treatments-xml/data/33/35/87/333587E1D14BFFC6FF64E245C45C7507.xml

<document id="61362329AF9E5DE3B858EC6B93E43298" ID="10.11646/zootaxa.4277.3.5SLASH11401" ID-CLB-Dataset="32822" ID-DOI="10.11646/zootaxa.4277.3.5" ID-GBIF-Dataset="6a00d899-3a3b-499f-a2d1-92fb0d526eea" ID-ISSN="1175-5326" ID-Zenodo-Dep="810721" ID-ZooBank="D8DB26BA-0646-48B4-8F1B-A63F6B77E2F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1497858852383" checkinUser="plazi" docAuthor="Castro-Huertas, Valentina &amp; Forero, Dimitri" docDate="2017" docId="333587E1D14BFFC6FF64E245C45C7507" docLanguage="en" docName="zootaxa.4277.3.5SLASH11401.pdf" docOrigin="Zootaxa 4277 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liaghinella tuberculata Castro-Huertas &amp; Forero 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="411" masterDocId="CF0CFF99D14EFFCAFFF3E659C100715C" masterDocTitle="Small range distributions in the high Andes: two new species of Liaghinella (Hemiptera: Heteroptera: Reduviidae: Emesinae) from Colombia" masterLastPageNumber="412" masterPageNumber="399" pageNumber="404" updateTime="1698447222784" updateUser="ExternalLinkService">
<mods:mods id="EA66F8B695644A5CE3DA971A542DDDC1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="ED57C60A7AF45D9A23E210C04F51D0B3">
<mods:title id="3E89B9FD6CA88BEE201384865A7FA3A9">Small range distributions in the high Andes: two new species of Liaghinella (Hemiptera: Heteroptera: Reduviidae: Emesinae) from Colombia</mods:title>
</mods:titleInfo>
<mods:name id="95B0468D1B311282D114DB1C28BE4883" type="personal">
<mods:role id="D1394A0F411BEFF8B935B03BAC65CF80">
<mods:roleTerm id="7C948C717B7BFCF6C4B84F486F6317B5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3F06096B7FC6137CE07FB26B2A8CADAA">Castro-Huertas, Valentina</mods:namePart>
</mods:name>
<mods:name id="1CE80F9B1710F51F9E4DC2E9064C2FED" type="personal">
<mods:role id="018B73E24D30C98C4DCC848319560E80">
<mods:roleTerm id="DC36F77FD09A1FD5CA076922EC3A5F6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B6E886BD95DB5681DB16C5280309CFE1">Forero, Dimitri</mods:namePart>
</mods:name>
<mods:typeOfResource id="91D0253B70162F5E7414401EB05C6EC1">text</mods:typeOfResource>
<mods:relatedItem id="7ABF341275E21D8045BA289CBF652A55" type="host">
<mods:titleInfo id="4BE83D6AB8698D011620C59CD4F660A7">
<mods:title id="B9BA984C08BDFD560E861AA3C190CEC6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DBB806A645984867B82A176660E780D6">
<mods:date id="2888E6EDF22CD9A5B187D6D50D7721C8">2017</mods:date>
<mods:detail id="F1A7B1C1A49DE9D83D8C64BD21142F05" type="volume">
<mods:number id="C770437E31420D68AE987C627587217F">4277</mods:number>
</mods:detail>
<mods:detail id="D277E0B055E984455755D5BD5AB592EB" type="issue">
<mods:number id="1B58FBB70E93C3C91BBDE25001CFC30D">3</mods:number>
</mods:detail>
<mods:extent id="EE74812EAB84A429F4E04DAEEF8EA664" unit="page">
<mods:start id="0558D7D63689744FBA3C312BA185D49E">399</mods:start>
<mods:end id="E3A6D010B3D5F83F4BB1AD5434FD429E">412</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="E1D1763350DEFF57C779872787EFB101">journal article</mods:classification>
<mods:identifier id="B4081A49A20CD109CF947A25FEFF1D62" type="CLB-Dataset">32822</mods:identifier>
<mods:identifier id="D808E6FBA63527EC029D2BB7D5B051D9" type="DOI">10.11646/zootaxa.4277.3.5</mods:identifier>
<mods:identifier id="38C910AD652CD430B4020F80A008E36A" type="GBIF-Dataset">6a00d899-3a3b-499f-a2d1-92fb0d526eea</mods:identifier>
<mods:identifier id="833EFD2CA4DD67EB9FFB17996F7A8D61" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1B85C35B5EC380B2CFCDBECFDEC8DCD3" type="Zenodo-Dep">810721</mods:identifier>
<mods:identifier id="5E5609BE55B35B7F2337D65723228CC3" type="ZooBank">D8DB26BA-0646-48B4-8F1B-A63F6B77E2F6</mods:identifier>
</mods:mods>
<treatment id="333587E1D14BFFC6FF64E245C45C7507" ID-DOI="http://doi.org/10.5281/zenodo.6020778" ID-GBIF-Taxon="131891916" ID-Zenodo-Dep="6020778" LSID="urn:lsid:plazi:treatment:333587E1D14BFFC6FF64E245C45C7507" httpUri="http://treatment.plazi.org/id/333587E1D14BFFC6FF64E245C45C7507" lastPageId="12" lastPageNumber="411" pageId="5" pageNumber="404">
<subSubSection id="F386657CD14BFFCFFF64E245C02E750B" pageId="5" pageNumber="404" type="nomenclature">
<paragraph id="BB2336F7D14BFFCFFF64E245C25F756A" blockId="5.[151,863,1052,1111]" box="[151,863,1052,1078]" pageId="5" pageNumber="404">
<heading id="E06B819BD14BFFCFFF64E245C25F756A" bold="true" box="[151,863,1052,1078]" fontSize="11" level="1" pageId="5" pageNumber="404" reason="1">
<emphasis id="89E8EAE5D14BFFCFFF64E245C25F756A" bold="true" box="[151,863,1052,1078]" pageId="5" pageNumber="404">
<taxonomicName id="7C9C4D74D14BFFCFFF64E245C0B4756A" ID-CoL="6Q782" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[151,436,1052,1078]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="404" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D14BFFCFFF64E245C0B4756A" bold="true" box="[151,436,1052,1078]" italics="true" pageId="5" pageNumber="404">Liaghinella tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED14BFFCFFE4FE245C319756A" box="[444,537,1052,1078]" pageId="5" pageNumber="404" rank="species">sp. nov.</taxonomicNameLabel>
Forero &amp; Castro-Huertas
</emphasis>
</heading>
</paragraph>
<paragraph id="BB2336F7D14BFFCFFF64E267C02E750B" blockId="5.[151,863,1052,1111]" box="[151,302,1086,1111]" pageId="5" pageNumber="404">
(
<figureCitation id="23A72A72D14BFFCFFF6CE267C00B750B" box="[159,267,1086,1111]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStartId-0="7.[151,250,1923,1945]" captionStartId-1="8.[151,250,1923,1945]" captionStartId-2="9.[151,250,1933,1955]" captionStartId-3="10.[151,250,1893,1915]" captionTargetBox-0="[226,1360,193,1902]" captionTargetBox-1="[321,1277,193,1909]" captionTargetBox-2="[258,1338,193,1921]" captionTargetBox-3="[226,1360,193,1871]" captionTargetId-0="figure@7.[226,1360,193,1902]" captionTargetId-1="figure@8.[321,1277,193,1909]" captionTargetId-2="figure@9.[258,1338,193,1921]" captionTargetId-3="figure@10.[226,1360,193,1871]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." captionText-1="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." captionText-2="FIGURE 5. Liaghinella tuberculata sp. nov., female paratype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and prothorax, lateral view. E. Prothorax, lateral view. F. Head, lateral view, arrow indicates clypeus." captionText-3="FIGURE 6. Liaghinella tuberculata sp. nov., female genitalia, left gcx 8 removed. A. Ventral view. B. Lateral view. C. Tergite 8 and 9, caudal view. D. Gonocoxa eight, ventral view. Abbreviations: bc, bursa copulatrix; gap 8, gonapophysis eight; gap 9, gonapophysis nine; gcx 8, gonocoxa eight; gcx 9, gonocoxa nine; gpl, gonoplac; gr, ring gland; lap, latero anterior prolongation of the gonocoxa eight; mpm 8, medial posterior margin of gonocoxa eight; mpm 9, medial posterior margin of gonocoxa nine; T 8, tergite 8; T 9, tergite 9." httpUri-0="https://zenodo.org/record/810727/files/figure.png" httpUri-1="https://zenodo.org/record/810729/files/figure.png" httpUri-2="https://zenodo.org/record/810731/files/figure.png" httpUri-3="https://zenodo.org/record/810733/files/figure.png" pageId="5" pageNumber="404">Figs. 3‒6</figureCitation>
, 
<figureCitation id="23A72A72D14BFFCFFEE5E266C025750B" box="[278,293,1087,1111]" captionStart="FIGURE 7" captionStartId="11.[151,250,2002,2024]" captionTargetBox="[262,1343,717,1968]" captionTargetId="figure@11.[193,1398,661,1988]" captionTargetPageId="11" captionText="FIGURE 7. Distribution map of Liaghinella species found in Colombia based on the examination of adult material." httpUri="https://zenodo.org/record/810735/files/figure.png" pageId="5" pageNumber="404">7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F386657CD14BFFCFFF64E2DFC299742B" pageId="5" pageNumber="404" type="materials_examined">
<paragraph id="BB2336F7D14BFFCFFF64E2DFC299742B" blockId="5.[151,1437,1158,2011]" pageId="5" pageNumber="404">
<materialsCitation id="0BF43CAAD14BFFCFFF64E2DFC1F57457" ID-GBIF-Occurrence="1569232901" collectingDate="2013-04-08" collectingDateMax="2013-04-12" collectingDateMin="2013-04-08" collectionCode="MPUJ, ENT" collectorName="D. Forero &amp; Forero &amp; Castro-Huertas" country="Colombia" elevation="2221" latitude="4.6148667" location="Reserva Chicaque" longLatPrecision="1" longitude="-74.31129" municipality="Male" pageId="5" pageNumber="404" specimenCount="1" stateProvince="Cundinamarca" typeStatus="holotype">
<emphasis id="89E8EAE5D14BFFCFFF64E2DFC00D75C3" bold="true" box="[151,269,1158,1183]" pageId="5" pageNumber="404">
<typeStatus id="64278855D14BFFCFFF64E2DFC00875C3" box="[151,264,1158,1183]" pageId="5" pageNumber="404" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectingMunicipality id="5B47AC8DD14BFFCFFEEBE2DFC05175C3" box="[280,337,1158,1183]" pageId="5" pageNumber="404">
<emphasis id="89E8EAE5D14BFFCFFEEBE2DFC05175C3" box="[280,337,1158,1183]" italics="true" pageId="5" pageNumber="404">Male</emphasis>
</collectingMunicipality>
: 
<collectingCountry id="C38B7667D14BFFCFFE97E2DEC0FE75C3" box="[356,510,1159,1183]" name="Colombia" pageId="5" pageNumber="404">COLOMBIA</collectingCountry>
: 
<collectingRegion id="7958F815D14BFFCFFDFFE2DFC3B375C3" box="[524,691,1158,1183]" country="Colombia" name="Cundinamarca" pageId="5" pageNumber="404">Cundinamarca</collectingRegion>
, 
<location id="BE43602CD14BFFCFFD32E2DEC29375C3" LSID="urn:lsid:plazi:treatment:333587E1D14BFFC6FF64E245C45C7507:BE43602CD14BFFCFFD32E2DEC29375C3" box="[705,915,1158,1183]" country="Colombia" latitude="4.6148667" longLatPrecision="1" longitude="-74.31129" municipality="Male" name="Reserva Chicaque" pageId="5" pageNumber="404" stateProvince="Cundinamarca">Reserva Chicaque</location>
, refugio, 
<geoCoordinate id="DEA85030D14BFFCFFBF6E2DFC58675C3" box="[1029,1158,1158,1183]" direction="north" orientation="latitude" pageId="5" pageNumber="404" precision="1" value="4.6148667">4°36,892´N</geoCoordinate>
<geoCoordinate id="DEA85030D14BFFCFFB62E2DFC42675C3" box="[1169,1318,1158,1183]" direction="west" orientation="longitude" pageId="5" pageNumber="404" precision="1" value="-74.31129">74°18,677´W</geoCoordinate>
, 
<elevation id="30B1D1C4D14BFFCFFAC6E2DEC49775C2" box="[1333,1431,1159,1183]" metricMagnitude="3" metricUnit="m" metricValue="2.221" pageId="5" pageNumber="404" unit="m" value="2221.0">
<quantity id="7C649B12D14BFFCFFAC6E2DEC49775C2" box="[1333,1431,1159,1183]" metricMagnitude="3" metricUnit="m" metricValue="2.221" pageId="5" pageNumber="404" unit="m" value="2221.0">2,221 m</quantity>
</elevation>
, 
<collectingDate id="DF66E9DFD14BFFCFFF64E2F2C040759F" box="[151,320,1194,1219]" pageId="5" pageNumber="404" value="2013-04-08" valueMax="2013-04-12" valueMin="2013-04-08">
<date id="CF221037D14BFFCFFF64E2F2C040759F" box="[151,320,1194,1219]" pageId="5" pageNumber="404" value="2013-04-08" valueMax="2013-04-12" valueMin="2013-04-08">8‒12 abr 2013</date>
</collectingDate>
, 
<collectorName id="16695321D14BFFCFFEBEE2F2C0BC759E" box="[333,444,1195,1218]" pageId="5" pageNumber="404">D. Forero</collectorName>
leg. / cerca al refugio bajo 
<taxonomicName id="7C9C4D74D14BFFCFFD0EE2F5C23B759F" box="[765,827,1196,1219]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="404" phylum="Mollusca" rank="genus">
<emphasis id="89E8EAE5D14BFFCFFD0EE2F5C23B759F" box="[765,827,1196,1219]" italics="true" pageId="5" pageNumber="404">Ficus</emphasis>
</taxonomicName>
, en hojarasca, noche [near to refuge, under 
<taxonomicName id="7C9C4D74D14BFFCFFACBE2F5C476759F" box="[1336,1398,1196,1219]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="5" pageNumber="404" phylum="Mollusca" rank="genus">
<emphasis id="89E8EAE5D14BFFCFFACBE2F5C476759F" box="[1336,1398,1196,1219]" italics="true" pageId="5" pageNumber="404">Ficus</emphasis>
</taxonomicName>
, in forest litter, at night] / 
<taxonomicName id="7C9C4D74D14BFFCFFE56E297C3B175BB" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[421,689,1230,1255]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="404" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D14BFFCFFE56E297C3B175BB" box="[421,689,1230,1255]" italics="true" pageId="5" pageNumber="404">Liaghinella tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED14BFFCFFD4FE297C21975BB" box="[700,793,1230,1255]" pageId="5" pageNumber="404" rank="species">
<emphasis id="89E8EAE5D14BFFCFFD4FE297C21975BB" bold="true" box="[700,793,1230,1255]" pageId="5" pageNumber="404">sp. nov.</emphasis>
</taxonomicNameLabel>
<collectorName id="16695321D14BFFCFFCD7E296C26F75BA" box="[804,879,1231,1254]" pageId="5" pageNumber="404">Forero</collectorName>
&amp; 
<collectorName id="16695321D14BFFCFFC69E296C54475BB" box="[922,1092,1231,1255]" pageId="5" pageNumber="404">Castro-Huertas</collectorName>
2017 / 
<collectionCode id="DD8DAE32D14BFFCFFB50E296C5EB75BA" box="[1187,1259,1231,1254]" pageId="5" pageNumber="404">MPUJ</collectionCode>
_ 
<collectionCode id="DD8DAE32D14BFFCFFB0AE296C42F75BA" box="[1273,1327,1231,1254]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="5" pageNumber="404">ENT</collectionCode>
0 0 10584 (
<collectionCode id="DD8DAE32D14BFFCFFF6CE2AAC1ED7457" box="[159,237,1267,1291]" pageId="5" pageNumber="404">MPUJ</collectionCode>
)
</materialsCitation>
. 
<emphasis id="89E8EAE5D14BFFCFFEF3E2ABC0817457" bold="true" box="[256,385,1266,1291]" pageId="5" pageNumber="404">Paratypes:</emphasis>
3 
<emphasis id="89E8EAE5D14BFFCFFE6CE2ABC3067457" box="[415,518,1266,1291]" italics="true" pageId="5" pageNumber="404">Females.</emphasis>
Same locality data / MPUJ_ENT 0 0 10771 / MPUJ_ENT 0 0 29637 / MPUJ_ENT 0 0 46844 (MPUJ); 2 
<emphasis id="89E8EAE5D14BFFCFFE77E34FC0C87473" box="[388,456,1302,1327]" italics="true" pageId="5" pageNumber="404">Males</emphasis>
. Same locality data / MPUJ_ENT 0 0 29636 / MPUJ_ENT 0 0 29638 (MPUJ); 1 
<emphasis id="89E8EAE5D14BFFCFFAA8E34FC4957473" box="[1371,1429,1302,1327]" italics="true" pageId="5" pageNumber="404">Male</emphasis>
. Same locality data / MPUJ_ENT 0 0 29639 (AMNH); 1 
<emphasis id="89E8EAE5D14BFFCFFCF7E363C23E740F" box="[772,830,1338,1363]" italics="true" pageId="5" pageNumber="404">Male</emphasis>
. Same locality data / MPUJ_ENT 0 0 29635 (NMNH). 
<emphasis id="89E8EAE5D14BFFCFFF64E307C0D9742B" bold="true" box="[151,473,1374,1399]" pageId="5" pageNumber="404">Other specimens examined</emphasis>
: 
<emphasis id="89E8EAE5D14BFFCFFE14E306C3BB742B" box="[487,699,1374,1399]" italics="true" pageId="5" pageNumber="404">Last instar nymph:</emphasis>
Same locality data.
</paragraph>
</subSubSection>
<subSubSection id="F386657CD14BFFCFFF34E3DBC33B775B" pageId="5" pageNumber="404" type="reference_group">
<paragraph id="BB2336F7D14BFFCFFF34E3DBC33B775B" blockId="5.[151,1437,1158,2011]" pageId="5" pageNumber="404">
<materialsCitation id="0BF43CAAD14BFFCFFF34E3DBC33B775B" ID-GBIF-Occurrence="1569232904" collectingDate="2015-01-23" collectionCode="MPUJ, ENT" collectorName="E. Tulande" country="Colombia" elevation="2250" latitude="4.61725" location="Reserva Chicaque" longLatPrecision="1" longitude="-74.31395" municipality="Male" pageId="5" pageNumber="404" specimenCount="1" stateProvince="Cundinamarca">
<taxonomicName id="7C9C4D74D14BFFCFFF34E3DBC36474C7" ID-CoL="6PV82" authority="Forero, 2007" authorityName="Forero" authorityYear="2007" box="[199,612,1410,1435]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="404" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D14BFFCFFF34E3DBC0AB74C7" box="[199,427,1410,1435]" italics="true" pageId="5" pageNumber="404">Liaghinella andina</emphasis>
<bibRefCitation id="DF0D4B06D14BFFCFFE33E3DAC36474C7" author="Forero" box="[448,612,1411,1435]" pageId="5" pageNumber="404" refString="Forero, D. (2007) Description of a new species of Liaghinella (Hemiptera: Heteroptera: Reduviidae: Emesinae) from the Colombian Andes, with notes on its feeding habits and conservation status. Zootaxa, 1502, 55 - 68." type="journal article" year="2007">Forero, 2007</bibRefCitation>
</taxonomicName>
: 
<collectingMunicipality id="5B47AC8DD14BFFCFFD8FE3DBC3B574C7" box="[636,693,1410,1435]" pageId="5" pageNumber="404">
<emphasis id="89E8EAE5D14BFFCFFD8FE3DBC3B574C7" box="[636,693,1410,1435]" italics="true" pageId="5" pageNumber="404">Male</emphasis>
</collectingMunicipality>
: 
<collectingCountry id="C38B7667D14BFFCFFD21E3DAC26C74C7" box="[722,876,1411,1435]" name="Colombia" pageId="5" pageNumber="404">COLOMBIA</collectingCountry>
: 
<collectingRegion id="7958F815D14BFFCFFC76E3DBC52B74C7" box="[901,1067,1410,1435]" country="Colombia" name="Cundinamarca" pageId="5" pageNumber="404">Cundinamarca</collectingRegion>
, 
<location id="BE43602CD14BFFCFFBB7E3DAC42074C7" LSID="urn:lsid:plazi:treatment:333587E1D14BFFC6FF64E245C45C7507:BE43602CD14BFFCFFBB7E3DAC42074C7" box="[1092,1312,1410,1435]" country="Colombia" latitude="4.61725" longLatPrecision="1" longitude="-74.31395" municipality="Male" name="Reserva Chicaque" pageId="5" pageNumber="404" stateProvince="Cundinamarca">Reserva Chicaque</location>
, robledal, 0 
<geoCoordinate id="DEA85030D14BFFCFFF57E3FFC03F74E3" box="[164,319,1446,1471]" direction="north" orientation="latitude" pageId="5" pageNumber="404" precision="1" value="4.61725">4.6172500 °N</geoCoordinate>
<geoCoordinate id="DEA85030D14BFFCFFEB5E3FFC0F574E3" box="[326,501,1446,1471]" direction="west" orientation="longitude" pageId="5" pageNumber="404" precision="1" value="-74.31395">74.3139500 °W</geoCoordinate>
, 
<elevation id="30B1D1C4D14BFFCFFE0CE3FFC35874E3" box="[511,600,1446,1471]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="404" unit="m" value="2250.0">
<quantity id="7C649B12D14BFFCFFE0CE3FFC35874E3" box="[511,600,1446,1471]" metricMagnitude="3" metricUnit="m" metricValue="2.25" pageId="5" pageNumber="404" unit="m" value="2250.0">2,250m</quantity>
</elevation>
, 
<collectingDate id="DF66E9DFD14BFFCFFD91E3FEC3ED74E3" box="[610,749,1446,1471]" pageId="5" pageNumber="404" value="2015-01-23">
<date id="CF221037D14BFFCFFD91E3FEC3ED74E3" box="[610,749,1446,1471]" pageId="5" pageNumber="404" value="2015-01-23">23 ene 2015</date>
</collectingDate>
, 
<collectorName id="16695321D14BFFCFFD04E3FEC27174E3" box="[759,881,1446,1471]" pageId="5" pageNumber="404">E. Tulande</collectorName>
/ colecta manual, bajo hojarasca de 
<taxonomicName id="7C9C4D74D14BFFCFFAF0E3FEC46174E2" ID-CoL="8W2YL" authorityName="C. Linnaeus" authorityYear="1753" box="[1283,1377,1447,1470]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="404" phylum="Tracheophyta" rank="genus">
<emphasis id="89E8EAE5D14BFFCFFAF0E3FEC46174E2" box="[1283,1377,1447,1470]" italics="true" pageId="5" pageNumber="404">Quercus</emphasis>
</taxonomicName>
sp. y 
<taxonomicName id="7C9C4D74D14BFFCFFF64E393C1E174BF" ID-CoL="3R8R" box="[151,225,1482,1507]" class="Insecta" family="Clusiidae" genus="Clusia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D14BFFCFFF64E393C1E174BF" box="[151,225,1482,1507]" italics="true" pageId="5" pageNumber="404">Clusia</emphasis>
</taxonomicName>
sp., colecta nocturna [manual collecting, under forest litter of 
<taxonomicName id="7C9C4D74D14BFFCFFC5DE392C50C74BE" ID-CoL="8W2YL" authorityName="C. Linnaeus" authorityYear="1753" box="[942,1036,1483,1506]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="404" phylum="Tracheophyta" rank="genus">
<emphasis id="89E8EAE5D14BFFCFFC5DE392C50C74BE" box="[942,1036,1483,1506]" italics="true" pageId="5" pageNumber="404">Quercus</emphasis>
</taxonomicName>
spp. and 
<taxonomicName id="7C9C4D74D14BFFCFFB8FE393C5C574BF" ID-CoL="3R8R" box="[1148,1221,1482,1507]" class="Insecta" family="Clusiidae" genus="Clusia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="404" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D14BFFCFFB8FE393C5C574BF" box="[1148,1221,1482,1507]" italics="true" pageId="5" pageNumber="404">Clusia</emphasis>
</taxonomicName>
sp., night collect], 23h. / 
<collectionCode id="DD8DAE32D14BFFCFFF2EE3B6C025775A" box="[221,293,1519,1542]" pageId="5" pageNumber="404">MPUJ</collectionCode>
_ 
<collectionCode id="DD8DAE32D14BFFCFFEC0E3B6C069775A" box="[307,361,1519,1542]" country="Uganda" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="5" pageNumber="404">ENT</collectionCode>
0 0 47555 (
<collectionCode id="DD8DAE32D14BFFCFFE11E3B6C330775B" box="[482,560,1519,1543]" pageId="5" pageNumber="404">MPUJ</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="F386657CD14BFFCFFF34E04BC2D077CB" pageId="5" pageNumber="404" type="diagnosis">
<paragraph id="BB2336F7D14BFFCFFF34E04BC2D077CB" blockId="5.[151,1437,1158,2011]" pageId="5" pageNumber="404">
<emphasis id="89E8EAE5D14BFFCFFF34E04BC0417777" bold="true" box="[199,321,1554,1579]" pageId="5" pageNumber="404">Diagnosis.</emphasis>
Recognized by a pair of medial tubercles on the posterior margin of the pronotum (
<figureCitation id="23A72A72D14BFFCFFAE1E04BC4617777" box="[1298,1377,1554,1579]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="5" pageNumber="404">Figs. 3</figureCitation>
C‒E; 4C‒E), meso- and metanotum with three longitudinal carinae and posterior margin of the meso- and metanotum with paired short tubercles on lateral carinae (
<figureCitation id="23A72A72D14BFFCFFD6BE003C3E4772F" box="[664,740,1626,1651]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="5" pageNumber="404">Figs. 3</figureCitation>
A, 5B), clypeus with a short spine (
<figureCitation id="23A72A72D14BFFCFFB82E003C5B3772F" box="[1137,1203,1626,1651]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="5" pageNumber="404">Fig. 3</figureCitation>
F, 5F), posterolateral angles of dorsal laterotergites produced as a short tubercle (
<figureCitation id="23A72A72D14BFFCFFCC2E027C27E77CB" box="[817,894,1662,1687]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="5" pageNumber="404">Figs. 3</figureCitation>
B, 5B).
</paragraph>
</subSubSection>
<subSubSection id="F386657CD14BFFCCFF34E0FBC0DD770C" lastPageId="6" lastPageNumber="405" pageId="5" pageNumber="404" type="description">
<paragraph id="BB2336F7D14BFFCFFF34E0FBC2777783" blockId="5.[151,1437,1158,2011]" pageId="5" pageNumber="404">
<emphasis id="89E8EAE5D14BFFCFFF34E0FBC30E77E7" bold="true" box="[199,526,1698,1723]" pageId="5" pageNumber="404">Description. Apterous male</emphasis>
(total length, 
<typeStatus id="64278855D14BFFCFFD59E0FBC20C77E7" box="[682,780,1698,1723]" pageId="5" pageNumber="404" type="holotype">holotype</typeStatus>
<quantity id="7C649B12D14BFFCFFCE1E0FBC27877E6" box="[786,888,1698,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="5" pageNumber="404" unit="mm" value="11.6">11.6 mm</quantity>
; head length, 
<quantity id="7C649B12D14BFFCFFBE0E0FBC56C77E6" box="[1043,1132,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="404" unit="mm" value="1.5">1.5 mm</quantity>
; prothorax length, 
<quantity id="7C649B12D14BFFCFFACCE0FAC49877E6" box="[1343,1432,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="404" unit="mm" value="1.9">1.9 mm</quantity>
; meso- and metathorax length, 
<quantity id="7C649B12D14BFFCFFE18E09EC3447782" box="[491,580,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="5" pageNumber="404" unit="mm" value="2.2">2.2 mm</quantity>
; abdomen length, 
<quantity id="7C649B12D14BFFCFFCE2E09FC26B7783" box="[785,875,1734,1759]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="5" pageNumber="404" unit="mm" value="5.9">5.9 mm</quantity>
).
</paragraph>
<paragraph id="BB2336F7D14BFFCCFF34E0B3C3097040" blockId="5.[151,1437,1158,2011]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="405" pageId="5" pageNumber="404">
<emphasis id="89E8EAE5D14BFFCFFF34E0B3C010765F" bold="true" box="[199,272,1770,1795]" pageId="5" pageNumber="404">Color.</emphasis>
<emphasis id="89E8EAE5D14BFFCFFEE8E0B3C061765F" box="[283,353,1770,1795]" italics="true" pageId="5" pageNumber="404">Head:</emphasis>
Ventrally dark brown, dorsally with several pale brown spots (
<figureCitation id="23A72A72D14BFFCFFBB8E0B3C592765F" box="[1099,1170,1770,1795]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="5" pageNumber="404">Fig. 3</figureCitation>
F). Clypeus spine dark brown, first and second segments of the labium basally dark brown and distally whitish, third segment of the labium dark brown. Scapus pale brown and distally dark brown, flagellomeres dark brown. 
<emphasis id="89E8EAE5D14BFFCFFB6FE16BC5ED7617" box="[1180,1261,1842,1867]" italics="true" pageId="5" pageNumber="404">Thorax</emphasis>
: Ventrally dark brown, dorsally pale brown, laterally with several whitish spots (
<figureCitation id="23A72A72D14BFFCFFC84E10FC2C57633" box="[887,965,1878,1903]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="5" pageNumber="404">Figs. 3</figureCitation>
C‒D); procoxal cavity pale brown, meso- and metacoxal cavities dark brown with a whitish fine line on the posterior margin; proleg dark brown with several whitish portions, procoxae ventrally dark brown and dorsally pale brown, protrochanter dark brown, profemur ventrally dark brown and dorsally pale brown with some spots whitish laterally; protibia dark brown, medial and basally whitish; protarsus dark brown basally yellow; meso- and meta coxae and trochanter dark brown, meso- and metafemur dark brown with a several dispersal spots whitish, meso- and meta tibiae pale brown with several dark brown dispersal spots, meso-and metatarsi dark brown. 
<emphasis id="89E8EAE5D148FFCCFCB2E686C2B371A4" box="[833,947,223,248]" italics="true" pageId="6" pageNumber="405">Abdomen:</emphasis>
Pale brown with several whitish spots; pygophore pale brown (
<figureCitation id="23A72A72D148FFCCFE51E75DC0E47040" box="[418,484,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="6" pageNumber="405">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="BB2336F7D148FFCCFF34E77EC5C77520" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E77EC03C701C" bold="true" box="[199,316,295,320]" pageId="6" pageNumber="405">Vestiture:</emphasis>
Body covered with numerous setiferous tubercles. 
<emphasis id="89E8EAE5D148FFCCFC42E77EC2ED701C" box="[945,1005,295,320]" italics="true" pageId="6" pageNumber="405">Head</emphasis>
: Dorsally with setiferous tubercles, ventrally with dense setae; antennal scapus with numerous, short setae; labium glabrous. 
<emphasis id="89E8EAE5D148FFCCFB22E715C44A7039" bold="true" box="[1233,1354,332,357]" pageId="6" pageNumber="405">Structure.</emphasis>
<emphasis id="89E8EAE5D148FFCCFAA4E715C4947039" box="[1367,1428,332,357]" italics="true" pageId="6" pageNumber="405">Head</emphasis>
: Elongated, interocular sulcus deep, clypeus with a short spine; eyes globose in dorsal view, elongate ovoid in lateral view (
<figureCitation id="23A72A72D148FFCCFEDCE7CDC07470F0" box="[303,372,404,429]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="6" pageNumber="405">Fig. 3</figureCitation>
F); antenna slender, scapus longer than others segments; first flagellomere shortest segment; first visible labial segment short, second visible labial segment swollen, about as long as second segment, third visible segment slender, about twice the length of the fourth segment. 
<emphasis id="89E8EAE5D148FFCCFC50E785C2F470A9" box="[931,1012,476,501]" italics="true" pageId="6" pageNumber="405">Thorax</emphasis>
: Pronotum longer than wide, about as long as meso- and metanotum together; lateral angles of the anterior margin with rounded tubercles (
<figureCitation id="23A72A72D148FFCCFAFFE7A6C4597344" box="[1292,1369,511,536]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="6" pageNumber="405">Figs. 3</figureCitation>
C‒E), posterior margin with a pair of conspicuous medial tubercles directed dorsad; meso- and metanotum with three longitudinal carinae, posterior margin of meso- and metanotum with paired short tubercles each on lateral carinae; stridulitrum narrow. 
<emphasis id="89E8EAE5D148FFCCFE8EE434C0B273D8" box="[381,434,621,644]" italics="true" pageId="6" pageNumber="405">Legs</emphasis>
: Proleg distinctly wider and shorter than meso- and metaleg; procoxa elongate, profemur about as long as three times the protibiae and wider; profemur with posteroventral series composed by spiniform process and setae, and located near to the base of article (
<figureCitation id="23A72A72D148FFCCFCEFE4EDC25D7390" box="[796,861,692,717]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="6" pageNumber="405">Fig. 3</figureCitation>
D), profemur with anteroventral series compose by a row of setae; protibia ventrally with a row of triangular denticles; protibial comb present; protarsus one segmented about as long as protibia and with two claws; meso- and metalegs larger and slender. 
<emphasis id="89E8EAE5D148FFCCFAD9E4A5C4947249" box="[1322,1428,764,789]" italics="true" pageId="6" pageNumber="405">Abdomen</emphasis>
: elongate, abdominal tergites I‒VI with a short median tubercle on the posterior margin; dorsal laterotergites with projections on lateroposterior margin produced dorsad (
<figureCitation id="23A72A72D148FFCCFCF4E51DC2497200" box="[775,841,836,861]" captionStart="FIGURE 3" captionStartId="7.[151,250,1923,1945]" captionTargetBox="[226,1360,193,1902]" captionTargetId="figure@7.[226,1360,193,1902]" captionTargetPageId="7" captionText="FIGURE 3. Liaghinella tuberculata sp. nov., male holotype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="6" pageNumber="405">Fig. 3</figureCitation>
B), distal margin of tergite VII acute. 
<emphasis id="89E8EAE5D148FFCCFB03E51DC4957201" box="[1264,1429,836,861]" italics="true" pageId="6" pageNumber="405">Male genitalia</emphasis>
: Pygophore elongated and ventrally produced (
<figureCitation id="23A72A72D148FFCCFD44E53EC20A72DC" box="[695,778,871,896]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Figs. 4</figureCitation>
C‒E); posterior margin of the pygophore with a medial process (
<emphasis id="89E8EAE5D148FFCCFF0EE5D4C02D72F8" box="[253,301,909,932]" italics="true" pageId="6" pageNumber="405">mpp</emphasis>
), dorsal margin concave (
<figureCitation id="23A72A72D148FFCCFDA9E5D5C3A072F8" box="[602,672,908,933]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Fig. 4</figureCitation>
E), process nearly vertical with a broad basal caudad protuberance (
<figureCitation id="23A72A72D148FFCCFF6CE5F6C1E37294" box="[159,227,943,968]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Fig. 4</figureCitation>
C); genital opening (
<emphasis id="89E8EAE5D148FFCCFE3AE5E8C0E57294" box="[457,485,945,968]" italics="true" pageId="6" pageNumber="405">go</emphasis>
) and anterior opening (
<emphasis id="89E8EAE5D148FFCCFD1EE5E8C2097294" box="[749,777,945,968]" italics="true" pageId="6" pageNumber="405">ao</emphasis>
) separated by a wide transverse bridge (
<emphasis id="89E8EAE5D148FFCCFB3EE5F6C5E67294" box="[1229,1254,943,968]" italics="true" pageId="6" pageNumber="405">br</emphasis>
) (
<figureCitation id="23A72A72D148FFCCFB0CE5F6C4427294" box="[1279,1346,943,968]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Fig. 4</figureCitation>
D); area surrounding paramere socket (
<emphasis id="89E8EAE5D148FFCCFE1FE58CC30572B0" box="[492,517,981,1004]" italics="true" pageId="6" pageNumber="405">ps</emphasis>
) with long, delicate setae (
<figureCitation id="23A72A72D148FFCCFCC1E58DC27572B0" box="[818,885,980,1005]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Fig. 4</figureCitation>
E). Paramere elongated and curved in their apical portion; body of the paramere with uniform diameter; apical curved portion cylindrical and apex acute, lateroventrally with long setae on distal region (
<figureCitation id="23A72A72D148FFCCFD49E245C3FE7568" box="[698,766,1052,1077]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Fig. 4</figureCitation>
G). Arms of articulatory apparatus (
<emphasis id="89E8EAE5D148FFCCFB64E244C5BA7568" box="[1175,1210,1053,1076]" italics="true" pageId="6" pageNumber="405">apt</emphasis>
) strongly curved in lateral view (
<figureCitation id="23A72A72D148FFCCFEC7E266C07C7504" box="[308,380,1087,1112]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Fig. 4</figureCitation>
G); dorsal phallothecal sclerite elongated; dorsoapically with a drop-like notch. Aedeagus symmetrical, cylindrical; endosoma with numerous tubercle-shaped sclerites (
<emphasis id="89E8EAE5D148FFCCFBF2E23CC51F7520" box="[1025,1055,1125,1148]" italics="true" pageId="6" pageNumber="405">tss</emphasis>
) (
<figureCitation id="23A72A72D148FFCCFBC4E23DC5857520" box="[1079,1157,1124,1149]" captionStart="FIGURE 4" captionStartId="8.[151,250,1923,1945]" captionTargetBox="[321,1277,193,1909]" captionTargetId="figure@8.[321,1277,193,1909]" captionTargetPageId="8" captionText="FIGURE 4. Liaghinella tuberculata sp. nov., male genitalia. A ‒ B. Sternite eight, A. ventral view. B. lateral view. C ‒ E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F ‒ G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites." httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="6" pageNumber="405">Figs. 4</figureCitation>
F‒G).
</paragraph>
<paragraph id="BB2336F7D148FFCCFF34E2DEC4057599" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E2DEC08B75FC" bold="true" box="[199,395,1159,1184]" pageId="6" pageNumber="405">Apterous female</emphasis>
(
<figureCitation id="23A72A72D148FFCCFE68E2DEC0E775FC" box="[411,487,1159,1184]" captionStart="FIGURE 5" captionStartId="9.[151,250,1933,1955]" captionTargetBox="[258,1338,193,1921]" captionTargetId="figure@9.[258,1338,193,1921]" captionTargetPageId="9" captionText="FIGURE 5. Liaghinella tuberculata sp. nov., female paratype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and prothorax, lateral view. E. Prothorax, lateral view. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810731/files/figure.png" pageId="6" pageNumber="405">Figs. 5</figureCitation>
A‒F) (total length, 
<quantity id="7C649B12D148FFCCFD31E2D1C22975FC" box="[706,809,1160,1184]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="6" pageNumber="405" unit="mm" value="12.3">12.3 mm</quantity>
; head length, 
<quantity id="7C649B12D148FFCCFC36E2D1C51F75FC" box="[965,1055,1160,1184]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="405" unit="mm" value="2.8">2.8 mm</quantity>
; prothorax length, 
<quantity id="7C649B12D148FFCCFB00E2DEC44C75FC" box="[1267,1356,1159,1184]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="6" pageNumber="405" unit="mm" value="3.5">3.5 mm</quantity>
; meso- and metathorax length, 
<quantity id="7C649B12D148FFCCFE6CE2F5C0F87598" box="[415,504,1196,1220]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="6" pageNumber="405" unit="mm" value="2.2">2.2 mm</quantity>
; abdomen length, 
<quantity id="7C649B12D148FFCCFD36E2F5C21F7598" box="[709,799,1196,1221]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="405" unit="mm" value="6.5">6.5 mm</quantity>
). Similar to male, slightly larger and wider.
</paragraph>
<paragraph id="BB2336F7D148FFCCFF34E296C5797451" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E296C01075B4" bold="true" box="[199,272,1231,1256]" pageId="6" pageNumber="405">Color.</emphasis>
Very similar to male’s except as follows. 
<emphasis id="89E8EAE5D148FFCCFD15E296C23775B4" box="[742,823,1231,1256]" italics="true" pageId="6" pageNumber="405">Thorax</emphasis>
: Dark brown, dorsally with several whitish spots (
<figureCitation id="23A72A72D148FFCCFA9CE296C1A57451" captionStart="FIGURE 5" captionStartId="9.[151,250,1933,1955]" captionTargetBox="[258,1338,193,1921]" captionTargetId="figure@9.[258,1338,193,1921]" captionTargetPageId="9" captionText="FIGURE 5. Liaghinella tuberculata sp. nov., female paratype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and prothorax, lateral view. E. Prothorax, lateral view. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810731/files/figure.png" pageId="6" pageNumber="405">Fig. 5</figureCitation>
E). 
<emphasis id="89E8EAE5D148FFCCFF3EE2ADC0407451" box="[205,320,1268,1293]" italics="true" pageId="6" pageNumber="405">Abdomen:</emphasis>
Dark brown with several spots whitish, abdominal tergite IX pale brown.
</paragraph>
<paragraph id="BB2336F7D148FFCCFF34E34EC0DD770C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E34EC03F746C" bold="true" box="[199,319,1303,1328]" pageId="6" pageNumber="405">Structure.</emphasis>
<emphasis id="89E8EAE5D148FFCCFEBFE34EC0B6746C" box="[332,438,1303,1328]" italics="true" pageId="6" pageNumber="405">Abdomen</emphasis>
: laterotergites with posterolateral dorsad projections larger than on the male (
<figureCitation id="23A72A72D148FFCCFA90E34EC1A47409" captionStart="FIGURE 5" captionStartId="9.[151,250,1933,1955]" captionTargetBox="[258,1338,193,1921]" captionTargetId="figure@9.[258,1338,193,1921]" captionTargetPageId="9" captionText="FIGURE 5. Liaghinella tuberculata sp. nov., female paratype. A ‒ B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and prothorax, lateral view. E. Prothorax, lateral view. F. Head, lateral view, arrow indicates clypeus." httpUri="https://zenodo.org/record/810731/files/figure.png" pageId="6" pageNumber="405">Figs. 5</figureCitation>
A‒B). 
<emphasis id="89E8EAE5D148FFCCFF09E365C0C07409" box="[250,448,1340,1365]" italics="true" pageId="6" pageNumber="405">Female genitalia</emphasis>
: Tergite 8 (
<emphasis id="89E8EAE5D148FFCCFDAFE365C37A7408" box="[604,634,1340,1364]" italics="true" pageId="6" pageNumber="405">T8</emphasis>
) oval, with the caudal margin (
<emphasis id="89E8EAE5D148FFCCFBFEE365C5527408" box="[1037,1106,1340,1364]" italics="true" pageId="6" pageNumber="405">mpm8</emphasis>
) rounded, medially with a longitudinal carina; tergite 9 (
<emphasis id="89E8EAE5D148FFCCFDF3E339C31D7424" box="[512,541,1376,1400]" italics="true" pageId="6" pageNumber="405">T9</emphasis>
) nearly rectangular with the distal margin (
<emphasis id="89E8EAE5D148FFCCFBC7E339C5787424" box="[1076,1144,1376,1400]" italics="true" pageId="6" pageNumber="405">mpm9</emphasis>
) emarginated (
<figureCitation id="23A72A72D148FFCCFADCE306C4797424" box="[1327,1401,1375,1400]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[226,1360,193,1871]" captionTargetId="figure@10.[226,1360,193,1871]" captionTargetPageId="10" captionText="FIGURE 6. Liaghinella tuberculata sp. nov., female genitalia, left gcx 8 removed. A. Ventral view. B. Lateral view. C. Tergite 8 and 9, caudal view. D. Gonocoxa eight, ventral view. Abbreviations: bc, bursa copulatrix; gap 8, gonapophysis eight; gap 9, gonapophysis nine; gcx 8, gonocoxa eight; gcx 9, gonocoxa nine; gpl, gonoplac; gr, ring gland; lap, latero anterior prolongation of the gonocoxa eight; mpm 8, medial posterior margin of gonocoxa eight; mpm 9, medial posterior margin of gonocoxa nine; T 8, tergite 8; T 9, tergite 9." httpUri="https://zenodo.org/record/810733/files/figure.png" pageId="6" pageNumber="405">Fig. 6</figureCitation>
C); gonocoxa 8 (
<emphasis id="89E8EAE5D148FFCCFED9E3DDC05F74C0" box="[298,351,1412,1436]" italics="true" pageId="6" pageNumber="405">gcx8</emphasis>
) nearly quadrangular, short and wide, with the lateral anterior area produced into a prolongation (
<emphasis id="89E8EAE5D148FFCCFF53E3FEC1C4749C" box="[160,196,1447,1472]" italics="true" pageId="6" pageNumber="405">lap</emphasis>
) sinuous apically (
<figureCitation id="23A72A72D148FFCCFE5EE3FEC0F6749C" box="[429,502,1447,1472]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[226,1360,193,1871]" captionTargetId="figure@10.[226,1360,193,1871]" captionTargetPageId="10" captionText="FIGURE 6. Liaghinella tuberculata sp. nov., female genitalia, left gcx 8 removed. A. Ventral view. B. Lateral view. C. Tergite 8 and 9, caudal view. D. Gonocoxa eight, ventral view. Abbreviations: bc, bursa copulatrix; gap 8, gonapophysis eight; gap 9, gonapophysis nine; gcx 8, gonocoxa eight; gcx 9, gonocoxa nine; gpl, gonoplac; gr, ring gland; lap, latero anterior prolongation of the gonocoxa eight; mpm 8, medial posterior margin of gonocoxa eight; mpm 9, medial posterior margin of gonocoxa nine; T 8, tergite 8; T 9, tergite 9." httpUri="https://zenodo.org/record/810733/files/figure.png" pageId="6" pageNumber="405">Fig. 6</figureCitation>
D); gonapophysis 8 (
<emphasis id="89E8EAE5D148FFCCFD07E3F1C22D749C" box="[756,813,1448,1472]" italics="true" pageId="6" pageNumber="405">gap8</emphasis>
) nearly triangular with some sparse setae medially; gonocoxa 9 (
<emphasis id="89E8EAE5D148FFCCFEDBE395C05D74B8" box="[296,349,1484,1508]" italics="true" pageId="6" pageNumber="405">gcx9</emphasis>
) elongated; gonapophysis 9 (
<emphasis id="89E8EAE5D148FFCCFD54E395C3E074B8" box="[679,736,1484,1508]" italics="true" pageId="6" pageNumber="405">gap9</emphasis>
) small and elongated; gonoplac (
<emphasis id="89E8EAE5D148FFCCFBA7E395C57774B9" box="[1108,1143,1484,1509]" italics="true" pageId="6" pageNumber="405">gpl</emphasis>
) rounded, fused medially, with a subapical ventral projection and longer setae on the apical margin (
<figureCitation id="23A72A72D148FFCCFC25E3B6C5197754" box="[982,1049,1519,1544]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[226,1360,193,1871]" captionTargetId="figure@10.[226,1360,193,1871]" captionTargetPageId="10" captionText="FIGURE 6. Liaghinella tuberculata sp. nov., female genitalia, left gcx 8 removed. A. Ventral view. B. Lateral view. C. Tergite 8 and 9, caudal view. D. Gonocoxa eight, ventral view. Abbreviations: bc, bursa copulatrix; gap 8, gonapophysis eight; gap 9, gonapophysis nine; gcx 8, gonocoxa eight; gcx 9, gonocoxa nine; gpl, gonoplac; gr, ring gland; lap, latero anterior prolongation of the gonocoxa eight; mpm 8, medial posterior margin of gonocoxa eight; mpm 9, medial posterior margin of gonocoxa nine; T 8, tergite 8; T 9, tergite 9." httpUri="https://zenodo.org/record/810733/files/figure.png" pageId="6" pageNumber="405">Fig. 6</figureCitation>
A); bursa copulatrix (
<emphasis id="89E8EAE5D148FFCCFAF8E3B6C4267754" box="[1291,1318,1519,1544]" italics="true" pageId="6" pageNumber="405">bc</emphasis>
) elongate, ventral medial surface of bursa copulatrix transversely striated; with ring gland (
<emphasis id="89E8EAE5D148FFCCFBCFE04CC5557770" box="[1084,1109,1557,1580]" italics="true" pageId="6" pageNumber="405">gr</emphasis>
) present, situated transverse across to the bursa (
<figureCitation id="23A72A72D148FFCCFE85E06EC0B9770C" box="[374,441,1591,1616]" captionStart="FIGURE 6" captionStartId="10.[151,250,1893,1915]" captionTargetBox="[226,1360,193,1871]" captionTargetId="figure@10.[226,1360,193,1871]" captionTargetPageId="10" captionText="FIGURE 6. Liaghinella tuberculata sp. nov., female genitalia, left gcx 8 removed. A. Ventral view. B. Lateral view. C. Tergite 8 and 9, caudal view. D. Gonocoxa eight, ventral view. Abbreviations: bc, bursa copulatrix; gap 8, gonapophysis eight; gap 9, gonapophysis nine; gcx 8, gonocoxa eight; gcx 9, gonocoxa nine; gpl, gonoplac; gr, ring gland; lap, latero anterior prolongation of the gonocoxa eight; mpm 8, medial posterior margin of gonocoxa eight; mpm 9, medial posterior margin of gonocoxa nine; T 8, tergite 8; T 9, tergite 9." httpUri="https://zenodo.org/record/810733/files/figure.png" pageId="6" pageNumber="405">Fig. 6</figureCitation>
A).
</paragraph>
</subSubSection>
<subSubSection id="F386657CD148FFCCFF34E005C5F277C4" pageId="6" pageNumber="405" type="etymology">
<paragraph id="BB2336F7D148FFCCFF34E005C5F277C4" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E005C04E7729" bold="true" box="[199,334,1628,1653]" pageId="6" pageNumber="405">Etymology.</emphasis>
The specific name 
<taxonomicName id="7C9C4D74D148FFCCFDCFE005C3BC7729" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[572,700,1628,1653]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="89E8EAE5D148FFCCFDCFE005C3BC7729" box="[572,700,1628,1653]" italics="true" pageId="6" pageNumber="405">tuberculata</emphasis>
</taxonomicName>
is taken from the Latin “tuberculum” (protuberance, tubercle) because of the conspicuous paired tubercles of the posterior margin of the pronotum of this species.
</paragraph>
</subSubSection>
<subSubSection id="F386657CD148FFCCFF34E0FDC5E877BC" pageId="6" pageNumber="405" type="biology_ecology">
<paragraph id="BB2336F7D148FFCCFF34E0FDC5E877BC" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E0FDC02777E1" bold="true" box="[199,295,1700,1725]" pageId="6" pageNumber="405">Biology.</emphasis>
The specimens were founded under forest litter and in very humid conditions. The specimens were collected when active, exclusively at night, similar to what was found for 
<taxonomicName id="7C9C4D74D148FFCCFC21E090C5E477BC" authority="Forero 2007" authorityName="Forero" authorityYear="2007" box="[978,1252,1735,1760]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D148FFCCFC21E090C53D77BC" box="[978,1085,1735,1760]" italics="true" pageId="6" pageNumber="405">L. andina</emphasis>
(
<bibRefCitation id="DF0D4B06D148FFCCFBBFE091C5DC77BC" author="Forero" box="[1100,1244,1736,1760]" pageId="6" pageNumber="405" refString="Forero, D. (2007) Description of a new species of Liaghinella (Hemiptera: Heteroptera: Reduviidae: Emesinae) from the Colombian Andes, with notes on its feeding habits and conservation status. Zootaxa, 1502, 55 - 68." type="journal article" year="2007">Forero 2007</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F386657CD148FFCCFF34E0B5C45C7610" pageId="6" pageNumber="405" type="distribution">
<paragraph id="BB2336F7D148FFCCFF34E0B5C45C7610" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E0B5C0607659" bold="true" box="[199,352,1772,1797]" pageId="6" pageNumber="405">Distribution.</emphasis>
Specimens of 
<taxonomicName id="7C9C4D74D148FFCCFDFEE0B4C3AD7659" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[525,685,1772,1797]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="89E8EAE5D148FFCCFDFEE0B4C3AD7659" box="[525,685,1772,1797]" italics="true" pageId="6" pageNumber="405">L. tuberculata</emphasis>
</taxonomicName>
were recently collected in one of the few relicts of high Andean forest near to urban areas (
<figureCitation id="23A72A72D148FFCCFE3DE156C3157674" box="[462,533,1807,1832]" captionStart="FIGURE 7" captionStartId="11.[151,250,2002,2024]" captionTargetBox="[262,1343,717,1968]" captionTargetId="figure@11.[193,1398,661,1988]" captionTargetPageId="11" captionText="FIGURE 7. Distribution map of Liaghinella species found in Colombia based on the examination of adult material." httpUri="https://zenodo.org/record/810735/files/figure.png" pageId="6" pageNumber="405">Fig. 7</figureCitation>
). “Parque Natural Chicaque” is a private reserve with 300 hectares of the wet montane forest with an altitudinal gradient between 2,000 to 2,700 meters (Jardin Botánico de 
<bibRefCitation id="DF0D4B06D148FFCCFB4FE16DC44F7610" author="Bogota" box="[1212,1359,1844,1868]" pageId="6" pageNumber="405" refString="Jardin Botanico de Bogota (1998) Guia Ecologica Parque Natural Chicaque. Jardin Botanico de Bogota Jose Celestino Mutis, 1, 1 - 27." type="journal article" year="1998">
<collectingRegion id="7958F815D148FFCCFB4FE16DC40D7610" box="[1212,1293,1844,1868]" country="Colombia" name="Bogota" pageId="6" pageNumber="405">Bogotá</collectingRegion>
1998
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F386657CD148FFC6FF34E10EC45C7507" lastPageId="12" lastPageNumber="411" pageId="6" pageNumber="405" type="discussion">
<paragraph id="BB2336F7D148FFCCFF34E10EC01F7681" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="405">
<emphasis id="89E8EAE5D148FFCCFF34E10EC04C762C" bold="true" box="[199,332,1879,1904]" pageId="6" pageNumber="405">Discussion.</emphasis>
This new species is very similar to 
<taxonomicName id="7C9C4D74D148FFCCFCFBE100C2B9762C" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[776,953,1879,1904]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="heldamariae" status="sp. nov.">
<emphasis id="89E8EAE5D148FFCCFCFBE100C2B9762C" box="[776,953,1879,1904]" italics="true" pageId="6" pageNumber="405">L. heldamariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED148FFCCFC37E10EC523762C" box="[964,1059,1879,1904]" pageId="6" pageNumber="405" rank="species">
<emphasis id="89E8EAE5D148FFCCFC37E10EC523762C" bold="true" box="[964,1059,1879,1904]" pageId="6" pageNumber="405">sp. nov.</emphasis>
</taxonomicNameLabel>
, but can be separated from the characters stated in the key above. It is also related to 
<taxonomicName id="7C9C4D74D148FFCCFD00E124C25F76C9" authorityName="Forero" authorityYear="2007" box="[755,863,1916,1941]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D148FFCCFD00E124C25F76C9" box="[755,863,1916,1941]" italics="true" pageId="6" pageNumber="405">L. andina</emphasis>
</taxonomicName>
and 
<taxonomicName id="7C9C4D74D148FFCCFC67E124C2E576C8" authorityName="Wygodzinsky" authorityYear="1966" box="[916,997,1916,1940]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="405" phylum="Arthropoda" rank="species" species="farri">
<emphasis id="89E8EAE5D148FFCCFC67E124C2E576C8" box="[916,997,1916,1940]" italics="true" pageId="6" pageNumber="405">L. farri</emphasis>
</taxonomicName>
, but several differences, particularly in the structure of head, structure of the thorax, and male and female genitalia, as stated above, allow distinguishing this species.
</paragraph>
<caption id="EFE3667FD149FFCDFF64E1DAC47A768A" httpUri="https://zenodo.org/record/810727/files/figure.png" pageId="7" pageNumber="406" targetBox="[226,1360,193,1902]" targetPageId="7">
<paragraph id="BB2336F7D149FFCDFF64E1DAC47A768A" blockId="7.[151,1435,1923,2006]" pageId="7" pageNumber="406">
<emphasis id="89E8EAE5D149FFCDFF64E1DAC01676C4" bold="true" box="[151,278,1923,1945]" pageId="7" pageNumber="406">FIGURE 3.</emphasis>
<taxonomicName id="7C9C4D74D149FFCDFED3E1DAC31276C5" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[288,530,1923,1945]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="406" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D149FFCDFED3E1DAC31276C5" box="[288,530,1923,1945]" italics="true" pageId="7" pageNumber="406">Liaghinella tuberculata</emphasis>
</taxonomicName>
<emphasis id="89E8EAE5D149FFCDFDEFE1DDC36F76C5" bold="true" box="[540,623,1924,1945]" pageId="7" pageNumber="406">
<taxonomicNameLabel id="92DB579ED149FFCDFDEFE1DDC36F76C5" box="[540,623,1924,1945]" pageId="7" pageNumber="406" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male holotype. A‒B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and thorax, lateral view. E. Prothorax, lateral view; arrow the pronotal process on distal margin. F. Head, lateral view, arrow indicates clypeus.
</paragraph>
</caption>
<caption id="EFE3667FD146FFC2FF64E1DAC58576A8" httpUri="https://zenodo.org/record/810729/files/figure.png" pageId="8" pageNumber="407" targetBox="[321,1277,193,1909]" targetPageId="8">
<paragraph id="BB2336F7D146FFC2FF64E1DAC58576A8" blockId="8.[151,1436,1923,2036]" pageId="8" pageNumber="407">
<emphasis id="89E8EAE5D146FFC2FF64E1DAC01776C4" bold="true" box="[151,279,1923,1945]" pageId="8" pageNumber="407">FIGURE 4.</emphasis>
<taxonomicName id="7C9C4D74D146FFC2FED1E1DAC31576C5" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[290,533,1923,1945]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="407" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D146FFC2FED1E1DAC31576C5" box="[290,533,1923,1945]" italics="true" pageId="8" pageNumber="407">Liaghinella tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED146FFC2FDECE1DAC37476C4" box="[543,628,1923,1944]" pageId="8" pageNumber="407" rank="species">
<emphasis id="89E8EAE5D146FFC2FDECE1DAC37476C4" bold="true" box="[543,628,1923,1944]" pageId="8" pageNumber="407">sp. nov.</emphasis>
</taxonomicNameLabel>
, male genitalia. A‒B. Sternite eight, A. ventral view. B. lateral view. C‒E. Pygophore, C. right lateral view. E. dorsal view, F. caudal view. F‒G. Extended phallus, G. dorsal view, H. right lateral view. H. Paramere, lateral view. Abbreviations: ao, anterior opening; apt, articulatory apparatus; br, transverse bridge; go, genital opening; mpp, medial process of the pygophore; ps, paramere socket; tss, tubercle-shaped sclerites.
</paragraph>
</caption>
<caption id="EFE3667FD147FFC3FF64E1D4C2B476BC" httpUri="https://zenodo.org/record/810731/files/figure.png" pageId="9" pageNumber="408" targetBox="[258,1338,193,1921]" targetPageId="9">
<paragraph id="BB2336F7D147FFC3FF64E1D4C2B476BC" blockId="9.[151,1435,1933,2016]" pageId="9" pageNumber="408">
<emphasis id="89E8EAE5D147FFC3FF64E1D4C01576FE" bold="true" box="[151,277,1933,1955]" pageId="9" pageNumber="408">FIGURE 5.</emphasis>
<taxonomicName id="7C9C4D74D147FFC3FEEDE1D4C30F76FF" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[286,527,1933,1955]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="408" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D147FFC3FEEDE1D4C30F76FF" box="[286,527,1933,1955]" italics="true" pageId="9" pageNumber="408">Liaghinella tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED147FFC3FDEBE1D4C36A76FE" box="[536,618,1933,1954]" pageId="9" pageNumber="408" rank="species">
<emphasis id="89E8EAE5D147FFC3FDEBE1D4C36A76FE" bold="true" box="[536,618,1933,1954]" pageId="9" pageNumber="408">sp. nov.</emphasis>
</taxonomicNameLabel>
, female paratype. A‒B. Habitus, A. dorsal view; B. lateral view. C. Pro and - mesothorax, dorsal view, arrow points to the pronotal process on posterior margin of prothorax. D. Head and prothorax, lateral view. E. Prothorax, lateral view. F. Head, lateral view, arrow indicates clypeus.
</paragraph>
</caption>
<caption id="EFE3667FD144FFC0FF64E13CC0A276A9" httpUri="https://zenodo.org/record/810733/files/figure.png" pageId="10" pageNumber="409" targetBox="[226,1360,193,1871]" targetPageId="10">
<paragraph id="BB2336F7D144FFC0FF64E13CC0A276A9" blockId="10.[151,1436,1892,2037]" pageId="10" pageNumber="409">
<emphasis id="89E8EAE5D144FFC0FF64E13CC0137626" bold="true" box="[151,275,1893,1915]" pageId="10" pageNumber="409">FIGURE 6.</emphasis>
<taxonomicName id="7C9C4D74D144FFC0FEE9E13DC3097626" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[282,521,1892,1914]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="409" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D144FFC0FEE9E13DC3097626" box="[282,521,1892,1914]" italics="true" pageId="10" pageNumber="409">Liaghinella tuberculata</emphasis>
</taxonomicName>
<emphasis id="89E8EAE5D144FFC0FDFCE13CC3677626" bold="true" box="[527,615,1893,1914]" pageId="10" pageNumber="409">
<taxonomicNameLabel id="92DB579ED144FFC0FDFCE13CC3627626" box="[527,610,1893,1914]" pageId="10" pageNumber="409" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female genitalia, left gcx8 removed. A. Ventral view. B. Lateral view. C. Tergite 8 and 9, caudal view. D. Gonocoxa eight, ventral view. Abbreviations: bc, bursa copulatrix; gap8, gonapophysis eight; gap9, gonapophysis nine; gcx8, gonocoxa eight; gcx9, gonocoxa nine; gpl, gonoplac; gr, ring gland; lap, latero anterior prolongation of the gonocoxa eight; mpm8, medial posterior margin of gonocoxa eight; mpm9, medial posterior margin of gonocoxa nine; T8, tergite 8; T9, tergite 9.
</paragraph>
</caption>
<paragraph id="BB2336F7D145FFC1FF34E6CEC2ED733C" blockId="11.[151,1437,151,608]" pageId="11" pageNumber="410">
The male and female genitalia in 
<taxonomicName id="7C9C4D74D145FFC1FDBFE6CEC3CD71EC" authorityName="Wygodzinsky" authorityYear="1966" box="[588,717,151,176]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D145FFC1FDBFE6CEC3CD71EC" box="[588,717,151,176]" italics="true" pageId="11" pageNumber="410">Liaghinella</emphasis>
</taxonomicName>
have valuable taxonomic information to separate species. The medial process of the pygophore is spiniform in 
<taxonomicName id="7C9C4D74D145FFC1FD40E6E4C2F57189" authority="Wygodzinsky 1966" authorityName="Wygodzinsky" authorityYear="1966" box="[691,1013,188,213]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="farri">
<emphasis id="89E8EAE5D145FFC1FD40E6E4C2037188" box="[691,771,188,212]" italics="true" pageId="11" pageNumber="410">L. farri</emphasis>
(
<bibRefCitation id="DF0D4B06D145FFC1FCE2E6E5C2ED7189" author="Wygodzinsky" box="[785,1005,188,213]" pageId="11" pageNumber="410" refString="Wygodzinsky, P. (1966) A monograph of the Emesinae (Reduviidae, Hemiptera). Bulletin of the American Museum of Natural History, 133, 1 - 614." type="journal article" year="1966">Wygodzinsky 1966</bibRefCitation>
)
</taxonomicName>
, rectangular with a straight apex in 
<taxonomicName id="7C9C4D74D145FFC1FA76E6E4C08871A4" authority="Forero 2007" authorityName="Forero" authorityYear="2007" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D145FFC1FA76E6E4C1E571A4" italics="true" pageId="11" pageNumber="410">L. andina</emphasis>
(
<bibRefCitation id="DF0D4B06D145FFC1FF00E6B9C08071A4" author="Forero" box="[243,384,224,248]" pageId="11" pageNumber="410" refString="Forero, D. (2007) Description of a new species of Liaghinella (Hemiptera: Heteroptera: Reduviidae: Emesinae) from the Colombian Andes, with notes on its feeding habits and conservation status. Zootaxa, 1502, 55 - 68." type="journal article" year="2007">Forero 2007</bibRefCitation>
)
</taxonomicName>
and rectangular with the apex concave in 
<taxonomicName id="7C9C4D74D145FFC1FC93E6B8C2FD71A4" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[864,1021,223,248]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D145FFC1FC93E6B8C2FD71A4" box="[864,1021,223,248]" italics="true" pageId="11" pageNumber="410">L. tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED145FFC1FBF7E686C55E71A4" box="[1028,1118,223,248]" pageId="11" pageNumber="410" rank="species">
<emphasis id="89E8EAE5D145FFC1FBF7E686C55E71A4" bold="true" box="[1028,1118,223,248]" pageId="11" pageNumber="410">sp. nov.</emphasis>
</taxonomicNameLabel>
The apex of the paramere is strongly curved in 
<taxonomicName id="7C9C4D74D145FFC1FE82E75CC0DF7041" authorityName="Forero" authorityYear="2007" box="[369,479,260,285]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D145FFC1FE82E75CC0DF7041" box="[369,479,260,285]" italics="true" pageId="11" pageNumber="410">L. andina</emphasis>
</taxonomicName>
and 
<emphasis id="89E8EAE5D145FFC1FDEAE75CC3747040" box="[537,628,260,284]" italics="true" pageId="11" pageNumber="410">
<taxonomicName id="7C9C4D74D145FFC1FDEAE75CC3707040" authorityName="Wygodzinsky" authorityYear="1966" box="[537,624,260,284]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="farri">L. farri</taxonomicName>
.
</emphasis>
The anterior and posterior opening of the pygophore are narrow in 
<taxonomicName id="7C9C4D74D145FFC1FA76E75CC1E5701C" authorityName="Forero" authorityYear="2007" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D145FFC1FA76E75CC1E5701C" italics="true" pageId="11" pageNumber="410">L. andina</emphasis>
</taxonomicName>
and wider in 
<taxonomicName id="7C9C4D74D145FFC1FE8EE770C31A701C" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[381,538,295,320]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="89E8EAE5D145FFC1FE8EE770C31A701C" box="[381,538,295,320]" italics="true" pageId="11" pageNumber="410">L. tuberculata</emphasis>
</taxonomicName>
. The particular sclerotizations of the endosoma are different between the known males: in 
<taxonomicName id="7C9C4D74D145FFC1FEFAE714C05C7038" authorityName="Wygodzinsky" authorityYear="1966" box="[265,348,332,356]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="farri">
<emphasis id="89E8EAE5D145FFC1FEFAE714C05C7038" box="[265,348,332,356]" italics="true" pageId="11" pageNumber="410">L. farri</emphasis>
</taxonomicName>
are small and more homogeneous, whereas in 
<taxonomicName id="7C9C4D74D145FFC1FC8DE714C2EC7039" authorityName="Forero" authorityYear="2007" box="[894,1004,332,357]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D145FFC1FC8DE714C2EC7039" box="[894,1004,332,357]" italics="true" pageId="11" pageNumber="410">L. andina</emphasis>
</taxonomicName>
and 
<taxonomicName id="7C9C4D74D145FFC1FBD4E714C5C77039" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[1063,1223,332,357]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D145FFC1FBD4E714C5C77039" box="[1063,1223,332,357]" italics="true" pageId="11" pageNumber="410">L. tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED145FFC1FB22E715C42D7039" box="[1233,1325,332,357]" pageId="11" pageNumber="410" rank="species">
<emphasis id="89E8EAE5D145FFC1FB22E715C42D7039" bold="true" box="[1233,1325,332,357]" pageId="11" pageNumber="410">sp. nov.</emphasis>
</taxonomicNameLabel>
are more strongly sclerotized and less homogenous. In the female genitalia, the structure of the tergites 8 and 9 are characteristic at least in 
<taxonomicName id="7C9C4D74D145FFC1FE50E7CCC30E70F1" authorityName="Forero" authorityYear="2007" box="[419,526,404,429]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D145FFC1FE50E7CCC30E70F1" box="[419,526,404,429]" italics="true" pageId="11" pageNumber="410">L. andina</emphasis>
</taxonomicName>
, 
<taxonomicName id="7C9C4D74D145FFC1FDEFE7CCC3C670F1" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[540,710,404,429]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="heldamariae" status="sp. nov.">
<emphasis id="89E8EAE5D145FFC1FDEFE7CCC3C670F1" box="[540,710,404,429]" italics="true" pageId="11" pageNumber="410">L. heldamariae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED145FFC1FD3EE7CDC22670F1" box="[717,806,404,429]" pageId="11" pageNumber="410" rank="species">
<emphasis id="89E8EAE5D145FFC1FD3EE7CDC22670F1" bold="true" box="[717,806,404,429]" pageId="11" pageNumber="410">sp. nov.</emphasis>
</taxonomicNameLabel>
, and 
<taxonomicName id="7C9C4D74D145FFC1FC91E7CCC2FF70F1" authority="Castro-Huertas &amp; Forero, 2017" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[866,1023,404,429]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="tuberculata" status="sp. nov.">
<emphasis id="89E8EAE5D145FFC1FC91E7CCC2FF70F1" box="[866,1023,404,429]" italics="true" pageId="11" pageNumber="410">L. tuberculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="92DB579ED145FFC1FBF5E7CDC55F70F1" box="[1030,1119,404,429]" pageId="11" pageNumber="410" rank="species">
<emphasis id="89E8EAE5D145FFC1FBF5E7CDC55F70F1" bold="true" box="[1030,1119,404,429]" pageId="11" pageNumber="410">sp. nov.</emphasis>
</taxonomicNameLabel>
All these attributes found in the Colombian species point to a marked difference between 
<taxonomicName id="7C9C4D74D145FFC1FCCDE7E0C28E708C" authorityName="Wygodzinsky" authorityYear="1966" box="[830,910,440,464]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="farri">
<emphasis id="89E8EAE5D145FFC1FCCDE7E0C28E708C" box="[830,910,440,464]" italics="true" pageId="11" pageNumber="410">L. farri</emphasis>
</taxonomicName>
and the Colombian species. The 
<typeStatus id="64278855D145FFC1FB0CE7EEC461708C" box="[1279,1377,439,464]" pageId="11" pageNumber="410" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="7C9C4D74D145FFC1FA76E7E0C1CA70A8" authorityName="Wygodzinsky" authorityYear="1966" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="species" species="farri">
<emphasis id="89E8EAE5D145FFC1FA76E7E0C1CA70A8" italics="true" pageId="11" pageNumber="410">L. farri</emphasis>
</taxonomicName>
is not well preserved (D.Forero, pers. obs), thus a reexamination of it will not be very helpful, in particular with regard to the male genitalia. Only further collecting for additional specimens and sexes, coupled with a phylogenetic analysis of all related taxa will clarify the relationships among species and will help to decide if the species from 
<collectingCountry id="C38B7667D145FFC1FEDFE41EC09A733C" box="[300,410,583,608]" name="Colombia" pageId="11" pageNumber="410">Colombia</collectingCountry>
are really congeneric with the species from 
<collectingCountry id="C38B7667D145FFC1FC78E41EC2E9733C" box="[907,1001,583,608]" name="Jamaica" pageId="11" pageNumber="410">Jamaica</collectingCountry>
.
</paragraph>
<caption id="EFE3667FD145FFC1FF64E18BC42E76B4" box="[151,1326,2002,2024]" httpUri="https://zenodo.org/record/810735/files/figure.png" pageId="11" pageNumber="410" targetBox="[262,1343,717,1968]" targetPageId="11">
<paragraph id="BB2336F7D145FFC1FF64E18BC42E76B4" blockId="11.[151,1326,2002,2024]" box="[151,1326,2002,2024]" pageId="11" pageNumber="410">
<emphasis id="89E8EAE5D145FFC1FF64E18BC01276BB" bold="true" box="[151,274,2002,2024]" pageId="11" pageNumber="410">FIGURE 7.</emphasis>
Distribution map of 
<taxonomicName id="7C9C4D74D145FFC1FE14E18BC35C76B4" authorityName="Wygodzinsky" authorityYear="1966" box="[487,604,2002,2024]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="410" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D145FFC1FE14E18BC35C76B4" box="[487,604,2002,2024]" italics="true" pageId="11" pageNumber="410">Liaghinella</emphasis>
</taxonomicName>
species found in Colombia based on the examination of adult material.
</paragraph>
</caption>
<paragraph id="BB2336F7D142FFC6FF34E6CEC2837039" blockId="12.[151,1437,151,1115]" pageId="12" pageNumber="411">
<emphasis id="89E8EAE5D142FFC6FF34E6CEC27E71EC" bold="true" box="[199,894,151,176]" pageId="12" pageNumber="411">
Small range distributions of 
<taxonomicName id="7C9C4D74D142FFC6FDEBE6CEC39E71EC" authorityName="Wygodzinsky" authorityYear="1966" box="[536,670,151,176]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D142FFC6FDEBE6CEC39E71EC" bold="true" box="[536,670,151,176]" italics="true" pageId="12" pageNumber="411">Liaghinella</emphasis>
</taxonomicName>
in the high 
<taxonomicName id="7C9C4D74D142FFC6FCDDE6CEC27971EC" authorityName="Armenteras et al." authorityYear="2003" box="[814,889,151,176]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="412" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
.
</emphasis>
Biogeographic hypotheses of 
<taxonomicName id="7C9C4D74D142FFC6FB29E6C1C45E71F3" authorityName="Latreille" authorityYear="1810" box="[1242,1374,152,175]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
have focused on the groups with medical and economic importance. Within the Cimicomorpha, 
<taxonomicName id="7C9C4D74D142FFC6FB4EE6E5C4457189" authorityName="Jeannel" authorityYear="1919" box="[1213,1349,188,213]" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="subFamily" subFamily="Triatominae">Triatominae</taxonomicName>
among 
<taxonomicName id="7C9C4D74D142FFC6FF64E686C01771A4" authorityName="Latreille" authorityYear="1807" box="[151,279,223,248]" class="Insecta" family="Reduviidae" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="family">Reduviidae</taxonomicName>
have been the focus of studies about distributional patterns (
<bibRefCitation id="DF0D4B06D142FFC6FBF2E686C5E671A4" author="Ferrari" box="[1025,1254,223,248]" pageId="12" pageNumber="411" refString="Ferrari, A., Barao, K., Michels, F. &amp; Grazia, J. (2015) Classification and Biogeography of Neotropical True Bugs. In: Panizzi, A. R. &amp; Grazia, J. (Eds.), True Bugs (Heteroptera) of the Neotropics. 1 st Edition. Entomology in Focus, 2, pp. 57 ‾ 87. https: // doi. org / 10.1007 / 978 - 94 - 017 - 9861 - 7 _ 3" type="journal article" year="2015">
Ferrari 
<emphasis id="89E8EAE5D142FFC6FBAEE6B8C59571A4" box="[1117,1173,223,248]" italics="true" pageId="12" pageNumber="411">et al</emphasis>
. 2015
</bibRefCitation>
). However, the distribution of the 
<taxonomicName id="7C9C4D74D142FFC6FE83E75CC0F47040" authorityName="Latreille" authorityYear="1810" box="[368,500,261,284]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
fauna in the Neotropics is still poorly known, particularly on the high 
<taxonomicName id="7C9C4D74D142FFC6FAC6E75DC47C7041" authorityName="Armenteras et al." authorityYear="2003" box="[1333,1404,260,285]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="412" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
in 
<collectingCountry id="C38B7667D142FFC6FF64E77EC00A701C" box="[151,266,295,320]" name="Colombia" pageId="12" pageNumber="411">Colombia</collectingCountry>
. This lack of information hinders the ability to identify and prioritize critical areas for conservation efforts and to understand the evolution of these areas in the 
<taxonomicName id="7C9C4D74D142FFC6FCC7E715C27F7039" authorityName="Armenteras et al." authorityYear="2003" box="[820,895,332,357]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="412" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
.
</paragraph>
<paragraph id="BB2336F7D142FFC6FF34E736C47F724B" blockId="12.[151,1437,151,1115]" pageId="12" pageNumber="411">
Species of 
<taxonomicName id="7C9C4D74D142FFC6FEBEE736C0CE70D4" authorityName="Wygodzinsky" authorityYear="1966" box="[333,462,367,392]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D142FFC6FEBEE736C0CE70D4" box="[333,462,367,392]" italics="true" pageId="12" pageNumber="411">Liaghinella</emphasis>
</taxonomicName>
found 
<collectingCountry id="C38B7667D142FFC6FDDBE736C39670D4" box="[552,662,367,392]" name="Colombia" pageId="12" pageNumber="411">Colombia</collectingCountry>
have small range distributions on the eastern Colombian 
<taxonomicName id="7C9C4D74D142FFC6FABDE736C49970D4" authorityName="Armenteras et al." authorityYear="2003" box="[1358,1433,367,392]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="412" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
. 
<bibRefCitation id="DF0D4B06D142FFC6FF64E7C1C03B70F3" author="Harvey" box="[151,315,407,432]" pageId="12" pageNumber="411" refString="Harvey, M. S. (2002) Short-range endemism amongst the Australian fauna: some examples from non-marine environments. Invertebrate Systematics, 16, 555 - 570." type="journal article" year="2002">Harvey (2002)</bibRefCitation>
proposed that less than 
<quantity id="7C649B12D142FFC6FDB9E7CEC3C270F3" box="[586,706,406,431]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="411" unit="km" value="10.0">10.000 km</quantity>
2 can qualify as a short-range endemic group, as is apparently the case with the Andean 
<taxonomicName id="7C9C4D74D142FFC6FE6AE7E3C31A708F" authorityName="Wygodzinsky" authorityYear="1966" box="[409,538,442,467]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D142FFC6FE6AE7E3C31A708F" box="[409,538,442,467]" italics="true" pageId="12" pageNumber="411">Liaghinella</emphasis>
</taxonomicName>
species. 
<taxonomicName id="7C9C4D74D142FFC6FD75E7E3C25F708F" authorityName="Forero" authorityYear="2007" box="[646,863,442,467]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D142FFC6FD75E7E3C25F708F" box="[646,863,442,467]" italics="true" pageId="12" pageNumber="411">Liaghinella andina</emphasis>
</taxonomicName>
is found in two localities less than 50 kilometers apart, 
<taxonomicName id="7C9C4D74D142FFC6FF2FE7B9C08870AB" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[220,392,478,503]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="heldamariae">
<emphasis id="89E8EAE5D142FFC6FF2FE7B9C08870AB" box="[220,392,478,503]" italics="true" pageId="12" pageNumber="411">L. heldamariae</emphasis>
</taxonomicName>
is found again in two localities set apart by about 130 kilometers, and 
<taxonomicName id="7C9C4D74D142FFC6FB6EE7B9C43A70AB" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[1181,1338,478,503]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="89E8EAE5D142FFC6FB6EE7B9C43A70AB" box="[1181,1338,478,503]" italics="true" pageId="12" pageNumber="411">L. tuberculata</emphasis>
</taxonomicName>
is found only in its 
<typeStatus id="64278855D142FFC6FEE3E45AC0407347" box="[272,320,515,539]" pageId="12" pageNumber="411">type</typeStatus>
locality (
<figureCitation id="23A72A72D142FFC6FE58E45BC0F07347" box="[427,496,514,539]" captionStart="FIGURE 7" captionStartId="11.[151,250,2002,2024]" captionTargetBox="[262,1343,717,1968]" captionTargetId="figure@11.[193,1398,661,1988]" captionTargetPageId="11" captionText="FIGURE 7. Distribution map of Liaghinella species found in Colombia based on the examination of adult material." httpUri="https://zenodo.org/record/810735/files/figure.png" pageId="12" pageNumber="411">Fig. 7</figureCitation>
). At least two species of 
<taxonomicName id="7C9C4D74D142FFC6FCFEE45BC28E7347" authorityName="Wygodzinsky" authorityYear="1966" box="[781,910,514,539]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D142FFC6FCFEE45BC28E7347" box="[781,910,514,539]" italics="true" pageId="12" pageNumber="411">Liaghinella</emphasis>
</taxonomicName>
are always in the same or very close localities for most of its range. This might be considered as sympatric, but in fact 
<taxonomicName id="7C9C4D74D142FFC6FC4CE471C55D7363" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[959,1117,550,575]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="89E8EAE5D142FFC6FC4CE471C55D7363" box="[959,1117,550,575]" italics="true" pageId="12" pageNumber="411">L. tuberculata</emphasis>
</taxonomicName>
and 
<taxonomicName id="7C9C4D74D142FFC6FB67E471C4007363" authorityName="Forero" authorityYear="2007" box="[1172,1280,550,575]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D142FFC6FB67E471C4007363" box="[1172,1280,550,575]" italics="true" pageId="12" pageNumber="411">L. andina</emphasis>
</taxonomicName>
are not found at the same altitude in Chicaque, with 
<taxonomicName id="7C9C4D74D142FFC6FDB3E415C3AB733F" authorityName="Forero" authorityYear="2007" box="[576,683,586,611]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D142FFC6FDB3E415C3AB733F" box="[576,683,586,611]" italics="true" pageId="12" pageNumber="411">L. andina</emphasis>
</taxonomicName>
always higher than 
<taxonomicName id="7C9C4D74D142FFC6FC79E415C526733F" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[906,1062,586,611]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="tuberculata">
<emphasis id="89E8EAE5D142FFC6FC79E415C526733F" box="[906,1062,586,611]" italics="true" pageId="12" pageNumber="411">L. tuberculata</emphasis>
</taxonomicName>
. It is not clear if this difference is due to an association to particular habitats, but at least with the data at hand it might not be the case. 
<taxonomicName id="7C9C4D74D142FFC6FAE8E437C1E573F7" authorityName="Forero" authorityYear="2007" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D142FFC6FAE8E437C1E573F7" italics="true" pageId="12" pageNumber="411">Liaghinella andina</emphasis>
</taxonomicName>
was described from a high Andean forest, and the habitat of the specimen from Chicaque is an Oak forest (
<taxonomicName id="7C9C4D74D142FFC6FF53E4EEC07E7393" box="[160,382,694,719]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="411" phylum="Tracheophyta" rank="species" species="humboldtii">
<emphasis id="89E8EAE5D142FFC6FF53E4EEC07E7393" box="[160,382,694,719]" italics="true" pageId="12" pageNumber="411">Quercus humboldtii</emphasis>
</taxonomicName>
). Similarly, 
<taxonomicName id="7C9C4D74D142FFC6FDFFE4E1C3B87393" authorityName="Castro-Huertas &amp; Forero" authorityYear="2017" box="[524,696,694,719]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="heldamariae">
<emphasis id="89E8EAE5D142FFC6FDFFE4E1C3B87393" box="[524,696,694,719]" italics="true" pageId="12" pageNumber="411">L. heldamariae</emphasis>
</taxonomicName>
is found in paramo in 
<collectingCountry id="C38B7667D142FFC6FC4BE4E1C5377393" box="[952,1079,696,719]" name="Montserrat" pageId="12" pageNumber="411">Monserrate</collectingCountry>
at about 3,000 meters, whereas 
<taxonomicName id="7C9C4D74D142FFC6FF64E485C00573AF" authorityName="Forero" authorityYear="2007" box="[151,261,730,755]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="species" species="andina">
<emphasis id="89E8EAE5D142FFC6FF64E485C00573AF" box="[151,261,730,755]" italics="true" pageId="12" pageNumber="411">L. andina</emphasis>
</taxonomicName>
is found nearby but at a lower altitude in a high Andean forest. Therefore, we argue that although all these species are found in relatively nearby localities, they are separated by the altitude in which they are found.
</paragraph>
<paragraph id="BB2336F7D142FFC6FF34E57AC3E072B3" blockId="12.[151,1437,151,1115]" pageId="12" pageNumber="411">
In 
<collectingCountry id="C38B7667D142FFC6FF1AE57BC05A7267" box="[233,346,802,827]" name="Colombia" pageId="12" pageNumber="411">Colombia</collectingCountry>
, only a broad-shouldered water strider heteropteran (
<taxonomicName id="7C9C4D74D142FFC6FC34E57BC5297267" box="[967,1065,802,827]" class="Insecta" family="Veliidae" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="family">Veliidae</taxonomicName>
: 
<taxonomicName id="7C9C4D74D142FFC6FBC4E57BC1D17203" authority="Padilla &amp; Moreira 2013" authorityName="Mayr" authorityYear="1865" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D142FFC6FBC4E57BC5B67267" box="[1079,1206,802,827]" italics="true" pageId="12" pageNumber="411">Rhagovelia</emphasis>
, 
<bibRefCitation id="DF0D4B06D142FFC6FB3BE57BC1D17203" author="Padilla" pageId="12" pageNumber="411" refString="Padilla-Gil, D. &amp; Moreira, F. (2013) Checklist, taxonomy and distribution of the Rhagovelia Mayr, 1865 (Hemiptera: Heteroptera: Veliidae) of the Americas. Zootaxa, 3640 (3), 409 - 424." type="journal article" year="2013">Padilla &amp; Moreira 2013</bibRefCitation>
</taxonomicName>
) has been documented with respect to altitudinal ranges. Studies with carabid beetles indicate that the high rates of microendemism in the high 
<taxonomicName id="7C9C4D74D142FFC6FDCAE533C38072DF" authorityName="Armenteras et al." authorityYear="2003" box="[569,640,874,899]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="412" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
of 
<collectingCountry id="C38B7667D142FFC6FD59E533C20772DF" box="[682,775,874,899]" name="Ecuador" pageId="12" pageNumber="411">Ecuador</collectingCountry>
are associated with climate, humid areas, volcanism, and others variables, resulting in conditions for the diversification of a highly specialized montane fauna (
<bibRefCitation id="DF0D4B06D142FFC6FAFCE5D6C49872FB" author="Moret" box="[1295,1432,910,935]" pageId="12" pageNumber="411" refString="Moret, P. (2005) Los coleopteros Carabidae del paramo en los Andes del Ecuador. Sistematica, ecologia y biogeografia. Monografia 2. Pontificia Universidad Catolica del Ecuador, Centro de Biodiversidad y Ambiente, Quito, 306 pp." type="book" year="2005">Moret 2005</bibRefCitation>
, 
<bibRefCitation id="DF0D4B06D142FFC6FF64E5EAC1D47297" author="Moret" box="[151,212,947,971]" pageId="12" pageNumber="411" refString="Moret, P. (2009) Altitudinal distribution, diversity and endemicity of Carabidae (Coleoptera) in the paramos of Ecuadorian Andes, Annales de la Societe entomologique de France, 45 (4), 500 - 510." type="journal article" year="2009">2009</bibRefCitation>
). This could be also the case for some groups in 
<taxonomicName id="7C9C4D74D142FFC6FD1EE5EAC2747296" authorityName="Latreille" authorityYear="1810" box="[749,884,947,970]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
, highlighting the importance of Andean forests in diversification processes of Colombian 
<taxonomicName id="7C9C4D74D142FFC6FDA7E581C3DC72B3" authorityName="Latreille" authorityYear="1810" box="[596,732,984,1007]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Heteroptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="order">Heteroptera</taxonomicName>
.
</paragraph>
<paragraph id="BB2336F7D142FFC6FF3CE5A3C45C7507" blockId="12.[151,1437,151,1115]" pageId="12" pageNumber="411">
If further collecting show that the high Andean species of 
<taxonomicName id="7C9C4D74D142FFC6FC94E5A3C2E8754F" authorityName="Wygodzinsky" authorityYear="1966" box="[871,1000,1018,1043]" class="Insecta" family="Reduviidae" genus="Liaghinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="411" phylum="Arthropoda" rank="genus">
<emphasis id="89E8EAE5D142FFC6FC94E5A3C2E8754F" box="[871,1000,1018,1043]" italics="true" pageId="12" pageNumber="411">Liaghinella</emphasis>
</taxonomicName>
have in fact small distribution ranges, this will affect negatively the extinction probabilities of their populations because they will be more susceptible to climate change (
<bibRefCitation id="DF0D4B06D142FFC6FEBDE21BC3217507" author="Prather" box="[334,545,1090,1115]" pageId="12" pageNumber="411" refString="Prather, C. M., Pelini, S. L., Laws, A., Rivest, E., Woltz, M., Bloch, C. P., Del Toro, I., Ho, C. - K., Kominoski, J., Newbold, T. A. S., Parsons, S. &amp; Joern, A. (2013) Invertebrates, ecosystem services and climate change. Biological Reviews, 88, 327 - 348." type="journal article" year="2013">
Prather 
<emphasis id="89E8EAE5D142FFC6FE5BE21DC0E07507" box="[424,480,1090,1115]" italics="true" pageId="12" pageNumber="411">et al.</emphasis>
2013
</bibRefCitation>
) and anthropogenic transformations of the landscape affecting the 
<taxonomicName id="7C9C4D74D142FFC6FAFEE21BC4587507" authorityName="Armenteras et al." authorityYear="2003" box="[1293,1368,1090,1115]" class="Insecta" family="Cixiidae" genus="Andes" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="412" phylum="Arthropoda" rank="genus">Andes</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>