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<document id="D23BF8DE097549902BF9FFAC4DDF61E0" ID-DOI="10.26879/747" ID-ISSN="1094-8074" ID-Zenodo-Dep="10961544" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1712855360717" checkinUser="felipe" docAuthor="Butler, Kaylene, Travouillon, Kenny J., Price, Gilbert J., Archer, Michael &amp; Hand, Suzanne J." docDate="2018" docId="8F33035AFD05FFE22284E403D758F9CF" docLanguage="en" docName="PalaeontolElectron.21.1.8A.pdf" docOrigin="Palaeontologia Electronica (8 A) 21 (1)" docSource="http://dx.doi.org/10.26879/747" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.3:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="3" docTitle="Ganawamaya aediculis Cooke 1992" docType="treatment" docVersion="5" lastPageNumber="17" masterDocId="730A7B22FD09FFF2211FE213D44AFFB7" masterDocTitle="Revision of Oligo-Miocene kangaroos, Ganawamaya and Nambaroo (Marsupialia: Macropodiformes, Balbaridae)" masterLastPageNumber="58" masterPageNumber="1" pageNumber="13" updateTime="1714000781735" updateUser="carolina" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="E76F5692234E405332480DB8B352AFA4">Revision of Oligo-Miocene kangaroos, Ganawamaya and Nambaroo (Marsupialia: Macropodiformes, Balbaridae)</mods:title>
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<mods:namePart id="8B4CDCFD9FD770FA4A3571DFD05366E0">Butler, Kaylene</mods:namePart>
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<mods:namePart id="907FB8A1700DA0D4626E4BA90203856E">Travouillon, Kenny J.</mods:namePart>
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<mods:namePart id="06F4EAEC9FB9DB2BBCD21F1899DBE6FF">Price, Gilbert J.</mods:namePart>
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<mods:date id="A8FBDD573A6A921688E7F187D143E5B4">2018</mods:date>
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<mods:title id="1BD1293F28DED2B98ECE79A5CD613C57">8 A</mods:title>
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<treatment id="8F33035AFD05FFE22284E403D758F9CF" LSID="urn:lsid:plazi:treatment:8F33035AFD05FFE22284E403D758F9CF" httpUri="http://treatment.plazi.org/id/8F33035AFD05FFE22284E403D758F9CF" lastPageId="16" lastPageNumber="17" pageId="12" pageNumber="13">
<subSubSection id="4F80E1C7FD05FFFE2284E403D107F99D" box="[923,1357,1552,1578]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="0725B24CFD05FFFE2284E403D107F99D" blockId="12.[923,1357,1552,1610]" box="[923,1357,1552,1578]" pageId="12" pageNumber="13">
<heading id="5C6D0520FD05FFFE2284E403D107F99D" box="[923,1357,1552,1578]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="13" reason="7">
<taxonomicName id="C09AC9CFFD05FFFE2284E403D107F99D" authority="Cooke, 1992" authorityName="Cooke" authorityYear="1992" box="[923,1357,1552,1578]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD05FFFE2284E403D0E5F99D" box="[923,1199,1552,1578]" italics="true" pageId="12" pageNumber="13">Ganawamaya aediculis</emphasis>
<bibRefCitation id="630BCFBDFD05FFFE25A9E403D107F99D" author="Cooke, B. N." box="[1206,1357,1552,1578]" journalOrPublisher="Alcheringa" pageId="12" pageNumber="13" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke, 1992</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4F80E1C7FD05FFFE255CE422D0EFF9FD" box="[1091,1189,1585,1610]" pageId="12" pageNumber="13" type="description">
<paragraph id="0725B24CFD05FFFE255CE422D0EFF9FD" blockId="12.[923,1357,1552,1610]" box="[1091,1189,1585,1610]" pageId="12" pageNumber="13">
<figureCitation id="9FA1AEC9FD05FFFE255CE422D0EFF9FD" box="[1091,1189,1585,1610]" captionStart="FIGURE 4" captionStartId="14.[197,288,1466,1489]" captionTargetBox="[222,1429,187,1416]" captionTargetId="figure-128@14.[199,1434,162,1449]" captionTargetPageId="14" captionText="FIGURE 4. Ganawamaya aediculis partial cranium (QM F58658) in ventral view (1) and left lateral view (2). Abbreviations: I1, first upper incisor; ioc, infraorbital canal; iof, infraorbital foramen; M1M4, upper first to fourth molar; mx, maxilla; P3, upper third premolar; pmx, premaxilla." figureDoi="http://doi.org/10.5281/zenodo.10961552" httpUri="https://zenodo.org/record/10961552/files/figure.png" pageId="12" pageNumber="13">Figure 4</figureCitation>
</paragraph>
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<paragraph id="0725B24CFD05FFFE2257E448D7DEF963" blockId="12.[840,1441,1627,1909]" pageId="12" pageNumber="13">
<materialsCitation id="B7F2B811FD05FFFE2257E448D7DEF963" collectionCode="QM" country="Australia" location="White Hunter Site" municipality="Riversleigh" pageId="12" pageNumber="13" specimenCode="F16843" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="35EE6E5EFD05FFFE2257E448D78BF9C3" bold="true" box="[840,961,1627,1652]" pageId="12" pageNumber="13">
<typeStatus id="D8210CEEFD05FFFE2257E448D7F7F9C3" box="[840,957,1627,1652]" pageId="12" pageNumber="13" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="618B2A89FD05FFFE22D5E448D7BFF9C2" box="[970,1013,1627,1653]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="12" pageNumber="13" type="Museum">QM</collectionCode>
<specimenCode id="573C1A37FD05FFFE22E1E448D017F9C3" box="[1022,1117,1627,1652]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="12" pageNumber="13" type="Museum">F16843</specimenCode>
, right dentary with p3, m1 m4 from
<location id="0245E497FD05FFFE22D9E468D0FAF922" LSID="urn:lsid:plazi:treatment:8F33035AFD05FFE22284E403D758F9CF:0245E497FD05FFFE22D9E468D0FAF922" box="[966,1200,1659,1685]" country="Australia" municipality="Riversleigh" name="White Hunter Site" pageId="12" pageNumber="13" stateProvince="Queensland">White Hunter Site</location>
,
<collectingMunicipality id="E7412836FD05FFFE25DAE468D103F923" box="[1221,1353,1659,1684]" pageId="12" pageNumber="13">Riversleigh</collectingMunicipality>
WHA, northwestern
<collectingRegion id="C55E7CAEFD05FFFE251CE488D0DDF902" box="[1027,1175,1691,1717]" country="Australia" name="Queensland" pageId="12" pageNumber="13">Queensland</collectingRegion>
,
<collectingCountry id="7F8DF2DCFD05FFFE25A5E488D168F903" box="[1210,1314,1691,1716]" name="Australia" pageId="12" pageNumber="13">Australia</collectingCountry>
(
<bibRefCitation id="630BCFBDFD05FFFE2455E488D7C3F963" author="Cooke, B. N." journalOrPublisher="Alcheringa" pageId="12" pageNumber="13" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke, 1992</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="0725B24CFD05FFFF2257E4C8D165F9CD" blockId="12.[840,1441,1627,1909]" lastBlockId="13.[840,1440,1601,1914]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<emphasis id="35EE6E5EFD05FFFE2257E4C8D01EF943" bold="true" box="[840,1108,1755,1780]" pageId="12" pageNumber="13">Referred specimens.</emphasis>
Gillespie's Gully Site:
<collectionCode id="618B2A89FD05FFFE246AE4C8D1EAF942" box="[1397,1440,1755,1781]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="12" pageNumber="13" type="Museum">QM</collectionCode>
F30076, right dentary with i1, p3, m1m4. LSO Site:
<collectionCode id="618B2A89FD05FFFE2295E508D7FFF882" box="[906,949,1819,1845]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="12" pageNumber="13" type="Museum">QM</collectionCode>
F58658, right maxilla with P3, M1M4;
<collectionCode id="618B2A89FD05FFFE2257E528D739F8E2" box="[840,883,1851,1877]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="12" pageNumber="13" type="Museum">QM</collectionCode>
F30298, right dentary with p3, m1m4;
<collectionCode id="618B2A89FD05FFFE246AE528D1EAF8E2" box="[1397,1440,1851,1877]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="12" pageNumber="13" type="Museum">QM</collectionCode>
F31463, left dentary with p3. White Hunter Site:
<collectionCode id="618B2A89FD04FFFF21DFE452D4A1F9EC" box="[192,235,1601,1627]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F58660, left dentary with p3, m1m4;
<collectionCode id="618B2A89FD04FFFF23F2E452D752F9EC" box="[749,792,1601,1627]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F58661, isolated m2m3;
<collectionCode id="618B2A89FD04FFFF20EBE472D655F9CC" box="[500,543,1633,1659]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F58662, isolated p3;
<collectionCode id="618B2A89FD04FFFF21DFE492D4A1F92C" box="[192,235,1665,1691]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F19584, left dentary with p3, m1, m
<quantity id="C0621FA9FD04FFFF23BFE492D684F92D" box="[672,718,1665,1690]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="14" unit="in" value="3.0">3 in</quantity>
crypt;
<collectionCode id="618B2A89FD04FFFF21DFE4B2D4A1F90C" box="[192,235,1697,1723]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F19605, right dentary with m4;
<collectionCode id="618B2A89FD04FFFF2360E4B2D6E0F90C" box="[639,682,1697,1723]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F19876, left dentary with m2m4;
<collectionCode id="618B2A89FD04FFFF20ECE4D2D654F96C" box="[499,542,1729,1755]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F19878, left dentary with dp3, unerupted p3, m1;
<collectionCode id="618B2A89FD04FFFF2311E4F2D673F94C" box="[526,569,1761,1787]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F19993, right dentary with m3, broken m4;
<collectionCode id="618B2A89FD04FFFF20EBE512D655F8AC" box="[500,543,1793,1819]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F19994, left dentary with m
<quantity id="C0621FA9FD04FFFF2012E532D573F88D" box="[269,313,1825,1850]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="13" pageNumber="14" unit="in" value="3.0">3 in</quantity>
crypt;
<collectionCode id="618B2A89FD04FFFF2095E532D5FFF88C" box="[394,437,1825,1851]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F20146, right isolated broken m2, m3;
<collectionCode id="618B2A89FD04FFFF2038E552D518F8EC" box="[295,338,1857,1883]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F20633, Left maxilla with m2m4;
<collectionCode id="618B2A89FD04FFFF23F2E552D752F8EC" box="[749,792,1857,1883]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F23354, left maxilla with M1M2, broken M3;
<collectionCode id="618B2A89FD04FFFF23F2E572D752F8CC" box="[749,792,1889,1915]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F31182, left dentary with p3, m1m3, broken m4;
<collectionCode id="618B2A89FD04FFFF2257E472D739F9CC" box="[840,883,1633,1659]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="13" pageNumber="14" type="Museum">QM</collectionCode>
F57791, left maxilla with P3, M1M2.
</paragraph>
</subSubSection>
<caption id="53E5E2C4FD04FFFF21DCE773D552F9BD" ID-DOI="http://doi.org/10.5281/zenodo.10961550" ID-Zenodo-Dep="10961550" httpUri="https://zenodo.org/record/10961550/files/figure.png" pageId="13" pageNumber="14" startId="13.[195,286,1376,1399]" targetBox="[221,1431,175,1304]" targetPageId="13" targetType="figure">
<paragraph id="0725B24CFD04FFFF21DCE773D552F9BD" blockId="13.[195,1439,1376,1546]" pageId="13" pageNumber="14">
<emphasis id="35EE6E5EFD04FFFF21DCE773D572FAC0" bold="true" box="[195,312,1376,1399]" pageId="13" pageNumber="14">FIGURE 3.</emphasis>
Occlusal view of lower dentary and line drawing of first molar for juvenile (
<emphasis id="35EE6E5EFD04FFFF255BE773D01BFAC0" bold="true" box="[1092,1105,1376,1399]" pageId="13" pageNumber="14">1</emphasis>
), adult (
<emphasis id="35EE6E5EFD04FFFF25B6E773D0FCFAC0" bold="true" box="[1193,1206,1376,1399]" pageId="13" pageNumber="14">2</emphasis>
), worn adult (
<emphasis id="35EE6E5EFD04FFFF2458E773D11EFAC0" bold="true" box="[1351,1364,1376,1399]" pageId="13" pageNumber="14">3</emphasis>
) specimens of a single species from Faunal Zone B; and line drawings for juvenile m1 (
<emphasis id="35EE6E5EFD04FFFF2538E76ED07EFA23" bold="true" box="[1063,1076,1405,1428]" pageId="13" pageNumber="14">4</emphasis>
), adult m1 (
<emphasis id="35EE6E5EFD04FFFF25AAE76ED088FA23" bold="true" box="[1205,1218,1405,1428]" pageId="13" pageNumber="14">5</emphasis>
), and worn adult m1 (
<emphasis id="35EE6E5EFD04FFFF21D4E788D492FA05" bold="true" box="[203,216,1435,1458]" pageId="13" pageNumber="14">6</emphasis>
). The specimens were previously attributed to
<emphasis id="35EE6E5EFD04FFFF23C4E788D6A2FA05" bold="true" box="[731,744,1435,1458]" pageId="13" pageNumber="14">1</emphasis>
,
<taxonomicName id="C09AC9CFFD04FFFF23E7E788D7E0FA05" box="[760,938,1435,1458]" class="Mammalia" family="Balbaridae" genus="Nambaroo" kingdom="Animalia" order="Diprotodontia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="undefined-4">
<emphasis id="35EE6E5EFD04FFFF23E7E788D722FA05" box="[760,872,1435,1458]" italics="true" pageId="13" pageNumber="14">Nambaroo</emphasis>
sp. 4
</taxonomicName>
;
<emphasis id="35EE6E5EFD04FFFF22A8E788D78EFA05" bold="true" box="[951,964,1435,1458]" pageId="13" pageNumber="14">2</emphasis>
,
<taxonomicName id="C09AC9CFFD04FFFF22CBE788D0C9FA05" box="[980,1155,1435,1458]" class="Mammalia" family="Balbaridae" genus="Nambaroo" kingdom="Animalia" order="Diprotodontia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="undefined-2">
<emphasis id="35EE6E5EFD04FFFF22CBE788D00EFA05" box="[980,1092,1435,1458]" italics="true" pageId="13" pageNumber="14">Nambaroo</emphasis>
sp. 2
</taxonomicName>
by
<bibRefCitation id="630BCFBDFD04FFFF25B0E789D105FA05" author="Cooke, B. N." box="[1199,1359,1434,1458]" journalOrPublisher="Memoirs of the Queensland Museum" pageId="13" pageNumber="14" pagination="281 - 294" part="41" refId="ref19421" refString="Cooke, B. N. 1997 c. New Miocene bulungamayine kangaroos (Marsupialia, Potoroidae) from Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum, 41: 281 - 294." title="New Miocene bulungamayine kangaroos (Marsupialia, Potoroidae) from Riversleigh, northwestern Queensland" type="journal article" year="1997">Cooke (1997c)</bibRefCitation>
and
<emphasis id="35EE6E5EFD04FFFF2495E788D1DDFA05" bold="true" box="[1418,1431,1435,1458]" pageId="13" pageNumber="14">3</emphasis>
,
<taxonomicName id="C09AC9CFFD04FFFF21DCE7ABD663FA78" authority="Cooke (1992)" authorityName="Cooke" authorityYear="1992" box="[195,553,1464,1487]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="acris">
<emphasis id="35EE6E5EFD04FFFF21DCE7ABD5C5FA78" box="[195,399,1464,1487]" italics="true" pageId="13" pageNumber="14">Ganawamaya acris</emphasis>
<bibRefCitation id="630BCFBDFD04FFFF208AE7ABD663FA78" author="Cooke, B. N." box="[405,553,1464,1487]" journalOrPublisher="Alcheringa" pageId="13" pageNumber="14" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke (1992)</bibRefCitation>
</taxonomicName>
. Abbreviations: co, cristid obliqua; end, entoconid; hc, hypocingulid; hyd, hypoconid; i1, lower incisor; p3, third premolar; m1m4, lower molars one to four; med, metaconid; phc, posthypocristid, prd, protoconid.
</paragraph>
</caption>
<subSubSection id="4F80E1C7FD04FFFC2257E492D65BF8F7" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="0725B24CFD04FFFC2257E492D65BF8F7" blockId="13.[840,1440,1601,1914]" lastBlockId="14.[192,792,1607,1856]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
<emphasis id="35EE6E5EFD04FFFF2257E492D0F9F92D" bold="true" box="[840,1203,1665,1690]" pageId="13" pageNumber="14">Emended species diagnosis.</emphasis>
<taxonomicName id="C09AC9CFFD04FFFF25A5E492D737F90D" authorityName="Cooke" authorityYear="1992" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD04FFFF25A5E492D737F90D" italics="true" pageId="13" pageNumber="14">Ganawamaya aediculis</emphasis>
</taxonomicName>
differs from all other species of
<taxonomicName id="C09AC9CFFD04FFFF25E3E4B2D1EAF90C" authorityName="Cooke" authorityYear="1992" box="[1276,1440,1697,1723]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
<emphasis id="35EE6E5EFD04FFFF25E3E4B2D1EAF90C" box="[1276,1440,1697,1723]" italics="true" pageId="13" pageNumber="14">Ganawamaya</emphasis>
</taxonomicName>
in having the following unique combination of features: long and sinuous i1 with dorsal and ventral flanges; additional cuspid on posterior end of p3 below the occlusal margin; linear occlusal edge on p3; inclined ridge from the anteriormost cuspid on p3; prominent hypocristid on m1 and m2; reduced paraconid on m1; no hypoconulid; no marked convexities on the lateral margins of the interloph valley of the lower molars; m1 anterior cingulid not extending from paracristid right across to buccal surface of the tooth; poorly developed anterior cingulum of M1; poorly developed anterior cingulum on M1; stylar cusp C on M1 reduced and connected to the postparacrista.
</paragraph>
</subSubSection>
<caption id="53E5E2C4FD07FFFC21DAE7A9D697F9BB" ID-DOI="http://doi.org/10.5281/zenodo.10961552" ID-Zenodo-Dep="10961552" httpUri="https://zenodo.org/record/10961552/files/figure.png" pageId="14" pageNumber="15" startId="14.[197,288,1466,1489]" targetBox="[222,1429,187,1416]" targetPageId="14" targetType="figure">
<paragraph id="0725B24CFD07FFFC21DAE7A9D697F9BB" blockId="14.[197,1436,1466,1548]" pageId="14" pageNumber="15">
<emphasis id="35EE6E5EFD07FFFC21DAE7A9D571FA66" bold="true" box="[197,315,1466,1489]" pageId="14" pageNumber="15">FIGURE 4.</emphasis>
<taxonomicName id="C09AC9CFFD07FFFC205EE7A9D672FA66" authorityName="Cooke" authorityYear="1992" box="[321,568,1466,1489]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD07FFFC205EE7A9D672FA66" box="[321,568,1466,1489]" italics="true" pageId="14" pageNumber="15">Ganawamaya aediculis</emphasis>
</taxonomicName>
partial cranium (QM F58658) in ventral view (
<emphasis id="35EE6E5EFD07FFFC2503E7A9D063FA66" bold="true" box="[1052,1065,1466,1489]" pageId="14" pageNumber="15">1</emphasis>
) and left lateral view (
<emphasis id="35EE6E5EFD07FFFC240DE7A9D155FA66" bold="true" box="[1298,1311,1466,1489]" pageId="14" pageNumber="15">2</emphasis>
). Abbreviations: I1, first upper incisor; ioc, infraorbital canal; iof, infraorbital foramen; M1M4, upper first to fourth molar; mx, maxilla; P3, upper third premolar; pmx, premaxilla.
</paragraph>
</caption>
<subSubSection id="4F80E1C7FD07FFE22257E454D529FA8F" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15" type="description">
<paragraph id="0725B24CFD07FFFC2257E454D791F9D7" blockId="14.[840,987,1607,1632]" box="[840,987,1607,1632]" pageId="14" pageNumber="15">
<emphasis id="35EE6E5EFD07FFFC2257E454D791F9D7" bold="true" box="[840,987,1607,1632]" pageId="14" pageNumber="15">Description</emphasis>
</paragraph>
<paragraph id="0725B24CFD07FFFC229DE464D08CF907" blockId="14.[840,1440,1655,1904]" pageId="14" pageNumber="15">
Description follows
<bibRefCitation id="630BCFBDFD07FFFC2576E464D146F927" author="Cooke, B. N." box="[1129,1292,1655,1681]" journalOrPublisher="Alcheringa" pageId="14" pageNumber="15" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke (1992)</bibRefCitation>
but we note the following additional features.
</paragraph>
<paragraph id="0725B24CFD07FFFC2257E4A4D037F8A7" blockId="14.[840,1440,1655,1904]" pageId="14" pageNumber="15">
<emphasis id="35EE6E5EFD07FFFC2257E4A4D79FF967" bold="true" box="[840,981,1719,1744]" pageId="14" pageNumber="15">Premaxilla.</emphasis>
In lateral view, the maxillary-premaxillary suture extends ventrally to the posterior margin of the canine alveolus.
</paragraph>
<paragraph id="0725B24CFD07FFFD2257E504D60EFF55" blockId="14.[840,1440,1655,1904]" lastBlockId="15.[192,793,169,1890]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="35EE6E5EFD07FFFC2257E504D014F887" bold="true" box="[840,1118,1815,1840]" pageId="14" pageNumber="15">Maxilla and palatine.</emphasis>
The infraorbital canal is tube-like and positioned at the anterior extremity of the suborbital shelf. The sphenopalatine foramen is oval in shape, anteriorly positioned on the palatine, posterior to the infraorbital canal.
</paragraph>
<paragraph id="0725B24CFD06FFFD21DFE2FAD55EFD35" blockId="15.[192,793,169,1890]" pageId="15" pageNumber="16">
<emphasis id="35EE6E5EFD06FFFD21DFE2FAD5C6FEB5" bold="true" box="[192,396,233,258]" pageId="15" pageNumber="16">Upper dentition.</emphasis>
Description of the upper dentition of
<taxonomicName id="C09AC9CFFD06FFFD21FAE31BD5DFFE95" authorityName="Cooke" authorityYear="1992" box="[229,405,264,290]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD21FAE31BD5DFFE95" box="[229,405,264,290]" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
is based on
<collectionCode id="618B2A89FD06FFFD2356E31BD63EFE95" box="[585,628,264,290]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="15" pageNumber="16" type="Museum">QM</collectionCode>
F58658 and 20633. In ventral view (
<figureCitation id="9FA1AEC9FD06FFFD20CAE33AD671FEF5" box="[469,571,297,322]" captionStart="FIGURE 4" captionStartId="14.[197,288,1466,1489]" captionTargetBox="[222,1429,187,1416]" captionTargetId="figure-128@14.[199,1434,162,1449]" captionTargetPageId="14" captionText="FIGURE 4. Ganawamaya aediculis partial cranium (QM F58658) in ventral view (1) and left lateral view (2). Abbreviations: I1, first upper incisor; ioc, infraorbital canal; iof, infraorbital foramen; M1M4, upper first to fourth molar; mx, maxilla; P3, upper third premolar; pmx, premaxilla." figureDoi="http://doi.org/10.5281/zenodo.10961552" httpUri="https://zenodo.org/record/10961552/files/figure.png" pageId="15" pageNumber="16">Figure 4</figureCitation>
) the canine alveolus is larger than each of the three incisor alveoli. The I1 is preserved, lightly recurved and anteroventrally directed. The size of the incisor alveoli may indicate that I2 is the largest incisor, followed by I1 and then I3. The left and right I1 alveoli are separated by a wide diastema. In occlusal view, I1 is long, elongate and laterally compressed. I1 is slightly recurved in lateral view. No I2, I3 or C1 are preserved, however their presence is indicated by alveoli.
</paragraph>
<paragraph id="0725B24CFD06FFFD21E5E09AD698FC15" blockId="15.[192,793,169,1890]" pageId="15" pageNumber="16">The P3 is flexed anterobuccally such that is not in line with the molar row. The tooth is roughly rectangular in occlusal outline but is tapered anteriorly. Five cuspules are visible with the posteriormost cusp being the largest. Each cuspule has an associated transcrista. A prominent posterior lingual cuspule is present from which a well-developed lingual cingulum extends. In lateral view, the occlusal margin of the P3 is slightly concave.</paragraph>
<paragraph id="0725B24CFD06FFFD21E5E1BAD68CF955" blockId="15.[192,793,169,1890]" pageId="15" pageNumber="16">
In occlusal view, the M1 is roughly rectangular in outline. The protoloph and metaloph are roughly equal in height, however, the metaloph is narrower than the protoloph. The paracone is shorter but more massive than the protocone. A short, well-developed preparacrista extends from the paracone connecting to a well-developed anterior cingulum at the anterior end of the tooth. The anterior cingulum is bordered lingually by a faint forelink, representing a remnant preprotocrista. No precingulum is present. On the face of the paracone and lingual to the preprotocrista, a shallow concavity is present. The postparacrista is prominent and extends towards the interloph valley from the paracone. The postparacrista on M1 extends posteriorly to meet a small but distinct cusp. This cusp is interpreted to represent stylar cusp C in
<taxonomicName id="C09AC9CFFD06FFFD205CE7DAD5ADFA55" authorityName="Butler &amp; Travouillon &amp; Price &amp; Archer &amp; Hand" authorityYear="2018" baseAuthorityName="Butler &amp; Travouillon &amp; Price &amp; Archer &amp; Hand" baseAuthorityYear="2018" box="[323,487,1481,1506]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="gillespieae">
<emphasis id="35EE6E5EFD06FFFD205CE7DAD5ADFA55" box="[323,487,1481,1506]" italics="true" pageId="15" pageNumber="16">N. gillespieae</emphasis>
</taxonomicName>
by
<bibRefCitation id="630BCFBDFD06FFFD2306E7DAD6B2FA55" author="Kear, B. P. &amp; Cooke, B. N. &amp; Archer, M. &amp; Flannery, T. F." box="[537,760,1481,1506]" journalOrPublisher="Journal of Paleontology" pageId="15" pageNumber="16" pagination="1147 - 1167" part="81" refId="ref19952" refString="Kear, B. P., Cooke, B. N., Archer, M., and Flannery, T. F. 2007. Implications of a new species of the Oligo-Miocene kangaroo (Marsupialia: Macropodoidea) Nambaroo, from the Riversleigh World Heritage Area, Queensland, Australia. Journal of Paleontology, 81 (6): 1147 - 1167. https: // doi. org / 10.1666 / 04 - 218.1" title="Implications of a new species of the Oligo-Miocene kangaroo (Marsupialia: Macropodoidea) Nambaroo, from the Riversleigh World Heritage Area, Queensland, Australia" type="journal article" year="2007">Kear et al. (2007)</bibRefCitation>
. A poorly developed postprotocrista extends posteriorly to the interloph valley where it forms a midlink. The metacone is slightly taller than the metaconule. A premetacrista extends anteriorly to meet the postparacrista in the interloph valley. A prominent postmetacrista extends from the metacone to meet the postmetaconule crista. Both the neometaconule and postlink are absent.
</paragraph>
<paragraph id="0725B24CFD06FFFD21E5E4FAD020FE36" blockId="15.[192,793,169,1890]" lastBlockId="15.[840,1441,168,1890]" pageId="15" pageNumber="16">
The M2 is similar in morphology to M1 except as follows: it is slightly larger; the preparacrista is more buccally positioned; the anterior cingulum is wider; a precingulum is present lingual to the very faint forelink and borders the anterolingual margin of the tooth; the postparacrista is straighter anteriorly but meets the premetacrista more lingually in the interloph valley; the premetacrista is less developed; StC is present as a distinct cusp on the postparacrista only on
<collectionCode id="618B2A89FD06FFFD2550E35BD030FED5" box="[1103,1146,328,354]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="15" pageNumber="16" type="Museum">QM</collectionCode>
F20633 where it is, however, poorly defined.
</paragraph>
<paragraph id="0725B24CFD06FFFD229DE39AD160FE55" blockId="15.[840,1441,168,1890]" pageId="15" pageNumber="16">The M3 is similar in morphology to the M2 except as follows: StC is absent; the premetacrista, postprotocrista, and midlink are reduced.</paragraph>
<paragraph id="0725B24CFD06FFFD229DE3FAD7B1FD35" blockId="15.[840,1441,168,1890]" pageId="15" pageNumber="16">The M4 is similar in morphology to the M3 except as follows: the metaloph is markedly narrower than the protoloph; the postparacrista and premetacrista are reduced; the forelink is more lingually situated.</paragraph>
<paragraph id="0725B24CFD06FFE22257E09AD529FA8F" blockId="15.[840,1441,168,1890]" lastBlockId="16.[192,793,1246,1657]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="35EE6E5EFD06FFFD2257E09AD05DFD15" bold="true" box="[840,1047,649,674]" pageId="15" pageNumber="16">Lower dentition.</emphasis>
The description in
<bibRefCitation id="630BCFBDFD06FFFD25E2E09BD1EAFD15" author="Cooke, B. N." box="[1277,1440,648,674]" journalOrPublisher="Alcheringa" pageId="15" pageNumber="16" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke (1992)</bibRefCitation>
is sufficient except for as follows: The protostylid on the m1 is present on unworn adult specimens e.g.,
<collectionCode id="618B2A89FD06FFFD229CE0FBD7E4FCB5" box="[899,942,744,770]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="15" pageNumber="16" type="Museum">QM</collectionCode>
F31181; and there are five cuspids on p3.
<emphasis id="35EE6E5EFD06FFFD2257E11AD78AFC95" bold="true" box="[840,960,777,802]" pageId="15" pageNumber="16">Remarks.</emphasis>
<bibRefCitation id="630BCFBDFD06FFFD22D4E11BD015FC95" author="Cooke, B. N." box="[971,1119,776,802]" journalOrPublisher="Alcheringa" pageId="15" pageNumber="16" pagination="201 - 217" part="16" refId="ref19289" refString="Cooke, B. N. 1992. Primitive macropodids from Riversleigh, north-western Queensland. Alcheringa, 16 (3): 201 - 217. https: // doi. org / 10.1080 / 03115519208619119" title="Primitive macropodids from Riversleigh, north-western Queensland" type="journal article" year="1992">Cooke 1992</bibRefCitation>
described the
<typeStatus id="D8210CEEFD06FFFD2405E11AD135FC95" box="[1306,1407,777,802]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C09AC9CFFD06FFFD2257E13BD7B8FCF5" authorityName="Cooke" authorityYear="1992" box="[840,1010,808,834]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD2257E13BD7B8FCF5" box="[840,1010,808,834]" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
(
<collectionCode id="618B2A89FD06FFFD251EE13BD064FCF5" box="[1025,1070,808,834]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="15" pageNumber="16" type="Museum">QM</collectionCode>
F16843) as having six cuspids on p3 with five associated transcristids. However, upon further inspection we clarify that like in other
<taxonomicName id="C09AC9CFFD06FFFD2257E19BD01DFC15" box="[840,1111,904,930]" pageId="15" pageNumber="16">
<emphasis id="35EE6E5EFD06FFFD2257E19BD7A6FC15" box="[840,1004,904,930]" italics="true" pageId="15" pageNumber="16">Ganawamaya</emphasis>
species
</taxonomicName>
, only five cuspids and four transcristids appear to be present on the occlusal surface (
<figureCitation id="9FA1AEC9FD06FFFD22B0E1DAD053FC55" box="[943,1049,969,994]" captionStart="FIGURE 5" captionStartId="16.[197,288,1028,1051]" captionTargetBox="[197,795,166,997]" captionTargetId="figure-565@16.[197,795,162,997]" captionTargetPageId="16" captionText="FIGURE 5. Lower third premolar of Ganawamaya aediculis holotype (QM F16843) in buccal view (1) and lingual view (2). Black arrows indicate each cuspid along the occlusal surface of the p3. Grey arrows indicate the additional sixth cuspid present below the occlusal margin." figureDoi="http://doi.org/10.5281/zenodo.10961554" httpUri="https://zenodo.org/record/10961554/files/figure.png" pageId="15" pageNumber="16">Figure 5</figureCitation>
). However, an additional in
<taxonomicName id="C09AC9CFFD06FFFD2479E1DBD7FAFBB5" authorityName="Cooke" authorityYear="1992" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD2479E1DBD7FAFBB5" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
, previously interpreted a sixth cuspid at the posterior of the p3 on the
<typeStatus id="D8210CEEFD06FFFD25ABE61AD157FB95" box="[1204,1309,1033,1058]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
, is present below the occlusal margin. The presence of a posterior cuspid on the p3 bellow the occlusal margin may be a distinguishing feature of
<taxonomicName id="C09AC9CFFD06FFFD25EBE67BD1EAFB35" authorityName="Cooke" authorityYear="1992" box="[1268,1440,1128,1154]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD25EBE67BD1EAFB35" box="[1268,1440,1128,1154]" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
(
<figureCitation id="9FA1AEC9FD06FFFD2251E69AD7F3FB15" box="[846,953,1161,1186]" captionStart="FIGURE 5" captionStartId="16.[197,288,1028,1051]" captionTargetBox="[197,795,166,997]" captionTargetId="figure-565@16.[197,795,162,997]" captionTargetPageId="16" captionText="FIGURE 5. Lower third premolar of Ganawamaya aediculis holotype (QM F16843) in buccal view (1) and lingual view (2). Black arrows indicate each cuspid along the occlusal surface of the p3. Grey arrows indicate the additional sixth cuspid present below the occlusal margin." figureDoi="http://doi.org/10.5281/zenodo.10961554" httpUri="https://zenodo.org/record/10961554/files/figure.png" pageId="15" pageNumber="16">Figure 5</figureCitation>
). We propose that all
<emphasis id="35EE6E5EFD06FFFD25D8E69BD116FB15" box="[1223,1372,1160,1186]" italics="true" pageId="15" pageNumber="16">Ganamwaya</emphasis>
have five cuspids along the occlusal row of the p3 while
<taxonomicName id="C09AC9CFFD06FFFD2257E6DBD7BCFB56" authorityName="Cooke" authorityYear="1992" box="[840,1014,1224,1249]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD2257E6DBD7BCFB56" box="[840,1014,1224,1249]" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
has an additional posterior cuspid below the occlusal row. On a number of specimens attributed here to
<taxonomicName id="C09AC9CFFD06FFFD253CE71BD09AFA96" authorityName="Cooke" authorityYear="1992" box="[1059,1232,1288,1313]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD253CE71BD09AFA96" box="[1059,1232,1288,1313]" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
, the presence of this cuspid cannot be confirmed as the posterior end of the p3 has been obscured and worn by a slightly overlapping m1, making it difficult to determine whether this additional cusp is diagnostic of all
<taxonomicName id="C09AC9CFFD06FFFD2273E7BBD05DFA76" authorityName="Cooke" authorityYear="1992" box="[876,1047,1448,1473]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD06FFFD2273E7BBD05DFA76" box="[876,1047,1448,1473]" italics="true" pageId="15" pageNumber="16">Gan. aediculis</emphasis>
</taxonomicName>
. All specimens referable to species of
<taxonomicName id="C09AC9CFFD06FFFD22DEE7DBD02FFA55" authorityName="Cooke" authorityYear="1992" box="[961,1125,1480,1506]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="35EE6E5EFD06FFFD22DEE7DBD02FFA55" box="[961,1125,1480,1506]" italics="true" pageId="15" pageNumber="16">Ganawamaya</emphasis>
</taxonomicName>
(and some previously referred to
<taxonomicName id="C09AC9CFFD06FFFD22CCE7FBD005F9B6" baseAuthorityName="Flannery and Rich" baseAuthorityYear="1986" box="[979,1103,1512,1537]" class="Mammalia" family="Balbaridae" genus="Nambaroo" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
<emphasis id="35EE6E5EFD06FFFD22CCE7FBD005F9B6" box="[979,1103,1512,1537]" italics="true" pageId="15" pageNumber="16">Nambaroo</emphasis>
</taxonomicName>
) from Faunal Zone A, also exhibit variation in apparent cuspid morphology that can be attributed to differences in inter-proximal dental wear similar to that of specimens reassigned to
<taxonomicName id="C09AC9CFFD06FFFD22DFE47BD076F936" authorityName="Cooke" authorityYear="1992" box="[960,1084,1640,1665]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="acris">
<emphasis id="35EE6E5EFD06FFFD22DFE47BD076F936" box="[960,1084,1640,1665]" italics="true" pageId="15" pageNumber="16">Gan. acris</emphasis>
</taxonomicName>
. These variations include the presence, or apparent absence, of the protostylid on m1, the shape of the paracristid on m1 and the complexity of enamel ridges on i1. However, specimens from Faunal Zone A assemblages differ from those in Faunal Zone B in having a better defined posthypocristid on m1 and m2. Specimens previously attributed to N. sp. 8 (
<bibRefCitation id="630BCFBDFD06FFFD25BDE55BD10AF8D6" author="Cooke, B. N." box="[1186,1344,1864,1890]" pageId="15" pageNumber="16" refId="ref19326" refString="Cooke, B. N. 1996. Researches into fossil kangaroos and kangaroo evolution. Unpublished PhD Thesis, University of New South Wales, Sydney, Australia." type="book" year="1996">Cooke, 1996</bibRefCitation>
,
<bibRefCitation id="630BCFBDFD06FFFD2452E55BD1D0F8D6" author="Cooke, B. N." box="[1357,1434,1864,1889]" journalOrPublisher="Memoirs of the Queensland Museum" pageId="15" pageNumber="16" pagination="295 - 302" part="41" refId="ref19356" refString="Cooke, B. N. 1997 a. Biostratigraphic implications of fossil kangaroos at Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum, 41: 295 - 302." title="Biostratigraphic implications of fossil kangaroos at Riversleigh, northwestern Queensland" type="journal article" year="1997">1997a</bibRefCitation>
) are referred here to
<taxonomicName id="C09AC9CFFD19FFE220A2E6CDD620FB4F" authorityName="Cooke" authorityYear="1992" box="[445,618,1246,1272]" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD19FFE220A2E6CDD620FB4F" box="[445,618,1246,1272]" italics="true" pageId="16" pageNumber="17">Gan. aediculis</emphasis>
</taxonomicName>
because they lack features that would warrant separation as a distinct taxon.
</paragraph>
</subSubSection>
<caption id="53E5E2C4FD19FFE221DAE617D4A7FB19" ID-DOI="http://doi.org/10.5281/zenodo.10961554" ID-Zenodo-Dep="10961554" httpUri="https://zenodo.org/record/10961554/files/figure.png" pageId="16" pageNumber="17" startId="16.[197,288,1028,1051]" targetBox="[197,795,166,997]" targetPageId="16" targetType="figure">
<paragraph id="0725B24CFD19FFE221DAE617D4A7FB19" blockId="16.[197,788,1028,1198]" pageId="16" pageNumber="17">
<emphasis id="35EE6E5EFD19FFE221DAE617D576FBAB" bold="true" box="[197,316,1028,1052]" pageId="16" pageNumber="17">FIGURE 5.</emphasis>
Lower third premolar of
<taxonomicName id="C09AC9CFFD19FFE2235CE617D4BFFB8E" authorityName="Cooke" authorityYear="1992" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD19FFE2235CE617D4BFFB8E" italics="true" pageId="16" pageNumber="17">Ganawamaya aediculis</emphasis>
</taxonomicName>
holotype (QM F16843) in buccal view (
<emphasis id="35EE6E5EFD19FFE223BAE631D6F8FB8E" bold="true" box="[677,690,1058,1081]" pageId="16" pageNumber="17">1</emphasis>
) and lingual view (
<emphasis id="35EE6E5EFD19FFE22023E62CD503FBE1" bold="true" box="[316,329,1087,1110]" pageId="16" pageNumber="17">2</emphasis>
). Black arrows indicate each cuspid along the occlusal surface of the p3. Grey arrows indicate the additional sixth cuspid present below the occlusal margin.
</paragraph>
</caption>
<subSubSection id="4F80E1C7FD19FFE221DFE72CD758F9CF" pageId="16" pageNumber="17" type="multiple">
<paragraph id="0725B24CFD19FFE221DFE72CD758F9CF" blockId="16.[192,793,1246,1657]" pageId="16" pageNumber="17">
<emphasis id="35EE6E5EFD19FFE221DFE72CD587FAEF" bold="true" box="[192,461,1343,1368]" pageId="16" pageNumber="17">Age and distribution.</emphasis>
The
<typeStatus id="D8210CEEFD19FFE22312E72CD638FAEF" box="[525,626,1343,1368]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
of
<taxonomicName id="C09AC9CFFD19FFE22385E72DD4BFFACF" authorityName="Cooke" authorityYear="1992" class="Mammalia" family="Balbaridae" genus="Ganawamaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diprotodontia" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="aediculis">
<emphasis id="35EE6E5EFD19FFE22385E72DD4BFFACF" italics="true" pageId="16" pageNumber="17">Gan. aediculis</emphasis>
</taxonomicName>
is from White Hunter Site, Riversleigh WHA, northwestern
<collectingRegion id="C55E7CAEFD19FFE2207AE76DD5B3FA2F" box="[357,505,1406,1432]" country="Australia" name="Queensland" pageId="16" pageNumber="17">Queensland</collectingRegion>
. The White Hunter Site is interpreted to be part of Faunal Zone A, which is interpreted to be late Oligocene in age (
<bibRefCitation id="630BCFBDFD19FFE223B9E7ACD566FA4F" author="Archer, M. &amp; Godthelp, H. &amp; Hand, S. J. &amp; Megirian, D." journalOrPublisher="Australian Zoologist" pageId="16" pageNumber="17" pagination="29 - 65" part="25" refId="ref18573" refString="Archer, M., Godthelp, H., Hand, S. J., and Megirian, D. 1989. Fossil mammals of Riversleigh, northwestern Queensland: preliminary overview of biostratigraphy, correlation and environmental change. Australian Zoologist, 25 (2): 29 - 65. https: // doi. org / 10.7882 / AZ. 1989.001" title="Fossil mammals of Riversleigh, northwestern Queensland: preliminary overview of biostratigraphy, correlation and environmental change" type="journal article" year="1989">Archer et al., 1989</bibRefCitation>
,
<bibRefCitation id="630BCFBDFD19FFE22026E7CCD532FA4F" author="Archer, M. &amp; Hand, S. J. &amp; Godthelp, H. &amp; Creaser, P." box="[313,376,1503,1528]" editor="Aguilar, J. P. &amp; Legendre, S. &amp; Michaux, J." journalOrPublisher="Institut de Montpellier" pageId="16" pageNumber="17" pagination="131 - 152" refId="ref18639" refString="Archer, M., Hand, S. J., Godthelp, H., and Creaser, P. 1997. Correlation of the Cainozoic sediments of the Riversleigh World Heritage Fossil Property, Queensland, Australia, p. 131 - 152. In Aguilar, J. P., Legendre, S., and Michaux, J. (eds.), Actes du Congr es BiochroM' 97, Memoires et Travaux. L'ecole Pratique des Hautes Etudes, Institut de Montpellier, Montpellier, France." title="Correlation of the Cainozoic sediments of the Riversleigh World Heritage Fossil Property, Queensland, Australia" type="journal article" volumeTitle="Actes du Congr es BiochroM' 97, Memoires et Travaux. L'ecole Pratique des Hautes Etudes" year="1997">1997</bibRefCitation>
;
<bibRefCitation id="630BCFBDFD19FFE2209AE7CCD6EEFA4F" author="Myers, T. &amp; Archer, M." box="[389,676,1503,1528]" journalOrPublisher="Memoirs of the Queensland Museum" pageId="16" pageNumber="17" pagination="379 - 392" part="41" refId="ref20353" refString="Myers, T. and Archer, M. 1997. Kuterintja ngama (Marsupialia, Ilariidae): a revised systematic analysis based on material from the late Oligocene of Riversleigh, northwestern Queensland. Memoirs of the Queensland Museum, 41: 379 - 392." title="Kuterintja ngama (Marsupialia, Ilariidae): a revised systematic analysis based on material from the late Oligocene of Riversleigh, northwestern Queensland" type="journal article" year="1997">Myers and Archer 1997</bibRefCitation>
;
<bibRefCitation id="630BCFBDFD19FFE223AEE7CCD53DF9AF" author="Travouillon, K. J. &amp; Archer, M. &amp; Hand, S. J. &amp; Godthelp, H." journalOrPublisher="Alcheringa: An Australasian Journal of Palaeontology, 30 (supplement" pageId="16" pageNumber="17" pagination="323 - 349" part="1" refId="ref20674" refString="Travouillon, K. J., Archer, M., Hand, S. J., and Godthelp, H. 2006. Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, northwestern Queensland. Alcheringa: An Australasian Journal of Palaeontology, 30 (supplement 1): 323 - 349. https: // doi. org / 10.1080 / 03115510608619590" title="Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, northwestern Queensland" type="journal article" year="2006">Travouillon et al., 2006</bibRefCitation>
,
<bibRefCitation id="630BCFBDFD19FFE2209BE7ECD58BF9AF" author="Travouillon, K. J. &amp; Escarguel, G. &amp; Legendre, S. &amp; Archer, M. &amp; Hand, S. J." box="[388,449,1535,1560]" journalOrPublisher="Paleobiology" pageId="16" pageNumber="17" pagination="696 - 709" part="37" refId="ref20827" refString="Travouillon, K. J., Escarguel, G., Legendre, S., Archer, M., and Hand, S. J. 2011. The use of MSR (Minimum Sample Richness) for sample assemblage comparisons. Paleobiology, 37 (4): 696 - 709. https: // doi. org / 10.1666 / 09050.1" title="The use of MSR (Minimum Sample Richness) for sample assemblage comparisons" type="journal article" year="2011">2011</bibRefCitation>
;
<bibRefCitation id="630BCFBDFD19FFE220D2E7ECD6E1F9AF" author="Arena, D. A. &amp; Travouillon, K. J. &amp; Beck, R. M. D. &amp; Black, K. &amp; Gillespie, A. K. &amp; Myers, T. J. &amp; Archer, M. &amp; Hand, S. J." box="[461,683,1535,1560]" journalOrPublisher="Lethaia" pageId="16" pageNumber="17" pagination="43 - 60" part="49" refId="ref18734" refString="Arena, D. A., Travouillon, K. J., Beck, R. M. D., Black, K., Gillespie, A. K., Myers, T. J., Archer, M., and Hand, S. J. 2015. Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia. Lethaia, 49: 43 - 60. https: // doi. org / 10.1111 / let. 12131" title="Mammalian lineages and the biostratigraphy and biochronology of Cenozoic faunas from the Riversleigh World Heritage Area, Australia" type="journal article" year="2015">Arena et al., 2015</bibRefCitation>
). Several other referred specimens are also from sites considered to belong to Faunal Zone A: Gillespie's Gully and LSO Site (
<bibRefCitation id="630BCFBDFD19FFE220B2E44CD6F5F9CF" author="Travouillon, K. J. &amp; Archer, M. &amp; Hand, S. J. &amp; Godthelp, H." box="[429,703,1631,1656]" journalOrPublisher="Alcheringa: An Australasian Journal of Palaeontology, 30 (supplement" pageId="16" pageNumber="17" pagination="323 - 349" part="1" refId="ref20674" refString="Travouillon, K. J., Archer, M., Hand, S. J., and Godthelp, H. 2006. Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, northwestern Queensland. Alcheringa: An Australasian Journal of Palaeontology, 30 (supplement 1): 323 - 349. https: // doi. org / 10.1080 / 03115510608619590" title="Multivariate analyses of Cenozoic mammalian faunas from Riversleigh, northwestern Queensland" type="journal article" year="2006">Travouillon et al., 2006</bibRefCitation>
,
<bibRefCitation id="630BCFBDFD19FFE223D7E44CD74DF9CF" author="Travouillon, K. J. &amp; Escarguel, G. &amp; Legendre, S. &amp; Archer, M. &amp; Hand, S. J." box="[712,775,1631,1656]" journalOrPublisher="Paleobiology" pageId="16" pageNumber="17" pagination="696 - 709" part="37" refId="ref20827" refString="Travouillon, K. J., Escarguel, G., Legendre, S., Archer, M., and Hand, S. J. 2011. The use of MSR (Minimum Sample Richness) for sample assemblage comparisons. Paleobiology, 37 (4): 696 - 709. https: // doi. org / 10.1666 / 09050.1" title="The use of MSR (Minimum Sample Richness) for sample assemblage comparisons" type="journal article" year="2011">2011</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>