKS:
<taxonomicName id="C0CC27C2FFB2B12C3973C833B2BF6A42" authorityName="Lourenco" authorityYear="2014" box="[275,483,1055,1079]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFB2B12C3973C833B2BF6A42" box="[275,483,1055,1079]" italics="true" pageId="120" pageNumber="120">Rhopalurus brejo</emphasis>
</taxonomicName>
was poorly described in an obscure journal with distorted plates. The original description and illustrations barely permit us to determine to which genus this taxon belongs. Although the holotype and only known specimen of
<taxonomicName id="C0CC27C2FFB2B12C39CDC8E9B1516AA9" authorityName="Lourenco" authorityYear="2014" box="[429,525,1220,1244]" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFB2B12C39CDC8E9B1516AA9" box="[429,525,1220,1244]" italics="true" pageId="120" pageNumber="120">R. brejo</emphasis>
</taxonomicName>
was not directly examined during the present investigation, its identity was verified by photographs received from the MNHN (and now publicly available at https://science.mnhn.fr/institution/ mnhn/collection/rs/). Examination thereof, together with the limited data provided in the original description, leaves no doubt that
<taxonomicName id="C0CC27C2FFB2B12C3A05C980B3FE6B90" authorityName="Lourenco" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFB2B12C3A05C980B3FE6B90" italics="true" pageId="120" pageNumber="120">R. brejo</emphasis>
</taxonomicName>
is conspecific with
<taxonomicName id="C0CC27C2FFB2B12C39E9C9E1B2B16B90" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[393,493,1485,1509]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFB2B12C39E9C9E1B2B16B90" box="[393,493,1485,1509]" italics="true" pageId="120" pageNumber="120">T. lacrau</emphasis>
</taxonomicName>
.
<taxonomicName id="C0CC27C2FFB2B12C399DC9E1B3FE6873" authorityName="Lourenco" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFB2B12C399DC9E1B3FE6873" italics="true" pageId="120" pageNumber="120">Rhopalurus brejo</emphasis>
</taxonomicName>
shares with
<taxonomicName id="C0CC27C2FFB2B12C3952C9C3B2C96873" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[306,405,1518,1542]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFB2B12C3952C9C3B2C96873" box="[306,405,1518,1542]" italics="true" pageId="120" pageNumber="120">T. lacrau</emphasis>
</taxonomicName>
the presence of distinct pedipalp carinae and eight subrows of denticles in the median denticle row of the chela fixed finger. We therefore propose the following synonym:
<taxonomicName id="C0CC27C2FFB2B12C395CCA5EB3FC68DE" authority="Lourenco, 2014" authorityName="Lourenco" authorityYear="2014" class="Arachnida" family="Buthidae" genus="Rhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="brejo">
<emphasis id="35B88053FFB2B12C395CCA5EB2A368FF" box="[316,511,1650,1674]" italics="true" pageId="120" pageNumber="120">Rhopalurus brejo</emphasis>
<bibRefCitation id="635D21B0FFB2B12C3A6ACA5EB3FC68DE" author="Lourenco" firstAuthor="Lourenco" journalOrPublisher="Acta Biologica Paranaense" pageId="120" pageNumber="120" pagination="69 - 76" part="43" refId="ref68287" refString="Lourenco, W. R. 2014. The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species. Acta Biologica Paranaense 43: 69 - 76." title="The genus Rhopalurus Thorell, 1876 (Scorpiones: Buthidae) in northeast Brazil; a possible case of a vicariant species" type="journal article" year="2014">Lourenço, 2014</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="C0CC27C2FFB2B12C38A2CABFB29168DE" authorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" authorityYear="2017" baseAuthorityName="Lauren A. Esposito &amp; Humberto Y. Yamaguti &amp; Cláudio A. Souza &amp; Ricardo Pinto-Da-Rocha &amp; Lorenzo Prendini" baseAuthorityYear="2017" box="[194,461,1683,1707]" class="Arachnida" family="Buthidae" genus="Troglorhopalurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Scorpiones" pageId="120" pageNumber="120" phylum="Arthropoda" rank="species" species="lacrau">
<emphasis id="35B88053FFB2B12C38A2CABFB29168DE" box="[194,461,1683,1707]" italics="true" pageId="120" pageNumber="120">Troglorhopalurus lacrau</emphasis>
</taxonomicName>
(
<bibRefCitation id="635D21B0FFB2B12C39BDCABFB23F68B9" author="Lourenco and Pinto-da-Rocha" firstAuthor="Lourenco" journalOrPublisher="Biogeographica (Paris) 73" pageId="120" pageNumber="120" pagination="181 - 191" part="4" refId="ref68476" refString="Lourenco, W. R., and R. Pinto-da-Rocha. 1997. A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species. Biogeographica (Paris) 73 (4): 181 - 191." title="A reappraisal of the geographic distribution of the genus Rhopalurus Thorell (Scorpiones, Buthidae) and description of two new species" type="journal article" year="1997">Lourenço and Pinto-da-Rocha, 1997</bibRefCitation>
), syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="4FD60FCAFFB2B12C3AACCCC8B7796C7C" pageId="120" pageNumber="120" type="materials_examined">
<paragraph id="07735C41FFB2B12C3AACCCC8B7796C7C" blockId="120.[684,1214,228,521]" pageId="120" pageNumber="120">
MATERIAL EXAMINED:
<emphasis id="35B88053FFB2B12C3B91CCC8B73E6E89" bold="true" box="[1009,1122,228,252]" pageId="120" pageNumber="120">BRAZIL:</emphasis>
<emphasis id="35B88053FFB2B12C3C12CCC8B7EA6E89" box="[1138,1206,228,252]" italics="true" pageId="120" pageNumber="120">Bahía</emphasis>
: Município Itaeté: trail connecting caves Lapa do Bode and Gruta Escondida, 12°569.1″S 41°0356.2″W, 21.i.2007, C.I. Mattoni, R. Pinto-da-Rocha and H.Y. Yamaguti, under rocks, 2 ♀ (AMNH), 1 subad. ♀, 4 juv. (AMCC [LP 7637]); Lapa do Bode Cave, 24.i.2007, C.I. Mattoni, R. Pinto-da-Rocha and H. Yamaguti, 1 subad. ♀ (AMCC [LP 10211]).
</paragraph>
</subSubSection>
</treatment>
</document>