treatments-xml/data/25/50/87/255087E5FFEE49221DE9FCC1FA1B2087.xml
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<document id="5F211AF95D34D44C32E86A8E88359779" ID-CLB-Dataset="8165" ID-DOI="10.11646/zootaxa.4926.4.3" ID-GBIF-Dataset="07abd153-3d09-4820-82dd-6991f1d14ced" ID-ISSN="1175-5326" ID-Zenodo-Dep="4529625" ID-ZooBank="C92BA48D-4493-4388-A24D-545B7FD1C19E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612956496051" checkinUser="plazi" docAuthor="Tian, Mingyi, Huang, Xinglong &amp; Li, Chenliang" docDate="2021" docId="255087E5FFEE49221DE9FCC1FA1B2087" docLanguage="en" docName="zootaxa.4926.4.3.pdf" docOrigin="Zootaxa 4926 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Guizhaphaenopsodes solidior Tian &amp; X. Huang 2021, n. sp." docType="treatment" docVersion="7" lastPageNumber="530" masterDocId="D969FF9DFFE8492B1D7EFA24FF822454" masterDocTitle="Contribution to the knowledge of subterranean ground beetles from eastern Wuling Mountains, China (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="534" masterPageNumber="521" pageNumber="527" updateTime="1698928636667" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="CE3E164C88C1FD8D46071DA8B65A9AFC">Contribution to the knowledge of subterranean ground beetles from eastern Wuling Mountains, China (Coleoptera: Carabidae: Trechinae)</mods:title>
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<mods:namePart id="67170E809ADF4D5C42B109AF5F5A4EBC">Tian, Mingyi</mods:namePart>
<mods:affiliation id="C72FD34340EAE835EF9C9B4AA57A0E49">Department of Entomology, College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou, 510642, China</mods:affiliation>
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<mods:namePart id="DFC941D0E7BBAC7A9970846D6EB38C60">Huang, Xinglong</mods:namePart>
<mods:affiliation id="B6A1C4EF840126945D9643E2F1142F84">College of Biology and Environmental Sciences, Jishou University, Jishou 416000, China</mods:affiliation>
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<mods:namePart id="D2C977D3A659D3EBAA92E40D353B3C35">Li, Chenliang</mods:namePart>
<mods:affiliation id="5C3D5C9569642B37E2427D48492EFFEC">Hubei Board Nature, Central China Normal University, 430079, Wuhan, China</mods:affiliation>
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<treatment id="255087E5FFEE49221DE9FCC1FA1B2087" ID-DOI="http://doi.org/10.5281/zenodo.4546191" ID-GBIF-Taxon="177422225" ID-Zenodo-Dep="4546191" LSID="urn:lsid:plazi:treatment:255087E5FFEE49221DE9FCC1FA1B2087" httpUri="http://treatment.plazi.org/id/255087E5FFEE49221DE9FCC1FA1B2087" lastPageId="9" lastPageNumber="530" pageId="6" pageNumber="527">
<subSubSection id="E5E36578FFEE492D1DE9FCC1FCAC2354" box="[151,814,1765,1792]" pageId="6" pageNumber="527" type="nomenclature">
<paragraph id="AD4636F3FFEE492D1DE9FCC1FCAC2354" blockId="6.[151,814,1765,1829]" box="[151,814,1765,1792]" pageId="6" pageNumber="527">
<heading id="F60E819FFFEE492D1DE9FCC1FCAC2354" bold="true" box="[151,814,1765,1792]" fontSize="11" level="1" pageId="6" pageNumber="527" reason="1">
<emphasis id="9F8DEAE1FFEE492D1DE9FCC1FCAC2354" bold="true" box="[151,814,1765,1792]" pageId="6" pageNumber="527">
<taxonomicName id="6AF94D70FFEE492D1DE9FCC1FD5D2354" authority="Tian &amp; X. Huang" authorityName="Tian &amp; X. Huang" authorityYear="2021" box="[151,735,1765,1792]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="solidior" status="sp. nov.">
<emphasis id="9F8DEAE1FFEE492D1DE9FCC1FE792354" bold="true" box="[151,507,1765,1792]" italics="true" pageId="6" pageNumber="527">Guizhaphaenopsodes solidior</emphasis>
Tian &amp; X. Huang
</taxonomicName>
,
<taxonomicNameLabel id="84BE579AFFEE492D1F95FCC2FCAC2354" box="[747,814,1766,1792]" pageId="6" pageNumber="527" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E5E36578FFEE492D1DE9FD2EFEAC2371" box="[151,302,1802,1829]" pageId="6" pageNumber="527" type="description">
<paragraph id="AD4636F3FFEE492D1DE9FD2EFEAC2371" blockId="6.[151,814,1765,1829]" box="[151,302,1802,1829]" pageId="6" pageNumber="527">
(
<figureCitation id="35C22A76FFEE492D1DE1FD2EFF6F2370" box="[159,237,1802,1828]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1367]" captionTargetBox="[210,1378,181,1320]" captionTargetId="figure-23@2.[210,1378,181,1320]" captionTargetPageId="2" captionText="FIGURE 1. A distribution map showing the locality caves of the three new species described in the present paper (Tangle Dong: Xiangxius jiangi Tian &amp; X. Huang, n. gen., n. sp. and Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp.; Qizimei Dong: Sinotroglodytes hefengensis Tian &amp; Li, n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4529629" httpUri="https://zenodo.org/record/4529629/files/figure.png" pageId="6" pageNumber="527">Figs. 1</figureCitation>
,
<figureCitation id="35C22A76FFEE492D1D85FD2EFEA42371" box="[251,294,1802,1829]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="7.[151,250,1903,1927]" captionStartId-1="8.[151,250,2010,2034]" captionStartId-2="9.[151,250,1031,1055]" captionTargetBox-0="[202,1394,381,1857]" captionTargetBox-1="[169,1416,892,1974]" captionTargetBox-2="[151,1428,345,1008]" captionTargetId-0="figure-92@7.[151,1435,344,1879]" captionTargetId-1="figure-359@8.[164,1424,885,1987]" captionTargetId-2="figure-112@9.[151,1436,345,1008]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURE 5. Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp., habitus" captionText-1="FIGURE 6. Ventral head (A) and elytral chaetotaxy (B) of Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp." captionText-2="FIGURE 7. Male genitalia of Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp. (A. median lobe and parameres, lateral view; B. apical lobe of median lobe, dorsal view)" figureDoi-0="http://doi.org/10.5281/zenodo.4529637" figureDoi-1="http://doi.org/10.5281/zenodo.4529639" figureDoi-2="http://doi.org/10.5281/zenodo.4529641" httpUri-0="https://zenodo.org/record/4529637/files/figure.png" httpUri-1="https://zenodo.org/record/4529639/files/figure.png" httpUri-2="https://zenodo.org/record/4529641/files/figure.png" pageId="6" pageNumber="527">57</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E5E36578FFEE492D1DE9FD76FA0023C4" pageId="6" pageNumber="527" type="materials_examined">
<paragraph id="AD4636F3FFEE492D1DE9FD76FA0023C4" blockId="6.[151,1436,1873,2009]" pageId="6" pageNumber="527">
<emphasis id="9F8DEAE1FFEE492D1DE9FD76FEC6233F" bold="true" box="[151,324,1873,1900]" pageId="6" pageNumber="527">Type material.</emphasis>
<materialsCitation id="1D913CAEFFEE492D1C37FD76FA0023C4" ID-GBIF-Occurrence="3037449305" collectingDate="2018-01-15" collectionCode="SCAU" collectorName="Xinglong Huang" elevation="270" latitude="28.305832" location="Jishou" longLatPrecision="20" longitude="109.65417" municipality="cave Tangle Dong" pageId="6" pageNumber="527" specimenCount="1" stateProvince="Hunan" typeStatus="holotype">
<typeStatus id="72428851FFEE492D1C37FD76FE332338" box="[329,433,1874,1900]" pageId="6" pageNumber="527" type="holotype">Holotype</typeStatus>
male,
<collectingMunicipality id="4D22AC89FFEE492D1C84FD76FD472338" box="[506,709,1874,1900]" pageId="6" pageNumber="527">cave Tangle Dong</collectingMunicipality>
, suburb of
<location id="A8266028FFEE492D1E40FD76FC0A2338" LSID="urn:lsid:plazi:treatment:255087E5FFEE49221DE9FCC1FA1B2087:A8266028FFEE492D1E40FD76FC0A2338" box="[830,904,1874,1900]" latitude="28.305832" longLatPrecision="20" longitude="109.65417" municipality="cave Tangle Dong" name="Jishou" pageId="6" pageNumber="527" stateProvince="Hunan">Jishou</location>
,
<collectorName id="000C5325FFEE492D1EECFD76FBA92338" box="[914,1067,1874,1900]" pageId="6" pageNumber="527">Xiangxi Tujia</collectorName>
&amp;
<collectorName id="000C5325FFEE492D1933FD76FA1A2338" box="[1101,1432,1874,1900]" pageId="6" pageNumber="527">Miao Autonomous Prefecture</collectorName>
, west
<collectingRegion id="6F3DF811FFEE492D1DACFD52FE0B23C4" box="[210,393,1910,1936]" country="China" name="Hunan" pageId="6" pageNumber="527">Hunan Province</collectingRegion>
,
<geoCoordinate id="C8CD5034FFEE492D1CEBFD52FD9C23C4" box="[405,542,1910,1936]" degrees="28" direction="north" minutes="18" orientation="latitude" pageId="6" pageNumber="527" precision="15" seconds="21" value="28.305832">28°1821”N</geoCoordinate>
/
<geoCoordinate id="C8CD5034FFEE492D1F51FD52FD4123C5" box="[559,707,1910,1937]" degrees="109" direction="east" minutes="39" orientation="longitude" pageId="6" pageNumber="527" precision="15" seconds="15" value="109.65417">109°3915”E</geoCoordinate>
,
<quantity id="6A019B16FFEE492D1FB1FD52FC9423C4" box="[719,790,1910,1936]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="527" unit="m" value="270.0">
<elevation id="26D4D1C0FFEE492D1FB1FD52FC9423C4" box="[719,790,1910,1936]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="6" pageNumber="527" unit="m" value="270.0">270 m</elevation>
</quantity>
in altitude,
<date id="D9471033FFEE492D1EE5FD52FB8F23C5" box="[923,1037,1910,1937]" pageId="6" pageNumber="527" value="2018-01-15">
<collectingDate id="C903E9DBFFEE492D1EE5FD52FB8F23C5" box="[923,1037,1910,1937]" pageId="6" pageNumber="527" value="2018-01-15">2018-I-15</collectingDate>
</date>
, leg.
<collectorName id="000C5325FFEE492D1937FD52FA8523C4" box="[1097,1287,1910,1936]" pageId="6" pageNumber="527">Xinglong Huang</collectorName>
, in
<collectionCode id="CBE8AE36FFEE492D184EFD52FAFC23C4" box="[1328,1406,1910,1936]" pageId="6" pageNumber="527">SCAU</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="E5E36578FFEE492D1DB9FDBDFA81238C" pageId="6" pageNumber="527" type="diagnosis">
<paragraph id="AD4636F3FFEE492D1DB9FDBDFA81238C" blockId="6.[151,1436,1873,2009]" pageId="6" pageNumber="527">
<emphasis id="9F8DEAE1FFEE492D1DB9FDBDFEC023E7" bold="true" box="[199,322,1945,1971]" pageId="6" pageNumber="527">Diagnosis.</emphasis>
A medium-sized semi-aphaenopsian species, somewhat looking like a
<taxonomicName id="6AF94D70FFEE492D1923FDBEFA9B23E0" authorityName="Vigna Taglianti" authorityYear="1997" box="[1117,1305,1946,1972]" class="Insecta" family="Carabidae" genus="Guizhaphaenops" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="9F8DEAE1FFEE492D1923FDBEFA9B23E0" box="[1117,1305,1946,1972]" italics="true" pageId="6" pageNumber="527">Guizhaphaenops</emphasis>
</taxonomicName>
species but the body more robust, partly pigmented, with strong antennae extending to about apical 1/5 of elytra.
</paragraph>
</subSubSection>
<subSubSection id="E5E36578FFEF49221DB9FAB2FE1D20DF" lastPageId="9" lastPageNumber="530" pageId="7" pageNumber="528" type="description">
<paragraph id="AD4636F3FFEF492C1DB9FAB2FC3724E5" blockId="7.[151,1437,150,321]" box="[199,949,150,177]" pageId="7" pageNumber="528">
<emphasis id="9F8DEAE1FFEF492C1DB9FAB2FEDB24E4" bold="true" box="[199,345,150,176]" pageId="7" pageNumber="528">Description.</emphasis>
Length:
<quantity id="6A019B16FFEF492C1CC0FAB2FD9424E5" box="[446,534,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="528" unit="mm" value="6.5">6.5 mm</quantity>
, width:
<quantity id="6A019B16FFEF492C1F0EFAB2FD4B24E5" box="[624,713,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="528" unit="mm" value="2.2">2.2 mm</quantity>
. Habitus as in
<figureCitation id="35C22A76FFEF492C1E13FAB2FC3224E5" box="[877,944,150,177]" captionStart="FIGURE 5" captionStartId="7.[151,250,1903,1927]" captionTargetBox="[202,1394,381,1857]" captionTargetId="figure-92@7.[151,1435,344,1879]" captionTargetPageId="7" captionText="FIGURE 5. Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp., habitus" figureDoi="http://doi.org/10.5281/zenodo.4529637" httpUri="https://zenodo.org/record/4529637/files/figure.png" pageId="7" pageNumber="528">Fig. 5</figureCitation>
.
</paragraph>
<paragraph id="AD4636F3FFEF492C1DB9FA9EFC7E2514" blockId="7.[151,1437,150,321]" pageId="7" pageNumber="528">
Head, pronotum, 1
<superScript id="5A8C9BBBFFEF492C1CE2FA9EFE342480" attach="none" box="[412,438,186,212]" fontSize="6" pageId="7" pageNumber="528">st</superScript>
8
<superScript id="5A8C9BBBFFEF492C1CBCFA9EFE4D249C" attach="left" box="[450,463,186,200]" fontSize="6" pageId="7" pageNumber="528">th</superScript>
antennomeres, femora and tibiae, undersides of head and thoraxes dark brown, elytra reddish-brown, palps, 9
<superScript id="5A8C9BBBFFEF492C1CDFFAFAFE3C24AC" attach="none" box="[417,446,222,248]" fontSize="6" pageId="7" pageNumber="528">th</superScript>
11
<superScript id="5A8C9BBBFFEF492C1CA9FAFAFE6624B8" attach="left" box="[471,484,222,236]" fontSize="6" pageId="7" pageNumber="528">th</superScript>
antennomeres and tarsi yellow. Body smooth and glabrous, but elytra with a minute and sparse pubescence. Microsculpture: isodiametric meshes on vertex, striate on frons, pronotum and elytra. Fore body (head plus pronotum including mandibles) distinctly shorter than elytra.
</paragraph>
<caption id="F986667BFFEF492C1DE9FD4BFC6F23D3" ID-DOI="http://doi.org/10.5281/zenodo.4529637" ID-Zenodo-Dep="4529637" box="[151,1005,1903,1928]" httpUri="https://zenodo.org/record/4529637/files/figure.png" pageId="7" pageNumber="528" startId="7.[151,250,1903,1927]" targetBox="[202,1394,381,1857]" targetPageId="7">
<paragraph id="AD4636F3FFEF492C1DE9FD4BFC6F23D3" blockId="7.[151,1005,1903,1928]" box="[151,1005,1903,1928]" pageId="7" pageNumber="528">
<emphasis id="9F8DEAE1FFEF492C1DE9FD4BFE9123D3" bold="true" box="[151,275,1903,1927]" pageId="7" pageNumber="528">FIGURE 5.</emphasis>
<taxonomicName id="6AF94D70FFEF492C1C67FD4BFC8223DC" authority="Tian &amp; X. Huang" authorityName="Tian &amp; X. Huang" authorityYear="2021" box="[281,768,1903,1928]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="solidior" status="n. gen., n. sp.">
<emphasis id="9F8DEAE1FFEF492C1C67FD4BFDC523D3" box="[281,583,1903,1927]" italics="true" pageId="7" pageNumber="528">Guizhaphaenopsodes solidior</emphasis>
Tian &amp; X. Huang
</taxonomicName>
,
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<emphasis id="9F8DEAE1FFEF492C1E75FD4BFCD023D3" bold="true" box="[779,850,1903,1927]" pageId="7" pageNumber="528">n. gen.</emphasis>
,
<emphasis id="9F8DEAE1FFEF492C1E20FD4BFC1A23D3" bold="true" box="[862,920,1903,1927]" pageId="7" pageNumber="528">n. sp.</emphasis>
</taxonomicNameLabel>
, habitus
</paragraph>
</caption>
<paragraph id="AD4636F3FFEF49231DB9FD91FE6125F9" blockId="7.[151,1437,1973,2036]" lastBlockId="8.[151,1437,151,861]" lastPageId="8" lastPageNumber="529" pageId="7" pageNumber="528">
Head (
<figureCitation id="35C22A76FFEF492C1C6AFD91FEE72384" box="[276,357,1973,2000]" captionStart="FIGURE 5" captionStartId="7.[151,250,1903,1927]" captionTargetBox="[202,1394,381,1857]" captionTargetId="figure-92@7.[151,1435,344,1879]" captionTargetPageId="7" captionText="FIGURE 5. Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp., habitus" figureDoi="http://doi.org/10.5281/zenodo.4529637" httpUri="https://zenodo.org/record/4529637/files/figure.png" pageId="7" pageNumber="528">Figs. 5</figureCitation>
,
<figureCitation id="35C22A76FFEF492C1C0AFD91FE152384" box="[372,407,1973,2000]" captionStart="FIGURE 6" captionStartId="8.[151,250,2010,2034]" captionTargetBox="[169,1416,892,1974]" captionTargetId="figure-359@8.[164,1424,885,1987]" captionTargetPageId="8" captionText="FIGURE 6. Ventral head (A) and elytral chaetotaxy (B) of Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.4529639" httpUri="https://zenodo.org/record/4529639/files/figure.png" pageId="7" pageNumber="528">6A</figureCitation>
) moderately elongate, longer than wide, HLm/HW = 1.9, HLl/HW = 1.6; nearly parallelsided, widest at about middle of head excluding mandibles; front and vertex convex; frontal furrows well-defined and complete, strongly divergent posteriorly; the posterior supraorbital pores located at ending point of frontal furrows; clypeus quadrisetose, labrum transverse, almost straight in the front margin, 6-setose; mandibles developed and moderately curved at apices; tooth short and broad, blunt at tip, about half as long as the lateral lobes; ligula adnated with paraglossae, 12-setose; palps long and slender, the 2
<superScript id="5A8C9BBBFFE049231E09FB26FC052544" attach="left" box="[887,903,258,272]" fontSize="6" pageId="8" pageNumber="529">nd</superScript>
labial palpomere about 1.2 times as long as 3
<superScript id="5A8C9BBBFFE0492318F9FB26FA172544" attach="none" box="[1415,1429,258,272]" fontSize="6" pageId="8" pageNumber="529">rd</superScript>
; 3
<superScript id="5A8C9BBBFFE049231DDBFB02FF312560" attach="left" box="[165,179,294,308]" fontSize="6" pageId="8" pageNumber="529">rd</superScript>
maxillary palpomere 1.25 times as long as 4
<superScript id="5A8C9BBBFFE049231FD8FB02FD312560" attach="none" box="[678,691,294,308]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
; suborbital pores close to neck constriction; antennae thick though filiform, pubescent from 2
<superScript id="5A8C9BBBFFE049231CBEFB6EFE622531" attach="none" box="[448,480,330,357]" fontSize="6" pageId="8" pageNumber="529">nd</superScript>
11
<superScript id="5A8C9BBBFFE049231C87FB6EFD84250C" attach="left" box="[505,518,330,344]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
antennomeres, antennomere 3 the longest, almost twice as long as scape; relative length of each antennomere as follows: 1
<superScript id="5A8C9BBBFFE049231F1AFB4BFDED2529" attach="left" box="[612,623,367,381]" fontSize="6" pageId="8" pageNumber="529">st</superScript>
(1.0), 2
<superScript id="5A8C9BBBFFE049231FBBFB4AFD572528" attach="left" box="[709,725,366,380]" fontSize="6" pageId="8" pageNumber="529">nd</superScript>
(1.0), 3
<superScript id="5A8C9BBBFFE049231E51FB4AFCBF2528" attach="left" box="[815,829,366,380]" fontSize="6" pageId="8" pageNumber="529">rd</superScript>
(1.9), 4
<superScript id="5A8C9BBBFFE049231EEBFB4AFC202528" attach="left" box="[917,930,366,380]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1.5), 5
<superScript id="5A8C9BBBFFE049231E85FB4AFB8A2528" attach="left" box="[1019,1032,366,380]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1. 3), 6
<superScript id="5A8C9BBBFFE04923191BFB4AFBF02528" attach="left" box="[1125,1138,366,380]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1.2), 7
<superScript id="5A8C9BBBFFE0492319B9FB4AFB562528" attach="left" box="[1223,1236,366,380]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1.1), 8
<superScript id="5A8C9BBBFFE049231853FB4AFAB82528" attach="left" box="[1325,1338,366,380]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1.0), 9
<superScript id="5A8C9BBBFFE0492318EEFB4AFA1F2528" attach="none" box="[1424,1437,366,380]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1.0), 10
<superScript id="5A8C9BBBFFE049231D89FBB6FE8625F4" attach="left" box="[247,260,402,416]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(0.9) and 11
<superScript id="5A8C9BBBFFE049231CF1FBB6FE1E25F4" attach="left" box="[399,412,402,416]" fontSize="6" pageId="8" pageNumber="529">th</superScript>
(1. 2).
</paragraph>
<paragraph id="AD4636F3FFE049231DB9FB93FC4526D1" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="529">
Pronotum (
<figureCitation id="35C22A76FFE049231C39FB93FE0E2585" box="[327,396,438,465]" captionStart="FIGURE 5" captionStartId="7.[151,250,1903,1927]" captionTargetBox="[202,1394,381,1857]" captionTargetId="figure-92@7.[151,1435,344,1879]" captionTargetPageId="7" captionText="FIGURE 5. Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp., habitus" figureDoi="http://doi.org/10.5281/zenodo.4529637" httpUri="https://zenodo.org/record/4529637/files/figure.png" pageId="8" pageNumber="529">Fig. 5</figureCitation>
) wider than head, PW/HW = 1.3, as long as head excluding mandibles, much longer than wide, PL/PW = 1.1; lateral margins gently expanded, narrowly and evenly bordered throughout, strongly reflexed at base, widest at about middle; base and front unbordered, almost straight, the former slightly wider than the latter; both front and hind angles obtuse, though more or less rectangular; anterior latero-marginal pores located apical 2/11 of pronotum; frontal transverse impression faint, whereas basal one well-defined, basal foveae large but shallow; middle line well-marked; disc moderately convex. Scutellum small.
</paragraph>
<paragraph id="AD4636F3FFE049231DB9F8ABFC372741" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="529">
Elytra (
<figureCitation id="35C22A76FFE049231C62F8ABFEE326FD" box="[284,353,654,681]" captionStart="FIGURE 5" captionStartId="7.[151,250,1903,1927]" captionTargetBox="[202,1394,381,1857]" captionTargetId="figure-92@7.[151,1435,344,1879]" captionTargetPageId="7" captionText="FIGURE 5. Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp., habitus" figureDoi="http://doi.org/10.5281/zenodo.4529637" httpUri="https://zenodo.org/record/4529637/files/figure.png" pageId="8" pageNumber="529">Fig. 5</figureCitation>
,
<figureCitation id="35C22A76FFE049231C13F8ABFE0D26FE" box="[365,399,655,682]" captionStart="FIGURE 6" captionStartId="8.[151,250,2010,2034]" captionTargetBox="[169,1416,892,1974]" captionTargetId="figure-359@8.[164,1424,885,1987]" captionTargetPageId="8" captionText="FIGURE 6. Ventral head (A) and elytral chaetotaxy (B) of Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.4529639" httpUri="https://zenodo.org/record/4529639/files/figure.png" pageId="8" pageNumber="529">6B</figureCitation>
) much longer than wide, EL/EW = 1.6, wider than pronotum, EW/PW = 1.9; unbordered at base; striae shallowly depressed, intervals slightly convex; basal pores distant from scutellum, anterior and posterior dorsal pores on the 3
<superScript id="5A8C9BBBFFE049231C02F8F2FE0826B0" attach="left" box="[380,394,726,740]" fontSize="6" pageId="8" pageNumber="529">rd</superScript>
striae at about 1/4 and 2/3 of elytra from base respectively; each group of marginal umbilicate series well separated from the other, apical group composed of 4 pores.
</paragraph>
<paragraph id="AD4636F3FFE049231DB9F93BFE0E2709" blockId="8.[151,1437,151,861]" pageId="8" pageNumber="529">Legs moderately long and densely pubescent; fore and middle tibiae longitudinally grooved externally, whereas smooth in hind tibiae.</paragraph>
<caption id="F986667BFFE049231DE9FDFEFAF223A7" ID-DOI="http://doi.org/10.5281/zenodo.4529639" ID-Zenodo-Dep="4529639" box="[151,1392,2010,2035]" httpUri="https://zenodo.org/record/4529639/files/figure.png" pageId="8" pageNumber="529" startId="8.[151,250,2010,2034]" targetBox="[169,1416,892,1974]" targetPageId="8">
<paragraph id="AD4636F3FFE049231DE9FDFEFAF223A7" blockId="8.[151,1392,2010,2035]" box="[151,1392,2010,2035]" pageId="8" pageNumber="529">
<emphasis id="9F8DEAE1FFE049231DE9FDFEFE9123A6" bold="true" box="[151,275,2010,2034]" pageId="8" pageNumber="529">FIGURE 6.</emphasis>
Ventral head (A) and elytral chaetotaxy (B) of
<taxonomicName id="6AF94D70FFE049231F8CFDFEFB5B23A7" authority="Tian &amp; X. Huang" authorityName="Tian &amp; X. Huang" authorityYear="2021" box="[754,1241,2010,2035]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="species" species="solidior" status="n. gen., n. sp.">
<emphasis id="9F8DEAE1FFE049231F8CFDFEFBA223A6" box="[754,1056,2010,2034]" italics="true" pageId="8" pageNumber="529">Guizhaphaenopsodes solidior</emphasis>
Tian &amp; X. Huang
</taxonomicName>
,
<taxonomicNameLabel id="84BE579AFFE04923199DFDFFFAF223A7" box="[1251,1392,2011,2035]" pageId="8" pageNumber="529" rank="species">
<emphasis id="9F8DEAE1FFE04923199DFDFFFAA823A7" bold="true" box="[1251,1322,2011,2035]" pageId="8" pageNumber="529">n. gen.</emphasis>
, n. sp.
</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="AD4636F3FFE149221DB9FAB3FBE924E5" blockId="9.[151,1436,151,321]" box="[199,1131,151,177]" pageId="9" pageNumber="530">Ventrites IVVI each with a pair of paramedial setae, VII bisetose apically in male.</paragraph>
<paragraph id="AD4636F3FFE149221DB9FA9FFA192515" blockId="9.[151,1436,151,321]" pageId="9" pageNumber="530">
Male genitalia (
<figureCitation id="35C22A76FFE149221C09FA9FFE3E2481" box="[375,444,187,213]" captionStart="FIGURE 7" captionStartId="9.[151,250,1031,1055]" captionTargetBox="[151,1428,345,1008]" captionTargetId="figure-112@9.[151,1436,345,1008]" captionTargetPageId="9" captionText="FIGURE 7. Male genitalia of Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp. (A. median lobe and parameres, lateral view; B. apical lobe of median lobe, dorsal view)" figureDoi="http://doi.org/10.5281/zenodo.4529641" httpUri="https://zenodo.org/record/4529641/files/figure.png" pageId="9" pageNumber="530">Fig. 7</figureCitation>
): The median lobe of aedeagus slightly arcuate at median portion, rounded blunt at apex; basal opening quite large, with a round and large sagittal aileron; inner sac provided with a thick and long copulatory piece, which is about 1/4 as long as aedeagus; in dorsal view, apical lobe longer than wide, widened at apex; parameres well-developed and elongated, the right one as long as the left, each armed with four long setae at apex.
</paragraph>
<caption id="F986667BFFE149221DE9FE23FD522017" ID-DOI="http://doi.org/10.5281/zenodo.4529641" ID-Zenodo-Dep="4529641" httpUri="https://zenodo.org/record/4529641/files/figure.png" pageId="9" pageNumber="530" startId="9.[151,250,1031,1055]" targetBox="[151,1428,345,1008]" targetPageId="9">
<paragraph id="AD4636F3FFE149221DE9FE23FD522017" blockId="9.[151,1437,1031,1091]" pageId="9" pageNumber="530">
<emphasis id="9F8DEAE1FFE149221DE9FE23FE91204B" bold="true" box="[151,275,1031,1055]" pageId="9" pageNumber="530">FIGURE 7.</emphasis>
Male genitalia of
<taxonomicName id="6AF94D70FFE149221CB1FE23FC3A204B" authority="Tian &amp; X. Huang" authorityName="Tian &amp; X. Huang" authorityYear="2021" box="[463,952,1031,1055]" class="Insecta" family="Carabidae" genus="Guizhaphaenopsodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="solidior" status="n. gen., n. sp.">
<emphasis id="9F8DEAE1FFE149221CB1FE23FD7F204B" box="[463,765,1031,1055]" italics="true" pageId="9" pageNumber="530">Guizhaphaenopsodes solidior</emphasis>
Tian &amp; X. Huang
</taxonomicName>
,
<taxonomicNameLabel id="84BE579AFFE149221EBCFE23FBD2204B" box="[962,1104,1031,1055]" pageId="9" pageNumber="530" rank="species">
<emphasis id="9F8DEAE1FFE149221EBCFE23FB88204B" bold="true" box="[962,1034,1031,1055]" pageId="9" pageNumber="530">n. gen.</emphasis>
,
<emphasis id="9F8DEAE1FFE149221969FE23FBD2204B" bold="true" box="[1047,1104,1031,1055]" pageId="9" pageNumber="530">n. sp.</emphasis>
</taxonomicNameLabel>
(A. median lobe and parameres, lateral view; B. apical lobe of median lobe, dorsal view)
</paragraph>
</caption>
<paragraph id="AD4636F3FFE149221DB9FE55FE1D20DF" blockId="9.[199,1433,1137,1236]" box="[199,415,1137,1163]" pageId="9" pageNumber="530">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="E5E36578FFE149221DB9FEB1FCA320E4" box="[199,801,1173,1200]" pageId="9" pageNumber="530" type="etymology">
<paragraph id="AD4636F3FFE149221DB9FEB1FCA320E4" blockId="9.[199,1433,1137,1236]" box="[199,801,1173,1200]" pageId="9" pageNumber="530">
<emphasis id="9F8DEAE1FFE149221DB9FEB1FECC20FB" bold="true" box="[199,334,1173,1199]" pageId="9" pageNumber="530">Etymology.</emphasis>
To refer to the strong body and antennae.
</paragraph>
</subSubSection>
<subSubSection id="E5E36578FFE149221DB9FE9DFA1B2087" box="[199,1433,1209,1236]" pageId="9" pageNumber="530" type="distribution">
<paragraph id="AD4636F3FFE149221DB9FE9DFA1B2087" blockId="9.[199,1433,1137,1236]" box="[199,1433,1209,1236]" pageId="9" pageNumber="530">
<emphasis id="9F8DEAE1FFE149221DB9FE9DFEE32087" bold="true" box="[199,353,1209,1235]" pageId="9" pageNumber="530">Distribution.</emphasis>
<collectingCountry id="D5EE7663FFE149221C19FE9DFE282087" box="[359,426,1209,1235]" name="China" pageId="9" pageNumber="530">China</collectingCountry>
(
<collectingRegion id="6F3DF811FFE149221CC7FE9DFD882087" box="[441,522,1209,1235]" country="China" name="Hunan" pageId="9" pageNumber="530">Hunan</collectingRegion>
). Known only from Tangle Dong, sympatric with
<taxonomicName id="6AF94D70FFE149221947FE9EFB732087" authority="Tian &amp; Huang &amp; Li, 2021" authorityName="Tian &amp; Huang &amp; Li" authorityYear="2021" box="[1081,1265,1210,1235]" class="Insecta" family="Carabidae" genus="Xiangxius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="species" species="jiangi" status="sp. nov.">
<emphasis id="9F8DEAE1FFE149221947FE9EFB732087" box="[1081,1265,1210,1235]" italics="true" pageId="9" pageNumber="530">Xiangxius jiangi</emphasis>
</taxonomicName>
<emphasis id="9F8DEAE1FFE149221986FE9DFAB52087" bold="true" box="[1272,1335,1209,1235]" pageId="9" pageNumber="530">
<taxonomicNameLabel id="84BE579AFFE149221986FE9DFAB52087" box="[1272,1335,1209,1235]" pageId="9" pageNumber="530" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="35C22A76FFE149221839FE9DFA0A2087" box="[1351,1416,1209,1235]" captionStart="FIGURE 1" captionStartId="2.[151,250,1343,1367]" captionTargetBox="[210,1378,181,1320]" captionTargetId="figure-23@2.[210,1378,181,1320]" captionTargetPageId="2" captionText="FIGURE 1. A distribution map showing the locality caves of the three new species described in the present paper (Tangle Dong: Xiangxius jiangi Tian &amp; X. Huang, n. gen., n. sp. and Guizhaphaenopsodes solidior Tian &amp; X. Huang, n. gen., n. sp.; Qizimei Dong: Sinotroglodytes hefengensis Tian &amp; Li, n. sp.)." figureDoi="http://doi.org/10.5281/zenodo.4529629" httpUri="https://zenodo.org/record/4529629/files/figure.png" pageId="9" pageNumber="530">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>