191 lines
21 KiB
XML
191 lines
21 KiB
XML
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<mods:title id="A543F7EC056248C99E627A399C6B83EC">Revision of Oligo-Miocene kangaroos, Ganawamaya and Nambaroo (Marsupialia: Macropodiformes, Balbaridae)</mods:title>
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<mods:namePart id="A1B5A99EF232AC9400A7E9CCB54418D5">Butler, Kaylene</mods:namePart>
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<mods:namePart id="F131DF12F901CBAFC554CE07D4BB39ED">Travouillon, Kenny J.</mods:namePart>
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<mods:namePart id="BDA4AD6771CAF9717A24ABE8294E50AD">Price, Gilbert J.</mods:namePart>
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<mods:namePart id="C89D6E4F23FEE15526FEF1CD03F48D48">Archer, Michael</mods:namePart>
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<mods:namePart id="3ED74B7CEB990AF57989AA02A7C01F46">Hand, Suzanne J.</mods:namePart>
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<mods:title id="A0709F8F3EB373AF8E883D6AEF653191">Palaeontologia Electronica</mods:title>
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<mods:date id="37B15151B1B8BC380695EC2BA2F73544">2018</mods:date>
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<mods:title id="DB800CD82A2D97D3409B2DDFAA60828D">8 A</mods:title>
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<subSubSection id="4F80E1C7FD0AFFF1200BE35BD68EFED5" box="[276,708,328,354]" pageId="3" pageNumber="4" type="nomenclature">
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<paragraph id="0725B24CFD0AFFF1200BE35BD68EFED5" blockId="3.[195,786,168,354]" box="[276,708,328,354]" pageId="3" pageNumber="4">
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Genus
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<taxonomicName id="C09AC9CFFD0AFFF12075E35BD68EFED5" authority="Cooke, 1992" authorityName="Cooke" authorityYear="1992" box="[362,708,328,354]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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GANAWAMAYA
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<bibRefCitation id="630BCFBDFD0AFFF12332E35BD68EFED5" author="Cooke, B. N." box="[557,708,328,354]" journalOrPublisher="Alcheringa" pageId="3" pageNumber="4" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke, 1992</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD0AFFF121DFE360D580FE1B" pageId="3" pageNumber="4" type="type_taxon">
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<paragraph id="0725B24CFD0AFFF121DFE360D580FE1B" blockId="3.[192,793,371,1900]" pageId="3" pageNumber="4">
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<emphasis id="35EE6E5EFD0AFFF121DFE360D53EFE3A" bold="true" box="[192,372,371,397]" pageId="3" pageNumber="4">Type Species.</emphasis>
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<taxonomicName id="C09AC9CFFD0AFFF1209FE360D75EFE3B" authority="Cooke, 1992" authorityName="Cooke" authorityYear="1992" box="[384,788,371,397]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="acris">
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<emphasis id="35EE6E5EFD0AFFF1209FE360D623FE3B" box="[384,617,371,397]" italics="true" pageId="3" pageNumber="4">Ganawamaya acris</emphasis>
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<bibRefCitation id="630BCFBDFD0AFFF1236AE360D75EFE3B" author="Cooke, B. N." box="[629,788,371,397]" journalOrPublisher="Alcheringa" pageId="3" pageNumber="4" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke, 1992</bibRefCitation>
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</taxonomicName>
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, by original description.
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD0AFFF121DFE3A0D64CFC9B" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph id="0725B24CFD0AFFF121DFE3A0D64CFC9B" blockId="3.[192,793,371,1900]" pageId="3" pageNumber="4">
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<emphasis id="35EE6E5EFD0AFFF121DFE3A0D629FE7B" bold="true" box="[192,611,435,461]" pageId="3" pageNumber="4">Emended Generic Diagnosis.</emphasis>
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Species of
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<taxonomicName id="C09AC9CFFD0AFFF121DFE3C0D52EFE5A" authorityName="Cooke" authorityYear="1992" box="[192,356,467,493]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF121DFE3C0D52EFE5A" box="[192,356,467,493]" italics="true" pageId="3" pageNumber="4">Ganawamaya</emphasis>
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</taxonomicName>
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differ from all other balbarids in having the following unique combination of features: a poorly developed hypocingulid; a lack of a neometaconule and postlink; a short buccally flexed p3 with five cuspids along the occlusal margin; pronounced lingual cingulum on P3; a well-developed posterobuccal transcrista on P3; a rectilinear P3; a poorly developed preprotocrista; a postprotocrista that extends into the interloph valley; a pronounced nuchal crest; and well-developed postorbital processes.
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</paragraph>
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</subSubSection>
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<subSubSection id="4F80E1C7FD0AFFF121DFE120D160FD95" pageId="3" pageNumber="4" type="discussion">
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<paragraph id="0725B24CFD0AFFF121DFE120D58DFA5B" blockId="3.[192,793,371,1900]" pageId="3" pageNumber="4">
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<emphasis id="35EE6E5EFD0AFFF121DFE120D572FCFB" bold="true" box="[192,312,819,844]" pageId="3" pageNumber="4">Remarks.</emphasis>
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While species of
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<taxonomicName id="C09AC9CFFD0AFFF12316E120D6E7FCFA" authorityName="Cooke" authorityYear="1992" box="[521,685,819,845]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF12316E120D6E7FCFA" box="[521,685,819,845]" italics="true" pageId="3" pageNumber="4">Ganawamaya</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="C09AC9CFFD0AFFF123F6E120D54EFCDB" class="Mammalia" family="Balbaridae" genus="Balbaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF123F6E120D54EFCDB" italics="true" pageId="3" pageNumber="4">Balbaroo</emphasis>
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</taxonomicName>
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all have a pronounced nuchal crest, it is less developed in
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<taxonomicName id="C09AC9CFFD0AFFF12077E160D646FC3A" authorityName="Cooke" authorityYear="1992" box="[360,524,883,909]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF12077E160D646FC3A" box="[360,524,883,909]" italics="true" pageId="3" pageNumber="4">Ganawamaya</emphasis>
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</taxonomicName>
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. However, it is better developed than in several macropodiform groups (e.g., macropodines). Species of
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<taxonomicName id="C09AC9CFFD0AFFF1236BE1A0D752FC7A" authorityName="Cooke" authorityYear="1992" box="[628,792,947,973]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF1236BE1A0D752FC7A" box="[628,792,947,973]" italics="true" pageId="3" pageNumber="4">Ganawamaya</emphasis>
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</taxonomicName>
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also have a more gracile cranium and smaller molars than those of
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<taxonomicName id="C09AC9CFFD0AFFF1231FE1E0D621FBBB" box="[512,619,1011,1036]" class="Mammalia" family="Balbaridae" genus="Balbaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF1231FE1E0D621FBBB" box="[512,619,1011,1036]" italics="true" pageId="3" pageNumber="4">Balbaroo</emphasis>
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</taxonomicName>
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. Species of
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<taxonomicName id="C09AC9CFFD0AFFF121DFE600D52EFB9A" authorityName="Cooke" authorityYear="1992" box="[192,356,1043,1069]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF121DFE600D52EFB9A" box="[192,356,1043,1069]" italics="true" pageId="3" pageNumber="4">Ganawamaya</emphasis>
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</taxonomicName>
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differ from species of
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<taxonomicName id="C09AC9CFFD0AFFF12363E600D6B2FB9B" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[636,760,1043,1068]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF12363E600D6B2FB9B" box="[636,760,1043,1068]" italics="true" pageId="3" pageNumber="4">Nambaroo</emphasis>
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</taxonomicName>
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in having a rectilinear p3 as opposed to a more plagiaulacoid form, a less developed preprotocrista, and a postprotocrista on the M1 that extends into the interloph valley and in lacking a neometaconule and postlink. Herein ‘plagiaulacoid’ is used in this study to refer to a shorter tooth with a recurved crown and many cuspids and transcristae, while ‘rectilinear’ refers to a tooth with a straight crown and less cuspids and transcristae than premolars with a plagiaulacoid form. For example, the p3 of
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<collectionCode id="618B2A89FD0AFFF121DFE760D4B1FA3B" box="[192,251,1395,1420]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34978" lsid="urn:lsid:biocol.org:col:34978" name="Museum Victoria" pageId="3" pageNumber="4" type="Museum">NMV</collectionCode>
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P157559, referred to
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<taxonomicName id="C09AC9CFFD0AFFF120E2E760D6CCFA3B" box="[509,646,1395,1420]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="tarrinyeri">
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<emphasis id="35EE6E5EFD0AFFF120E2E760D6CCFA3B" box="[509,646,1395,1420]" italics="true" pageId="3" pageNumber="4">N. tarrinyeri</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="630BCFBDFD0AFFF123AFE760D5C9FA1B" author="Flannery, T. & Rich, T. H. V." journalOrPublisher="Journal of Paleontology" pageId="3" pageNumber="4" pagination="418 - 447" part="60" refId="ref19628" refString="Flannery, T. and Rich, T. H. V. 1986. Macropodoids from the Middle Miocene Namba Formation, South Australia, and the homology of some dental structures in kangaroos. Journal of Paleontology, 60 (2): 418 - 447. https: // doi. org / 10.1017 / S 0022336000021958" title="Macropodoids from the Middle Miocene Namba Formation, South Australia, and the homology of some dental structures in kangaroos" type="journal article" year="1986">Flannery and Rich (1986)</bibRefCitation>
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, has a recurved crown with seven cuspids with associated transcristae anterior to the large posterior cuspid.
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</paragraph>
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<paragraph id="0725B24CFD0AFFF121E5E7E0D160FD95" blockId="3.[192,793,371,1900]" lastBlockId="3.[840,1441,168,546]" pageId="3" pageNumber="4">
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In this study, we also examined the
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<typeStatus id="D8210CEEFD0AFFF123ACE7E0D752F9BB" box="[691,792,1523,1548]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
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and sole specimen of
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<taxonomicName id="C09AC9CFFD0AFFF120FCE400D485F9FB" authority=", QM F" authorityName="QM F" class="Mammalia" family="Balbaridae" genus="Galanarla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="tessellata">
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<emphasis id="35EE6E5EFD0AFFF120FCE400D697F99B" box="[483,733,1555,1580]" italics="true" pageId="3" pageNumber="4">Galanarla tessellata,</emphasis>
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QM F
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</taxonomicName>
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10644. The distinguishing features of
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<taxonomicName id="C09AC9CFFD0AFFF1238FE420D4A0F9DB" class="Mammalia" family="Balbaridae" genus="Galanarla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="tessellata">
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<emphasis id="35EE6E5EFD0AFFF1238FE420D4A0F9DB" italics="true" pageId="3" pageNumber="4">Gal. tessellata</emphasis>
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</taxonomicName>
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proposed by
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<bibRefCitation id="630BCFBDFD0AFFF12090E440D6DAF9DB" author="Flannery, T. & Archer, M. & Plane, M." box="[399,656,1619,1644]" journalOrPublisher="BMR Journal of Australian Geology & Geophysics" pageId="3" pageNumber="4" pagination="287 - 302" part="7" refId="ref19571" refString="Flannery, T., Archer, M., and Plane, M. 1982. Middle Miocene kangaroos (Macropodoidea, Marsupialia) from three localities in northern Australian, with a description of two new subfamilies. BMR Journal of Australian Geology & Geophysics, 7 (4): 287 - 302." title="Middle Miocene kangaroos (Macropodoidea, Marsupialia) from three localities in northern Australian, with a description of two new subfamilies" type="journal article" year="1982">Flannery et al. (1982)</bibRefCitation>
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are shared with species of
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<taxonomicName id="C09AC9CFFD0AFFF12083E460D652F93B" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[412,536,1651,1676]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
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<emphasis id="35EE6E5EFD0AFFF12083E460D652F93B" box="[412,536,1651,1676]" italics="true" pageId="3" pageNumber="4">Nambaroo</emphasis>
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</taxonomicName>
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and
|
||
<taxonomicName id="C09AC9CFFD0AFFF12373E460D75AF93A" authorityName="Cooke" authorityYear="1992" box="[620,784,1651,1677]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="genus">
|
||
<emphasis id="35EE6E5EFD0AFFF12373E460D75AF93A" box="[620,784,1651,1677]" italics="true" pageId="3" pageNumber="4">Ganawamaya</emphasis>
|
||
</taxonomicName>
|
||
. Those include: lophodont molars, a convex ventral margin of the dentary, a small buccal opening for the masseteric canal, a p3 flexed buccally out of alignment with the molar row. The
|
||
<typeStatus id="D8210CEEFD0AFFF12343E4E0D68BF8BB" box="[604,705,1779,1804]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="C09AC9CFFD0AFFF123F8E4E0D57AF89B" class="Mammalia" family="Balbaridae" genus="Galanarla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="tessellata">
|
||
<emphasis id="35EE6E5EFD0AFFF123F8E4E0D57AF89B" italics="true" pageId="3" pageNumber="4">Gal. tessellata</emphasis>
|
||
</taxonomicName>
|
||
is a heavily damaged jaw with two very worn molars. It is closest to
|
||
<taxonomicName id="C09AC9CFFD0AFFF12309E520D68BF8FB" box="[534,705,1843,1868]" pageId="3" pageNumber="4" rank="species" species="aediculis">
|
||
<emphasis id="35EE6E5EFD0AFFF12309E520D68BF8FB" box="[534,705,1843,1868]" italics="true" pageId="3" pageNumber="4">Gan. aediculis</emphasis>
|
||
</taxonomicName>
|
||
in size and morphology. However, because most diagnostic features distinguishing balbarids (those of p3 and m1) are not preserved, its relationship to other balbarid taxa cannot, at present, be determined. However, the type locality for
|
||
<emphasis id="35EE6E5EFD0AFFF125BDE31BD11BFE95" box="[1186,1361,264,290]" italics="true" pageId="3" pageNumber="4">
|
||
<taxonomicName id="C09AC9CFFD0AFFF125BDE31BD107FE95" box="[1186,1357,264,290]" class="Mammalia" family="Balbaridae" genus="Galanarla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="tessellata">Gal. tessellata</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Riversleigh’s D Site, is no longer accessible for fossil collection. Additionally, more informative specimens may in the future be recovered from other Depositional Phase 1 deposits such as Hiatus and White Hunter Sites (e.g.,
|
||
<bibRefCitation id="630BCFBDFD0AFFF1259FE3BAD12EFE75" author="Arena, D. A. & Travouillon, K. J. & Beck, R. M. D. & Black, K. & Gillespie, A. K. & Myers, T. J. & Archer, M. & Hand, S. J." box="[1152,1380,425,450]" journalOrPublisher="Lethaia" pageId="3" pageNumber="4" pagination="43 - 60" part="49" refId="ref18734" refString="Arena, D. A., Travouillon, K. J., Beck, R. M. D., Black, K., Gillespie, A. K., Myers, T. J., Archer, M., and Hand, S. J. 2015. Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia. Lethaia, 49: 43 - 60. https: // doi. org / 10.1111 / let. 12131" title="Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia" type="journal article" year="2015">Arena et al., 2015</bibRefCitation>
|
||
). We suggest
|
||
<taxonomicName id="C09AC9CFFD0AFFF122AEE3DBD012FE55" box="[945,1112,456,482]" pageId="3" pageNumber="4" rank="species" species="tesselata">
|
||
<emphasis id="35EE6E5EFD0AFFF122AEE3DBD012FE55" box="[945,1112,456,482]" italics="true" pageId="3" pageNumber="4">Gal. tesselata</emphasis>
|
||
</taxonomicName>
|
||
might be considered as a
|
||
<emphasis id="35EE6E5EFD0AFFF12257E3FAD7B6FDB5" box="[840,1020,489,514]" italics="true" pageId="3" pageNumber="4">nomen dubium</emphasis>
|
||
as the
|
||
<typeStatus id="D8210CEEFD0AFFF1257EE3FAD08CFDB5" box="[1121,1222,489,514]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
specimen cannot be confidently ascribed to a single taxon.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |