254 lines
33 KiB
XML
254 lines
33 KiB
XML
<document id="BEF7BBF0C4FCB3C5AB84246CE277538B" ID-CLB-Dataset="32293" ID-DOI="10.1206/0003-0090-415.1.1" ID-GBIF-Dataset="22b02e67-634f-4e71-a8fa-3392aff63985" ID-Handle="2246/6723" ID-Zenodo-Dep="851382" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1503785870183" checkinUser="admin" docAuthor="Lauren A. Esposito, Humberto Y. Yamaguti, Cláudio A. Souza, Ricardo Pinto-Da-Rocha & Lorenzo Prendini" docDate="2017" docId="8F65ED57FF94B10B38B0CD9BB12D6B41" docLanguage="en" docName="BulAmeMusNatHis.2017.415.1-63.pdf.imf" docOrigin="Bulletin of the American Museum of Natural History 415" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Physoctonus striatus Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini 2017, sp. nov." docType="treatment" docUuid="5E21CA29-855B-4708-8C0B-18A85E6B09E1" docUuidSource="ZooBank" docVersion="20" lastPageNumber="95" masterDocId="735C952FFFCAB1543860CC2CB35C6E75" masterDocTitle="Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="94" updateTime="1698467378382" updateUser="ExternalLinkService">
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<mods:title id="84A2B6661AFE1F4C6AC20941FB086FC2">Systematic Revision of the Neotropical Club-Tailed Scorpions, Physoctonus, Rhopalurus, and Troglorhopalurus, Revalidation of Heteroctenus, and Descriptions of Two New Genera and Three New Species (Buthidae: Rhopalurusinae)</mods:title>
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<mods:roleTerm id="BCC026C10080F935DF3384C68B19D7CD">Author</mods:roleTerm>
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<mods:namePart id="5AA15D24F8F9F5EDCB734EC327B5B6EE">Lauren A. Esposito</mods:namePart>
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<mods:roleTerm id="ED8E6D02D5E3F0E1846EA11AC59CA729">Author</mods:roleTerm>
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<mods:namePart id="2297F8176B9A61021438EB47189976B1">Humberto Y. Yamaguti</mods:namePart>
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<mods:roleTerm id="22A13D8387F302E0A80C4F95E6D6AB25">Author</mods:roleTerm>
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<mods:namePart id="F2AC2E9F3A85456A9F07907604B6A638">Cláudio A. Souza</mods:namePart>
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<mods:namePart id="D8F7F9A5ACC0E78B57E1DAC11C2CC607">Ricardo Pinto-Da-Rocha</mods:namePart>
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<mods:roleTerm id="8EEE2032EC889FB5B461AC81C10F4E2A">Author</mods:roleTerm>
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<mods:namePart id="63F1282BF373ABEA1A69548A9D19D02B">Lorenzo Prendini</mods:namePart>
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<mods:typeOfResource id="8059AB50627EBD393CC223E5038C795C">text</mods:typeOfResource>
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<mods:date id="87F0FDE387369FD8D55675407F32F6A5">2017</mods:date>
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<mods:number id="BD551058152A24B7E1A043C4C60D8FD2">2017-06-26</mods:number>
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<treatment id="8F65ED57FF94B10B38B0CD9BB12D6B41" ID-DOI="http://doi.org/10.5281/zenodo.4610690" ID-GBIF-Taxon="132953595" ID-Zenodo-Dep="4610690" LSID="urn:lsid:zoobank.org:act:5E21CA29-855B-4708-8C0B-18A85E6B09E1" httpUri="http://treatment.plazi.org/id/8F65ED57FF94B10B38B0CD9BB12D6B41" lastPageId="95" lastPageNumber="95" pageId="94" pageNumber="94">
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<subSubSection id="4FD60FCAFF94B10A38B0CD9BB1496FBA" box="[208,533,439,464]" pageId="94" pageNumber="94" type="nomenclature">
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<paragraph id="07735C41FF94B10A38B0CD9BB1496FBA" blockId="94.[208,533,439,464]" box="[208,533,439,464]" pageId="94" pageNumber="94">
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<heading id="5C3BEB2DFF94B10A38B0CD9BB1496FBA" box="[208,533,439,464]" centered="true" fontSize="10" level="4" pageId="94" pageNumber="94" reason="4">
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<emphasis id="35B88053FF94B10A38B0CD9BB2E16FA5" bold="true" box="[208,445,439,464]" pageId="94" pageNumber="94">
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<taxonomicName id="C0CC27C2FF94B10A38B0CD9BB2EA6FA5" ID-CoL="4HHR9" authority="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[208,438,439,464]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
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<emphasis id="35B88053FF94B10A38B0CD9BB2EA6FA5" bold="true" box="[208,438,439,464]" italics="true" pageId="94" pageNumber="94">Physoctonus striatus</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="2E8B3D28FF94B10A39A4CD94B1496FBA" box="[452,533,440,463]" pageId="94" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="4FD60FCAFF94B10A38FBCDCAB1106F8B" box="[155,588,486,510]" pageId="94" pageNumber="94" type="description">
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<paragraph id="07735C41FF94B10A38FBCDCAB1106F8B" blockId="94.[155,588,486,510]" box="[155,588,486,510]" pageId="94" pageNumber="94">
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<figureCitation id="9FF740C4FF94B10A38FBCDCAB2496F8B" box="[155,277,486,510]" captionStart="FIG" captionStartId="24.[107,149,1416,1437]" captionTargetBox="[138,1207,234,1374]" captionTargetId="figure@24.[107,1211,224,1383]" captionTargetPageId="24" captionText="FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C.L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses)." httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="94" pageNumber="94">
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<figureCitation id="9FF740C4FF94B10A38FBCDCAB25C6F8B" box="[155,256,486,510]" captionStart="FIG. 9" captionStartId="24.[107,149,1416,1437]" captionTargetBox="[138,1207,234,1374]" captionTargetId="figure@24.[107,1211,224,1383]" captionTargetPageId="24" captionText="FIG. 9. Map of northeastern South America, plotting known locality records of six species of Rhopalurusinae Bücherl, 1971: A. Ischnotelson guanambiensis (Lenarducci et al., 2005), comb. nov. (star), Ischnotelson peruassu, sp. nov. (triangle), Physoctonus striatus, sp. nov. (circle), Troglorhopalurus lacrau (Lourenço and Pinto-da- Rocha, 1997), comb. nov (diamonds). B. Physoctonus debilis (C. L. Koch, 1840) (squares), Troglorhopalurus translucidus Lourenço et al., 2004 (crosses)." httpUri="https://zenodo.org/record/851400/files/figure.png" pageId="94" pageNumber="94">Figures 9</figureCitation>
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A
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</figureCitation>
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,
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<figureCitation id="9FF740C4FF94B10A3940CDCAB2106F8B" box="[288,332,486,510]" captionStart="FIG" captionStartId="34.[109,151,1628,1649]" captionTargetBox="[208,1116,224,1599]" captionTargetId="figure@34.[109,1212,224,1599]" captionTargetPageId="34" captionText="FIG. 16. Rhopalurusinae Bücherl, 1971, carapace, dorsal aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♀ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Guatire, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 1 mm (A, B), 2 mm (C–F)." httpUri="https://zenodo.org/record/851414/files/figure.png" pageId="94" pageNumber="94">16B</figureCitation>
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,
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<figureCitation id="9FF740C4FF94B10A3937CDCAB2DA6F8B" box="[343,390,486,510]" captionStart="FIG" captionStartId="36.[106,149,1153,1174]" captionTargetBox="[106,634,224,1125]" captionTargetId="figure@36.[106,634,224,1125]" captionTargetPageId="36" captionText="FIG. 17. Rhopalurusinae Bücherl, 1971, pedipalp chela movable finger, dorsal aspect, illustrating dentition (denticle rows). A. Heteroctenus junceus (Herbst, 1800), comb. nov., ♀ (AMNH). B. Ischnotelson peruassu, sp. nov., paratype ♀ (MZSP). C. Jaguajir rochae (Borelli, 1910), comb. nov., ♂ (MZSP). D. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). E. Rhopalurus ochoai, sp. nov., paratype ♀ (AMNH). F. Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997), comb. nov., ♀ (MZSP). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851416/files/figure.png" pageId="94" pageNumber="94">17D</figureCitation>
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,
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<figureCitation id="9FF740C4FF94B10A39F2CDCAB2E26F8B" box="[402,446,486,510]" captionStart="FIG" captionStartId="40.[108,151,1490,1511]" captionTargetBox="[108,1212,224,1461]" captionTargetId="figure@40.[108,1212,224,1462]" captionTargetPageId="40" captionText="FIG. 20. Rhopalurusinae Bücherl, 1971, sternum, pectines and sternite III, ventral aspect. A. Physoctonus debilis (C.L. Koch, 1840), ♂ (MZSP). B. Physoctonus striatus, sp. nov., paratype ♂ (MZSP). C. Rhopalurus caribensis Teruel and Roncallo, 2008, ♂ (AMNH). D. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Laguna Canaima, Venezuela. E. Rhopalurus laticauda Thorell, 1876, ♂ (AMNH), Puerto Colombia, Venezuela. F. Rhopalurus ochoai, sp. nov., holotype ♂ (AMNH). Scale bars = 2 mm." httpUri="https://zenodo.org/record/851422/files/figure.png" pageId="94" pageNumber="94">20B</figureCitation>
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,
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<figureCitation id="9FF740C4FF94B10A39A9CDCAB2AB6F8B" box="[457,503,486,510]" captionStart="FIG" captionStartId="88.[108,150,1167,1188]" captionTargetBox="[108,1212,224,1137]" captionTargetId="figure@88.[108,1212,224,1137]" captionTargetPageId="88" captionText="FIG. 48. Physoctonus Mello-Leitão, 1934, pedipalp chela, retrolateral aspect. A. P. debilis (C.L. Koch, 1840), ♂ (MZSP). B. P. debilis (C.L. Koch, 1840), ♀ (MZSP). C. P. striatus, sp. nov., ♂ (MZSP). Scale bars = 1 mm." httpUri="https://zenodo.org/record/851478/files/figure.png" pageId="94" pageNumber="94">48C</figureCitation>
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;
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<tableCitation id="4A4E69FAFF94B10A3A63CDCAB1106F8B" box="[515,588,486,510]" captionStart="TABLE 3" captionStartId="69.[610,690,228,252]" captionTargetBox="[131,1171,439,1728]" captionText="TABLE 3 Meristic data for type material of Ischnotelson peruassu, sp. n., Physoctonus striatus, sp. n., and Physoctonus debilis (C. L. Koch, 1840). Measurements follow Prendini (2000, 2003, 2004). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri="http://table.plazi.org/id/53B30CC9FF8FB1113A02CCC8B0CF6FEA" pageId="94" pageNumber="94" tableUuid="53B30CC9FF8FB1113A02CCC8B0CF6FEA">table 3</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4FD60FCAFF94B10A380BCE30B3A86C23" pageId="94" pageNumber="94" type="reference_group">
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<paragraph id="07735C41FF94B10A380BCE30B3A86C23" blockId="94.[107,620,540,598]" pageId="94" pageNumber="94">
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<taxonomicName id="C0CC27C2FF94B10A380BCE30B1716C41" ID-CoL="4HHR8" authority=": Porto et al., 2010: 295" authorityName="Porto et al." authorityPageNumber="295" authorityYear="2010" box="[107,557,540,564]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
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<emphasis id="35B88053FF94B10A380BCE30B2686C41" box="[107,308,540,564]" italics="true" pageId="94" pageNumber="94">Physoctonus debilis</emphasis>
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:
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<bibRefCitation id="635D21B0FF94B10A3922CE30B1716C41" author="Porto" box="[322,557,540,564]" etAl="et al." firstAuthor="Porto" journalOrPublisher="Check List" pageId="94" pageNumber="94" pagination="292 - 297" part="6" refId="ref70138" refString="Porto, T. J., T. K. Brazil, and R. M. Lira-da-Silva. 2010. Scorpions, state of Bahia, northeastern Brazil. Check List 6 (2): 292 - 297." title="Scorpions, state of Bahia, northeastern Brazil" type="journal article" year="2010">Porto et al., 2010: 295</bibRefCitation>
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</taxonomicName>
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,
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<tableCitation id="4A4E69FAFF94B10A3A59CE30B3F46C23" captionStart="TABLE 1" captionStartId="6.[609,689,226,250]" captionTargetBox="[115,1198,353,998]" captionText="TABLE 1 Revised classification of the New World buthid Subfamily Rhopalurusinae Bücherl, 1971 (excluding Centruroides Marx, 1890), with countries of distribution." httpUri="http://table.plazi.org/id/53B30CC9FFCCB1523A01CCCEB0B66F30" pageId="94" pageNumber="94" tableUuid="53B30CC9FFCCB1523A01CCCEB0B66F30">table 1</tableCitation>
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(part).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="4FD60FCAFF94B10A38EBCE56B1096D63" pageId="94" pageNumber="94" type="materials_examined">
|
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<paragraph id="07735C41FF94B10A38EBCE56B1096D63" blockId="94.[107,637,634,1749]" pageId="94" pageNumber="94">
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TYPE MATERIAL:
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<emphasis id="35B88053FF94B10A393ECE56B2966CE7" bold="true" box="[350,458,634,658]" pageId="94" pageNumber="94">BRAZIL:</emphasis>
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<emphasis id="35B88053FF94B10A39B8CE56B1456CE7" box="[472,537,634,658]" italics="true" pageId="94" pageNumber="94">Bahía</emphasis>
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: Xique- Xique, 10°49′60″S 42°43′60″W: Holotype ♂ (MZSP 30869/AMCC [LP 9681]), 2.x.2008, T.J. Porto. Paratype ♂ (MZSP 31128/AMCC [LP 9950]), same data except 3.x.2008, T.J. Porto.
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</paragraph>
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</subSubSection>
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<subSubSection id="4FD60FCAFF94B10A38EBCF33B22B6BDE" pageId="94" pageNumber="94" type="diagnosis">
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<paragraph id="07735C41FF94B10A38EBCF33B22B6BDE" blockId="94.[107,637,634,1749]" pageId="94" pageNumber="94">
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DIAGNOSIS:
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<taxonomicName id="C0CC27C2FF94B10A397CCF33B1556D42" authority="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & CLOSE Lorenzo Prendini Scorpion Systematics Research Group, 2017" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[284,521,799,823]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="striatus" status="sp. nov.">
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<emphasis id="35B88053FF94B10A397CCF33B1556D42" box="[284,521,799,823]" italics="true" pageId="94" pageNumber="94">Physoctonus striatus</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="2E8B3D28FF94B10A3A7BCF0CB1246D42" box="[539,632,800,823]" pageId="94" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
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, differs from its sister species,
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<taxonomicName id="C0CC27C2FF94B10A39A4CF6DB17E6D2D" box="[452,546,832,856]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
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<emphasis id="35B88053FF94B10A39A4CF6DB17E6D2D" box="[452,546,832,856]" italics="true" pageId="94" pageNumber="94">P. debilis</emphasis>
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</taxonomicName>
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, as follows. The mesosomal tergites of
|
||
<taxonomicName id="C0CC27C2FF94B10A3983CF4EB10F6D0C" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[483,595,866,889]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF94B10A3983CF4EB10F6D0C" box="[483,595,866,889]" italics="true" pageId="94" pageNumber="94">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
are dark, with transverse bands of infuscation, whereas the tergites of
|
||
<taxonomicName id="C0CC27C2FF94B10A3915CF88B28F6DCE" box="[373,467,931,955]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF94B10A3915CF88B28F6DCE" box="[373,467,931,955]" italics="true" pageId="94" pageNumber="94">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
are uniformly pale, except for the infuscate dorsomedian carinae, which form a thin stripe longitudinally. The carinae of the carapace, pedipalps, and metasoma are less pronounced in
|
||
<emphasis id="35B88053FF94B10A3A45C804B3D16A15" italics="true" pageId="94" pageNumber="94">
|
||
<taxonomicName id="C0CC27C2FF94B10A3A45C804B12C6A4A" box="[549,624,1064,1087]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="stria">P. stria</taxonomicName>
|
||
- tus
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName id="C0CC27C2FF94B10A3896C865B20B6A14" box="[246,343,1097,1121]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF94B10A3896C865B20B6A14" box="[246,343,1097,1121]" italics="true" pageId="94" pageNumber="94">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
: for example, the lateral inframedian carinae of metasomal segments II and III are restricted to the posterior third of the segment in
|
||
<taxonomicName id="C0CC27C2FF94B10A3944C881B2C96AB1" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[292,405,1197,1220]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF94B10A3944C881B2C96AB1" box="[292,405,1197,1220]" italics="true" pageId="94" pageNumber="94">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
, but complete in
|
||
<taxonomicName id="C0CC27C2FF94B10A3A0AC881B3E86A90" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF94B10A3A0AC881B3E86A90" italics="true" pageId="94" pageNumber="94">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
; and the retromedian and secondary accessory carinae of the pedipalp chela manus are vestigial to absent in
|
||
<taxonomicName id="C0CC27C2FF94B10A39F2C93CB15E6B52" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[402,514,1296,1319]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF94B10A39F2C93CB15E6B52" box="[402,514,1296,1319]" italics="true" pageId="94" pageNumber="94">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
, but complete and granular in
|
||
<taxonomicName id="C0CC27C2FF94B10A3904C91DB29F6B3D" box="[356,451,1328,1352]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF94B10A3904C91DB29F6B3D" box="[356,451,1328,1352]" italics="true" pageId="94" pageNumber="94">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
. Pedipalp chela fixed finger trichobothrium
|
||
<emphasis id="35B88053FF94B10A39D1C97DB2916B1C" box="[433,461,1361,1385]" italics="true" pageId="94" pageNumber="94">db</emphasis>
|
||
is aligned with trichobothrium
|
||
<emphasis id="35B88053FF94B10A394CC95FB21D6BFF" box="[300,321,1395,1418]" italics="true" pageId="94" pageNumber="94">et</emphasis>
|
||
in
|
||
<taxonomicName id="C0CC27C2FF94B10A390FC95FB2B86BFF" authorityName="Lauren A. Esposito & Humberto Y. Yamaguti & Cláudio A. Souza & Ricardo Pinto-Da-Rocha & Lorenzo Prendini" authorityYear="2017" box="[367,484,1395,1418]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="striatus">
|
||
<emphasis id="35B88053FF94B10A390FC95FB2B86BFF" box="[367,484,1395,1418]" italics="true" pageId="94" pageNumber="94">P. striatus</emphasis>
|
||
</taxonomicName>
|
||
but situated distal to
|
||
<emphasis id="35B88053FF94B10A38B1C9B8B3B96BDE" box="[209,229,1428,1451]" italics="true" pageId="94" pageNumber="94">et</emphasis>
|
||
in
|
||
<taxonomicName id="C0CC27C2FF94B10A396DC9B8B2306BDE" box="[269,364,1427,1451]" class="Arachnida" family="Buthidae" genus="Physoctonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="94" pageNumber="94" phylum="Arthropoda" rank="species" species="debilis">
|
||
<emphasis id="35B88053FF94B10A396DC9B8B2306BDE" box="[269,364,1427,1451]" italics="true" pageId="94" pageNumber="94">P. debilis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF94B10A38EBC999B24F687A" pageId="94" pageNumber="94" type="etymology">
|
||
<paragraph id="07735C41FF94B10A38EBC999B24F687A" blockId="94.[107,637,634,1749]" pageId="94" pageNumber="94">ETYMOLOGY: The species name refers to the transverse bands of infuscation on the mesosomal tergites.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF94B10B38EBCA34B1036A71" lastPageId="95" lastPageNumber="95" pageId="94" pageNumber="94" type="description">
|
||
<paragraph id="07735C41FF94B10A38EBCA34B25068C1" blockId="94.[107,637,634,1749]" pageId="94" pageNumber="94">
|
||
DESCRIPTION: The following description is based on the holotype male unless otherwise noted (for measurements, see
|
||
<tableCitation id="4A4E69FAFF94B10A39B6CA76B1746807" box="[470,552,1626,1650]" captionStart="TABLE 3" captionStartId="69.[610,690,228,252]" captionTargetBox="[131,1171,439,1728]" captionText="TABLE 3 Meristic data for type material of Ischnotelson peruassu, sp. n., Physoctonus striatus, sp. n., and Physoctonus debilis (C. L. Koch, 1840). Measurements follow Prendini (2000, 2003, 2004). 1 Sum of carapace, tergites I – VII, metasomal segments I – V, and telson; 2 sum of metasomal segments I – V and telson; 3 measured from base of condyle to tip of fixed finger." httpUri="http://table.plazi.org/id/53B30CC9FF8FB1113A02CCC8B0CF6FEA" pageId="94" pageNumber="94" tableUuid="53B30CC9FF8FB1113A02CCC8B0CF6FEA">table 3</tableCitation>
|
||
). Only characters that differ from the generic description are noted.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A38EBCA91B11368A1" blockId="94.[107,637,634,1749]" box="[139,591,1725,1749]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A38EBCA91B25668A0" box="[139,266,1725,1749]" italics="true" pageId="94" pageNumber="94">Total length</emphasis>
|
||
: Small scorpions, 22–26 mm.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACCCC9B0326D64" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACCCC9B0526E88" box="[716,782,229,253]" italics="true" pageId="94" pageNumber="94">Color:</emphasis>
|
||
Base color dark yellow to light brown. Carapace immaculate except interocular surface infuscate, forming dark triangle, almost reaching anterior margin, and thin, dark line of infuscation around margins. Tergites each with infuscate median carina forming a thin dark line longitudinally, with transverse band of infuscation, and a dark line of infuscation around margins. Coxosternal region, pectines, and sternites immaculate, dark yellow. Metasomal segments I–IV, dorsal surfaces immaculate, similar in color to carapace and tergites; II–IV, ventral surfaces infuscate, forming a longitudinal stripe; V completely infuscate. Telson vesicle similar to metasomal segment V dorsal surface, aculeus almost black. Chelicerae, pedipalps, and legs brown, entirely infuscate.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACCF37B0536A6E" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACCF37B06C6D47" box="[716,816,795,818]" italics="true" pageId="94" pageNumber="94">Carapace</emphasis>
|
||
: Pentagonal in shape, anterior width approximately two thirds posterior width. Median ocular tubercle low; median sulci shallow; lateral ocular carinae continuous with posterior median carinae. Carinae obsolete, finely granular and barely distinguishable from surface granulation; anterior margin with large round granules.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACC809B0AC6B71" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACC809B06C6A48" box="[716,816,1061,1085]" italics="true" pageId="94" pageNumber="94">Pedipalps</emphasis>
|
||
: Pedipalp femoral and patellar carinae coarsely granular; chelal carinae more finely granular (fig. 48C). Chela fixed finger trichobothrium
|
||
<emphasis id="35B88053FF94B10A3A8BC8A4B0596AD5" box="[747,773,1160,1184]" italics="true" pageId="94" pageNumber="94">db</emphasis>
|
||
aligned with trichobothrium
|
||
<emphasis id="35B88053FF94B10A3C35C8A5B7346AD5" box="[1109,1128,1161,1184]" italics="true" pageId="94" pageNumber="94">et</emphasis>
|
||
. Retromedian carina vestigial, restricted to distal quarter of chela manus; secondary accessory carina of chela manus absent.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACC922B0726BDF" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACC922B1A66B50" box="[716,762,1294,1317]" italics="true" pageId="94" pageNumber="94">Legs</emphasis>
|
||
: Legs III and IV, tibial spurs absent; I–IV, surfaces carinate; basitarsi each with simple prolateral pedal spur; telotarsi each with distinct pro- and retroventral rows of fine, acuminate macrosetae.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACC998B7EC6B98" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACC998B07A6BBE" box="[716,806,1460,1483]" italics="true" pageId="94" pageNumber="94">Sternum</emphasis>
|
||
: Subtriangular. Median longitudinal sulcus shallow anteriorly, deep, wide posteriorly.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACC9DAB0056824" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACC9DAB0C8687B" box="[716,916,1526,1550]" italics="true" pageId="94" pageNumber="94">Genital operculum</emphasis>
|
||
: Genital opercula suboval, completely divided longitudinally; genital papillae present (♂).
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10A3AACCA77B7D068E1" blockId="94.[684,1214,229,1750]" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACCA77B0406807" box="[716,796,1627,1650]" italics="true" pageId="94" pageNumber="94">Pectines</emphasis>
|
||
: Tooth count, 15/15 (♂). Pectinal plate trapezoid, with deep anterior notch (fig. 20B).
|
||
</paragraph>
|
||
<paragraph id="07735C41FF94B10B3AACCAB2B2136C59" blockId="94.[684,1214,229,1750]" lastBlockId="95.[107,637,229,1332]" lastPageId="95" lastPageNumber="95" pageId="94" pageNumber="94">
|
||
<emphasis id="35B88053FF94B10A3AACCAB2B06668C0" box="[716,826,1694,1717]" italics="true" pageId="94" pageNumber="94">Mesosoma</emphasis>
|
||
: Tergites IV–VI slightly wider than than I–III; I–VI unicarinate, dorsosubmedian carinae absent, dorsomedian carina reduced to posterior half on I–VI. Tergite VII pentacarinate, dorsomedian carina reduced. Sternites III–VI smooth, carinae absent or obsolete; sternite III, lateral margins not forming smooth, raised carina, ventromedian carina not elevated anteriorly, ventrosubmedian surfaces not forming paired depressions, smooth; sternite VII with four granular carinae; respiratory spiracles (stigmata) width more than 5× length.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF95B10B38EBCE1BB2D66D0B" blockId="95.[107,637,229,1332]" pageId="95" pageNumber="95">
|
||
<emphasis id="35B88053FF95B10B38EBCE1BB3A76C3B" box="[139,251,567,590]" italics="true" pageId="95" pageNumber="95">Metasoma</emphasis>
|
||
: Metasomal segments I–III each with 10 carinae, IV with eight carinae, V with five carinae; lateral inframedian carinae complete on segment I, restricted to posterior third of segment on II and III, absent on IV and V. Ventral intercarinal surfaces of segments I–IV coarsely granular, dorsal and lateral surfaces weakly granular or smooth, segment V smooth. Metasoma increasing slightly in width posteriorly, segments IV and V wider than I in adult male.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF95B10B38EBCFABB2756DB4" blockId="95.[107,637,229,1332]" pageId="95" pageNumber="95">
|
||
<emphasis id="35B88053FF95B10B38EBCFABB38C6DEA" box="[139,208,903,927]" italics="true" pageId="95" pageNumber="95">Telson</emphasis>
|
||
: Vesicle pentacarinate; subaculear tubercle vestigial.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF95B10B38EBCFE7B15C6D96" blockId="95.[107,637,229,1332]" box="[139,512,971,995]" pageId="95" pageNumber="95">
|
||
<emphasis id="35B88053FF95B10B38EBCFE7B23F6D96" box="[139,355,971,995]" italics="true" pageId="95" pageNumber="95">Hemispermatophore</emphasis>
|
||
: As for genus.
|
||
</paragraph>
|
||
<paragraph id="07735C41FF95B10B38EBCFC1B1036A71" blockId="95.[107,637,229,1332]" box="[139,607,1004,1029]" pageId="95" pageNumber="95">
|
||
<emphasis id="35B88053FF95B10B38EBCFC1B2066A70" box="[139,346,1005,1029]" italics="true" pageId="95" pageNumber="95">Sexual dimorphism</emphasis>
|
||
: Females are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF95B10B38EBC823B24E6A1F" pageId="95" pageNumber="95" type="distribution">
|
||
<paragraph id="07735C41FF95B10B38EBC823B24E6A1F" blockId="95.[107,637,229,1332]" pageId="95" pageNumber="95">DISTRIBUTION: This species is known only from the type locality in the Brazilian state of Bahía (fig. 9A).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4FD60FCAFF95B10B38EBC858B12D6B41" pageId="95" pageNumber="95" type="biology_ecology">
|
||
<paragraph id="07735C41FF95B10B38EBC858B12D6B41" blockId="95.[107,637,229,1332]" pageId="95" pageNumber="95">
|
||
ECOLOGY: The type locality is situated in the Brazilian caatinga, a very dry environment with sandy soil. Specimens were collected under stones during the day or with UV light detection at night. The habitus is consistent with the lapidicolous ecomorphotype (
|
||
<bibRefCitation id="635D21B0FF95B10B39D0C930B10A6B41" author="Prendini" box="[432,598,1308,1332]" firstAuthor="Prendini" journalOrPublisher="Journal of Arachnology" pageId="95" pageNumber="95" pagination="173 - 188" part="29" refId="ref70328" refString="Prendini, L. 2001 a. Further additions to the scorpion fauna of Trinidad and Tobago. Journal of Arachnology 29 (2): 173 - 188." title="Further additions to the scorpion fauna of Trinidad and Tobago" type="journal article" year="2001">Prendini, 2001</bibRefCitation>
|
||
b).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |