298 lines
43 KiB
XML
298 lines
43 KiB
XML
<document id="E78564147A0AAE0084FED8EBBCE0F7D6" ID-DOI="10.26879/747" ID-ISSN="1094-8074" ID-Zenodo-Dep="10961544" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712855360717" checkinUser="felipe" docAuthor="Butler, Kaylene, Travouillon, Kenny J., Price, Gilbert J., Archer, Michael & Hand, Suzanne J." docDate="2018" docId="8F33035AFD18FFE122B4E528D63EFA35" docLanguage="en" docName="PalaeontolElectron.21.1.8A.pdf" docOrigin="Palaeontologia Electronica (8 A) 21 (1)" docSource="http://dx.doi.org/10.26879/747" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Ganawamaya couperi Butler, Travouillon, Price, Archer & Hand, 2018, comb. nov." docType="treatment" docVersion="5" lastPageNumber="20" masterDocId="730A7B22FD09FFF2211FE213D44AFFB7" masterDocTitle="Revision of Oligo-Miocene kangaroos, Ganawamaya and Nambaroo (Marsupialia: Macropodiformes, Balbaridae)" masterLastPageNumber="58" masterPageNumber="1" pageNumber="18" updateTime="1714000781735" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:titleInfo id="67A56EA8499782C9947237B147E19B2C">
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<mods:title id="0F439D3D317206270470C3AA677F8EA8">Revision of Oligo-Miocene kangaroos, Ganawamaya and Nambaroo (Marsupialia: Macropodiformes, Balbaridae)</mods:title>
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<mods:namePart id="362490BBC3B79BB3081F3A96AB4E517C">Butler, Kaylene</mods:namePart>
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<mods:roleTerm id="2C1D8CECB1AE1CB2C5CEE514474A2275">Author</mods:roleTerm>
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<mods:namePart id="75DFD0B7111620928CB4868D4A8C9391">Travouillon, Kenny J.</mods:namePart>
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<mods:namePart id="85605F70694FD4CC105DFC2F1FBDA3B9">Price, Gilbert J.</mods:namePart>
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<mods:namePart id="2969C493D53591904FAABC35E40C01D6">Archer, Michael</mods:namePart>
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<mods:namePart id="0ABEC5B93EA2373A1BCA398BA45C6386">Hand, Suzanne J.</mods:namePart>
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<mods:typeOfResource id="415A48A89652D85F5D2A5C546931B203">text</mods:typeOfResource>
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<mods:title id="453FE3DD796BE86579D5D46B275955CC">Palaeontologia Electronica</mods:title>
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<mods:part id="DC10D71B92CAB8ED2E3A3E1D01874719">
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<mods:date id="2CA530FCC344E94B46527A1C5EC750A1">2018</mods:date>
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<mods:title id="300B1DD8AD4F91EA8B060A76C48AFBBA">8 A</mods:title>
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<mods:number id="76682F520C8DC98CD034483BBBA82814">2018-03-31</mods:number>
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<mods:number id="DEA877DDA0EA3FF6BE05BBCD2C646A1A">21</mods:number>
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<mods:number id="8ED3637F52E12C46DD17AB7DCCB00617">1</mods:number>
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<mods:url id="A1507403CC5FCA0C783186CCB6F11AE9">http://dx.doi.org/10.26879/747</mods:url>
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<mods:classification id="99164757E4B71007ED9A2F0721F71CFA">journal article</mods:classification>
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<mods:identifier id="12E50CF47B33B9B4ABD7089350461FD0" type="DOI">10.26879/747</mods:identifier>
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<mods:identifier id="5BD1ABECB3BADDF2D70E0E1E3B612A52" type="ISSN">1094-8074</mods:identifier>
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<mods:identifier id="C3553FF7299F18722F5CA10F2ED788BC" type="Zenodo-Dep">10961544</mods:identifier>
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<treatment id="8F33035AFD18FFE122B4E528D63EFA35" LSID="urn:lsid:plazi:treatment:8F33035AFD18FFE122B4E528D63EFA35" httpUri="http://treatment.plazi.org/id/8F33035AFD18FFE122B4E528D63EFA35" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
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<subSubSection id="4F80E1C7FD18FFE322B4E528D17DF8E2" box="[939,1335,1851,1877]" pageId="17" pageNumber="18" type="nomenclature">
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<paragraph id="0725B24CFD18FFE322B4E528D17DF8E2" blockId="17.[939,1335,1851,1909]" box="[939,1335,1851,1877]" pageId="17" pageNumber="18">
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<heading id="5C6D0520FD18FFE322B4E528D17DF8E2" box="[939,1335,1851,1877]" centered="true" fontSize="10" level="2" pageId="17" pageNumber="18" reason="7">
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<taxonomicName id="C09AC9CFFD18FFE322B4E528D0E4F8E2" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" box="[939,1198,1851,1877]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="couperi" status="comb. nov.">
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<emphasis id="35EE6E5EFD18FFE322B4E528D0E4F8E2" box="[939,1198,1851,1877]" italics="true" pageId="17" pageNumber="18">Ganawamaya couperi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2EDDD325FD18FFE325AAE52FD17DF8E2" box="[1205,1335,1852,1877]" pageId="17" pageNumber="18" rank="species">comb. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="4F80E1C7FD18FFE3255CE54FD0EFF8C2" box="[1091,1189,1884,1909]" pageId="17" pageNumber="18" type="description">
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<paragraph id="0725B24CFD18FFE3255CE54FD0EFF8C2" blockId="17.[939,1335,1851,1909]" box="[1091,1189,1884,1909]" pageId="17" pageNumber="18">
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<figureCitation id="9FA1AEC9FD18FFE3255CE54FD0EFF8C2" box="[1091,1189,1884,1909]" captionStart="FIGURE 6" captionStartId="17.[200,291,1426,1449]" captionTargetBox="[216,1383,201,1372]" captionTargetId="figure-154@17.[192,1427,165,1405]" captionTargetPageId="17" captionText="FIGURE 6. Cast of lower dentition of Ganawamaya couperi comb. nov. (UCMP 57334) (1-3) and cast of upper dentition of Gan. couperi comb. nov. (UCMP 57340) (4-6) from the Ngapakaldi Local Fauna, South Australia. Right lateral view (1 and 4), left lateral view (2 and 5), and occlusal view (3 and 6). Abbreviations: C, canine, I1, first upper incisor; i1, first lower incisor, M1–M4, upper first to fourth molar; m1–m3, lower first to third molar, P3, upper third premolar; p3, lower third premolar." figureDoi="http://doi.org/10.5281/zenodo.10961556" httpUri="https://zenodo.org/record/10961556/files/figure.png" pageId="17" pageNumber="18">Figure 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD1BFFE021DFE2B5D551FEF5" pageId="18" pageNumber="19" type="reference_group">
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<paragraph id="0725B24CFD1BFFE021DFE2B5D758FF0A" blockId="18.[192,793,166,962]" box="[192,786,166,189]" pageId="18" pageNumber="19">
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v* 1997
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<taxonomicName id="C09AC9CFFD1BFFE02021E2B5D649FF0A" authorityName="Cooke" authorityYear="1997" box="[318,515,166,189]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="couperi">
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<emphasis id="35EE6E5EFD1BFFE02021E2B5D649FF0A" box="[318,515,166,189]" italics="true" pageId="18" pageNumber="19">Nambaroo couperi</emphasis>
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</taxonomicName>
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; Cooke, p. 270, figs. 1, 2.
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</paragraph>
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<paragraph id="0725B24CFD1BFFE021DFE2DAD551FEF5" blockId="18.[192,793,166,962]" pageId="18" pageNumber="19">
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<materialsCitation id="B7F2B811FD1BFFE021DFE2DAD551FEF5" collectionCode="QM" country="Australia" location="White Hunter Site" municipality="Riversleigh" pageId="18" pageNumber="19" specimenCode="F30401" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
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<emphasis id="35EE6E5EFD1BFFE021DFE2DAD573FF55" bold="true" box="[192,313,201,226]" pageId="18" pageNumber="19">
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<typeStatus id="D8210CEEFD1BFFE021DFE2DAD57FFF55" box="[192,309,201,226]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="618B2A89FD1BFFE0205BE2DBD525FF55" box="[324,367,200,226]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="18" pageNumber="19" type="Museum">QM</collectionCode>
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<specimenCode id="573C1A37FD1BFFE02065E2DAD592FF55" box="[378,472,201,226]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="18" pageNumber="19" type="Museum">F30401</specimenCode>
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, right dentary with p3 and m1–m4 from
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<location id="0245E497FD1BFFE0207FE2FAD673FEB5" LSID="urn:lsid:plazi:treatment:8F33035AFD18FFE122B4E528D63EFA35:0245E497FD1BFFE0207FE2FAD673FEB5" box="[352,569,232,258]" country="Australia" municipality="Riversleigh" name="White Hunter Site" pageId="18" pageNumber="19" stateProvince="Queensland">White Hunter Site</location>
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,
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<collectingMunicipality id="E7412836FD1BFFE0235AE2FAD683FEB5" box="[581,713,233,258]" pageId="18" pageNumber="19">Riversleigh</collectingMunicipality>
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WHA, northwestern
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<collectingRegion id="C55E7CAEFD1BFFE02064E31BD645FE95" box="[379,527,264,290]" country="Australia" name="Queensland" pageId="18" pageNumber="19">Queensland</collectingRegion>
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,
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<collectingCountry id="7F8DF2DCFD1BFFE0232DE31AD6D0FE95" box="[562,666,265,290]" name="Australia" pageId="18" pageNumber="19">Australia</collectingCountry>
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(
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<bibRefCitation id="630BCFBDFD1BFFE023DEE31BD55AFEF5" author="Cooke, B. N." journalOrPublisher="Memoirs of the Queensland Museum" pageId="18" pageNumber="19" pagination="269 - 280" part="41" refId="ref19388" refString="Cooke, B. N. 1997 b. Two new balbarine kangaroos and lower molar evolution within the subfamily. Memoirs of the Queensland Museum, 41: 269 - 280." title="Two new balbarine kangaroos and lower molar evolution within the subfamily" type="journal article" year="1997">Cooke, 1997b</bibRefCitation>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD1BFFE021DFE35AD632FDB5" pageId="18" pageNumber="19" type="materials_examined">
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<paragraph id="0725B24CFD1BFFE021DFE35AD632FDB5" blockId="18.[192,793,166,962]" pageId="18" pageNumber="19">
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<emphasis id="35EE6E5EFD1BFFE021DFE35AD59AFED5" bold="true" box="[192,464,329,354]" pageId="18" pageNumber="19">Referred specimens.</emphasis>
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The following specimens are referred based on casts of the original specimens. Ngapakaldi Local Fauna,
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<collectingRegion id="C55E7CAEFD1BFFE02349E39BD75FFE15" box="[598,789,392,418]" country="Australia" name="South Australia" pageId="18" pageNumber="19">South Australia</collectingRegion>
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:
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<collectionCode id="618B2A89FD1BFFE021DFE3BBD544FE75" box="[192,270,424,450]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
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57340, left maxilla with P3, M1–M4;
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<collectionCode id="618B2A89FD1BFFE023D5E3BBD752FE75" box="[714,792,424,450]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
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57337 right M1–M2 and right p3, m1–m3;
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<collectionCode id="618B2A89FD1BFFE023D6E3DBD75DFE55" box="[713,791,456,482]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
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57334, right dentary with p3, m1–m3.
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD1BFFE021DFE01AD55FFC75" pageId="18" pageNumber="19" type="diagnosis">
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<paragraph id="0725B24CFD1BFFE021DFE01AD55FFC75" blockId="18.[192,793,166,962]" pageId="18" pageNumber="19">
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<emphasis id="35EE6E5EFD1BFFE021DFE01AD61DFD95" bold="true" box="[192,599,521,546]" pageId="18" pageNumber="19">Emended species diagnosis.</emphasis>
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<taxonomicName id="C09AC9CFFD1BFFE0236BE01BD55DFDF5" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="couperi" status="comb. nov.">
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<emphasis id="35EE6E5EFD1BFFE0236BE01BD55DFDF5" italics="true" pageId="18" pageNumber="19">Ganawamaya couperi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2EDDD325FD1BFFE0203FE03AD5E8FDF5" box="[288,418,553,578]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
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differs from all other species of
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<taxonomicName id="C09AC9CFFD1BFFE021DFE05BD52EFDD5" authorityName="Cooke" authorityYear="1992" box="[192,356,584,610]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD1BFFE021DFE05BD52EFDD5" box="[192,356,584,610]" italics="true" pageId="18" pageNumber="19">Ganawamaya</emphasis>
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</taxonomicName>
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in having the following combination of features: i1 long and narrow with no ventral flange; a hypoconulid present posterior to the entoconid on the buccal side of the m1; prominent posthypocristid on the m1–m4; marked convexities on the lateral margins of m1–m4; anterior cingulum on M1 is well developed; stylar cusp C is a prominent blade-like convexity buccal to the postparacrista; generally smaller molar size; no additional cuspid on the posterior end of the p3 below the occlusal margin; a poorly developed posterolingual cusp on the P3.
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD1BFFE121DFE1CAD6FBFF55" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
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<paragraph id="0725B24CFD1BFFE021DFE1CAD518FC45" blockId="18.[192,338,985,1010]" box="[192,338,985,1010]" pageId="18" pageNumber="19">
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<emphasis id="35EE6E5EFD1BFFE021DFE1CAD518FC45" bold="true" box="[192,338,985,1010]" pageId="18" pageNumber="19">Description</emphasis>
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</paragraph>
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<paragraph id="0725B24CFD1BFFE021DFE61AD64DFA15" blockId="18.[192,793,1033,1890]" pageId="18" pageNumber="19">
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<emphasis id="35EE6E5EFD1BFFE021DFE61AD5C6FB95" bold="true" box="[192,396,1033,1058]" pageId="18" pageNumber="19">Upper dentition.</emphasis>
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Description of the upper dentition of
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<taxonomicName id="C09AC9CFFD1BFFE021FDE63BD534FBF5" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" box="[226,382,1064,1090]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="couperi" status="comb. nov.">
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<emphasis id="35EE6E5EFD1BFFE021FDE63BD534FBF5" box="[226,382,1064,1090]" italics="true" pageId="18" pageNumber="19">Gan. couperi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="2EDDD325FD1BFFE02094E63AD65AFBF5" box="[395,528,1065,1090]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
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is based on casts of
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<collectionCode id="618B2A89FD1BFFE021DFE65BD544FBD5" box="[192,270,1096,1122]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
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57340 (
|
||
<figureCitation id="9FA1AEC9FD1BFFE02065E65AD5ACFBD5" box="[378,486,1097,1122]" captionStart="FIGURE 6" captionStartId="17.[200,291,1426,1449]" captionTargetBox="[216,1383,201,1372]" captionTargetId="figure-154@17.[192,1427,165,1405]" captionTargetPageId="17" captionText="FIGURE 6. Cast of lower dentition of Ganawamaya couperi comb. nov. (UCMP 57334) (1-3) and cast of upper dentition of Gan. couperi comb. nov. (UCMP 57340) (4-6) from the Ngapakaldi Local Fauna, South Australia. Right lateral view (1 and 4), left lateral view (2 and 5), and occlusal view (3 and 6). Abbreviations: C, canine, I1, first upper incisor; i1, first lower incisor, M1–M4, upper first to fourth molar; m1–m3, lower first to third molar, P3, upper third premolar; p3, lower third premolar." figureDoi="http://doi.org/10.5281/zenodo.10961556" httpUri="https://zenodo.org/record/10961556/files/figure.png" pageId="18" pageNumber="19">Figure 6</figureCitation>
|
||
) and 57337. In occlusal view, I1 is large, elongate, and laterally compressed. In lateral view, I1 is slightly recurved, with a low but distinct posterobuccal cusp present. The crown of I1 sits much higher than that of I2 and I3. In occlusal view, I2 is oval in shape but much longer than it is wide. The crown is completely flat from wear. I3 is shorter but wider than I2. I3 is distinctly bilobed buccally, with the anterior lobe being larger than the posterior lob. Lingually, the crown has been flattened by wear.
|
||
</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE021E5E7BAD749F9F5" blockId="18.[192,793,1033,1890]" pageId="18" pageNumber="19">The canines sit directly posterior to the incisors with no intervening diastemata. They are tall and reach a height equal to that of the crowns of I2-3. They are recurved and are slightly laterally inclined. A large diastema separates C1 from P3.</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE021E5E45AD570F8D5" blockId="18.[192,793,1033,1890]" pageId="18" pageNumber="19">In occlusal view, the occlusal margin of P3 is anterobuccally flexed and is out of alignment with M1. The P3 is blade-like, elliptical in shape, and tapered anteriorly. Six cuspules, each with associated lingual and buccal transcristae, are present along the occlusal margin. The posteriormost lingual and buccal transcristae are well developed as lateral blades, with a poorly developed posterolingual cusp.</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE0229DE2B5D07AFC68" blockId="18.[840,1441,166,1887]" pageId="18" pageNumber="19">
|
||
The M1 is bilophodont, low crowned and roughly rectangular in outline. The metaloph is narrower than the protoloph. However, both are subequal in height. The paracone is shorter but more massive than the protocone. A short, well-developed preparacrista extends from the paracone to the anterior end of the tooth where it meets a reduced anterior cingulum. The anterior cingulum is bordered lingually by the forelink (remnant of the preprotocrista). No precingulum is present. A shallow concavity is present on the anterior face of the paracone lingual to the preparacrista. The postparacrista extends from the paracone posteriorly towards the interloph valley on
|
||
<collectionCode id="618B2A89FD1BFFE025DFE055D145FDD7" box="[1216,1295,582,608]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
|
||
57340. The postparacrista is less developed on
|
||
<collectionCode id="618B2A89FD1BFFE025E8E075D10FFD37" box="[1271,1349,614,640]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
|
||
57337. A prominent blade-like convexity is evident buccal to the postparacrista which is interpreted as stylar cusp C. The postprotocrista is weakly developed but extends posteriorly to the interloph valley where it forms a midlink. The metacone is taller than the metaconule. A prominent premetacrista extends into the interloph valley where it meets the postparacrista. A well-developed postmetacrista extends posteriorly from the metacone to meet the postmetaconule crista. The neometaconule and postlink are absent.
|
||
</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE0229DE1F5D7D0FB08" blockId="18.[840,1441,166,1887]" pageId="18" pageNumber="19">
|
||
The M2 is similar in morphology to the M1 except as follows: it is larger; its protocone is taller; a long precingulum borders the anterior flank of the tooth from the forelink to the lingual side of the tooth; the premetacrista is less well developed; the postprotocrista and midlink are reduced on
|
||
<collectionCode id="618B2A89FD1BFFE0244DE695D1EAFB17" box="[1362,1440,1158,1184]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
|
||
57337.
|
||
</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE0229DE6D5D160FA88" blockId="18.[840,1441,166,1887]" pageId="18" pageNumber="19">The M3 is similar in morphology to the M2 except as follows: the preparacrista is more buccally situated; the postparacrista, premetacrista, postprotocrista, and midlink are reduced.</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE0229DE755D7ABF9A8" blockId="18.[840,1441,166,1887]" pageId="18" pageNumber="19">The M4 is similar in morphology to M3 except as follows: the metaloph is markedly shorter than the protoloph; the postparacrista is reduced; StC is absent; the postprotocrista and midlink are reduced; the forelink is more lingually situated, equalling the size of the anterior cingulum and precingulum.</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE02257E435D14BF968" blockId="18.[840,1441,166,1887]" pageId="18" pageNumber="19">
|
||
<emphasis id="35EE6E5EFD1BFFE02257E435D05CF988" bold="true" box="[840,1046,1574,1599]" pageId="18" pageNumber="19">Lower dentition.</emphasis>
|
||
The description of
|
||
<taxonomicName id="C09AC9CFFD1BFFE025E3E435D7D5F9E8" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="couperi" status="comb. nov.">
|
||
<emphasis id="35EE6E5EFD1BFFE025E3E435D7D5F9E8" italics="true" pageId="18" pageNumber="19">Ganawamaya couperi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2EDDD325FD1BFFE022B6E455D066F9E8" box="[937,1068,1606,1631]" pageId="18" pageNumber="19" rank="species">comb. nov.</taxonomicNameLabel>
|
||
is as stated by
|
||
<bibRefCitation id="630BCFBDFD1BFFE025F1E455D1EAF9E8" author="Cooke, B. N." box="[1262,1440,1606,1632]" journalOrPublisher="Memoirs of the Queensland Museum" pageId="18" pageNumber="19" pagination="269 - 280" part="41" refId="ref19388" refString="Cooke, B. N. 1997 b. Two new balbarine kangaroos and lower molar evolution within the subfamily. Memoirs of the Queensland Museum, 41: 269 - 280." title="Two new balbarine kangaroos and lower molar evolution within the subfamily" type="journal article" year="1997">Cooke (1997b)</bibRefCitation>
|
||
except as follows: the i1, which is preserved in
|
||
<collectionCode id="618B2A89FD1BFFE02257E495D7DCF917" box="[840,918,1670,1696]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP</collectionCode>
|
||
57334, is long and narrow and rises to the level of the molar row. A dorsal enamel flange is present but there is no ventral flange.
|
||
</paragraph>
|
||
<paragraph id="0725B24CFD1BFFE1229DE4F5D6FBFF55" blockId="18.[840,1441,166,1887]" lastBlockId="19.[192,793,168,1410]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
|
||
The protostylid of the m1is tall in
|
||
<specimenCode id="573C1A37FD1BFFE0244DE4F5D7DCF8A8" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP 10600</specimenCode>
|
||
, a juvenile specimen, and is also present although more worn on the holotype and
|
||
<specimenCode id="573C1A37FD1BFFE0244EE535D7DCF8E8" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="18" pageNumber="19" type="University or college">UCMP 57337</specimenCode>
|
||
. The portion of the missing protostylid is represented by a flat wear facet on
|
||
<specimenCode id="573C1A37FD1AFFE12375E2BBD75EFF75" box="[618,788,168,194]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="19" pageNumber="20" type="University or college">UCMP 57334</specimenCode>
|
||
, which represents an even older individual.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4F80E1C7FD1AFFE121DFE2FAD62EFBB5" pageId="19" pageNumber="20" type="discussion">
|
||
<paragraph id="0725B24CFD1AFFE121DFE2FAD62EFBB5" blockId="19.[192,793,168,1410]" pageId="19" pageNumber="20">
|
||
<emphasis id="35EE6E5EFD1AFFE121DFE2FAD572FEB5" bold="true" box="[192,312,233,258]" pageId="19" pageNumber="20">Remarks.</emphasis>
|
||
<bibRefCitation id="630BCFBDFD1AFFE12057E2FAD6C4FEB5" author="Flannery, T. & Rich, T. H. V." box="[328,654,233,258]" journalOrPublisher="Journal of Paleontology" pageId="19" pageNumber="20" pagination="418 - 447" part="60" refId="ref19628" refString="Flannery, T. and Rich, T. H. V. 1986. Macropodoids from the Middle Miocene Namba Formation, South Australia, and the homology of some dental structures in kangaroos. Journal of Paleontology, 60 (2): 418 - 447. https: // doi. org / 10.1017 / S 0022336000021958" title="Macropodoids from the Middle Miocene Namba Formation, South Australia, and the homology of some dental structures in kangaroos" type="journal article" year="1986">Flannery and Rich (1986)</bibRefCitation>
|
||
diagnosed species of the genus
|
||
<taxonomicName id="C09AC9CFFD1AFFE120D6E31AD60FFE95" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[457,581,265,290]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE120D6E31AD60FFE95" box="[457,581,265,290]" italics="true" pageId="19" pageNumber="20">Nambaroo</emphasis>
|
||
</taxonomicName>
|
||
as exhibiting the following features: well-developed lophs and lophids, m1 with a markedly compressed trigonid, a protostylid, posthypocristid on m1, and absence of a well-developed posterior cingulid on the lower molars. Our study demonstrates, in particular, that the protostylid is present in species of
|
||
<taxonomicName id="C09AC9CFFD1AFFE123B3E3DBD54BFDB5" authorityName="Cooke" authorityYear="1992" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE123B3E3DBD54BFDB5" italics="true" pageId="19" pageNumber="20">Ganawamaya</emphasis>
|
||
</taxonomicName>
|
||
as well as
|
||
<taxonomicName id="C09AC9CFFD1AFFE12097E3FAD64EFDB5" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[392,516,489,514]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE12097E3FAD64EFDB5" box="[392,516,489,514]" italics="true" pageId="19" pageNumber="20">Nambaroo</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="C09AC9CFFD1AFFE1230BE3FBD752FDB5" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" box="[532,792,488,514]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="couperi" status="comb. nov.">
|
||
<emphasis id="35EE6E5EFD1AFFE1230BE3FBD752FDB5" box="[532,792,488,514]" italics="true" pageId="19" pageNumber="20">Ganawamaya couperi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2EDDD325FD1AFFE121DFE01AD502FD95" box="[192,328,521,546]" pageId="19" pageNumber="20" rank="species">comb. nov.</taxonomicNameLabel>
|
||
differs from species of
|
||
<taxonomicName id="C09AC9CFFD1AFFE12366E01AD6BFFD95" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[633,757,521,546]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE12366E01AD6BFFD95" box="[633,757,521,546]" italics="true" pageId="19" pageNumber="20">Nambaroo</emphasis>
|
||
</taxonomicName>
|
||
in having a rectilinear rather than a third premolar with a plagiaulacoid form, a less developed preprotocrista and a postprotocrista that extends into the interloph valley on M1. These features are present in the specimen previously attributed by
|
||
<bibRefCitation id="630BCFBDFD1AFFE123D4E0BBD556FD55" author="Cooke, B. N." journalOrPublisher="Memoirs of the Queensland Museum" pageId="19" pageNumber="20" pagination="269 - 280" part="41" refId="ref19388" refString="Cooke, B. N. 1997 b. Two new balbarine kangaroos and lower molar evolution within the subfamily. Memoirs of the Queensland Museum, 41: 269 - 280." title="Two new balbarine kangaroos and lower molar evolution within the subfamily" type="journal article" year="1997">Cooke (1997b)</bibRefCitation>
|
||
to
|
||
<taxonomicName id="C09AC9CFFD1AFFE1205FE0DAD5F3FD55" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" box="[320,441,713,738]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="couperi">
|
||
<emphasis id="35EE6E5EFD1AFFE1205FE0DAD5F3FD55" box="[320,441,713,738]" italics="true" pageId="19" pageNumber="20">N. couperi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectionCode id="618B2A89FD1AFFE120D8E0DBD64CFD55" box="[455,518,712,738]" pageId="19" pageNumber="20">QMF</collectionCode>
|
||
30401) and for this reason we contend that
|
||
<taxonomicName id="C09AC9CFFD1AFFE120DDE0FBD617FCB5" authorityName="Butler & Travouillon & Price & Archer & Hand" authorityYear="2018" baseAuthorityName="Butler & Travouillon & Price & Archer & Hand" baseAuthorityYear="2018" box="[450,605,744,770]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="couperi" status="comb. nov.">
|
||
<emphasis id="35EE6E5EFD1AFFE120DDE0FBD617FCB5" box="[450,605,744,770]" italics="true" pageId="19" pageNumber="20">Gan. couperi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="2EDDD325FD1AFFE12377E0FAD6A8FCB5" box="[616,738,745,770]" pageId="19" pageNumber="20" rank="species">comb.nov.</taxonomicNameLabel>
|
||
is a species of
|
||
<taxonomicName id="C09AC9CFFD1AFFE1205CE11BD5ADFC95" authorityName="Cooke" authorityYear="1992" box="[323,487,776,802]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE1205CE11BD5ADFC95" box="[323,487,776,802]" italics="true" pageId="19" pageNumber="20">Ganawamaya</emphasis>
|
||
</taxonomicName>
|
||
rather than
|
||
<emphasis id="35EE6E5EFD1AFFE12363E11AD74BFC95" box="[636,769,777,802]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="C09AC9CFFD1AFFE12363E11AD6B6FC95" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[636,764,777,802]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">Nambaroo</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It is currently unclear whether the protostylid is lost due to wear in other species of
|
||
<taxonomicName id="C09AC9CFFD1AFFE12359E15AD688FCD5" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[582,706,841,866]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE12359E15AD688FCD5" box="[582,706,841,866]" italics="true" pageId="19" pageNumber="20">Nambaroo</emphasis>
|
||
</taxonomicName>
|
||
. Large upper canines are also preserved in
|
||
<collectionCode id="618B2A89FD1AFFE1236FE17BD6F4FC35" box="[624,702,872,898]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="19" pageNumber="20" type="University or college">UCMP</collectionCode>
|
||
57340. Although canine alveoli have been described for
|
||
<taxonomicName id="C09AC9CFFD1AFFE121DFE1BAD65DFC75" authority="(Kear et al. 2007)" baseAuthorityName="Kear" baseAuthorityYear="2007" box="[192,535,937,962]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE121DFE1BAD576FC75" box="[192,316,937,962]" italics="true" pageId="19" pageNumber="20">Nambaroo</emphasis>
|
||
(
|
||
<bibRefCitation id="630BCFBDFD1AFFE12053E1BAD658FC75" author="Kear, B. P. & Cooke, B. N. & Archer, M. & Flannery, T. F." box="[332,530,937,962]" journalOrPublisher="Journal of Paleontology" pageId="19" pageNumber="20" pagination="1147 - 1167" part="81" refId="ref19952" refString="Kear, B. P., Cooke, B. N., Archer, M., and Flannery, T. F. 2007. Implications of a new species of the Oligo-Miocene kangaroo (Marsupialia: Macropodoidea) Nambaroo, from the Riversleigh World Heritage Area, Queensland, Australia. Journal of Paleontology, 81 (6): 1147 - 1167. https: // doi. org / 10.1666 / 04 - 218.1" title="Implications of a new species of the Oligo-Miocene kangaroo (Marsupialia: Macropodoidea) Nambaroo, from the Riversleigh World Heritage Area, Queensland, Australia" type="journal article" year="2007">Kear et al. 2007</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="C09AC9CFFD1AFFE12349E1BBD6B0FC75" authorityName="Cooke" authorityYear="1992" box="[598,762,936,962]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="19" pageNumber="20" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD1AFFE12349E1BBD6B0FC75" box="[598,762,936,962]" italics="true" pageId="19" pageNumber="20">Ganawamaya</emphasis>
|
||
</taxonomicName>
|
||
in our study, this specimen is the first to be described in which the canines are preserved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4F80E1C7FD1AFFE121DFE61AD63EFA35" pageId="19" pageNumber="20" type="multiple">
|
||
<paragraph id="0725B24CFD1AFFE121DFE61AD63EFA35" blockId="19.[192,793,168,1410]" pageId="19" pageNumber="20">
|
||
<emphasis id="35EE6E5EFD1AFFE121DFE61AD587FB95" bold="true" box="[192,461,1033,1058]" pageId="19" pageNumber="20">Age and distribution.</emphasis>
|
||
<materialsCitation id="B7F2B811FD1AFFE120C9E61AD63AFA35" collectorName="Faunal Zone A" country="Australia" location="Queensland" pageId="19" pageNumber="20" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
|
||
The
|
||
<typeStatus id="D8210CEEFD1AFFE12313E61AD63BFB95" box="[524,625,1033,1058]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
is from White Hunter Site, Riversleigh WHA, northwestern
|
||
<collectingRegion id="C55E7CAEFD1AFFE121DFE65BD51EFBD5" box="[192,340,1096,1122]" country="Australia" name="Queensland" pageId="19" pageNumber="20">Queensland</collectingRegion>
|
||
,
|
||
<collectingCountry id="7F8DF2DCFD1AFFE1207BE65AD59AFBD5" box="[356,464,1097,1122]" name="Australia" pageId="19" pageNumber="20">Australia</collectingCountry>
|
||
. This site is part of Riversleigh’s Depositional Phase 1 and contains species interpreted to represent
|
||
<collectorName id="AA6FD79AFD1AFFE120ECE69AD6F8FB15" box="[499,690,1161,1186]" pageId="19" pageNumber="20">Faunal Zone A</collectorName>
|
||
assemblages which have been interpreted to be late Oligocene in age (
|
||
<bibRefCitation id="630BCFBDFD1AFFE1209CE6DAD63BFB55" author="Archer, M. & Godthelp, H. & Hand, S. J. & Megirian, D." box="[387,625,1225,1250]" journalOrPublisher="Australian Zoologist" pageId="19" pageNumber="20" pagination="29 - 65" part="25" refId="ref18573" refString="Archer, M., Godthelp, H., Hand, S. J., and Megirian, D. 1989. Fossil mammals of Riversleigh, northwestern Queensland: preliminary overview of biostratigraphy, correlation and environmental change. Australian Zoologist, 25 (2): 29 - 65. https: // doi. org / 10.7882 / AZ. 1989.001" title="Fossil mammals of Riversleigh, northwestern Queensland: preliminary overview of biostratigraphy, correlation and environmental change" type="journal article" year="1989">Archer et al., 1989</bibRefCitation>
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,
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<bibRefCitation id="630BCFBDFD1AFFE1239EE6DAD68AFB55" author="Archer, M. & Hand, S. J. & Godthelp, H. & Creaser, P." box="[641,704,1225,1250]" editor="Aguilar, J. P. & Legendre, S. & Michaux, J." journalOrPublisher="Institut de Montpellier" pageId="19" pageNumber="20" pagination="131 - 152" refId="ref18639" refString="Archer, M., Hand, S. J., Godthelp, H., and Creaser, P. 1997. Correlation of the Cainozoic sediments of the Riversleigh World Heritage Fossil Property, Queensland, Australia, p. 131 - 152. In Aguilar, J. P., Legendre, S., and Michaux, J. (eds.), Actes du Congr es BiochroM' 97, Memoires et Travaux. L'ecole Pratique des Hautes Etudes, Institut de Montpellier, Montpellier, France." title="Correlation of the Cainozoic sediments of the Riversleigh World Heritage Fossil Property, Queensland, Australia" type="journal article" volumeTitle="Actes du Congr es BiochroM' 97, Memoires et Travaux. L'ecole Pratique des Hautes Etudes" year="1997">1997</bibRefCitation>
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;
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<bibRefCitation id="630BCFBDFD1AFFE123CFE6DAD5DEFAB5" author="Myers, T. & Archer, M." journalOrPublisher="Memoirs of the Queensland Museum" pageId="19" pageNumber="20" pagination="379 - 392" part="41" refId="ref20353" refString="Myers, T. and Archer, M. 1997. Kuterintja ngama (Marsupialia, Ilariidae): a revised systematic analysis based on material from the late Oligocene of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum, 41: 379 - 392." title="Kuterintja ngama (Marsupialia, Ilariidae): a revised systematic analysis based on material from the late Oligocene of Riversleigh, northwestern Queensland" type="journal article" year="1997">Myers and Archer 1997</bibRefCitation>
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;
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<bibRefCitation id="630BCFBDFD1AFFE120BAE6FAD682FAB5" author="Travouillon, K. J. & Archer, M. & Hand, S. J. & Godthelp, H." box="[421,712,1257,1282]" journalOrPublisher="Alcheringa: An Australasian Journal of Palaeontology, 30 (supplement" pageId="19" pageNumber="20" pagination="323 - 349" part="1" refId="ref20674" refString="Travouillon, K. J., Archer, M., Hand, S. J., and Godthelp, H. 2006. Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, northwestern Queensland. Alcheringa: An Australasian Journal of Palaeontology, 30 (supplement 1): 323 - 349. https: // doi. org / 10.1080 / 03115510608619590" title="Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, northwestern Queensland" type="journal article" year="2006">Travouillon et al., 2006</bibRefCitation>
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,
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<bibRefCitation id="630BCFBDFD1AFFE123C8E6FAD75EFAB5" author="Travouillon, K. J. & Escarguel, G. & Legendre, S. & Archer, M. & Hand, S. J." box="[727,788,1257,1282]" journalOrPublisher="Paleobiology" pageId="19" pageNumber="20" pagination="696 - 709" part="37" refId="ref20827" refString="Travouillon, K. J., Escarguel, G., Legendre, S., Archer, M., and Hand, S. J. 2011. The use of MSR (Minimum Sample Richness) for sample assemblage comparisons. Paleobiology, 37 (4): 696 - 709. https: // doi. org / 10.1666 / 09050.1" title="The use of MSR (Minimum Sample Richness) for sample assemblage comparisons" type="journal article" year="2011">2011</bibRefCitation>
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;
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<bibRefCitation id="630BCFBDFD1AFFE121DFE71AD5E0FA95" author="Arena, D. A. & Travouillon, K. J. & Beck, R. M. D. & Black, K. & Gillespie, A. K. & Myers, T. J. & Archer, M. & Hand, S. J." box="[192,426,1289,1314]" journalOrPublisher="Lethaia" pageId="19" pageNumber="20" pagination="43 - 60" part="49" refId="ref18734" refString="Arena, D. A., Travouillon, K. J., Beck, R. M. D., Black, K., Gillespie, A. K., Myers, T. J., Archer, M., and Hand, S. J. 2015. Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia. Lethaia, 49: 43 - 60. https: // doi. org / 10.1111 / let. 12131" title="Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia" type="journal article" year="2015">Arena et al., 2015</bibRefCitation>
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). The referred specimens are from the Ngapakaldi Quarry of the Etadunna Formation is interpreted to be late Oligocene (24.6 Ma) in age (
|
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<bibRefCitation id="630BCFBDFD1AFFE12054E77AD623FA35" author="Woodburne, M. O. & Macfadden, B. J. & Case, J. A. & Springer, M. S. & Pledge, N. S. & Power, J. D. & Woodburne, J. M. & Springer, K. B." box="[331,617,1385,1410]" journalOrPublisher="Journal of Vertebrate Paleontology" pageId="19" pageNumber="20" pagination="483 - 515" part="13" refId="ref21097" refString="Woodburne, M. O., Macfadden, B. J., Case, J. A., Springer, M. S., Pledge, N. S., Power, J. D., Woodburne, J. M., and Springer, K. B. 1993. Land mammal biostratigraphy and magnetostratigraphy of the Etadunna Formation (late Oligocene) of South Australia. Journal of Vertebrate Paleontology, 13 (4): 483 - 515. https: // doi. org / 10.1080 / 02724634.1994.10011527" title="Land mammal biostratigraphy and magnetostratigraphy of the Etadunna Formation (late Oligocene) of South Australia" type="journal article" year="1993">Woodburne et al., 1993</bibRefCitation>
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||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
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</treatment>
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</document> |