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<document id="E4E4ED0B6949E27F23D04DF7C5939243" ID-CLB-Dataset="21174" ID-DOI="10.11646/zootaxa.4629.3.1" ID-GBIF-Dataset="9419bc9b-dbe4-426c-a34e-f0034848ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995508" ID-ZooBank="1D47B452-0A40-4BC5-9BBA-C06DEB298C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598059654494" checkinUser="felipe" docAuthor="Silva, Fernando A. B. &amp; Valois, Marcely" docDate="2019" docId="9D12E5573A44FF8395F1F8D9FB7DC361" docLanguage="en" docName="zootaxa.4629.3.1.pdf" docOrigin="Zootaxa 4629 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scybalocanthon papaxibe Silva &amp; Valois 2019, new species" docType="treatment" docVersion="8" lastPageNumber="330" masterDocId="612B9D2F3A59FF9E9566F840FFF1C516" masterDocTitle="A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="330" updateTime="1698845243763" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3253D7DF7A3EC5550E0DBE8A291E5BA6">A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:namePart id="1FD693E7CC708BE8B4F6E0C16D708AE5">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="51C6C85ACAE5AFEED23280BD19C741EC">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br</mods:affiliation>
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<mods:namePart id="67011E132991675972A1B8EFA5F249E1">Valois, Marcely</mods:namePart>
<mods:affiliation id="7685F5DCD595074778A8BA645849E4BC">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<mods:date id="85EE75F9A67C27060A54DBC9AA32AA20">2019</mods:date>
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<mods:number id="19C93926972CADD604295638F030A6C4">2019-07-08</mods:number>
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<treatment id="9D12E5573A44FF8395F1F8D9FB7DC361" ID-DOI="http://doi.org/10.5281/zenodo.4335184" ID-GBIF-Taxon="167293231" ID-Zenodo-Dep="4335184" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361" httpUri="http://treatment.plazi.org/id/9D12E5573A44FF8395F1F8D9FB7DC361" lastPageNumber="330" pageId="29" pageNumber="330">
<subSubSection id="5DA107CA3A44FF8395F1F8D9FCD5C5A5" box="[151,804,153,180]" pageId="29" pageNumber="330" type="nomenclature">
<paragraph id="150454413A44FF8395F1F8D9FCD5C5A5" blockId="29.[151,804,153,216]" box="[151,804,153,180]" pageId="29" pageNumber="330">
<heading id="4E4CE32D3A44FF8395F1F8D9FCD5C5A5" bold="true" box="[151,804,153,180]" fontSize="11" level="1" pageId="29" pageNumber="330" reason="1">
<emphasis id="27CF88533A44FF8395F1F8D9FCD5C5A5" bold="true" box="[151,804,153,180]" pageId="29" pageNumber="330">
<taxonomicName id="D2BB2FC23A44FF8395F1F8D9FD77C5A5" ID-CoL="4W3LT" authority="Silva &amp; Valois" authorityName="Silva &amp; Valois" authorityYear="2019" box="[151,646,153,180]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="papaxibe" status="sp. nov.">
<emphasis id="27CF88533A44FF8395F1F8D9FE3CC5A2" bold="true" box="[151,461,153,180]" italics="true" pageId="29" pageNumber="330">Scybalocanthon papaxibe</emphasis>
Silva &amp; Valois
</taxonomicName>
,
<taxonomicNameLabel id="3CFC35283A44FF8397F2F8DAFCD5C5A5" box="[660,804,153,180]" pageId="29" pageNumber="330" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A44FF8395F1F8FDFE65C5CE" box="[151,404,189,216]" pageId="29" pageNumber="330" type="description">
<paragraph id="150454413A44FF8395F1F8FDFE65C5CE" blockId="29.[151,804,153,216]" box="[151,404,189,216]" pageId="29" pageNumber="330">
(
<figureCitation id="8D8048C43A44FF8395F9F8FDFF0FC5C1" box="[159,254,189,215]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="29" pageNumber="330">Figs. 1T</figureCitation>
,
<figureCitation id="8D8048C43A44FF83946CF8FDFEDCC5C1" box="[266,301,189,215]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">4R</figureCitation>
,
<figureCitation id="8D8048C43A44FF83945EF8FDFEACC5CE" box="[312,349,189,216]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="29" pageNumber="330">5Q</figureCitation>
,
<figureCitation id="8D8048C43A44FF83940FF8FDFE7DC5CE" box="[361,396,189,216]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="29" pageNumber="330">6D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A44FF8395F1F946FB9FC471" pageId="29" pageNumber="330" type="etymology">
<paragraph id="150454413A44FF8395F1F946FB9FC471" blockId="29.[151,1437,261,1656]" pageId="29" pageNumber="330">
<emphasis id="27CF88533A44FF8395F1F946FEEFC436" bold="true" box="[151,286,262,288]" pageId="29" pageNumber="330">Etymology.</emphasis>
The new specific name, “papaxibe” [papa = eater; xibé = cassava flour porridge], is a typical term in the Pará state in
<collectingCountry id="6DAC14D13A44FF839440F969FE9FC455" box="[294,366,297,323]" name="Brazil" pageId="29" pageNumber="330">Brazil</collectingCountry>
. This term is often attributed to people who are born in Pará due to the old habit of consuming a
<typeStatus id="CA00EAE33A44FF8395F1F90EFF36C47E" box="[151,199,334,360]" pageId="29" pageNumber="330">type</typeStatus>
of cassava flour porridge. The specific name should be treated as a Latin adjective.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A44FF8395A1F931FA84C62D" pageId="29" pageNumber="330" type="diagnosis">
<paragraph id="150454413A44FF8395A1F931FA84C62D" blockId="29.[151,1437,261,1656]" pageId="29" pageNumber="330">
<emphasis id="27CF88533A44FF8395A1F931FECBC49A" bold="true" box="[199,314,369,396]" pageId="29" pageNumber="330">Diagnosis</emphasis>
. Specimens of
<taxonomicName id="D2BB2FC23A44FF83948BF932FD83C49D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[493,626,369,396]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="papaxibe">
<emphasis id="27CF88533A44FF83948BF932FD83C49D" box="[493,626,369,396]" italics="true" pageId="29" pageNumber="330">S. papaxibe</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="D2BB2FC23A44FF839619F932FC07C49D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[895,1014,369,396]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="27CF88533A44FF839619F932FC07C49D" box="[895,1014,369,396]" italics="true" pageId="29" pageNumber="330">S. arnaudi</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A44FF839160F932FB2EC49D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[1030,1247,369,396]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="federicoescobari">
<emphasis id="27CF88533A44FF839160F932FB2EC49D" box="[1030,1247,369,396]" italics="true" pageId="29" pageNumber="330">S. federicoescobari</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A44FF839196F932FA89C49D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[1264,1400,370,396]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="27CF88533A44FF839196F932FA89C49D" box="[1264,1400,370,396]" italics="true" pageId="29" pageNumber="330">S. martinezi</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A44FF8390E1F932FEFEC4B9" baseAuthorityName="Kirsch" baseAuthorityYear="1873" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="pinopterus">
<emphasis id="27CF88533A44FF8390E1F932FEFEC4B9" italics="true" pageId="29" pageNumber="330">S. pinopterus</emphasis>
</taxonomicName>
, and
<taxonomicName id="D2BB2FC23A44FF83942DF9D6FE02C4B9" authorityName="Silva &amp; Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[331,499,405,432]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="uniplagiatus">
<emphasis id="27CF88533A44FF83942DF9D6FE02C4B9" box="[331,499,405,432]" italics="true" pageId="29" pageNumber="330">S. uniplagiatus</emphasis>
</taxonomicName>
by having the pronotum uniformly colored, or with one elliptical spot on the central portion (
<figureCitation id="8D8048C43A44FF839591F9F9FEA4C4C5" box="[247,341,441,467]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="29" pageNumber="330">Figs. 1T</figureCitation>
,
<figureCitation id="8D8048C43A44FF839406F9F9FE75C4C5" box="[352,388,441,467]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="29" pageNumber="330">2A</figureCitation>
,
<figureCitation id="8D8048C43A44FF8394F6F9F9FE5CC4C5" box="[400,429,441,467]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="29" pageNumber="330">1P</figureCitation>
,
<figureCitation id="8D8048C43A44FF8394DEF9F9FE2DC4C5" box="[440,476,441,467]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="29" pageNumber="330">2K</figureCitation>
,
<figureCitation id="8D8048C43A44FF839481F9F9FDDEC4C5" box="[487,559,441,467]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="29" pageNumber="330">1D, 1I</figureCitation>
); femora almost completely yellow or brown, with black spots only on the tips (
<figureCitation id="8D8048C43A44FF8395C6F99DFF0AC4E1" box="[160,251,477,503]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="29" pageNumber="330">Fig. 3G</figureCitation>
); eighth elytral stria with thin carina on anterior portion (
<figureCitation id="8D8048C43A44FF8396E3F99DFC11C4E1" box="[901,992,477,503]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="29" pageNumber="330">Fig. 3D</figureCitation>
); endophallus with bristles right beside the FLP sclerite (
<figureCitation id="8D8048C43A44FF83943DFA41FDA5C70A" box="[347,596,513,540]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="29" pageNumber="330">Figs. 5Q, N, W, D, H</figureCitation>
); and additional sclerite (
<collectingCountry id="6DAC14D13A44FF839612FA41FC6BC70D" box="[884,922,513,539]" name="American Samoa" pageId="29" pageNumber="330">AS</collectingCountry>
) absent (
<figureCitation id="8D8048C43A44FF839165FA41FB0DC70A" box="[1027,1276,513,540]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="29" pageNumber="330">Figs. 5Q, N, W, D, H</figureCitation>
). They can be distinguished from those of
<taxonomicName id="D2BB2FC23A44FF8394BCFA66FD80C729" baseAuthorityName="Kirsch" baseAuthorityYear="1873" box="[474,625,550,576]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="pinopterus">
<emphasis id="27CF88533A44FF8394BCFA66FD80C729" box="[474,625,550,576]" italics="true" pageId="29" pageNumber="330">S. pinopterus</emphasis>
</taxonomicName>
and
<taxonomicName id="D2BB2FC23A44FF8397CDFA66FCD2C729" authorityName="Silva &amp; Valois" authorityYear="2019" box="[683,803,549,576]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="27CF88533A44FF8397CDFA66FCD2C729" box="[683,803,549,576]" italics="true" pageId="29" pageNumber="330">S. arnaudi</emphasis>
</taxonomicName>
by the strongly asymmetrical parameres; left paramere with acute projection on dorsal margin and bilobate excavation on ventral margin (
<figureCitation id="8D8048C43A44FF83913EFA09FB40C775" box="[1112,1201,585,611]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Fig. 4R</figureCitation>
) (other species with slightly asymmetrical parameres, lacking acute projection and bilobate excavation on dorsal and ventral margins, respectively (
<figureCitation id="8D8048C43A44FF83944BFAD1FE58C7BA" box="[301,425,657,684]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Figs. 4S, E</figureCitation>
)); from those of
<taxonomicName id="D2BB2FC23A44FF839705FAD2FCFBC7BD" authorityName="Silva &amp; Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[611,778,657,684]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="uniplagiatus">
<emphasis id="27CF88533A44FF839705FAD2FCFBC7BD" box="[611,778,657,684]" italics="true" pageId="29" pageNumber="330">S. uniplagiatus</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A44FF839671FAD2FC6DC7BD" authorityName="Silva &amp; Valois" authorityYear="2019" box="[791,924,658,684]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="27CF88533A44FF839671FAD2FC6DC7BD" box="[791,924,658,684]" italics="true" pageId="29" pageNumber="330">S. martinezi</emphasis>
</taxonomicName>
, and
<taxonomicName id="D2BB2FC23A44FF8396BEFAD2FB5EC7BD" authorityName="Silva &amp; Valois" authorityYear="2019" box="[984,1199,657,684]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="federicoescobari">
<emphasis id="27CF88533A44FF8396BEFAD2FB5EC7BD" box="[984,1199,657,684]" italics="true" pageId="29" pageNumber="330">S. federicoescobari</emphasis>
</taxonomicName>
by the bilobate excavation on the ventral margin of the left paramere wider than deep in lateral view, extending along one-fourth of the paramere margin in lateral view (
<figureCitation id="8D8048C43A44FF83976DFA99FD90C7E5" box="[523,609,729,755]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Fig. 4R</figureCitation>
) (in
<taxonomicName id="D2BB2FC23A44FF8397F0FA9AFCEAC7E5" authorityName="Silva &amp; Valois" authorityYear="2019" box="[662,795,730,756]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="27CF88533A44FF8397F0FA9AFCEAC7E5" box="[662,795,730,756]" italics="true" pageId="29" pageNumber="330">S. martinezi</emphasis>
</taxonomicName>
the bilobate excavation is wider and deeper, extending at least one-half of the paramere margin (
<figureCitation id="8D8048C43A44FF839729FABDFD5BC601" box="[591,682,765,791]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Fig. 4O</figureCitation>
); in
<taxonomicName id="D2BB2FC23A44FF8397BBFABEFC74C601" authorityName="Silva &amp; Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[733,901,765,792]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="uniplagiatus">
<emphasis id="27CF88533A44FF8397BBFABEFC74C601" box="[733,901,765,792]" italics="true" pageId="29" pageNumber="330">S. uniplagiatus</emphasis>
</taxonomicName>
its excavation extending along one-third of the paramere margin (
<figureCitation id="8D8048C43A44FF839402FB61FE35C62D" box="[356,452,801,827]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Fig. 4W</figureCitation>
)); in
<taxonomicName id="D2BB2FC23A44FF839499FB62FD26C62D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[511,727,801,828]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="330" phylum="Arthropoda" rank="species" species="federicoescobari">
<emphasis id="27CF88533A44FF839499FB62FD26C62D" box="[511,727,801,828]" italics="true" pageId="29" pageNumber="330">S. federicoescobari</emphasis>
</taxonomicName>
its excavation is deeper than wide in lateral view (
<figureCitation id="8D8048C43A44FF839076FB61FAAFC62D" box="[1296,1374,801,827]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Fig. 4I</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A44FF8395A1FB05FBA1C1B2" pageId="29" pageNumber="330" type="description">
<paragraph id="150454413A44FF8395A1FB05FBA1C1B2" blockId="29.[151,1437,261,1656]" pageId="29" pageNumber="330">
<emphasis id="27CF88533A44FF8395A1FB05FE54C676" bold="true" box="[199,421,837,864]" pageId="29" pageNumber="330">Description. Body.</emphasis>
Oval-elongate. Surface opaque, completely microgranulate.
<emphasis id="27CF88533A44FF83913FFB05FB6CC676" bold="true" box="[1113,1181,837,864]" pageId="29" pageNumber="330">Color</emphasis>
. Head and elytra dark brown. Other parts yellow or light brown.
<emphasis id="27CF88533A44FF83970DFB2AFD39C692" bold="true" box="[619,712,874,900]" pageId="29" pageNumber="330">Length.</emphasis>
<quantity id="D243F9A43A44FF8397A8FB29FCA6C692" box="[718,855,873,900]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="9.5" metricValueMin="7.8" pageId="29" pageNumber="330" unit="mm" value="8.65" valueMax="9.5" valueMin="7.8">7.8–9.5 mm</quantity>
.
<emphasis id="27CF88533A44FF839606FB2AFC48C692" bold="true" box="[864,953,874,900]" pageId="29" pageNumber="330">Thorax</emphasis>
. Anterior angles of pronotum approximately 85°. Lateral margin regularly curved outward, not forming an angle at the middle portion.
<emphasis id="27CF88533A44FF8391C6FBCEFB1DC6BE" bold="true" box="[1184,1260,910,936]" pageId="29" pageNumber="330">Elytra</emphasis>
. Striae thin and shiny, weakly impressed, punctures inconspicuous. Eighth stria with a thin carina at the anterior portion.
<emphasis id="27CF88533A44FF839044FBF1FA6DC6DA" bold="true" box="[1314,1436,945,972]" pageId="29" pageNumber="330">Aedeagus.</emphasis>
Parameres strongly asymmetrical (
<figureCitation id="8D8048C43A44FF839779FB95FD88C6F9" box="[543,633,981,1007]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="29" pageNumber="330">Fig. 4R</figureCitation>
). Dorsal margin of right paramere curved inward, apex obliquely truncate. Ventral margin of right paramere substraight, with a rounded excavation at the basal portion. Dorsal margin of left paramere curved inward from the basal to medial portions, apical portion with a short and pointed projection. Ventral margin of left paramere substraight, with a bilobate excavation at the basal portion, apical portion pointed. SRP circular, with rounded handle-shaped extension (
<figureCitation id="8D8048C43A44FF83978DFC25FCB2C196" box="[747,835,1125,1152]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="29" pageNumber="330">Fig. 5Q</figureCitation>
). FLP short, comma-shaped, with three sets of bristles right beside it (
<figureCitation id="8D8048C43A44FF839427FCC9FE6BC1B2" box="[321,410,1161,1188]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="29" pageNumber="330">Fig. 5Q</figureCitation>
). A+SA with two superposed and elongate sclerites (
<figureCitation id="8D8048C43A44FF83968FFCC9FBB2C1B2" box="[1001,1091,1161,1188]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="29" pageNumber="330">Fig. 5Q</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A44FF8395A1FCEEFE41C339" pageId="29" pageNumber="330" type="materials_examined">
<paragraph id="150454413A44FF8395A1FCEEFE41C339" blockId="29.[151,1437,261,1656]" pageId="29" pageNumber="330">
<emphasis id="27CF88533A44FF8395A1FCEEFE87C1DE" bold="true" box="[199,374,1197,1224]" pageId="29" pageNumber="330">Type material.</emphasis>
<materialsCitation id="A5D35E1C3A44FF839418FCEEFE24C1FD" ID-GBIF-Occurrence="2847261328" collectingDate="1998-10-12" collectionCode="CEMT" collectorName="O.P. Young" country="Brazil" latitude="-7.8333335" location="Fazenda Marajoara" longLatPrecision="1302" longitude="-50.266666" municipality="Pau d'Arco" pageId="29" pageNumber="330" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="27CF88533A44FF839418FCEEFD9EC1DE" bold="true" box="[382,623,1197,1224]" pageId="29" pageNumber="330">
<typeStatus id="CA00EAE33A44FF839418FCEEFE1CC1DE" box="[382,493,1198,1224]" pageId="29" pageNumber="330" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="6DAC14D13A44FF83949DFCEDFD9AC1DE" box="[507,619,1197,1224]" name="Brazil" pageId="29" pageNumber="330">BRAZIL</collectingCountry>
:
</emphasis>
PARÁ:
<collectingMunicipality id="F560CE3B3A44FF8397B6FCEEFCA5C1D1" box="[720,852,1197,1224]" pageId="29" pageNumber="330">Pau d’Arco</collectingMunicipality>
,
<location id="1064029A3A44FF839607FCEDFBC8C1D1" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF839607FCEDFBC8C1D1" box="[865,1081,1197,1223]" country="Brazil" latitude="-7.8333335" longLatPrecision="1302" longitude="-50.266666" municipality="Pau d'Arco" name="Fazenda Marajoara" pageId="29" pageNumber="330">Fazenda Marajoara</location>
(
<geoCoordinate id="708F32863A44FF83912CFCEDFB43C1DE" box="[1098,1202,1197,1224]" degrees="50" direction="west" minutes="16" orientation="longitude" pageId="29" pageNumber="330" precision="925" value="-50.266666">50°16’W</geoCoordinate>
,
<geoCoordinate id="708F32863A44FF8391D8FCEDFAFEC1DE" box="[1214,1295,1197,1224]" degrees="7" direction="south" minutes="50" orientation="latitude" pageId="29" pageNumber="330" precision="925" value="-7.8333335">7°50’S</geoCoordinate>
),
<date id="610572813A44FF839043FCEDFA66C1D1" box="[1317,1431,1197,1223]" pageId="29" pageNumber="330" value="1998-10-12">
<collectingDate id="71418B693A44FF839043FCEDFA66C1D1" box="[1317,1431,1197,1223]" pageId="29" pageNumber="330" value="1998-10-12">12.x.1998</collectingDate>
</date>
,
<collectorName id="B84E31973A44FF8395F1FC92FF32C1FA" box="[151,195,1234,1260]" name="O.P. Young" pageId="29" pageNumber="330">P.Y.</collectorName>
Scheffler—
<specimenCount id="03BD9FC83A44FF839437FC91FE9DC1FD" box="[337,364,1233,1259]" pageId="29" pageNumber="330" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A44FF83941FFC91FE3CC1FD" box="[377,461,1233,1259]" pageId="29" pageNumber="330">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A5D35E1C3A44FF8394BBFC92FBA8C019" ID-GBIF-Occurrence="2847261307" collectingDate="1979-01" collectingDateMax="2016-01-26" collectingDateMin="1979-01" collectionCode="CEMT, CMNC" collectorName="O.P. Young" country="Brazil" county="Santarem" latitude="-3.7738888" location="Amazon" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" pageId="29" pageNumber="330" specimenCount="32" specimenCount-female="12" specimenCount-male="20" stateProvince="W" typeStatus="paratype">
<emphasis id="27CF88533A44FF8394BBFC92FCBAC1FA" bold="true" box="[477,843,1233,1260]" pageId="29" pageNumber="330">
<typeStatus id="CA00EAE33A44FF8394BBFC92FDA4C1FA" box="[477,597,1234,1260]" pageId="29" pageNumber="330" type="paratype">Paratypes</typeStatus>
[
<specimenCount id="03BD9FC83A44FF839707FC91FD3BC1FA" box="[609,714,1233,1260]" pageId="29" pageNumber="330" type="male">12 males</specimenCount>
,
<specimenCount id="03BD9FC83A44FF8397B2FC91FCB5C1FA" box="[724,836,1233,1260]" pageId="29" pageNumber="330" type="female">9 females</specimenCount>
]
</emphasis>
.
<emphasis id="27CF88533A44FF839630FC91FC3AC1FA" bold="true" box="[854,971,1233,1260]" pageId="29" pageNumber="330">
<collectingCountry id="6DAC14D13A44FF839630FC91FC37C1FA" box="[854,966,1233,1260]" name="Brazil" pageId="29" pageNumber="330">BRAZIL</collectingCountry>
:
</emphasis>
PARÁ: Redenção, Pinkaiti Aik (
<geoCoordinate id="708F32863A44FF839056FC91FA69C1FA" box="[1328,1432,1233,1260]" degrees="52" direction="west" minutes="02" orientation="longitude" pageId="29" pageNumber="330" precision="925" value="-52.033333">52°02’W</geoCoordinate>
,
<geoCoordinate id="708F32863A44FF8395F1FCB5FF19C006" box="[151,232,1269,1296]" degrees="7" direction="south" minutes="46" orientation="latitude" pageId="29" pageNumber="330" precision="925" value="-7.7666664">7°46’S</geoCoordinate>
),
<date id="610572813A44FF839598FCB5FEA4C019" box="[254,341,1269,1295]" pageId="29" pageNumber="330" value="1999-06">
<collectingDate id="71418B693A44FF839598FCB5FEA4C019" box="[254,341,1269,1295]" pageId="29" pageNumber="330" value="1999-06">vi.1999</collectingDate>
</date>
,
<collectorName id="B84E31973A44FF839405FCB6FE7EC006" box="[355,399,1270,1296]" name="O.P. Young" pageId="29" pageNumber="330">P.Y.</collectorName>
Scheffler—
<specimenCount id="03BD9FC83A44FF839740FCB5FDCDC019" box="[550,572,1269,1295]" pageId="29" pageNumber="330" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A44FF83972BFCB5FD50C019" box="[589,673,1269,1295]" pageId="29" pageNumber="330">CEMT</collectionCode>
); same data but
<date id="610572813A44FF83963BFCB5FC5AC019" box="[861,939,1269,1295]" pageId="29" pageNumber="330" value="1999-10">
<collectingDate id="71418B693A44FF83963BFCB5FC5AC019" box="[861,939,1269,1295]" pageId="29" pageNumber="330" value="1999-10">x.1999</collectingDate>
</date>
—
<specimenCount id="03BD9FC83A44FF8396ADFCB5FC19C019" box="[971,1000,1269,1295]" pageId="29" pageNumber="330" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A44FF83969FFCB5FBBCC019" box="[1017,1101,1269,1295]" pageId="29" pageNumber="330">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A44FF839104FCB5FB4FC025" ID-GBIF-Occurrence="2847261386" collectingDate="1979-01" collectingDateMax="2016-01-26" collectingDateMin="1979-01" collectionCode="CEMT, CMNC" collectorName="O.P. Young" country="Brazil" county="Santarem" latitude="-3.7738888" location="Amazon" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" pageId="29" pageNumber="330" specimenCount="32" specimenCount-female="12" specimenCount-male="20" stateProvince="W" typeStatus="paratype">
<collectingCounty id="FC652CCD3A44FF839104FCB5FB3EC019" box="[1122,1231,1269,1295]" pageId="29" pageNumber="330">Santarém</collectingCounty>
,
<collectingMunicipality id="F560CE3B3A44FF8391BAFCB6FA68C019" box="[1244,1433,1269,1296]" pageId="29" pageNumber="330">Reserva Tapajós</collectingMunicipality>
,
<location id="1064029A3A44FF8395F1FD5AFF07C022" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF8395F1FD5AFF07C022" box="[151,246,1306,1332]" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Amazon" pageId="29" pageNumber="330" stateProvince="W">Amazon</location>
rainforest (
<geoCoordinate id="708F32863A44FF839415FD59FDE3C022" box="[371,530,1305,1332]" pageId="29" pageNumber="330">
55°39’842”
<collectingRegion id="D77F9AA33A44FF839492FD59FDE3C022" box="[500,530,1305,1332]" country="Brazil" name="W" pageId="29" pageNumber="330">W</collectingRegion>
</geoCoordinate>
,
<geoCoordinate id="708F32863A44FF83977DFD59FD53C022" box="[539,674,1305,1332]" pageId="29" pageNumber="330">2°45’463”S</geoCoordinate>
),
<date id="610572813A44FF8397D3FD59FCC9C022" box="[693,824,1305,1332]" pageId="29" pageNumber="330" value="2008-12-25">
<collectingDate id="71418B693A44FF8397D3FD59FCC9C022" box="[693,824,1305,1332]" pageId="29" pageNumber="330" value="2008-12-25">25.xii.2008</collectingDate>
</date>
,
<collectingMethod id="CCFA2C563A44FF839627FD59FC75C025" box="[833,900,1305,1331]" pageId="29" pageNumber="330">pitfall</collectingMethod>
, R.Andrade—
<specimenCount id="03BD9FC83A44FF83915DFD59FBA1C025" box="[1083,1104,1305,1331]" pageId="29" pageNumber="330" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A44FF839138FD59FB43C025" box="[1118,1202,1305,1331]" pageId="29" pageNumber="330">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A44FF8391A5FD59FC0FC041" ID-GBIF-Occurrence="2847261439" collectingDate="1979-01" collectingDateMax="2016-01-26" collectingDateMin="1979-01" collectionCode="CEMT, CMNC" collectorName="O.P. Young" country="Brazil" county="Santarem" latitude="-3.7738888" location="Amazon" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" pageId="29" pageNumber="330" specimenCount="32" specimenCount-female="12" specimenCount-male="20" stateProvince="W" typeStatus="paratype">
<location id="1064029A3A44FF8391A5FD59FAA9C025" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF8391A5FD59FAA9C025" box="[1219,1368,1305,1331]" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Curionopolis" pageId="29" pageNumber="330" stateProvince="W">Curionópolis</location>
,
<location id="1064029A3A44FF839007FD59FF25C04E" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF839007FD59FF25C04E" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Serra Leste" pageId="29" pageNumber="330" stateProvince="W">Serra Leste</location>
(
<geoCoordinate id="708F32863A44FF839582FD7DFE84C041" box="[228,373,1341,1367]" degrees="49" direction="west" minutes="38" orientation="longitude" pageId="29" pageNumber="330" precision="15" seconds="14" value="-49.637222">49°38’14”W</geoCoordinate>
,
<geoCoordinate id="708F32863A44FF8394E7FD7DFE0BC04E" box="[385,506,1341,1368]" degrees="5" direction="south" minutes="58" orientation="latitude" pageId="29" pageNumber="330" precision="15" seconds="27" value="-5.974167">5°58’27”S</geoCoordinate>
),
<date id="610572813A44FF839776FD7DFD8DC041" box="[528,636,1341,1367]" pageId="29" pageNumber="330" value="2016-01-26">
<collectingDate id="71418B693A44FF839776FD7DFD8DC041" box="[528,636,1341,1367]" pageId="29" pageNumber="330" value="2016-01-26">26.i.2016</collectingDate>
</date>
,
<collectingMethod id="CCFA2C563A44FF8397EFFD7DFD3EC041" box="[649,719,1341,1367]" pageId="29" pageNumber="330">pitfall</collectingMethod>
, T. Portella—
<specimenCount id="03BD9FC83A44FF839609FD7DFC7FC041" box="[879,910,1341,1367]" pageId="29" pageNumber="330" type="male">4♂</specimenCount>
(
<collectionCode id="73AACC843A44FF8396F8FD7DFC03C041" box="[926,1010,1341,1367]" pageId="29" pageNumber="330">CEMT</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A44FF839160FD7DFF37C0B6" ID-GBIF-Occurrence="2847261618" collectingDate="1979-01" collectingDateMax="2016-01-26" collectingDateMin="1979-01" collectionCode="CEMT, CMNC" collectorName="O.P. Young" country="Brazil" county="Santarem" latitude="-3.7738888" location="Amazon" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" pageId="29" pageNumber="330" specimenCount="32" specimenCount-female="12" specimenCount-male="20" stateProvince="W" typeStatus="paratype">
<location id="1064029A3A44FF839160FD7DFBAFC041" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF839160FD7DFBAFC041" box="[1030,1118,1341,1367]" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Tucurui" pageId="29" pageNumber="330" stateProvince="W">Tucuruí</location>
(
<geoCoordinate id="708F32863A44FF839109FD7DFB1AC041" box="[1135,1259,1341,1367]" degrees="3" direction="south" minutes="46" orientation="latitude" pageId="29" pageNumber="330" precision="15" seconds="26" value="-3.7738888">3°46’26’’S</geoCoordinate>
,
<geoCoordinate id="708F32863A44FF83919EFD7DFA7EC041" box="[1272,1423,1341,1367]" degrees="49" direction="west" minutes="41" orientation="longitude" pageId="29" pageNumber="330" precision="15" seconds="19" value="-49.688614">49°41’19’’W</geoCoordinate>
),
<date id="610572813A44FF8395F1FD21FF10C06D" box="[151,225,1377,1403]" pageId="29" pageNumber="330" value="1979-01">
<collectingDate id="71418B693A44FF8395F1FD21FF10C06D" box="[151,225,1377,1403]" pageId="29" pageNumber="330" value="1979-01">i.1979</collectingDate>
</date>
, Alvarenga—
<specimenCount id="03BD9FC83A44FF8394E7FD21FE51C06D" box="[385,416,1377,1403]" pageId="29" pageNumber="330" type="male">2♂</specimenCount>
<specimenCount id="03BD9FC83A44FF8394CCFD21FE3CC06D" box="[426,461,1377,1403]" pageId="29" pageNumber="330" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A44FF8394B8FD21FDC6C06D" box="[478,567,1377,1403]" pageId="29" pageNumber="330">CMNC</collectionCode>
).
<collectingRegion id="D77F9AA33A44FF83972DFD21FD1DC06D" box="[587,748,1377,1403]" country="Brazil" name="Tocantins" pageId="29" pageNumber="330">TOCANTINS</collectingRegion>
,
<location id="1064029A3A44FF83979EFD21FC81C06D" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF83979EFD21FC81C06D" box="[760,880,1377,1403]" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Araguaina" pageId="29" pageNumber="330" stateProvince="W">Araguaína</location>
,
<date id="610572813A44FF83961BFD21FC09C06D" box="[893,1016,1377,1403]" pageId="29" pageNumber="330" value="2014-11-22">
<collectingDate id="71418B693A44FF83961BFD21FC09C06D" box="[893,1016,1377,1403]" pageId="29" pageNumber="330" value="2014-11-22">22.xi.2014</collectingDate>
</date>
,
<location id="1064029A3A44FF839163FD22FBB0C06A" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF839163FD22FBB0C06A" box="[1029,1089,1378,1404]" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Mata" pageId="29" pageNumber="330" stateProvince="W">Mata</location>
,
<location id="1064029A3A44FF839129FD22FB64C06A" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF839129FD22FB64C06A" box="[1103,1173,1378,1404]" country="Brazil" county="Santarem" latitude="-3.7738888" longLatPrecision="21" longitude="-49.688614" municipality="Reserva Tapajos" name="Marta" pageId="29" pageNumber="330" stateProvince="W">Marta</location>
, &amp; Sara—
<specimenCount id="03BD9FC83A44FF839070FD21FAC3C06D" box="[1302,1330,1377,1403]" pageId="29" pageNumber="330" type="male">1♂</specimenCount>
(MZUF- PA)
</materialsCitation>
;
<materialsCitation id="A5D35E1C3A44FF8395B6FDC5FE5AC339" ID-GBIF-Occurrence="2847261667" collectingDate="2014-08" collectingDateMax="2017-03-23" collectingDateMin="2014-08" collectionCode="MZUFPA" collectorName="Araguaina &amp; D. Moura" country="Brazil" county="Universidade Federal do Tocantins-Escola de Medicina Veterinaria" location="Universidade Federal Tocantins-Escola de Medicina Veterinaria" municipality="Mata de Galeria" pageId="29" pageNumber="330" specimenCount="10" specimenCount-female="6" specimenCount-male="4" typeStatus="paratype">
<collectorName id="B84E31973A44FF8395B6FDC5FEB9C089" box="[208,328,1413,1439]" pageId="29" pageNumber="330">Araguaína</collectorName>
,
<date id="610572813A44FF839435FDC5FE4BC089" box="[339,442,1413,1439]" pageId="29" pageNumber="330" value="2014-08">
<collectingDate id="71418B693A44FF839435FDC5FE4BC089" box="[339,442,1413,1439]" pageId="29" pageNumber="330" value="2014-08">viii.2014</collectingDate>
</date>
, M.L.S. Rippel—
<specimenCount id="03BD9FC83A44FF8397ECFDC5FD57C089" box="[650,678,1413,1439]" pageId="29" pageNumber="330" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A44FF8397D3FDC5FCDAC089" box="[693,811,1413,1439]" pageId="29" pageNumber="330">MZUFPA</collectionCode>
); Araguaína,
<collectingCounty id="FC652CCD3A44FF8396A6FDC5FE55C0D5" pageId="29" pageNumber="330">Universidade Federal do Tocantins-Escola de Medicina Veterinária</collectingCounty>
e Zoologia,
<collectingMunicipality id="F560CE3B3A44FF839754FDEAFD18C0D5" box="[562,745,1449,1476]" pageId="29" pageNumber="330">Mata de Galeria</collectingMunicipality>
, Corrego da vaca,
<date id="610572813A44FF8396DCFDE9FBC7C0D5" box="[954,1078,1449,1475]" pageId="29" pageNumber="330" value="2017-03-16">
<collectingDate id="71418B693A44FF8396DCFDE9FBC7C0D5" box="[954,1078,1449,1475]" pageId="29" pageNumber="330" value="2017-03-16">16.iii.2017</collectingDate>
</date>
,
<collectingMethod id="CCFA2C563A44FF839127FDE9FB75C0D5" box="[1089,1156,1449,1475]" pageId="29" pageNumber="330">pitfall</collectingMethod>
01,
<collectorName id="B84E31973A44FF8391D0FDEAFAD2C0D2" box="[1206,1315,1450,1476]" pageId="29" pageNumber="330">D. Moura</collectorName>
&amp; S. Dan-tas—
<specimenCount id="03BD9FC83A44FF8395BBFD8DFF0AC0F1" box="[221,251,1485,1511]" pageId="29" pageNumber="330" type="male">1♂</specimenCount>
<specimenCount id="03BD9FC83A44FF83959DFD8DFEE9C0F1" box="[251,280,1485,1511]" pageId="29" pageNumber="330" type="female">2♀</specimenCount>
(
<collectionCode id="73AACC843A44FF83944FFD8DFE6EC0F1" box="[297,415,1485,1511]" pageId="29" pageNumber="330">MZUFPA</collectionCode>
); Araguaína, área aberta,
<date id="610572813A44FF8397A2FD8DFCB1C0F1" box="[708,832,1485,1511]" pageId="29" pageNumber="330" value="2017-03-23">
<collectingDate id="71418B693A44FF8397A2FD8DFCB1C0F1" box="[708,832,1485,1511]" pageId="29" pageNumber="330" value="2017-03-23">23.iii.2017</collectingDate>
</date>
,
<collectingMethod id="CCFA2C563A44FF83962BFD8DFC61C0F1" box="[845,912,1485,1511]" pageId="29" pageNumber="330">pitfall</collectingMethod>
04,
<collectorName id="B84E31973A44FF8396A0FD8EFBC4C0FE" box="[966,1077,1486,1512]" pageId="29" pageNumber="330">D. Moura</collectorName>
&amp; S. Dantas—
<specimenCount id="03BD9FC83A44FF839189FD8DFAF8C0F1" box="[1263,1289,1485,1511]" pageId="29" pageNumber="330" type="female">2♀</specimenCount>
(
<collectionCode id="73AACC843A44FF83907CFD8DFA61C0F1" box="[1306,1424,1485,1511]" pageId="29" pageNumber="330">MZUFPA</collectionCode>
); Araguaína,
<location id="1064029A3A44FF83947DFDB1FC1DC31D" LSID="urn:lsid:plazi:treatment:9D12E5573A44FF8395F1F8D9FB7DC361:1064029A3A44FF83947DFDB1FC1DC31D" box="[283,1004,1521,1547]" country="Brazil" county="Universidade Federal do Tocantins-Escola de Medicina Veterinaria" municipality="Mata de Galeria" name="Universidade Federal Tocantins-Escola de Medicina Veterinaria" pageId="29" pageNumber="330">Universidade Federal Tocantins-Escola de Medicina Veterinária</location>
e Zoologia,
<date id="610572813A44FF83911AFDB1FB16C31D" box="[1148,1255,1521,1547]" pageId="29" pageNumber="330" value="2016-04-09">
<collectingDate id="71418B693A44FF83911AFDB1FB16C31D" box="[1148,1255,1521,1547]" pageId="29" pageNumber="330" value="2016-04-09">9.iv.2016</collectingDate>
</date>
,
<collectingMethod id="CCFA2C563A44FF839195FDB1FAC8C31D" box="[1267,1337,1521,1547]" pageId="29" pageNumber="330">pitfall</collectingMethod>
, S. Dan-tas—
<specimenCount id="03BD9FC83A44FF8395B3FE55FF07C339" box="[213,246,1557,1583]" pageId="29" pageNumber="330" type="male">2♂</specimenCount>
<specimenCount id="03BD9FC83A44FF83959AFE55FEEEC339" box="[252,287,1557,1583]" pageId="29" pageNumber="330" type="female">2♀</specimenCount>
(
<collectionCode id="73AACC843A44FF839448FE55FE55C339" box="[302,420,1557,1583]" pageId="29" pageNumber="330">MZUFPA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A44FF8395A1FE79FB7DC361" pageId="29" pageNumber="330" type="distribution">
<paragraph id="150454413A44FF8395A1FE79FB7DC361" blockId="29.[151,1437,261,1656]" pageId="29" pageNumber="330">
<emphasis id="27CF88533A44FF8395A1FE79FE90C342" bold="true" box="[199,353,1593,1620]" pageId="29" pageNumber="330">Distribution.</emphasis>
Known from
<collectingCountry id="6DAC14D13A44FF839761FE79FDBAC345" box="[519,587,1593,1619]" name="Brazil" pageId="29" pageNumber="330">Brazil</collectingCountry>
(Pará and
<collectingRegion id="D77F9AA33A44FF8397AAFE79FCCBC342" box="[716,826,1593,1620]" country="Brazil" name="Tocantins" pageId="29" pageNumber="330">Tocantins</collectingRegion>
) (
<figureCitation id="8D8048C43A44FF839632FE79FC40C342" box="[852,945,1593,1620]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="29" pageNumber="330">Fig. 6D</figureCitation>
).
<emphasis id="27CF88533A44FF8396A1FE79FB61C342" bold="true" box="[967,1168,1593,1620]" pageId="29" pageNumber="330">Endemism areas</emphasis>
:
<emphasis id="27CF88533A44FF8391C4FE79FA65C342" bold="true" box="[1186,1428,1593,1620]" pageId="29" pageNumber="330">Chacoan sub-region</emphasis>
: South-eastern Amazonian dominion: Xingu-Tapajós province (see
<bibRefCitation id="712A29B03A44FF8396E5FE1EFBD9C361" author="Morrone, J. J." box="[899,1064,1629,1656]" pageId="29" pageNumber="330" pagination="1 - 110" refId="ref47051" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
; fig. 12).
</paragraph>
</subSubSection>
</treatment>
</document>
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