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<mods:title id="9C7CB85377034A86D2740A5F27668B63">A revision of the deep-sea porter crabs of the genus Gordonopsis Guinot &amp; Richer de Forges, 1995 (Crustacea, Decapoda, Brachyura, Homolidae), with descriptions of five new species</mods:title>
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<heading id="5C6081C3A828BA26721FFC21D9AD4FFE" bold="true" box="[828,1437,925,949]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="278" reason="2">
<taxonomicName id="C0974D2CA828BA26721FFC21D9AD4FFE" ID-CoL="84GW2" authority="Ng, Padate &amp; Saravanane, 2019" authorityName="Ng, Padate &amp; Saravanane" authorityYear="2019" box="[828,1437,925,949]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="278" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="35E3EABDA828BA26721FFC21D9AD4FFE" bold="true" box="[828,1437,925,949]" pageId="11" pageNumber="278">
<emphasis id="35E3EABDA828BA26721FFC21D8134FFE" bold="true" box="[828,1059,925,949]" italics="true" pageId="11" pageNumber="278">Gordonopsis robusta</emphasis>
<bibRefCitation id="63064B5EA828BA26750FFC21D9AD4FFE" author="Ng PKL &amp; Padate VP &amp; Saravanane N" box="[1068,1437,925,949]" pageId="11" pageNumber="278" pagination="510 - 516" refId="ref18180" refString="Ng PKL, Padate VP &amp; Saravanane N (2019) Gordonopsis robusta, a new species of deep-sea porter crab (Crustacea: Brachyura: Homolidae) from the Andaman Sea, India. Raffles Bulletin of Zoology, 67: 510 - 516." type="journal article" year="2019">Ng, Padate &amp; Saravanane, 2019</bibRefCitation>
</emphasis>
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<paragraph id="072836AFA828BA267500FC01D8864F9E" blockId="11.[828,1437,925,981]" box="[1059,1206,957,981]" pageId="11" pageNumber="278">
(
<figureCitation id="9FAC2A2AA828BA267508FC01D8B64F9E" box="[1067,1158,957,981]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Figs. 12</figureCitation>
,
<figureCitation id="9FAC2A2AA828BA2675B0FC01D89E4F9E" box="[1171,1198,957,981]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="11" pageNumber="278">13</figureCitation>
)
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<paragraph id="072836AFA828BA26720EFC40D99D485A" blockId="11.[813,1453,1020,1042]" box="[813,1453,1020,1042]" pageId="11" pageNumber="278">
<treatmentCitationGroup id="27871181A828BA26720EFC40D99D485A" box="[813,1453,1020,1042]" pageId="11" pageNumber="278">
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<taxonomicName id="C0974D2CA828BA26720EFC40D97E4859" ID-CoL="84GW2" authority="Ng, Padate &amp; Saravanane, 2019: 511" authorityName="Ng, Padate &amp; Saravanane" authorityPageNumber="511" authorityYear="2019" box="[813,1358,1020,1042]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="278" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="35E3EABDA828BA26720EFC40DFC3485A" box="[813,1011,1020,1041]" italics="true" pageId="11" pageNumber="278">Gordonopsis robusta</emphasis>
<bibRefCitation id="63064B5EA828BA2672D4FC40D97E4859" author="Ng PKL &amp; Padate VP &amp; Saravanane N" box="[1015,1358,1020,1042]" pageId="11" pageNumber="278" pagination="510 - 516" refId="ref18180" refString="Ng PKL, Padate VP &amp; Saravanane N (2019) Gordonopsis robusta, a new species of deep-sea porter crab (Crustacea: Brachyura: Homolidae) from the Andaman Sea, India. Raffles Bulletin of Zoology, 67: 510 - 516." type="journal article" year="2019">Ng, Padate &amp; Saravanane, 2019: 511</bibRefCitation>
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, figs. 14.
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</paragraph>
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<subSubSection id="4F8D6524A828BA26720EFB86DF8148B9" pageId="11" pageNumber="278" type="materials_examined">
<paragraph id="072836AFA828BA26720EFB86DF8148B9" blockId="11.[813,1453,1082,1266]" pageId="11" pageNumber="278">
<emphasis id="35E3EABDA828BA26720EFB86D8284819" bold="true" box="[813,1048,1082,1106]" pageId="11" pageNumber="278">Material examined.</emphasis>
<typeStatus id="D82C880DA828BA267500FB86D8BF4819" box="[1059,1167,1082,1106]" pageId="11" pageNumber="278" type="holotype">Holotype</typeStatus>
: male (33.5 ×
<quantity id="C06F9B4AA828BA26741EFB86D9944819" box="[1341,1444,1082,1106]" metricMagnitude="-2" metricUnit="m" metricValue="4.49" pageId="11" pageNumber="278" unit="mm" value="44.9">44.9 mm</quantity>
) (CMLRE IO/SS/BRC/00082), Andaman Sea, FORVSS station 36708,
<geoCoordinate id="62A35068A828BA2672EEFBC6D81A48D9" box="[973,1066,1146,1170]" degrees="13.27" direction="north" orientation="latitude" pageId="11" pageNumber="278" precision="555" value="13.27">13.27°N</geoCoordinate>
,
<geoCoordinate id="62A35068A828BA267516FBC6D8BE48D9" box="[1077,1166,1146,1170]" degrees="93.26" direction="east" orientation="longitude" pageId="11" pageNumber="278" precision="555" value="93.26">93.26°E</geoCoordinate>
,
<quantity id="C06F9B4AA828BA2675B9FBC6D8D048D9" box="[1178,1248,1146,1170]" metricMagnitude="2" metricUnit="m" metricValue="6.35" pageId="11" pageNumber="278" unit="m" value="635.0">635 m</quantity>
, HSDT (CV), coll. S. S. Cubelio,
<date id="7329106FA828BA2672E6FB26D8A448F9" box="[965,1172,1178,1202]" pageId="11" pageNumber="278" value="2017-11-26">26 November 2017</date>
.
<typeStatus id="D82C880DA828BA2675BEFB27D8CF48F8" box="[1181,1279,1179,1203]" pageId="11" pageNumber="278" type="paratype">Paratype</typeStatus>
: 1 young female (15.6 ×
<quantity id="C06F9B4AA828BA2672A7FB06DFD64899" box="[900,998,1210,1234]" metricMagnitude="-2" metricUnit="m" metricValue="2.08" pageId="11" pageNumber="278" unit="mm" value="20.8">20.8 mm</quantity>
) (CMLRE IO/SS/BRC/00083), same data as holotype.
</paragraph>
</subSubSection>
<subSubSection id="4F8D6524A828BA3C720EFAA6D83D4B8E" lastPageId="17" lastPageNumber="284" pageId="11" pageNumber="278" type="diagnosis">
<paragraph id="072836AFA828BA3C720EFAA6D91D4BCE" blockId="11.[812,1453,1306,2098]" lastBlockId="17.[813,1452,1869,1925]" lastPageId="17" lastPageNumber="284" pageId="11" pageNumber="278">
<emphasis id="35E3EABDA828BA26720EFAA6DF944979" bold="true" box="[813,932,1306,1330]" pageId="11" pageNumber="278">Diagnosis.</emphasis>
<collectingRegion id="C553F84DA828BA26728DFAA6DFDA4979" box="[942,1002,1306,1330]" country="Maldives" name="Maale" pageId="11" pageNumber="278">Male</collectingRegion>
: carapace longitudinally ovate, width to length ratio 0.75, distinctly wider posteriorly than anteriorly; dorsal carapace surface with well-defined regions, separated by broad, deep grooves; lateral margin distinctly convex; dorsal parts with dense soft setae partially obscuring surface; lateral parts densely setose, completely obscuring surface (
<figureCitation id="9FAC2A2AA828BA267216FA66DFA249B9" box="[821,914,1498,1522]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Figs. 12</figureCitation>
AD, 13A, B). Rostrum relatively long, sharp, with 2 relatively long, obliquely directed pseudorostral spines, just shorter than rostrum (
<figureCitation id="9FAC2A2AA828BA26758AF9A6D9204A79" box="[1193,1296,1562,1586]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Supraorbital margin relatively narrow, C-shaped; pseudorostral spines directed obliquely laterally at angle of about 45° (
<figureCitation id="9FAC2A2AA828BA26745DF9E6DF6F4AD9" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Supraorbital margin relatively narrow, C-shaped; pseudorostral spines directed obliquely laterally at angle of about 45° (
<figureCitation id="9FAC2A2AA828BA2672EAF906D81E4A99" box="[969,1070,1722,1746]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Eyes with short ocular peduncle, cornea prominent; no discernible orbit (
<figureCitation id="9FAC2A2AA828BA2675C3F966D9744AB9" box="[1248,1348,1754,1778]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Hepatic region gently inflated, with short obliquely directed spine (
<figureCitation id="9FAC2A2AA828BA267216F8A6DFA64B79" box="[821,918,1818,1842]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Gastric region without spines (
<figureCitation id="9FAC2A2AA828BA2675CFF8A6D97D4B79" box="[1260,1357,1818,1842]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Gastric groove well marked, with distinct ovate gastric fossae just above (
<figureCitation id="9FAC2A2AA828BA26725EF8E6DFD04B39" box="[893,992,1882,1906]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Cardiac region swollen; branchial region inflated, with distinct branchio-cardiac grooves (
<figureCitation id="9FAC2A2AA828BA26741FF8C6D9904BD9" box="[1340,1440,1914,1938]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Latero-posterior tubercle strong, sharp (
<figureCitation id="9FAC2A2AA828BA2675FCF826D9744BF9" box="[1247,1348,1946,1970]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="11" pageNumber="278">Fig. 12C</figureCitation>
). Base of antenna with short spine. Antennal flagellum short, second and third articles thick, setose. Epistome triangular. Third maxilliped pediform, merus elongate with dentiform external angle (
<figureCitation id="9FAC2A2AA828BA267257F7A6DFE44479" box="[884,980,2074,2098]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="11" pageNumber="278">Fig. 13C</figureCitation>
). Chelipeds long; fingers short, less than half length of palm; surface gently rugose with 4 or 5 longitudinal rows of small granules, margins with sharp granules and short spines; carpus longitudinally ovate, outer margin with 810 short spines, distal edge with short spine, inner margin with 5 or 6 spines, dorsal surface with ridge lined with 35 short sharp tubercles or granules; merus triangular in cross-section, relatively long, curved, dorsal margin with 5 short spines and distal 2 spines bracketing chela, outer ventral margin with 1618 spines and spinules, inner ventral margin with 68 spines or spinules (
<figureCitation id="9FAC2A2AA833BA3D70BAF80EDE1F4B81" box="[409,559,1970,1994]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="16" pageNumber="283">Figs. 12A, E</figureCitation>
,
<figureCitation id="9FAC2A2AA833BA3D731DF80EDE5B4B81" box="[574,619,1970,1994]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">13F</figureCitation>
). Ambulatory legs relatively long; basis-ischium with small granules, not spines; P2 merus with 6 spines or sharp tubercles on dorsal margin (excluding 1 distal spine), ventral margin with 35 spinules or tubercles; P3 merus with 5 spines on dorsal margin (excluding 1 distal spine), ventral margin with 1 or 2 sharp tubercles; P4 merus not known (
<figureCitation id="9FAC2A2AA833BA3D75D6F96ED9B24AA1" box="[1269,1410,1746,1770]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="16" pageNumber="283">Figs. 12A, B</figureCitation>
,
<figureCitation id="9FAC2A2AA833BA3D74AEF96EDF6A4B41" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">13G, H</figureCitation>
). P5 merus slender, unarmed on all margins, just reaching gastric groove when folded anteriorly; carpus long, propodus short, enlarged, forming prominent pseudochela with stout, gently curved dactylus; occlusal margin of fixed finger with 8 spines, that of dactylus with 8 spines (
<figureCitation id="9FAC2A2AA833BA3D75D8F8CED9594BC1" box="[1275,1385,1906,1930]" captionStart="Fig" captionStartId="15.[113,148,2024,2046]" captionTargetBox="[111,1453,191,2008]" captionTargetId="figure-11@15.[111,1453,191,2008]" captionTargetPageId="15" captionText="Fig. 12. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="16" pageNumber="283">Figs. 12D</figureCitation>
,
<figureCitation id="9FAC2A2AA833BA3D7455F8CED9854BC1" box="[1398,1461,1906,1930]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">13I, J</figureCitation>
). Outer margins of P2P4 coxae smooth, unarmed (
<figureCitation id="9FAC2A2AA833BA3D747EF82EDF674B81" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">Fig. 13D, E</figureCitation>
). Telson triangular, with gently convex lateral margins and sharp tip (
<figureCitation id="9FAC2A2AA833BA3D7296F86ED80D4BA1" box="[949,1085,2002,2026]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">Fig. 13D, E</figureCitation>
). G1 short, stout, distal part gently upcurved, tip concave (
<figureCitation id="9FAC2A2AA833BA3D7562F84ED8964441" box="[1089,1190,2034,2058]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">Fig. 13K</figureCitation>
). G2 stout, tubular, length subequal to G1, distal portion cup-shaped (
<figureCitation id="9FAC2A2AA833BA3D75DDF7AED96C4461" box="[1278,1372,2066,2090]" captionStart="Fig" captionStartId="16.[136,171,1519,1541]" captionTargetBox="[136,1474,189,1503]" captionTargetId="figure-7@16.[136,1474,189,1503]" captionTargetPageId="16" captionText="Fig. 13. Gordonopsis robusta Ng, Padate &amp; Saravanane, 2019, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2." figureDoi="http://doi.org/10.5281/zenodo.4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283">Fig. 13L</figureCitation>
). Female: non-sexual characters similar to male except cheliped merus with 6 spines on upper margin, 8 short spines and tubercles on outer ventral margin, 5 or 6 sharp tubercles on inner ventral margin; P2 merus with 5 spines on dorsal margin, ventral margin with 1 sharp tubercle; P3 merus with 5 spines on dorsal margin, ventral margin with 1 sharp tubercle; P4 with 3 spines on dorsal margin, ventral margin unarmed; telson broadly triangular with convex lateral margins.
</paragraph>
<caption id="53E86627A82FBA2171ABF7ACDEFF4409" ID-DOI="http://doi.org/10.5281/zenodo.4576761" ID-Zenodo-Dep="4576761" httpUri="https://zenodo.org/record/4576761/files/figure.png" pageId="12" pageNumber="279" startId="12.[136,171,2064,2086]" targetBox="[178,1432,189,2048]" targetPageId="12">
<paragraph id="072836AFA82FBA2171ABF7ACDEFF4409" blockId="12.[136,1474,2064,2115]" pageId="12" pageNumber="279">
Fig. 9.
<taxonomicName id="C0974D2CA82FBA2171F2F7ACDDBE446E" authority="Ng &amp; Forges, 2020" authorityName="Ng &amp; Forges" authorityYear="2020" box="[209,398,2064,2085]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="279" phylum="Arthropoda" rank="species" species="alaini" status="sp. nov.">
<emphasis id="35E3EABDA82FBA2171F2F7ACDDBE446E" box="[209,398,2064,2085]" italics="true" pageId="12" pageNumber="279">Gordonopsis alaini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2ED057C6A82FBA2170BEF7ACDE26446D" box="[413,534,2064,2086]" pageId="12" pageNumber="279" rank="species">new species</taxonomicNameLabel>
, holotype male (15.4 × 10.0 mm) (MNHN-IU-2017-9049), Madagascar. A, overall habitus; B, dorsal view of carapace; C, dorso-frontal view of carapace.
</paragraph>
</caption>
<caption id="53E86627A82EBA207152F87ADDED447A" ID-DOI="http://doi.org/10.5281/zenodo.4576763" ID-Zenodo-Dep="4576763" httpUri="https://zenodo.org/record/4576763/files/figure.png" pageId="13" pageNumber="280" startId="13.[113,148,1990,2012]" targetBox="[113,1451,189,1974]" targetPageId="13">
<paragraph id="072836AFA82EBA207152F87ADDED447A" blockId="13.[113,1452,1989,2098]" pageId="13" pageNumber="280">
Fig. 10.
<taxonomicName id="C0974D2CA82EBA2071EBF87ADDB84B90" authority="Ng &amp; Forges, 2020" authorityName="Ng &amp; Forges" authorityYear="2020" box="[200,392,1990,2011]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="280" phylum="Arthropoda" rank="species" species="alaini" status="sp. nov.">
<emphasis id="35E3EABDA82EBA2071EBF87ADDB84B90" box="[200,392,1990,2011]" italics="true" pageId="13" pageNumber="280">Gordonopsis alaini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2ED057C6A82EBA2070BBF87ADE224B97" box="[408,530,1990,2012]" pageId="13" pageNumber="280" rank="species">new species</taxonomicNameLabel>
, holotype male (15.4 × 10.0 mm) (MNHN-IU-2017-9049), Madagascar. A, lateral view of cephalothorax; B, frontal view of cephalothorax; C, left third maxilliped; D, ventral view showing buccal cavity, epistome, antennae, and antennules; E, outer view of right carpus and chela; F, dorsal view of left cheliped; G, dorsal view of right cheliped; H, ventral view of right G1; I, ventral view of right G2.
</paragraph>
</caption>
<caption id="53E86627A82DBA2371ABF8D2DFB04B91" ID-DOI="http://doi.org/10.5281/zenodo.4576765" ID-Zenodo-Dep="4576765" httpUri="https://zenodo.org/record/4576765/files/figure.png" pageId="14" pageNumber="281" startId="14.[136,172,1902,1924]" targetBox="[136,1474,293,1886]" targetPageId="14">
<paragraph id="072836AFA82DBA2371ABF8D2DFB04B91" blockId="14.[136,1474,1902,2010]" pageId="14" pageNumber="281">
Fig. 11.
<taxonomicName id="C0974D2CA82DBA2371FCF8D2DD904BC8" authority="Ng &amp; Forges, 2020" authorityName="Ng &amp; Forges" authorityYear="2020" box="[223,416,1902,1923]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="281" phylum="Arthropoda" rank="species" species="alaini" status="sp. nov.">
<emphasis id="35E3EABDA82DBA2371FCF8D2DD904BC8" box="[223,416,1902,1923]" italics="true" pageId="14" pageNumber="281">Gordonopsis alaini</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="2ED057C6A82DBA23708CF8D2DE1A4BCF" box="[431,554,1902,1924]" pageId="14" pageNumber="281" rank="species">new species</taxonomicNameLabel>
, holotype male (15.4 × 10.0 mm) (MNHN-IU-2017-9049), Madagascar. A, dorsal view of carapace showing relative P5 length; B, ventral view of cephalothorax showing pleon and unarmed P2P4 coxae; C, ventral view of cephalothorax showing sternopleonal cavity and unarmed P2P4 coxae; D, pleon; EH, right P2P5, respectively (all to same scale); I, right P5 pseudochela (same scale as EH); J, left P5; K, left P5 pseudochela.
</paragraph>
</caption>
<caption id="53E86627A82CBA227152F854DEC7447C" ID-DOI="http://doi.org/10.5281/zenodo.4576767" ID-Zenodo-Dep="4576767" httpUri="https://zenodo.org/record/4576767/files/figure.png" pageId="15" pageNumber="282" startId="15.[113,148,2024,2046]" targetBox="[111,1453,191,2008]" targetPageId="15">
<paragraph id="072836AFA82CBA227152F854DEC7447C" blockId="15.[113,1451,2023,2103]" pageId="15" pageNumber="282">
Fig. 12.
<taxonomicName id="C0974D2CA82CBA2271E6F854DEEA4BB6" authority="Ng, Padate &amp; Saravanane, 2019" authorityName="Ng, Padate &amp; Saravanane" authorityYear="2019" box="[197,730,2024,2046]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="282" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="35E3EABDA82CBA2271E6F854DDA34BB6" box="[197,403,2024,2045]" italics="true" pageId="15" pageNumber="282">Gordonopsis robusta</emphasis>
<bibRefCitation id="63064B5EA82CBA2270B8F854DEEA4BB6" author="Ng PKL &amp; Padate VP &amp; Saravanane N" box="[411,730,2024,2046]" pageId="15" pageNumber="282" pagination="510 - 516" refId="ref18180" refString="Ng PKL, Padate VP &amp; Saravanane N (2019) Gordonopsis robusta, a new species of deep-sea porter crab (Crustacea: Brachyura: Homolidae) from the Andaman Sea, India. Raffles Bulletin of Zoology, 67: 510 - 516." type="journal article" year="2019">Ng, Padate &amp; Saravanane, 2019</bibRefCitation>
</taxonomicName>
. A, C, E, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea; B, D, paratype female (15.6 × 20.8 mm) (CMLRE), Andaman Sea. A, B, overall habitus; C, dorsal view of carapace; D, dorsal view of carapace showing relative P5 length; E, outer view of right chela.
</paragraph>
</caption>
<caption id="53E86627A833BA3D71ABFA53DE874A11" ID-DOI="http://doi.org/10.5281/zenodo.4576769" ID-Zenodo-Dep="4576769" httpUri="https://zenodo.org/record/4576769/files/figure.png" pageId="16" pageNumber="283" startId="16.[136,171,1519,1541]" targetBox="[136,1474,189,1503]" targetPageId="16">
<paragraph id="072836AFA833BA3D71ABFA53DE874A11" blockId="16.[136,1475,1519,1627]" pageId="16" pageNumber="283">
Fig. 13.
<taxonomicName id="C0974D2CA833BA3D71FAFA53DED04A4F" authority="Ng, Padate &amp; Saravanane, 2019" authorityName="Ng, Padate &amp; Saravanane" authorityYear="2019" box="[217,736,1519,1541]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="283" phylum="Arthropoda" rank="species" species="robusta">
<emphasis id="35E3EABDA833BA3D71FAFA53DD944A4F" box="[217,420,1519,1540]" italics="true" pageId="16" pageNumber="283">Gordonopsis robusta</emphasis>
<bibRefCitation id="63064B5EA833BA3D7089FA53DED04A4F" author="Ng PKL &amp; Padate VP &amp; Saravanane N" box="[426,736,1519,1541]" pageId="16" pageNumber="283" pagination="510 - 516" refId="ref18180" refString="Ng PKL, Padate VP &amp; Saravanane N (2019) Gordonopsis robusta, a new species of deep-sea porter crab (Crustacea: Brachyura: Homolidae) from the Andaman Sea, India. Raffles Bulletin of Zoology, 67: 510 - 516." type="journal article" year="2019">Ng, Padate &amp; Saravanane, 2019</bibRefCitation>
</taxonomicName>
, holotype male (33.5 × 44.9 mm) (CMLRE), Andaman Sea. A, dorso-lateral view of cephalothorax; B, frontal view of cephalothorax; C, right third maxilliped; D, E, ventral view of cephalothorax showing pleon and coxae; F, dorsal view of carpus of right cheliped; GI, left P3P5 meri and leg, respectively (all to same scale); J, left P5 pseudochela; K, ventral view of right G1; L, ventral view of right G2.
</paragraph>
</caption>
<caption id="53E86627A832BA3C7152F95FDFE84B5D" ID-DOI="http://doi.org/10.5281/zenodo.4576773" ID-Zenodo-Dep="4576773" httpUri="https://zenodo.org/record/4576773/files/figure.png" pageId="17" pageNumber="284" startId="17.[113,148,1763,1785]" targetBox="[113,1451,195,1748]" targetPageId="17">
<paragraph id="072836AFA832BA3C7152F95FDFE84B5D" blockId="17.[113,1452,1763,1814]" pageId="17" pageNumber="284">
Fig. 14.
<taxonomicName id="C0974D2CA832BA3C71E4F95FDEA94AB2" authority="Takeda &amp; Suyama, 2019" authorityName="Takeda &amp; Suyama" authorityYear="2019" box="[199,665,1763,1785]" class="Malacostraca" family="Homolidae" genus="Gordonopsis" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="284" phylum="Arthropoda" rank="species" species="pacifica">
<emphasis id="35E3EABDA832BA3C71E4F95FDDA74AB2" box="[199,407,1763,1785]" italics="true" pageId="17" pageNumber="284">Gordonopsis pacifica</emphasis>
<bibRefCitation id="63064B5EA832BA3C7083F95FDEA94AB2" author="Takeda M &amp; Suyama N" box="[416,665,1763,1785]" pageId="17" pageNumber="284" pagination="45 - 59" refId="ref18803" refString="Takeda M &amp; Suyama N (2019) Five species of crabs (Crustacea: Decapoda: Brachyura) from the depths off Okino-Torishima, the southernmost island in Japan. Bulletin of the National Museum of Nature and Science, Tokyo, Series A (Zoology), 45: 45 - 59." type="journal article" year="2019">Takeda &amp; Suyama, 2019</bibRefCitation>
</taxonomicName>
, holotype male (31.0 × 43.0 mm) (NSMT-Cr 26053), Okino-Torishima Island, Japan. A, overall habitus; B, dorsal view of carapace; C, dorso-frontal view of carapace.
</paragraph>
</caption>
<paragraph id="072836AFA832BA3C720EF811D83D4B8E" blockId="17.[813,1037,1965,1989]" box="[813,1037,1965,1989]" pageId="17" pageNumber="284">
<emphasis id="35E3EABDA832BA3C720EF811DFB44B8E" bold="true" box="[813,900,1965,1989]" pageId="17" pageNumber="284">Colour.</emphasis>
<collectingRegion id="C553F84DA832BA3C72AFF812DF844B8E" box="[908,948,1966,1989]" country="Maldives" name="Uthuru" pageId="17" pageNumber="284">Not</collectingRegion>
known.
</paragraph>
</subSubSection>
<subSubSection id="4F8D6524A832BA3C720EF852DF91446D" pageId="17" pageNumber="284" type="discussion">
<paragraph id="072836AFA832BA3C720EF852DF91446D" blockId="17.[813,1450,2029,2086]" pageId="17" pageNumber="284">
<emphasis id="35E3EABDA832BA3C720EF852DFAF444D" bold="true" box="[813,927,2030,2054]" pageId="17" pageNumber="284">Remarks.</emphasis>
See general discussion for comparisons with congeners.
</paragraph>
</subSubSection>
</treatment>
</document>