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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZBK="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt &amp; Liew, Thor-Seng" docDate="2017" docId="3301A013AA937836D472F9776624E312" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus perakensis Crosse 1879" docType="treatment" docVersion="5" lastPageNumber="37" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="36" updateTime="1668164728433" updateUser="ExternalLinkService">
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<mods:title>A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
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<paragraph pageId="35" pageNumber="36">
<taxonomicName authority="Crosse, 1879 a" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis Crosse, 1879a</taxonomicName>
Figures 7S, 24, 31V
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="reference_group">
<paragraph pageId="35" pageNumber="36">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
Crosse, 1879a: 206-208, plate 7, figure 7;
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="1 - 2" title="A brief collecting trip to Perak, West Malaysia: Part Two." volume="12" year="1998 b">Crosse 1879b</bibRefCitation>
: 339-340;
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Journal of the Asiatic Society of Bengal" pageId="52" pageNumber="53" pagination="299 - 316" title="The landshells of Perak." url="http://www.biodiversitylibrary.org/part/145559/summary" volume="55" year="1886">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1886
</bibRefCitation>
: 310;
<bibRefCitation author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="53" pageNumber="54" pagination="194 - 199" title="On the land operculate Mollusca collected during the &quot; Skeat Expedition &quot; to the Malay Peninsula in 1899 - 1900." url="http://www.biodiversitylibrary.org/part/145330/summary" volume="1903" year="1903">Sykes 1903</bibRefCitation>
: 195;
<bibRefCitation author="Tenison-Woods, JE" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="53" pageNumber="54" pagination="1003 - 1095" title="Malaysian land and freshwater mollusca." url="https://doi.org/10.5962/bhl.title.13026" volume="3" year="1888">Tension-Woods 1888</bibRefCitation>
: 1068;
<bibRefCitation author="Tielecke, H" journalOrPublisher="Archiv fuer Naturgeschichte: Zeitschrift fuer systematische Zoologie" pageId="53" pageNumber="54" pagination="317 - 371" title="Anatomie, Phylogenie und Tiergeographie der Cyclophoriden." volume="9" year="1940">Tielecke 1940</bibRefCitation>
: 345;
<bibRefCitation author="Davison, GWH" journalOrPublisher="Malayan Nature Journal" pageId="50" pageNumber="51" pagination="233 - 248" title="The terrestrial molluscan fauna of Temengor Forest Reserve, Hulu Perak, Malaysia." volume="48" year="1995">Davison 1995</bibRefCitation>
: 236;
<bibRefCitation author="Schilthuizen, M" journalOrPublisher="Malayan Nature Journal" pageId="53" pageNumber="54" pagination="351 - 354" title="A note on the ecology of West-Malaysian calcicolous snails (Mollusca: Gastropoda: Diplommatinidae, Cyclophoridae, Vertiginidae, Streptaxidae)." volume="53" year="1999">Schilthuizen et al. 1999</bibRefCitation>
: 353;
<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="49" pageNumber="50" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="https://doi.org/10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
: 2760;
<bibRefCitation author="Foon, JK" journalOrPublisher="ZooKeys" pageId="50" pageNumber="51" pagination="1 - 94" title="Diversity and biogeography of land snails (Mollusca, Gastropoda) in the limestone hills of Perak, Peninsular Malaysia." url="https://doi.org/10.3897/zookeys.682.12999" volume="682" year="2017">Foon et al. 2017</bibRefCitation>
: 13, figure 4F.
</paragraph>
<paragraph pageId="35" pageNumber="36">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Orthalycaeus) perakensis" order="Architaenioglossa" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="perakensis" subGenus="Orthalycaeus">Alycaeus (Orthalycaeus) perakensis</taxonomicName>
:
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="330 - 348" title="On the land and freshwater shells of Perak." url="https://doi.org/10.1111/j.1096-3642.1891.tb01757.x" volume="1891" year="1891">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
1891
</bibRefCitation>
: 342; Kobelt and
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
<bibRefCitation author="Kobelt, W" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="51" pageNumber="52" pagination="73 - 192" title="Catalog der gegenwaertig lebend bekannten Pneumonopomen." url="https://doi.org/10.5962/bhl.title.14877" volume="1" year="1897">1897</bibRefCitation>
: 150.
</paragraph>
<paragraph pageId="35" pageNumber="36">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Alycaeus) perakensis" order="Architaenioglossa" pageId="35" pageNumber="36" phylum="Mollusca" rank="species" species="perakensis" subGenus="Alycaeus">Alycaeus (Alycaeus) perakensis</taxonomicName>
:
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="51" pageNumber="52" title="Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen. In: Kobelt W (Ed.) In Verbindung mit der deutschen Zoologischen Gesellschaft herausgegeben von der Koeniglich Preussischen Akademie der Wissenschaften zu Berlin. Mollusca: Cyclophoridae. R." url="https://doi.org/10.5962/bhl.title.1227" year="1902">Kobelt 1902</bibRefCitation>
: 348;
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="51" pageNumber="52" pagination="25 - 37" title="A list of the Land and Fresh-water Mollusca of the Malay Peninsula with notes." url="http://www.jstor.org/stable/41559818" volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 35;
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="4 - 5" title="A brief collecting trip to Perak, West Malaysia: Part One." volume="12" year="1998 a">Chan 1998a</bibRefCitation>
: 4;
<bibRefCitation author="Chan, SY" journalOrPublisher="The Papustyla" pageId="49" pageNumber="50" pagination="1 - 2" title="A brief collecting trip to Perak, West Malaysia: Part Two." volume="12" year="1998 b">Chan 1998b</bibRefCitation>
: 1.
</paragraph>
<paragraph pageId="35" pageNumber="36">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="35" pageNumber="36" phylum="Mollusca" rank="subSpecies" species="perakensis" subSpecies="perakensis">Alycaeus perakensis perakensis</taxonomicName>
:
<bibRefCitation pageId="35" pageNumber="36">Maassen 2001</bibRefCitation>
: 23;
<bibRefCitation author="Tarruella, A" journalOrPublisher="Spira" pageId="53" pageNumber="54" pagination="71 - 76" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J. E. Gray, 1850." url="http://www.molluscat.com/assets/spira_4_12_7.pdf" volume="4" year="2011">
Tarruella and
<normalizedToken originalValue="Domènech">Domenech</normalizedToken>
2011
</bibRefCitation>
: 72.
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="type locality">
<paragraph pageId="35" pageNumber="36">Type locality.</paragraph>
<paragraph pageId="35" pageNumber="36">
Buket Pondong (= PRK 55 Gunung Pondok), Perak (
<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.7825">4°46'57&quot;N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="100.83361">100°50'01&quot;E</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="type material">
<paragraph pageId="35" pageNumber="36">Type material.</paragraph>
<paragraph pageId="35" pageNumber="36">Syntypes. Buket Pondong (= PRK 55 Gunung Pondok), Perak: MNHN IM-2000-31793/1 (Seen), MNHN IM-2000-31794/1 (Seen), MNHN IM-2000-31795/1 (Seen).</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="36" type="materials_examined">
<paragraph pageId="35" pageNumber="36">Other materials.</paragraph>
<paragraph pageId="35" pageNumber="36">
KDH 01 Bukit Baling, Kedah: BOR/MOL 6525/1, BOR/MOL 6854/5, BOR/MOL 8527/1, BOR/MOL 8528/2. KDH 02 Gunung Pulai, Kedah: BOR/MOL 6858/2. PRK 55 Gunung Pondok, Perak: BOR/MOL 272/1, BOR/MOL 6849/1, BOR/MOL 8349/29, BOR/MOL 8350/23, BOR/MOL 11479/46, BOR/MOL 11506/31, BOR/MOL 11539/50, BOR/MOL 11563/20, ZRC 1997.25/5, ZRC 1975.2.22.175-182/12, ZRC 1975.2.22.36-66/31, ANSP 423056/2. PRK 61 Gua Badak, Perak: ZRC 1975.2.22.160-174/15. PRK 62 Gunung Dayak, Perak: BOR/MOL 277/5, BOR/MOL 6201/3, ZRC 1975.2.22.127-159/33. PRK 64 Bukit Kepala Gajah, Perak: BOR/MOL 6852/1, BOR/MOL 8354/33, BOR/MOL 8355/5, BOR/MOL 10088/31, BOR/MOL 10124/15, BOR/MOL 10136/18, BOR/MOL 10166/21, BOR/MOL 13001/2. Mykarst-171 Tasik Temenggor Hill S2, Perak: BOR/MOL 6243/1. Hill 40 m east of Sungai Siput Hospital, Perak: BOR/MOL 271/1, BOR/MOL 6877/1, BOR/MOL 8385/24, BOR/MOL 8386/1. Limestone outcrop 50m from roadside, Kampung Pahit, Klian Intan, Perak: BOR/MOL 6874/1, BOR/MOL 8379/22, BOR/MOL 8380/9. 'Bat
<normalizedToken originalValue="Cave">Cave'</normalizedToken>
Hill, roadside towards Perlop 1 and 2 oil palm plantations, Perak: BOR/MOL 6878/1, BOR/MOL 8387/17, BOR/MOL 8388/5, BOR/MOL 9770/11, BOR/MOL 9799/3, BOR/MOL 9833/1, BOR/MOL 9877/4. Batu Kebelah, Kampung Kenang Baru, Jalong Tinggi, Perak: BOR/MOL 6879/1, BOR/MOL 8389/5, BOR/MOL 8390/5, BOR/MOL 9546/23, BOR/MOL 9565/2, BOR/MOL 9573/5, BOR/MOL 9585/17. Lenggong, Perak: ZRC 1997.24/4 specimens. Perak: RMNH 153573/3, RMNH 153574/2. Banang Pupo, Yala, Thailand: ANSP 446394/3.
</paragraph>
</subSubSection>
<subSubSection lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph pageId="35" pageNumber="36">Description.</paragraph>
<paragraph pageId="35" pageNumber="36">Protoconch. Smooth.</paragraph>
<paragraph pageId="35" pageNumber="36">Shell shape. Conical. Shell height: 5.98-7.27 mm. Shell width: 6.48-8.62 mm.</paragraph>
<paragraph pageId="35" pageNumber="36">
Spire. Spire height: 1.83-2.64 mm. Spire width: 2.80-3.39 mm. Number of whorls: up to 4
<normalizedToken originalValue="½">1/2</normalizedToken>
. Spire shape: oblong conical. Whorl periphery rounded. Umbilicus open.
</paragraph>
<paragraph pageId="35" pageNumber="36">
Whorl constriction. At about 4
<normalizedToken originalValue="¼">1/4</normalizedToken>
whorls posterior of protoconch.
</paragraph>
<paragraph pageId="35" pageNumber="36">Breathing tube. Length: 0.61-0.88 mm.</paragraph>
<paragraph pageId="35" pageNumber="36">Aperture and peristome. Aperture circular, expanded. Aperture height: 3.06-4.32 mm. Aperture width: 2.86-4.10 mm. Peristome double, not thickened, notched at suture. Peristome orientation 43-49° oblique with respect to the coiling axis.</paragraph>
<paragraph pageId="35" pageNumber="36">Spiral lines. Distinct. Regularly spaced. Approximately 17-35 lines per 1 mm.</paragraph>
<paragraph pageId="36" pageNumber="37">
<pageBreakToken pageId="36" pageNumber="37" start="start">Radial</pageBreakToken>
ribs running anterior of breathing tube. Radial ribs pronounced, evenly spaced. Approximately 8-14 ribs per 1 mm.
</paragraph>
<paragraph pageId="36" pageNumber="37">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced, white and thicker than those anterior of breathing tube, evenly spaced. Approximately 9-15 ribs per 1 mm.</paragraph>
<paragraph pageId="36" pageNumber="37">Radial ribs running posterior of breathing tube. Radial ribs pronounced, widely and regularly spaced, approximately 9-16 ribs per 1 mm. Prior to aperture, radial ribs become absent.</paragraph>
<paragraph pageId="36" pageNumber="37">Operculum. Concave, rounded. Exterior covered with thin calcareous layer, surface finely granulated with a scattered arrangement of calcareous spikes. Interior covered with proteinaceous coating, smooth.</paragraph>
<paragraph pageId="36" pageNumber="37">Shell colour. Whorls yellow. Peristome white.</paragraph>
<paragraph pageId="36" pageNumber="37">Living animal. Body cream-white. Head pink. Tentacles brown.</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="habitat">
<paragraph pageId="36" pageNumber="37">Habitat and ecology.</paragraph>
<paragraph pageId="36" pageNumber="37">Lives on both dry and wet limestone walls. Some individuals have been spotted on tree trunks and fallen logs beside limestone cliffs. In shady forests on limestone hills.</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="distribution">
<paragraph pageId="36" pageNumber="37">Distribution range and habitat.</paragraph>
<paragraph pageId="36" pageNumber="37">
Restricted to central and northern Perak as well as northeastern Kedah (Bukit Baling and Gunung Pulai). Elsewhere, in Yala, Thailand (
<bibRefCitation author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="53" pageNumber="54" pagination="194 - 199" title="On the land operculate Mollusca collected during the &quot; Skeat Expedition &quot; to the Malay Peninsula in 1899 - 1900." url="http://www.biodiversitylibrary.org/part/145330/summary" volume="1903" year="1903">Sykes 1903</bibRefCitation>
,
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="51" pageNumber="52" pagination="25 - 37" title="A list of the Land and Fresh-water Mollusca of the Malay Peninsula with notes." url="http://www.jstor.org/stable/41559818" volume="6" year="1928">Laidlaw 1928</bibRefCitation>
, ANSP 446394).
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="differential diagnosis">
<paragraph pageId="36" pageNumber="37">Differential diagnosis.</paragraph>
<paragraph pageId="36" pageNumber="37">
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
is most similar to
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus roebeleni" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="roebeleni">Alycaeus roebeleni</taxonomicName>
in conical shell shape but differs in being smaller (smaller by about 0.97 mm in shell height, 1.96 mm in shell width), aperture margins more circular and has a less expanded ultimate whorl.
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
is distinguished from
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus kapayanensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="kapayanensis">Alycaeus kapayanensis</taxonomicName>
and
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus kurauensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="kurauensis">Alycaeus kurauensis</taxonomicName>
sp. n. by its much larger shell (larger by about 1.43 mm in shell height, 2.32 mm in shell width), more expanded penultimate whorl, operculum exterior with spikes and more oblique peristome. Shells of
<taxonomicName lsidName="A. perakensis" pageId="36" pageNumber="37" rank="species" species="perakensis">A. perakensis</taxonomicName>
vary in the degree of expansion of the ultimate whorl.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="37" type="discussion">
<paragraph pageId="36" pageNumber="37">Discussion.</paragraph>
<paragraph pageId="36" pageNumber="37">
Historically, the lack of access to type specimens and poor original diagnosis have led to the misidentification of
<taxonomicName lsidName="A. selangoriensis" pageId="36" pageNumber="37" rank="species" species="selangoriensis">A. selangoriensis</taxonomicName>
sp. n.,
<taxonomicName lsidName="A. clementsi" pageId="36" pageNumber="37" rank="species" species="clementsi">A. clementsi</taxonomicName>
sp. n. and
<taxonomicName lsidName="A. kapayanensis" pageId="36" pageNumber="37" rank="species" species="kapayanensis">A. kapayanensis</taxonomicName>
as
<taxonomicName lsidName="A. perakensis" pageId="36" pageNumber="37" rank="species" species="perakensis">A. perakensis</taxonomicName>
(
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="The Bulletin of the Raffles Museum" pageId="51" pageNumber="52" pagination="35 - 41" title="Notes on the land-mollusca of the Batu Caves, Selangor, with descriptions of two new species." url="http://lkcnhm.nus.edu.sg/nus/pdf/PUBLICATION/Raffles%20Bulletin%20of%20Zoology/Past%20Volumes/RBZ%207(1)/07brm035-041.pdf" volume="7" year="1932">Laidlaw 1932</bibRefCitation>
, van
<bibRefCitation author="Benthem Jutting, van WSS" journalOrPublisher="Basteria" pageId="49" pageNumber="50" pagination="10 - 20" title="Some notes on land and freshwater mollusca of Malaya." url="http://natuurtijdschriften.nl/record/596402" volume="24" year="1960 a">Bethem Jutting 1960a</bibRefCitation>
,
<bibRefCitation author="Clements, R" journalOrPublisher="Biological Conservation" pageId="49" pageNumber="50" pagination="2751 - 2764" title="Using biogeographical patterns of endemic land snails to improve conservation planning for limestone karsts." url="https://doi.org/10.1016/j.biocon.2008.08.011" volume="141" year="2008">Clements et al. 2008</bibRefCitation>
).
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
shells are consistent in shell size and spire height but varies in the degree of expansion of the ultimate whorl with no particular geographic pattern.
<bibRefCitation author="Tielecke, H" journalOrPublisher="Archiv fuer Naturgeschichte: Zeitschrift fuer systematische Zoologie" pageId="53" pageNumber="54" pagination="317 - 371" title="Anatomie, Phylogenie und Tiergeographie der Cyclophoriden." volume="9" year="1940">Tielecke (1940)</bibRefCitation>
used
<taxonomicName lsidName="A. perakensis" pageId="36" pageNumber="37" rank="species" species="perakensis">A. perakensis</taxonomicName>
as a representative species for
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="genus">Alycaeus</taxonomicName>
in his comparison of cyclophorid reproductive anatomy.
<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="52" pageNumber="53" pagination="146 - 156" title="On a collection of land-shells from the Samui Islands, Gulf of Siam." url="http://www.biodiversitylibrary.org/item/97157215" volume="1894" year="1894">
<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
(1894)
</bibRefCitation>
considered
<taxonomicName lsidName="A. perakensis" pageId="36" pageNumber="37" rank="species" species="perakensis">A. perakensis</taxonomicName>
to be closely related to
<taxonomicName lsidName="A. roebeleni" pageId="36" pageNumber="37" rank="species" species="roebeleni">A. roebeleni</taxonomicName>
.
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
shells were also collected from Biserat and Banang Pupo, Yala, southern Thailand (
<bibRefCitation author="Sykes, ER" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="53" pageNumber="54" pagination="194 - 199" title="On the land operculate Mollusca collected during the &quot; Skeat Expedition &quot; to the Malay Peninsula in 1899 - 1900." url="http://www.biodiversitylibrary.org/part/145330/summary" volume="1903" year="1903">Sykes 1903</bibRefCitation>
,
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malayan Branch of the Royal Asiatic Society" pageId="51" pageNumber="52" pagination="25 - 37" title="A list of the Land and Fresh-water Mollusca of the Malay Peninsula with notes." url="http://www.jstor.org/stable/41559818" volume="6" year="1928">Laidlaw 1928</bibRefCitation>
, ANSP 446394).
</paragraph>
<caption pageId="36" pageNumber="37">
<paragraph pageId="36" pageNumber="37">
Figure 24.
<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="36" pageNumber="37" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
Crosse, 1879a.
<normalizedToken originalValue="AE">A-E</normalizedToken>
Shell of syntype, MNHN-IM-2000-31793
<normalizedToken originalValue="FJ">F-J</normalizedToken>
Shell of BOR/MOL 8350
<normalizedToken originalValue="KM">K-M</normalizedToken>
Close up of shell of BOR/MOL 8350
<normalizedToken originalValue="NQ">N-Q</normalizedToken>
Operculum of BOR/MOL 8349
<normalizedToken originalValue="RV">R-V</normalizedToken>
Shell of BOR/MOL 8355. Scale bars:
<normalizedToken originalValue="AJ">A-J</normalizedToken>
, K, L,
<normalizedToken originalValue="RV">R-V</normalizedToken>
1 mm; M,
<normalizedToken originalValue="NQ">N-Q</normalizedToken>
0.5 mm. Photographs
<normalizedToken originalValue="AE">A-E</normalizedToken>
by Manuel Caballer (reproduced herein under CC-BY 4.0 license, Project RECOLNAT, MNHN). Photographs
<normalizedToken originalValue="FV">F-V</normalizedToken>
by Junn Kitt Foon.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>