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171 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59143" ID-GBIF-Dataset="83d5b180-4e0b-44d1-ad83-67ad55ee0355" ID-GBIF-Taxon="182236449" ID-Pensoft-Pub="1860-0743-1-1" ID-Pensoft-UUID="BD006A3AD2C6516AB596CDA05E0F375D" ID-ZooBank="1D5553BA24894F2C9527B7E46B9B432B" ModsDocID="1860-0743-97-1-1" checkinTime="1621423297205" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Pall-Gergely, Barna, Sutcharit, Chirasak & Tongkerd, Piyoros" docDate="2021" docId="1C276FCEEFB257D1B5EA4376DB254B83" docLanguage="en" docName="ZoosystEvol 97(1): 1-20" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-01-05" docSource="http://dx.doi.org/10.3897/zse.97.59143" docTitle="Dicharax omissus" docType="treatment" docVersion="3" id="BD006A3AD2C6516AB596CDA05E0F375D" lastPageNumber="1" masterDocId="BD006A3AD2C6516AB596CDA05E0F375D" masterDocTitle="The operculate micro land snail genus Dicharax Kobelt & Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species" masterLastPageNumber="20" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1632012559486" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>The operculate micro land snail genus Dicharax Kobelt & Moellendorff, 1900 (Caenogastropoda, Alycaeidae) in Thailand, with description of new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pall-Gergely, Barna</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
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<mods:affiliation>Plant Protection Institute, Centre for Agricultural Research, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tongkerd, Piyoros</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9221-9293</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">piyorose@hotmail.com</mods:nameIdentifier>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-01-05</mods:number>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>20</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.97.59143</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.59143</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-1</mods:identifier>
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<mods:identifier type="ZooBank">1D5553BA24894F2C9527B7E46B9B432B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">BD006A3AD2C6516AB596CDA05E0F375D</mods:identifier>
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<treatment ID-GBIF-Taxon="182236449" LSID="urn:lsid:plazi:treatment:1C276FCEEFB257D1B5EA4376DB254B83" httpUri="http://treatment.plazi.org/id/1C276FCEEFB257D1B5EA4376DB254B83" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="1C276FCE-EFB2-57D1-B5EA-4376DB254B83" authority="(Godwin-Austen, 1914)" baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax omissus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="omissus">Dicharax omissus (Godwin-Austen, 1914)</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">Figs 6A</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">, 8A, B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus omissus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="omissus">Alycaeus omissus</taxonomicName>
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Godwin-Austen, 1914: 411, pl. 155, fig. 13. Type locality: Siam and Shan State boundary.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Godwin-Austen" authorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Chamalycaeus) omissus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="omissus" subGenus="Chamalycaeus">Alycaeus (Chamalycaeus) omissus</taxonomicName>
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-
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B20" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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: 231.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Kobelt & Moellendorff" authorityYear="1900" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Dicharax</taxonomicName>
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(?)
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<taxonomicName baseAuthorityName="Godwin-Austen" baseAuthorityYear="1914" class="Gastropoda" family="Cyclophoridae" genus="Dicharax" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Dicharax omissus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="omissus">Dicharax omissus</taxonomicName>
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-
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.981.53583" author="Pall-Gergely, B" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 220" refId="B47" refString="Pall-Gergely, B, Sajan, S, Tripathy, B, Meng, K, Takahiro, A, Ablett, J, 2020b. Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue. ZooKeys 981: 1 - 220, DOI: https://doi.org/10.3897/zookeys.981.53583" title="Genus-level revision of the Alycaeidae (Gastropoda: Cyclophoroidea), with an annotated species catalogue." url="https://doi.org/10.3897/zookeys.981.53583" volume="981" year="2020 b">
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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et al. 2020b
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</bibRefCitation>
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: 100.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
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NHMUK 1903.7.1.1228 from Siam & Shan boundary (2 shells; Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation collectingDate="2017-01-19" collectorName="C. Sutcharit" country="Thailand" latitude="19.574696" location="Tham Mae La Na" longLatPrecision="1" longitude="98.21594" specimenCount="1" stateProvince="Mae Hong Son Province">
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CUMZ 7426 from
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<location LSID="urn:lsid:plazi:treatment:1C276FCEEFB257D1B5EA4376DB254B83:91A9A8DC5A573BE14882BC98AD9289E6" country="Thailand" latitude="19.574696" longLatPrecision="1" longitude="98.21594" name="Tham Mae La Na" stateProvince="Mae Hong Son Province">Tham Mae La Na</location>
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,
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<collectingRegion country="Thailand" name="Mae Hong Son">Mae Hong Son Province</collectingRegion>
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,
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<collectingCountry name="Thailand">Thailand</collectingCountry>
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,
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<geoCoordinate degrees="19" direction="north" minutes="34" orientation="latitude" precision="1" seconds="28.9" value="19.574696">19°34'28.9"N</geoCoordinate>
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,
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<geoCoordinate degrees="98" direction="east" minutes="12" orientation="longitude" precision="1" seconds="57.4" value="98.21594">98°12'57.4"E</geoCoordinate>
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,
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<collectingDate value="2017-01-19">19 Jan. 2017</collectingDate>
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, coll.
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<collectorName>C. Sutcharit</collectorName>
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(1 shell; Figs
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Synoptic view of the eight Dicharax taxa in Thailand. A. Dicharax omissus, specimen CUMZ 7426 (gold plated); B. Dicharax pratatensis, specimen CUMZ 7427; C. Dicharax panhai sp. nov., paratype CUMZ 7429 / 2; D. Dicharax pongrati sp. nov., paratype CUMZ 7430 / 2; E. Dicharax notus, specimen CUMZ 7425; F. Dicharax burchi sp. nov., paratype CUMZ 7428 / 2; G. Dicharax borealis sp. nov., paratype CUMZ 5072 / 2; H. Dicharax cucullatus, specimen CUMZ 7421." figureDoi="10.3897/zse.97.59143.figure6" httpUri="https://binary.pensoft.net/fig/494370" pageId="0" pageNumber="1">6A</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8B</figureCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell very small (SH up to 1.7 mm, SW up to 2.7 mm), depressed-conical. Spire ca. 1/4 of shell height. R1 with regular, strong ribs; R2 shorter than R3; R2 with 35-36 ribs. R3 with blunt central swelling. Aperture round. Inner peristome without indentation, with beak-like basal protrusion in newly examined population. Outer peristome expanded, not reflected. Umbilicus round.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Border region between Thailand and Myanmar (
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<bibRefCitation author="Gude, GK" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="1" refId="B20" refString="Gude, GK, 1921. The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates. Taylor and Francis, London" title="The Fauna of British India including Ceylon and Burma. Mollusca - III. Land Operculates." year="1921">Gude 1921</bibRefCitation>
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), and Mae Hong Son Province, Thailand.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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The specimen from Mae Hong Son Province (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8B</figureCitation>
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) exhibits stronger R1 ribs and possesses a beak-like protrusion at the base of the aperture, which is absent in the syntype (Fig.
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. A, B. Dicharax omissus A. Syntype NHMUK 1903.7.1.1228; B. Specimen CUMZ 7426 from Tham Mae La Na, Mae Hong Son Province C, D. Dicharax pratatensis C. Holotype CUMZ Cy 001; D. Paratype CUMZ Cy 002 from Doi Chiang Dao, Chiang Mai Province. Close-up images of the aperture not to scale." figureDoi="10.3897/zse.97.59143.figure8" httpUri="https://binary.pensoft.net/fig/494372" pageId="0" pageNumber="1">8A</figureCitation>
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). Apart from these differences, this specimen is nearly identical to the syntype in terms of shell size and shape, and R2 and R3 length. In addition, the locality of this specimen lies only approximately 12 km from the boundary between Thailand and Shan State, Myanmar. We thus identify this specimen as
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<taxonomicName genus="D." lsidName="D. omissus" pageId="0" pageNumber="1" rank="species" species="omissus">
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<emphasis italics="true" pageId="0" pageNumber="1">D. omissus</emphasis>
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</taxonomicName>
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. The most similar species to
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<taxonomicName genus="D." lsidName="D. omissus" pageId="0" pageNumber="1" rank="species" species="omissus">
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<emphasis italics="true" pageId="0" pageNumber="1">D. omissus</emphasis>
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</taxonomicName>
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is
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<taxonomicName genus="D." lsidName="D. notus" pageId="0" pageNumber="1" rank="species" species="notus">
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<emphasis italics="true" pageId="0" pageNumber="1">D. notus</emphasis>
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</taxonomicName>
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, which is larger and has reflected outer peristome.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |