396 lines
44 KiB
XML
396 lines
44 KiB
XML
<document id="90096A5E4321DE4D25F0314414AE8B02" ID-CLB-Dataset="4959" ID-DOI="10.1080/00222933.2013.809491" ID-GBIF-Dataset="573cbdf3-37de-422a-9fca-9a34315d2cf9" ID-ISSN="1464-5262" ID-Zenodo-Dep="5199249" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628889267124" checkinUser="felipe" docAuthor="Hong, Yong, James, Samuel W. & Inkhavilay, Khamla" docDate="2013" docId="652DA5089058FFFBFECA3862D6EAB3AB" docLanguage="en" docName="JNatHist.48.5-6.251-256.pdf" docOrigin="Journal of Natural History 48 (5 - 6)" docSource="http://dx.doi.org/10.1080/00222933.2013.809491" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Amynthas hauyhowensis Hong and James 2013, sp. nov." docType="treatment" docVersion="5" lastPageNumber="254" masterDocId="9914DD70905BFFFEFF913A67D613B534" masterDocTitle="A new species of earthworm in Amynthas hexathecus species group (Clitellata: Megascolecidae) from the Nam Xam National Protected Area, Laos" masterLastPageNumber="256" masterPageNumber="251" pageNumber="252" updateTime="1699080211362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="402C531ABA0EEDA129AA247B866393A4">
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<mods:title id="9943F223753D6C2B89DA20D65E20692B">A new species of earthworm in Amynthas hexathecus species group (Clitellata: Megascolecidae) from the Nam Xam National Protected Area, Laos</mods:title>
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<mods:name id="36DCC8E57CDEE29F43E9A0C71AD8533F" type="personal">
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<mods:role id="D38FAF71222FB39623DA6F6D859895DA">
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<mods:roleTerm id="CE156FAB36FD0452F5C139AA3BC3C019">Author</mods:roleTerm>
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<mods:namePart id="58CD1341C8F85F30D40D77D688B7BB2A">Hong, Yong</mods:namePart>
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<mods:affiliation id="01A12722A6587596C703D114BD8820FB">Department of Agricultural Biology, College of Agriculture & Life Sciences, Chonbuk National University, Jeonju, Republic of Korea;</mods:affiliation>
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<mods:roleTerm id="AD3EC6EBFAEBD92D3D0B94C3A4DC3414">Author</mods:roleTerm>
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<mods:namePart id="2A9EB7CDCF35B9D29BC10597942F02F2">James, Samuel W.</mods:namePart>
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<mods:affiliation id="E684952B4ACECA5C0F91E317E258B684">Department of Biology, University of Iowa, Iowa City, Iowa, USA;</mods:affiliation>
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</mods:name>
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<mods:name id="FEF9121FEB98D19329AE35E00B3C73D2" type="personal">
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<mods:roleTerm id="9607414BB945858B10568B9E94B97FDC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="62A6497A63A84828072E99F7C31028DA">Inkhavilay, Khamla</mods:namePart>
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<mods:affiliation id="36850E7ABA0FF2F589B2BBBCA03BF8B3">Department of Biology, Faculty of Science, National University of Laos, Vientiane, Laos</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="2475D74EA48DBA50B3731A444821CC97">text</mods:typeOfResource>
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<mods:title id="4F3512598F5BC73653987360280240DD">Journal of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="87AC7F75ACC844D76C21296F565C6C20">
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<mods:date id="36FCDC28E3CE48AC6230A12F41E9327C">2013</mods:date>
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<mods:number id="94EB208B21D3CA93118EC963C3202E34">2013-12-10</mods:number>
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<mods:detail id="3A3FCD091A06A2BC58F703F9F14B6E56" type="volume">
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<mods:number id="E48E2E72253CB194E5BAD9BD31D8E0C9">48</mods:number>
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<mods:number id="8440628BBDF7C0C0FF6390B53947AD69">5 - 6</mods:number>
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<mods:start id="B6B005E83BFCFD1ED4F8649953B2719B">251</mods:start>
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<mods:url id="6D58C84FB1115A9D6418641C73DFF1B9">http://dx.doi.org/10.1080/00222933.2013.809491</mods:url>
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<mods:identifier id="F45297D38ABCF261868BD752704B3388" type="Zenodo-Dep">5199249</mods:identifier>
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<treatment id="652DA5089058FFFBFECA3862D6EAB3AB" ID-DOI="http://doi.org/10.5281/zenodo.5199231" ID-GBIF-Taxon="184192438" ID-Zenodo-Dep="5199231" LSID="urn:lsid:plazi:treatment:652DA5089058FFFBFECA3862D6EAB3AB" httpUri="http://treatment.plazi.org/id/652DA5089058FFFBFECA3862D6EAB3AB" lastPageId="5" lastPageNumber="254" pageId="3" pageNumber="252">
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<subSubSection id="A59E47959058FFFDFECA3862D597B729" box="[347,900,517,542]" pageId="3" pageNumber="252" type="nomenclature">
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<paragraph id="ED3B141E9058FFFDFECA3862D597B729" blockId="3.[347,900,453,573]" box="[347,900,517,542]" pageId="3" pageNumber="252">
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<heading id="B673A3729058FFFDFECA3862D597B729" box="[347,900,517,542]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="252" reason="2">
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<taxonomicName id="2A846F9D9058FFFDFECA3862D53BB729" authority="Hong and James" authorityName="Hong and James" authorityYear="2013" box="[347,808,517,542]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="3" pageNumber="252" phylum="Annelida" rank="species" species="hauyhowensis" status="sp. nov.">
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<emphasis id="DFF0C80C9058FFFDFECA3862D475B729" bold="true" box="[347,614,517,541]" italics="true" pageId="3" pageNumber="252">Amynthas hauyhowensis</emphasis>
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Hong and James
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</taxonomicName>
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<taxonomicNameLabel id="C4C375779058FFFDFCBE3861D597B729" box="[815,900,518,541]" pageId="3" pageNumber="252" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A59E47959058FFFDFDA53842D4B7B709" box="[564,676,549,573]" pageId="3" pageNumber="252" type="description">
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<paragraph id="ED3B141E9058FFFDFDA53842D4B7B709" blockId="3.[347,900,453,573]" box="[564,676,549,573]" pageId="3" pageNumber="252">
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(
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<figureCitation id="75BF089B9058FFFDFDAC3842D488B709" box="[573,667,549,573]" captionStart="Figure 1" captionStartId="4.[211,278,1664,1686]" captionTargetBox="[261,1124,161,1630]" captionTargetId="figure-5@4.[261,1124,161,1630]" captionTargetPageId="4" captionText="Figure 1. Amynthas hauyhowensis sp. nov. (A) ventral view; (B) spermathecae; (C) intestinal caeca; (D) prostate gland. Scale bars = 2.5 mm (A), 2 mm (B–D)." figureDoi="http://doi.org/10.5281/zenodo.5199251" httpUri="https://zenodo.org/record/5199251/files/figure.png" pageId="3" pageNumber="252">Figure 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="A59E47959058FFFDFF1E38E3D4B8B6FC" pageId="3" pageNumber="252" type="materials_examined">
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<paragraph id="ED3B141E9058FFFDFF1E38E3D73BB7AF" blockId="3.[143,1104,643,834]" box="[143,296,643,667]" pageId="3" pageNumber="252">
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<heading id="B673A3729058FFFDFF1E38E3D73BB7AF" box="[143,296,643,667]" fontSize="10" level="3" pageId="3" pageNumber="252" reason="5">
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<emphasis id="DFF0C80C9058FFFDFF1E38E3D73BB7AF" box="[143,296,643,667]" italics="true" pageId="3" pageNumber="252">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="ED3B141E9058FFFDFF1E38CCD72CB676" blockId="3.[143,1104,643,834]" pageId="3" pageNumber="252">
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<materialsCitation id="5DEC1E439058FFFDFF1E38CCD25FB637" ID-GBIF-Occurrence="3334359302" collectingDate="2004-09-02" collectionCode="BDNUL" country="Laos" county="Clitellate" elevation="470" latitude="20.01135" location="Nam Xam" longLatPrecision="1" longitude="104.6735" municipality="Xam Tay district" pageId="3" pageNumber="252" specimenCode="BDNUL 0017" specimenCount="1" stateProvince="Houaphan" typeStatus="holotype">
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<typeStatus id="323FAABC9058FFFDFF1E38CCD6EDB7F7" box="[143,254,683,707]" pageId="3" pageNumber="252" type="holotype">Holotype</typeStatus>
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:
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<collectingCounty id="045A6C929058FFFDFE9A38CCD767B7F7" box="[267,372,683,707]" pageId="3" pageNumber="252">Clitellate</collectingCounty>
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: (
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<specimenCode id="BD22BC659058FFFDFE1838CCD420B7F7" box="[393,563,683,707]" collectionCode="BDNUL" pageId="3" pageNumber="252">BDNUL 0017</specimenCode>
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):
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<collectingCountry id="9593548E9058FFFDFDD938CBD492B7F7" box="[584,641,684,707]" name="Laos" pageId="3" pageNumber="252">Laos</collectingCountry>
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,
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<collectingRegion id="2F40DAFC9058FFFDFD0138CCD565B7F7" box="[656,886,683,707]" country="Laos" name="Houaphan" pageId="3" pageNumber="252">Houaphan Province</collectingRegion>
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,
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<collectingMunicipality id="0D5F8E649058FFFDFC1438CBD258B7F7" box="[901,1099,683,708]" pageId="3" pageNumber="252">Xam Tay district</collectingMunicipality>
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,
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<location id="E85B42C59058FFFDFF1E38ABD714B7D7" LSID="urn:lsid:plazi:treatment:652DA5089058FFFBFECA3862D6EAB3AB:E85B42C59058FFFDFF1E38ABD714B7D7" box="[143,263,716,739]" country="Laos" county="Clitellate" latitude="20.01135" longLatPrecision="1" longitude="104.6735" municipality="Xam Tay district" name="Nam Xam" pageId="3" pageNumber="252" stateProvince="Houaphan">Nam Xam</location>
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NPA,
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<location id="E85B42C59058FFFDFEC338ACD459B7D7" LSID="urn:lsid:plazi:treatment:652DA5089058FFFBFECA3862D6EAB3AB:E85B42C59058FFFDFEC338ACD459B7D7" box="[338,586,715,739]" country="Laos" county="Clitellate" latitude="20.01135" longLatPrecision="1" longitude="104.6735" municipality="Xam Tay district" name="South Hauyhow town" pageId="3" pageNumber="252" stateProvince="Houaphan">South Hauyhow town</location>
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(
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<geoCoordinate id="88B072D99058FFFDFDC638ACD4E2B7D6" box="[599,753,711,739]" degrees="20" direction="north" minutes="00. 681" orientation="latitude" pageId="3" pageNumber="252" precision="1" value="20.01135">
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20
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<superScript id="1AF1B9569058FFFDFDE238A0D46FB7EC" attach="none" box="[627,636,711,728]" fontSize="7" pageId="3" pageNumber="252">◦</superScript>
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00. 681
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<superScript id="1AF1B9569058FFFDFD5C38A0D4C1B7EC" attach="left" box="[717,722,711,728]" fontSize="7" pageId="3" pageNumber="252">′</superScript>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="88B072D99058FFFDFD6D38ACD58AB7D7" box="[764,921,711,739]" degrees="104" direction="east" minutes="40.410" orientation="longitude" pageId="3" pageNumber="252" precision="1" value="104.6735">
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104
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<superScript id="1AF1B9569058FFFDFCB538A0D53EB7EC" attach="none" box="[804,813,711,728]" fontSize="7" pageId="3" pageNumber="252">◦</superScript>
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40.410
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<superScript id="1AF1B9569058FFFDFCE938A0D56EB7EC" attach="left" box="[888,893,711,728]" fontSize="7" pageId="3" pageNumber="252">′</superScript>
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E
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</geoCoordinate>
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),
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<quantity id="2A7CB9FB9058FFFDFC3C38ACD5E7B7D6" box="[941,1012,715,739]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="252" unit="m" value="470.0">
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<elevation id="66A9F32D9058FFFDFC3C38ACD5E7B7D6" box="[941,1012,715,739]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="3" pageNumber="252" unit="m" value="470.0">470 m</elevation>
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</quantity>
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, natural forest condition with dark soil on limestone,
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<date id="993A32DE9058FFFDFD1F388CD548B637" box="[654,859,747,771]" pageId="3" pageNumber="252" value="2004-09-02">
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<collectingDate id="897ECB369058FFFDFD1F388CD548B637" box="[654,859,747,771]" pageId="3" pageNumber="252" value="2004-09-02">2 September 2004</collectingDate>
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</date>
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; Monsri, Vanya colls
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</materialsCitation>
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.
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<materialsCitation id="5DEC1E439058FFFDFF1E396CD728B676" ID-GBIF-Occurrence="3334359301" collectingDate="2004-09-02" collectionCode="BDNUL" country="Laos" county="Clitellate" elevation="470" latitude="20.01135" location="Nam Xam" longLatPrecision="1" longitude="104.6735" municipality="Xam Tay district" pageId="3" pageNumber="252" specimenCode="BDNUL 0018, BDNUL 0019, BDNUL 0020" specimenCount="1" stateProvince="Houaphan" typeStatus="paratype">
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<typeStatus id="323FAABC9058FFFDFF1E396CD712B617" box="[143,257,779,803]" pageId="3" pageNumber="252" type="paratype">Paratypes</typeStatus>
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: three clitellates (
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<specimenCode id="BD22BC659058FFFDFE53396DD474B616" box="[450,615,778,802]" collectionCode="BDNUL" pageId="3" pageNumber="252">BDNUL 0018</specimenCode>
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,
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<specimenCode id="BD22BC659058FFFDFDE3396CD505B616" box="[626,790,778,802]" collectionCode="BDNUL" pageId="3" pageNumber="252">BDNUL 0019</specimenCode>
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,
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<specimenCode id="BD22BC659058FFFDFCB0396CD5D4B616" box="[801,967,778,802]" collectionCode="BDNUL" pageId="3" pageNumber="252">BDNUL 0020</specimenCode>
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): Same data as for holotype
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="ED3B141E9058FFFDFF1E39EED721B695" blockId="3.[143,683,905,968]" box="[143,306,905,929]" pageId="3" pageNumber="252">
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<heading id="B673A3729058FFFDFF1E39EED721B695" box="[143,306,905,929]" fontSize="10" level="3" pageId="3" pageNumber="252" reason="5">
|
||
<emphasis id="DFF0C80C9058FFFDFF1E39EED721B695" box="[143,306,905,929]" italics="true" pageId="3" pageNumber="252">Other material</emphasis>
|
||
</heading>
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||
</paragraph>
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||
<paragraph id="ED3B141E9058FFFDFF1E39D7D4B8B6FC" blockId="3.[143,683,905,968]" box="[143,683,944,968]" pageId="3" pageNumber="252">Fifty-three clitellates, same data as for holotype.</paragraph>
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</subSubSection>
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||
<subSubSection id="A59E47959058FFFDFF1E3E68D4C0B17B" box="[143,723,1039,1103]" pageId="3" pageNumber="252" type="etymology">
|
||
<paragraph id="ED3B141E9058FFFDFF1E3E68D714B113" blockId="3.[143,723,1039,1103]" box="[143,263,1039,1063]" pageId="3" pageNumber="252">
|
||
<heading id="B673A3729058FFFDFF1E3E68D714B113" box="[143,263,1039,1063]" fontSize="10" level="3" pageId="3" pageNumber="252" reason="5">
|
||
<emphasis id="DFF0C80C9058FFFDFF1E3E68D714B113" box="[143,263,1039,1063]" italics="true" pageId="3" pageNumber="252">Etymology</emphasis>
|
||
</heading>
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||
</paragraph>
|
||
<paragraph id="ED3B141E9058FFFDFF1E3E50D4C0B17B" blockId="3.[143,723,1039,1103]" box="[143,723,1079,1103]" pageId="3" pageNumber="252">
|
||
The species is named for its
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<typeStatus id="323FAABC9058FFFDFE583E50D7EAB17B" box="[457,505,1079,1103]" pageId="3" pageNumber="252">type</typeStatus>
|
||
locality, Hauyhow.
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="A59E47959058FFFDFF1E3EF1D79AB021" pageId="3" pageNumber="252" type="diagnosis">
|
||
<paragraph id="ED3B141E9058FFFDFF1E3EF1D6E8B199" blockId="3.[143,1103,1174,1301]" box="[143,251,1174,1197]" pageId="3" pageNumber="252">
|
||
<heading id="B673A3729058FFFDFF1E3EF1D6E8B199" box="[143,251,1174,1197]" fontSize="10" level="3" pageId="3" pageNumber="252" reason="5">
|
||
<emphasis id="DFF0C80C9058FFFDFF1E3EF1D6E8B199" box="[143,251,1174,1197]" italics="true" pageId="3" pageNumber="252">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="ED3B141E9058FFFDFF1E3EDAD79AB021" blockId="3.[143,1103,1174,1301]" pageId="3" pageNumber="252">
|
||
Spermathecal pores five pairs in 4
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<emphasis id="DFF0C80C9058FFFDFD9B3EDAD405B1E1" bold="true" box="[522,534,1213,1237]" italics="true" pageId="3" pageNumber="252">/</emphasis>
|
||
5–8
|
||
<emphasis id="DFF0C80C9058FFFDFDAF3EDAD459B1E1" bold="true" box="[574,586,1213,1237]" italics="true" pageId="3" pageNumber="252">/</emphasis>
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9, secondary male pore equatorial in XVIII at 11th setal lines, paired invaginated genital papillae postsetal on XVII anterior to male pores, caeca manicate.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A59E47959058FFFBFF1E3F3BD6C2B629" lastPageId="5" lastPageNumber="254" pageId="3" pageNumber="252" type="description">
|
||
<paragraph id="ED3B141E9058FFFDFF1E3F3BD71DB047" blockId="3.[143,1104,1372,1690]" box="[143,270,1372,1395]" pageId="3" pageNumber="252">
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||
<heading id="B673A3729058FFFDFF1E3F3BD71DB047" box="[143,270,1372,1395]" fontSize="10" level="3" pageId="3" pageNumber="252" reason="5">
|
||
<emphasis id="DFF0C80C9058FFFDFF1E3F3BD71DB047" box="[143,270,1372,1395]" italics="true" pageId="3" pageNumber="252">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="ED3B141E9058FFFDFF1E3FE4D710B3AE" blockId="3.[143,1104,1372,1690]" pageId="3" pageNumber="252">
|
||
Dimensions
|
||
<quantity id="2A7CB9FB9058FFFDFE8F3FE4D78BB0AF" box="[286,408,1411,1435]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" metricValueMax="8.0" metricValueMin="6.2" pageId="3" pageNumber="252" unit="mm" value="71.0" valueMax="80.0" valueMin="62.0">62–80 mm</quantity>
|
||
, by 4.0–
|
||
<quantity id="2A7CB9FB9058FFFDFE663FE4D45EB0AF" box="[503,589,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="3" pageNumber="252" unit="mm" value="4.4">4.4 mm</quantity>
|
||
at segment X,
|
||
<quantity id="2A7CB9FB9058FFFDFD6A3FE4D593B0AF" box="[763,896,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="3" pageNumber="252" unit="mm" value="4.0" valueMax="4.5" valueMin="3.5">3.5–4.5 mm</quantity>
|
||
at segment XXX,
|
||
<quantity id="2A7CB9FB9058FFFDFF1E3FC4D707B08F" box="[143,276,1443,1467]" metricMagnitude="-3" metricUnit="m" metricValue="4.45" metricValueMax="4.5" metricValueMin="4.4" pageId="3" pageNumber="252" unit="mm" value="4.45" valueMax="4.5" valueMin="4.4">4.4–4.5 mm</quantity>
|
||
at clitellum; body cylindrical, segments 59–78. Setae numbering 56–62 at VII, 50–62 at XX, 11–15 between male pores, slightly more widely spaced ventrally, setal formula AA: AB: ZZ: YZ = 3: 4: 3: 3 at XIII. Female pore single on XIV within a circular porophore
|
||
<quantity id="2A7CB9FB9058FFFDFE1B3C65D7F1B32E" box="[394,482,1538,1562]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="252" unit="mm" value="0.3">0.3 mm</quantity>
|
||
in diameter. Prostomium epilobic with tongue open. Very dark brown to black anterior–dorsal pigment, also pigmented in ventral I–IX, lighter on setal rings and post-clitellar segments, clitellum coffee coloured, formalin preservation. First dorsal pore at 12
|
||
<emphasis id="DFF0C80C9058FFFDFDBA3C05D424B34E" bold="true" box="[555,567,1634,1658]" italics="true" pageId="3" pageNumber="252">/</emphasis>
|
||
13. Clitellum annular XIV–XVI; setae invisible externally.
|
||
</paragraph>
|
||
<caption id="B9FB4496905FFFFAFF423CE7D57AB387" ID-DOI="http://doi.org/10.5281/zenodo.5199251" ID-Zenodo-Dep="5199251" httpUri="https://zenodo.org/record/5199251/files/figure.png" pageId="4" pageNumber="253" startId="4.[211,278,1664,1686]" targetBox="[261,1124,161,1630]" targetPageId="4">
|
||
<paragraph id="ED3B141E905FFFFAFF423CE7D57AB387" blockId="4.[211,1171,1664,1715]" pageId="4" pageNumber="253">
|
||
Figure 1.
|
||
<taxonomicName id="2A846F9D905FFFFAFEAF3CE7D422B3A2" authority="Hong & James & Inkhavilay, 2013" authorityName="Hong & James & Inkhavilay" authorityYear="2013" box="[318,561,1664,1686]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="4" pageNumber="253" phylum="Annelida" rank="species" species="hauyhowensis" status="sp. nov.">
|
||
<emphasis id="DFF0C80C905FFFFAFEAF3CE7D422B3A2" box="[318,561,1664,1686]" italics="true" pageId="4" pageNumber="253">Amynthas hauyhowensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4C37577905FFFFAFDA83CE6D49AB3A2" box="[569,649,1665,1686]" pageId="4" pageNumber="253" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(A) ventral view; (B) spermathecae; (C) intestinal caeca; (D) prostate gland. Scale bars = 2.5 mm (A), 2 mm (B–D).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="ED3B141E905EFFFBFF2B3AC0D43DB46B" blockId="5.[143,1104,167,797]" pageId="5" pageNumber="254">
|
||
Secondary male pores equatorial in XVIII at 11th setal line, 0.14–0.24 circumference apart ventrally; paired invaginated genital papillae postsetal on XVII anterior to male pores, both pores in transverse sphincter-like slits
|
||
<quantity id="2A7CB9FB905EFFFBFCC83A80D5A6B5CB" box="[857,949,231,255]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="254" unit="mm" value="0.3">0.3 mm</quantity>
|
||
in diameter. Primary male pores within shallow invagination confined to circular body wall muscle layer. Spermathecal pores five pairs in 4
|
||
<emphasis id="DFF0C80C905EFFFBFD1F3B40D489B40B" bold="true" box="[654,666,295,319]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
5–8
|
||
<emphasis id="DFF0C80C905EFFFBFD523B40D4DCB40B" bold="true" box="[707,719,295,319]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
9, conspicuous, 0.14–0.26 ventral circumference apart from each other.
|
||
</paragraph>
|
||
<paragraph id="ED3B141E905EFFFBFF2B3B00D511B7AA" blockId="5.[143,1104,167,797]" pageId="5" pageNumber="254">
|
||
Septa 5
|
||
<emphasis id="DFF0C80C905EFFFBFE813B00D70FB44B" bold="true" box="[272,284,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
6–6
|
||
<emphasis id="DFF0C80C905EFFFBFED53B00D743B44B" bold="true" box="[324,336,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
7 thick, 7
|
||
<emphasis id="DFF0C80C905EFFFBFE2E3B00D7D8B44B" bold="true" box="[447,459,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
8 thin, 8
|
||
<emphasis id="DFF0C80C905EFFFBFDBE3B00D428B44B" bold="true" box="[559,571,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
9–9
|
||
<emphasis id="DFF0C80C905EFFFBFDF23B00D47CB44B" bold="true" box="[611,623,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
10 absent, 10
|
||
<emphasis id="DFF0C80C905EFFFBFC993B00D507B44B" bold="true" box="[776,788,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
11–12
|
||
<emphasis id="DFF0C80C905EFFFBFCC73B00D571B44B" bold="true" box="[854,866,359,383]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
13 slightly muscular. Gizzard in VIII–X. Intestine begins in XV, lymph glands not found. Typhlosole small from XXVII. Intestinal caeca manicate, originating XXVII, extending anteriorly about to XXIII each containing four to eight small finger-shaped lobes. Hearts oesophageal in X–XII. Male sexual system holandric, testes, funnels in ventrally joined sacs in X, XI. Seminal vesicle large two pairs in XI, XII, small round dorsal lobes in shallow clefts. Prostates XVIII dense three to five lobes extending from XVII to XXII, duct large, muscular, coiled in two or three complete spiral clockwise left side, anticlockwise right side; from gland up to muscular spiral section, a short non-muscular duct section spirals in the reverse direction on each side.
|
||
</paragraph>
|
||
<paragraph id="ED3B141E905EFFFBFF2B38C1D6C2B629" blockId="5.[143,1104,167,797]" pageId="5" pageNumber="254">Ovaries in XIII. Paired spermathecae in V–IX, each ampulla ovate to blocky, duct shorter than ampulla. Diverticulum with short straight stalk, elongate chamber coiled and tightly bound into approximately oval outline, no nephridia on spermathecae ducts.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A59E4795905EFFFBFF1E3905D6EAB3AB" pageId="5" pageNumber="254" type="discussion">
|
||
<paragraph id="ED3B141E905EFFFBFF1E3905D6E2B64E" blockId="5.[143,1104,866,1695]" box="[143,241,866,890]" pageId="5" pageNumber="254">
|
||
<heading id="B673A372905EFFFBFF1E3905D6E2B64E" box="[143,241,866,890]" fontSize="10" level="3" pageId="5" pageNumber="254" reason="5">
|
||
<emphasis id="DFF0C80C905EFFFBFF1E3905D6E2B64E" box="[143,241,866,890]" italics="true" pageId="5" pageNumber="254">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="ED3B141E905EFFFBFF1E39EDD6EAB3AB" blockId="5.[143,1104,866,1695]" pageId="5" pageNumber="254">
|
||
We are considering the very shallow invagination of the male pores and genital papillae as belonging to
|
||
<taxonomicName id="2A846F9D905EFFFBFE3B39CDD40AB6F6" authorityName="Kinberg" authorityYear="1867" box="[426,537,938,962]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFE3B39CDD40AB6F6" box="[426,537,938,962]" italics="true" pageId="5" pageNumber="254">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
, because there is no resemblance to the coelomic copulatory pouches said to be held in common between
|
||
<taxonomicName id="2A846F9D905EFFFBFCBA39ADD58DB6D6" authorityName="Sims & Easton" authorityYear="1972" box="[811,926,970,994]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFCBA39ADD58DB6D6" box="[811,926,970,994]" italics="true" pageId="5" pageNumber="254">Metaphire</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2A846F9D905EFFFBFC4F39ADD243B6D6" box="[990,1104,970,994]" class="Clitellata" family="Megascolecidae" genus="Pheretima" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFC4F39ADD243B6D6" box="[990,1104,970,994]" italics="true" pageId="5" pageNumber="254">Pheretima</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="891569EF905EFFFBFF20398ED7DEB135" author="Sims RW & Easton EG" box="[177,461,1001,1025]" pageId="5" pageNumber="254" pagination="169 - 268" refId="ref2382" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linn Soc. 4: 169 - 268." type="journal article" year="1972">Sims and Easton (1972)</bibRefCitation>
|
||
. Furthermore those authors state that their
|
||
<taxonomicName id="2A846F9D905EFFFBFC71398DD25CB136" authorityName="Kinberg" authorityYear="1867" box="[992,1103,1002,1026]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFC71398DD25CB136" box="[992,1103,1002,1026]" italics="true" pageId="5" pageNumber="254">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
concept was simplified by deliberately excluding six species clearly clustering with
|
||
<taxonomicName id="2A846F9D905EFFFBFF1E3E4ED6EDB175" authorityName="Kinberg" authorityYear="1867" box="[143,254,1065,1089]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFF1E3E4ED6EDB175" box="[143,254,1065,1089]" italics="true" pageId="5" pageNumber="254">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
in their phenetic analysis, and transferring said species to
|
||
<taxonomicName id="2A846F9D905EFFFBFC443E4ED25BB175" authorityName="Sims & Easton" authorityYear="1972" box="[981,1096,1065,1089]" class="Clitellata" family="Megascolecidae" genus="Metaphire" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFC443E4ED25BB175" box="[981,1096,1065,1089]" italics="true" pageId="5" pageNumber="254">Metaphire</emphasis>
|
||
</taxonomicName>
|
||
. We see no reason to make it more so.
|
||
<taxonomicName id="2A846F9D905EFFFBFDC43E2ED571B155" authority="Hong & James & Inkhavilay, 2013" authorityName="Hong & James & Inkhavilay" authorityYear="2013" box="[597,866,1097,1121]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="hauyhowensis" status="sp. nov.">
|
||
<emphasis id="DFF0C80C905EFFFBFDC43E2ED571B155" box="[597,866,1097,1121]" italics="true" pageId="5" pageNumber="254">Amynthas hauyhowensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4C37577905EFFFBFCFC3E2DD5D6B155" box="[877,965,1098,1121]" pageId="5" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
|
||
bears character states in common with several Lao
|
||
<taxonomicName id="2A846F9D905EFFFBFDF83E0ED53DB1B5" box="[617,814,1129,1153]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="DFF0C80C905EFFFBFDF83E0ED4CBB1B5" box="[617,728,1129,1153]" italics="true" pageId="5" pageNumber="254">Amynthas</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="2A846F9D905EFFFBFC123E0DD217B1B5" box="[899,1028,1130,1153]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="aeruginosus">
|
||
<emphasis id="DFF0C80C905EFFFBFC123E0DD217B1B5" box="[899,1028,1130,1153]" italics="true" pageId="5" pageNumber="254">aeruginosus</emphasis>
|
||
</taxonomicName>
|
||
-group (spermathecal pores 7
|
||
<emphasis id="DFF0C80C905EFFFBFE163EEED780B195" bold="true" box="[391,403,1161,1185]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
8
|
||
<emphasis id="DFF0C80C905EFFFBFE303EEED7BEB195" bold="true" box="[417,429,1161,1185]" italics="true" pageId="5" pageNumber="254">/</emphasis>
|
||
9), notably the mirror-image coiled prostatic ducts and lack of lymph glands on the intestine (
|
||
<bibRefCitation id="891569EF905EFFFBFD953ECED492B1F5" author="Hong Y" box="[516,641,1193,1217]" pageId="5" pageNumber="254" pagination="599 - 609" refId="ref2288" refString="Hong Y. 2010. Four new earthworms of the Amynthas aeruginosus species group (Oligochaeta: Megascolecidae) from Nam Xam NBCA, Laos. Rev Suisse Zool. 117: 599 - 609." type="journal article" year="2010">Hong 2010</bibRefCitation>
|
||
).
|
||
<taxonomicName id="2A846F9D905EFFFBFD063ECED5B3B1F5" authority="Hong & James & Inkhavilay, 2013" authorityName="Hong & James & Inkhavilay" authorityYear="2013" box="[663,928,1193,1217]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="hauyhowensis" status="sp. nov.">
|
||
<emphasis id="DFF0C80C905EFFFBFD063ECED5B3B1F5" box="[663,928,1193,1217]" italics="true" pageId="5" pageNumber="254">Amynthas hauyhowensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C4C37577905EFFFBFC363ECED5E8B1F4" box="[935,1019,1193,1216]" pageId="5" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
|
||
keys to the
|
||
<taxonomicName id="2A846F9D905EFFFBFF2C3EAED725B1D5" baseAuthorityName="Benham" baseAuthorityYear="1896" box="[189,310,1225,1249]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="hexathecus">
|
||
<emphasis id="DFF0C80C905EFFFBFF2C3EAED725B1D5" box="[189,310,1225,1249]" italics="true" pageId="5" pageNumber="254">hexathecus</emphasis>
|
||
</taxonomicName>
|
||
group in
|
||
<bibRefCitation id="891569EF905EFFFBFE3F3EAFD4D4B1D4" author="Sims RW & Easton EG" box="[430,711,1224,1248]" pageId="5" pageNumber="254" pagination="169 - 268" refId="ref2382" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linn Soc. 4: 169 - 268." type="journal article" year="1972">Sims and Easton (1972)</bibRefCitation>
|
||
. No such Lao earthworm species have been recorded (
|
||
<bibRefCitation id="891569EF905EFFFBFE153E8FD4A5B034" author="Thai TB & Samphon K" box="[388,694,1256,1280]" pageId="5" pageNumber="254" pagination="86 - 89" refId="ref2575" refString="Thai TB, Samphon K. 1991 a. A list of discovered species of earthworms fron Laos. Sci Inf Hanoi Nat Ped Inst: 86 - 89. (Vietnamese)." type="book chapter" year="1991">Thai and Samphon 1991a</bibRefCitation>
|
||
). The position of spermathecae in V–IX is unusual in
|
||
<collectingCountry id="9593548E905EFFFBFEEE3F6ED7ABB014" box="[383,440,1289,1312]" name="Laos" pageId="5" pageNumber="254">Laos</collectingCountry>
|
||
, and very uncommon in
|
||
<taxonomicName id="2A846F9D905EFFFBFD7D3F6ED548B015" authorityName="Kinberg" authorityYear="1867" box="[748,859,1289,1313]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFD7D3F6ED548B015" box="[748,859,1289,1313]" italics="true" pageId="5" pageNumber="254">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
generally (
|
||
<bibRefCitation id="891569EF905EFFFBFC703F6FD70EB074" author="Sims RW & Easton EG" pageId="5" pageNumber="254" pagination="169 - 268" refId="ref2382" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linn Soc. 4: 169 - 268." type="journal article" year="1972">Sims and Easton 1972</bibRefCitation>
|
||
). Of the 10 species listed by
|
||
<bibRefCitation id="891569EF905EFFFBFDFF3F4FD592B074" author="Sims RW & Easton EG" box="[622,897,1320,1344]" pageId="5" pageNumber="254" pagination="169 - 268" refId="ref2382" refString="Sims RW, Easton EG. 1972. A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. Biol J Linn Soc. 4: 169 - 268." type="journal article" year="1972">Sims and Easton (1972)</bibRefCitation>
|
||
in the
|
||
<taxonomicName id="2A846F9D905EFFFBFC473F4FD25CB074" baseAuthorityName="Benham" baseAuthorityYear="1896" box="[982,1103,1320,1344]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="hexathecus">
|
||
<emphasis id="DFF0C80C905EFFFBFC473F4FD25CB074" box="[982,1103,1320,1344]" italics="true" pageId="5" pageNumber="254">hexathecus</emphasis>
|
||
</taxonomicName>
|
||
group, four are referable to
|
||
<taxonomicName id="2A846F9D905EFFFBFE463F2FD455B054" authorityName="Kinberg" authorityYear="1867" box="[471,582,1352,1376]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="genus">
|
||
<emphasis id="DFF0C80C905EFFFBFE463F2FD455B054" box="[471,582,1352,1376]" italics="true" pageId="5" pageNumber="254">Amynthas</emphasis>
|
||
</taxonomicName>
|
||
based on somatic characters. The remaining species with the same spermathecal location are
|
||
<taxonomicName id="2A846F9D905EFFFBFD223F0FD258B0B4" authority="(Gates, 1958)" baseAuthorityName="Gates" baseAuthorityYear="1958" box="[691,1099,1384,1408]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="brevicinctus">
|
||
<emphasis id="DFF0C80C905EFFFBFD223F0FD5B8B0B4" box="[691,939,1384,1408]" italics="true" pageId="5" pageNumber="254">Amynthas brevicinctus</emphasis>
|
||
(
|
||
<bibRefCitation id="891569EF905EFFFBFC2A3F0FD250B0B4" author="Gates GE" box="[955,1091,1384,1408]" pageId="5" pageNumber="254" pagination="1 - 33" refId="ref2191" refString="Gates GE. 1958. On some species of the Oriental earthworm genus Pheretima Kinberg, 1867, with a key to species reported from the Americas. Am Mus Novit. 1888: 1 - 33." type="journal article" year="1958">Gates, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2A846F9D905EFFFBFF1E3FEFD407B094" authority="(Gates, 1949)" baseAuthorityName="Gates" baseAuthorityYear="1949" box="[143,532,1416,1440]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="flustrellus">
|
||
<emphasis id="DFF0C80C905EFFFBFF1E3FEFD762B094" box="[143,369,1416,1440]" italics="true" pageId="5" pageNumber="254">Amynthas flustrellus</emphasis>
|
||
(
|
||
<bibRefCitation id="891569EF905EFFFBFE133FEFD41FB094" author="Gates GE" box="[386,524,1416,1440]" pageId="5" pageNumber="254" pagination="5 - 38" refId="ref2168" refString="Gates GE. 1949. On some earthworms from Perlis and Kedah. Bull Raffles Mus. 19: 5 - 38." type="journal article" year="1949">Gates, 1949</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2A846F9D905EFFFBFDB03FEFD5C0B094" authority="(Benham, 1896)" baseAuthorityName="Benham" baseAuthorityYear="1896" box="[545,979,1416,1440]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="hexathecus">
|
||
<emphasis id="DFF0C80C905EFFFBFDB03FEFD501B094" box="[545,786,1416,1440]" italics="true" pageId="5" pageNumber="254">Amynthas hexathecus</emphasis>
|
||
(
|
||
<bibRefCitation id="891569EF905EFFFBFCB23FEFD5D8B094" author="Benham WB" box="[803,971,1416,1440]" pageId="5" pageNumber="254" pagination="429 - 448" refId="ref2098" refString="Benham WB. 1896. Some earthworms from Celebes. Ann Mag Nat Hist (ser 6). 18: 429 - 448." type="journal article" year="1896">Benham, 1896</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2A846F9D905EFFFBFC713FEFD7EAB08B" authority="(Michaelsen, 1928)" baseAuthorityName="Michaelsen" baseAuthorityYear="1928" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="polytoreutus">
|
||
<emphasis id="DFF0C80C905EFFFBFC713FEFD705B0F4" italics="true" pageId="5" pageNumber="254">Amynthas polytoreutus</emphasis>
|
||
(
|
||
<bibRefCitation id="891569EF905EFFFBFEB63FCFD7E2B08B" author="Michaelsen W" box="[295,497,1447,1472]" pageId="5" pageNumber="254" pagination="1 - 60" refId="ref2364" refString="Michaelsen W. 1928. Die Oligochaten Borneos. Ark Zool. 20: 1 - 60." type="journal article" year="1928">Michaelsen, 1928</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2A846F9D905EFFFBFD973FCFD20BB08B" authority="(Goto and Hatai, 1898)" baseAuthorityName="Goto and Hatai" baseAuthorityYear="1898" box="[518,1048,1447,1472]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="scholasticus">
|
||
<emphasis id="DFF0C80C905EFFFBFD973FCFD4ECB0F4" box="[518,767,1448,1472]" italics="true" pageId="5" pageNumber="254">Amynthas scholasticus</emphasis>
|
||
(
|
||
<bibRefCitation id="891569EF905EFFFBFC9E3FC0D203B08B" author="Goto S & Hatai S" box="[783,1040,1447,1471]" pageId="5" pageNumber="254" pagination="65 - 78" refId="ref2229" refString="Goto S, Hatai S. 1898. New or imperfectly known species of earthworms. No. 1. Annot Zool Jap. 2: 65 - 78." type="journal article" year="1898">Goto and Hatai, 1898</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2A846F9D905EFFFBFF1E3FAFD43DB0EB" authority="(Beddard, 1899)" baseAuthorityName="Beddard" baseAuthorityYear="1899" box="[143,558,1479,1504]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="solomonis">
|
||
<emphasis id="DFF0C80C905EFFFBFF1E3FAFD763B0D4" box="[143,368,1480,1504]" italics="true" pageId="5" pageNumber="254">Amynthas solomonis</emphasis>
|
||
(
|
||
<bibRefCitation id="891569EF905EFFFBFEEF3FA0D434B0EB" author="Beddard FE" box="[382,551,1479,1503]" pageId="5" pageNumber="254" pagination="181 - 194" refId="ref2058" refString="Beddard FE. 1899. On a collection of earthworms from New Britain, the Solomon Islands, The New Hebrides and the Loyalty Islands. In A. Willey, Zool Res. 2: 181 - 194." type="journal article" year="1899">Beddard, 1899</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="2A846F9D905EFFFBFDA93FAFD53CB0D4" baseAuthorityName="Goto and Hatai" baseAuthorityYear="1898" box="[568,815,1480,1504]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="scholasticus">
|
||
<emphasis id="DFF0C80C905EFFFBFDA93FAFD53CB0D4" box="[568,815,1480,1504]" italics="true" pageId="5" pageNumber="254">Amynthas scholasticus</emphasis>
|
||
</taxonomicName>
|
||
has its last hearts in XIII, fewer setae and only four pairs of spermathecae in V–VIII.
|
||
<taxonomicName id="2A846F9D905EFFFBFCDF3F80D25BB0CB" baseAuthorityName="Gates" baseAuthorityYear="1958" box="[846,1096,1511,1535]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="brevicinctus">
|
||
<emphasis id="DFF0C80C905EFFFBFCDF3F80D25BB0CB" box="[846,1096,1511,1535]" italics="true" pageId="5" pageNumber="254">Amynthas brevicinctus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2A846F9D905EFFFBFF1E3C60D733B32B" baseAuthorityName="Beddard" baseAuthorityYear="1899" box="[143,288,1543,1567]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="solomonis">
|
||
<emphasis id="DFF0C80C905EFFFBFF1E3C60D6B1B32B" box="[143,162,1543,1567]" italics="true" pageId="5" pageNumber="254">A</emphasis>
|
||
.
|
||
<emphasis id="DFF0C80C905EFFFBFF253C60D733B32B" box="[180,288,1543,1567]" italics="true" pageId="5" pageNumber="254">solomonis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2A846F9D905EFFFBFEA33C60D7D2B32B" baseAuthorityName="Gates" baseAuthorityYear="1949" box="[306,449,1543,1567]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="flustrellus">
|
||
<emphasis id="DFF0C80C905EFFFBFEA33C60D756B32B" box="[306,325,1543,1567]" italics="true" pageId="5" pageNumber="254">A</emphasis>
|
||
.
|
||
<emphasis id="DFF0C80C905EFFFBFEC63C60D7D2B32B" box="[343,449,1543,1567]" italics="true" pageId="5" pageNumber="254">flustrellus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2A846F9D905EFFFBFD913C60D4BFB32B" baseAuthorityName="Michaelsen" baseAuthorityYear="1928" box="[512,684,1543,1567]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="polytoreutus">
|
||
<emphasis id="DFF0C80C905EFFFBFD913C60D400B32B" box="[512,531,1543,1567]" italics="true" pageId="5" pageNumber="254">A</emphasis>
|
||
.
|
||
<emphasis id="DFF0C80C905EFFFBFDB43C60D4BFB32B" box="[549,684,1543,1567]" italics="true" pageId="5" pageNumber="254">polytoreutus</emphasis>
|
||
</taxonomicName>
|
||
all have superficial male pores, and each differs in genital marking numbers and locations, with none being invaginated. The last two also have their last hearts in XIII.
|
||
<taxonomicName id="2A846F9D905EFFFBFD2E3C20D5A2B36B" baseAuthorityName="Benham" baseAuthorityYear="1896" box="[703,945,1607,1631]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="5" pageNumber="254" phylum="Annelida" rank="species" species="hexathecus">
|
||
<emphasis id="DFF0C80C905EFFFBFD2E3C20D5A2B36B" box="[703,945,1607,1631]" italics="true" pageId="5" pageNumber="254">Amynthas hexathecus</emphasis>
|
||
</taxonomicName>
|
||
has far more setae (∼
|
||
<quantity id="2A7CB9FB905EFFFBFF7E3C00D72BB34B" box="[239,312,1639,1663]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="254" unit="in" value="100.0">100 in</quantity>
|
||
post-clitellar segments) and genital papillae in XVII, XIX and XX (
|
||
<tableCitation id="A00621A5905EFFFBFF093CE0D6FAB3AB" box="[152,233,1671,1695]" captionStart="Table 1" captionStartId="6.[58,113,525,547]" captionText="Table 1. Comparison of species of the genus Amynthas from Nam Xam National Protected Area, Laos." pageId="5" pageNumber="254">Table 1</tableCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |