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<document id="CCDC2FBF8111F2C1158D9A217E91AF38" ID="10.11646/zootaxa.4173.6.2" ID-CLB-Dataset="38065" ID-DOI="10.11646/zootaxa.4173.6.2" ID-GBIF-Dataset="f9f36ad7-2af7-4b6f-bc33-37418b11470b" ID-ISSN="1175-5326" ID-Zenodo-Dep="254767" ID-ZooBank="AEBD61A6-37B5-4413-A9B1-7895D3EE53F4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476092631919" checkinUser="plazi" docAuthor="Moeseneder, Christian H. &amp; Hutchinson, Paul M." docDate="2016" docId="F805B047FFF4FFBBFF538E66FA49CCC5" docLanguage="en" docName="zootaxa.4173.6.2.pdf" docOrigin="Zootaxa 4173 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Navigator" docType="key" docVersion="9" lastPageNumber="536" masterDocId="043CC83FFFF2FFBDFFC48B7EFFD5CB0C" masterDocTitle="Navigator, a new endemic genus of Cetoniinae (Coleoptera: Scarabaeidae) from Australia, with descriptions of two new species and behavioural studies" masterLastPageNumber="556" masterPageNumber="530" pageNumber="536" updateTime="1698429497132" updateUser="ExternalLinkService">
<mods:mods id="6E08656A4395E836E77AA85388A456DA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="70F8AEE0D4BA52AB58728874DE53BE72">Navigator, a new endemic genus of Cetoniinae (Coleoptera: Scarabaeidae) from Australia, with descriptions of two new species and behavioural studies</mods:title>
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<mods:namePart id="F57F6B37F24BDAA96D021A3E9011A077">Moeseneder, Christian H.</mods:namePart>
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<mods:name id="99043DBB265F48D276DF34AE10639807" type="personal">
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<mods:namePart id="E05FEF634B399AA97B69D9820B070964">Hutchinson, Paul M.</mods:namePart>
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<mods:typeOfResource id="0281184A777934437DFB039CB4198405">text</mods:typeOfResource>
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<mods:title id="FE6874AE232997BFC8CA27ED94FCFE2E">Zootaxa</mods:title>
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<mods:date id="9C27419DC31A0830A53E6B79244F2E73">2016</mods:date>
<mods:detail id="697AB569F108DC6D2E0ABA8D85F36A55" type="volume">
<mods:number id="B4848248AD0C344418BBF88C5A1B932B">4173</mods:number>
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<mods:number id="0E642B2102A4948A28A5247924EFAFC1">6</mods:number>
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<mods:classification id="4FF60855C47D955091F48906D3203DB2">journal article</mods:classification>
<mods:identifier id="781D99A3D9AF7626C32D48655BDFA784" type="CLB-Dataset">38065</mods:identifier>
<mods:identifier id="75FFDE434113B2FBF984F8092076F450" type="DOI">10.11646/zootaxa.4173.6.2</mods:identifier>
<mods:identifier id="1314BA18869390D271D225F631C093CC" type="GBIF-Dataset">f9f36ad7-2af7-4b6f-bc33-37418b11470b</mods:identifier>
<mods:identifier id="1BA15265A9B853F1E617E05B2DF1D63D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D91585D80CC797F7D88EF33A0F003D9B" type="Zenodo-Dep">254767</mods:identifier>
<mods:identifier id="1684FC79B65EDD1B7F379E1EC679E203" type="ZooBank">AEBD61A6-37B5-4413-A9B1-7895D3EE53F4</mods:identifier>
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<treatment id="F805B047FFF4FFBBFF538E66FA49CCC5" ID-DOI="http://doi.org/10.5281/zenodo.6071753" ID-GBIF-Taxon="124997310" ID-Zenodo-Dep="6071753" LSID="urn:lsid:plazi:treatment:F805B047FFF4FFBBFF538E66FA49CCC5" httpUri="http://treatment.plazi.org/id/F805B047FFF4FFBBFF538E66FA49CCC5" lastPageNumber="536" pageId="6" pageNumber="536">
<subSubSection id="38B652DAFFF4FFBBFF538E66FE74CD0A" pageId="6" pageNumber="536" type="nomenclature">
<paragraph id="70130151FFF4FFBBFF538E66FDC1CE3D" blockId="6.[151,532,1303,1330]" box="[151,532,1303,1330]" pageId="6" pageNumber="536">
<heading id="2B5BB63DFFF4FFBBFF538E66FDC1CE3D" bold="true" box="[151,532,1303,1330]" fontSize="11" level="1" pageId="6" pageNumber="536" reason="1">
<emphasis id="42D8DD43FFF4FFBBFF538E66FDC1CE3D" bold="true" box="[151,532,1303,1330]" pageId="6" pageNumber="536">
Key to the species of
<taxonomicName id="B7AC7AD2FFF4FFBBFE588E69FDC1CE3D" ID-CoL="5XXJ" box="[412,532,1303,1329]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF4FFBBFE588E69FDC1CE3D" bold="true" box="[412,532,1303,1329]" italics="true" pageId="6" pageNumber="536">Navigator</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="70130151FFF4FFBBFF538E20FE74CD0A" blockId="6.[151,1437,1374,1543]" pageId="6" pageNumber="536">
Prerequisite for the use of this key is sex determination of the specimen. In males of
<emphasis id="42D8DD43FFF4FFBBFB838E21FA90CE7B" box="[1095,1349,1375,1399]" italics="true" pageId="6" pageNumber="536">
<taxonomicName id="B7AC7AD2FFF4FFBBFB838E21FB79CE7B" box="[1095,1196,1375,1399]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="fossor">N. fossor</taxonomicName>
,
<taxonomicName id="B7AC7AD2FFF4FFBBFB728E21FA90CE7B" box="[1206,1349,1375,1399]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="ruficornis">N. ruficornis</taxonomicName>
</emphasis>
, and
<taxonomicName id="B7AC7AD2FFF4FFBBFA468E21FF1CCE97" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF4FFBBFA468E21FF1CCE97" italics="true" pageId="6" pageNumber="536">N. pixii</emphasis>
</taxonomicName>
(
<taxonomicName id="B7AC7AD2FFF4FFBBFF1F8EFAFE86CE97" box="[219,339,1412,1435]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="interior">
<emphasis id="42D8DD43FFF4FFBBFF1F8EFAFE86CE97" box="[219,339,1412,1435]" italics="true" pageId="6" pageNumber="536">N. interior</emphasis>
</taxonomicName>
not yet known), males have a longitudinal abdominal impression or flat sternites, antennal club length longer than antennomeres 1–7, and non-spatulate metatibial internal spur. Females of all
<taxonomicName id="B7AC7AD2FFF4FFBBFB168ED9FA91CEB2" box="[1234,1348,1447,1470]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF4FFBBFB168ED9FA91CEB2" box="[1234,1348,1447,1470]" italics="true" pageId="6" pageNumber="536">Navigator</emphasis>
</taxonomicName>
species have medially convex sternites, antennal club length equal to or less than antennomeres 1–7, and spatulate metatibial internal spur.
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF4FFBBFF538D4BFA49CCC5" pageId="6" pageNumber="536" type="key">
<keyStep id="CB5819F4FFF4FFBBFF538D4BFA49CDAF" pageId="6" pageNumber="536">
<paragraph id="70130151FFF4FFBBFF538D4BFA49CD65" blockId="6.[151,1436,1589,1993]" pageId="6" pageNumber="536">
<keyLead id="CB5DA264FFF4FFBBFF538D4BFA49CD65" pageId="6" pageNumber="536">
1 Preclypeus shallow (
<figureCitation id="E8971DD4FFF4FFBBFE6A8D4BFE32CD47" box="[430,487,1589,1611]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
A, detail 5 and cross-section 6); clypeal lateral declivity absent (
<figureCitation id="E8971DD4FFF4FFBBFBB68D4BFB7ECD47" box="[1138,1195,1589,1611]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
A, details 2, 7). Female abdominal sternites distended (
<figureCitation id="E8971DD4FFF4FFBBFDCF8D2DFD94CD65" box="[523,577,1619,1641]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
E, detail 1); disc of pygidium visible in dorsal view (
<figureCitation id="E8971DD4FFF4FFBBFBF98D2DFBA6CD65" box="[1085,1139,1619,1641]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
D, detail 8)............... 2
</keyLead>
</paragraph>
<paragraph id="70130151FFF4FFBBFF538D0FFA49CDAF" blockId="6.[151,1436,1589,1993]" pageId="6" pageNumber="536">
<keyLead id="CB5DA264FFF4FFBBFF538D0FFA49CDAF" pageId="6" pageNumber="536">
– Preclypeus deep (
<figureCitation id="E8971DD4FFF4FFBBFE4B8D0EFE1DCD8A" box="[399,456,1648,1670]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
B, detail 2 and cross-section 3); clypeal lateral declivity present (
<figureCitation id="E8971DD4FFF4FFBBFB968D0EFB5ECD8A" box="[1106,1163,1648,1670]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
B, details 1 and 4). Female abdominal sternites not distended; pygidium not well visible in dorsal view (
<figureCitation id="E8971DD4FFF4FFBBFC7B8DF3FC20CDAF" box="[959,1013,1677,1699]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
D, detail 9)......................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="CB5819F4FFF4FFBBFF538DD5FA49CC5F" pageId="6" pageNumber="536">
<paragraph id="70130151FFF4FFBBFF538DD5FA49CDF7" blockId="6.[151,1436,1589,1993]" pageId="6" pageNumber="536">
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<specimenCount id="66AACAD8FFF4FFBBFF538DD5FEC7CDCD" box="[151,274,1707,1729]" pageId="6" pageNumber="536" type="male">2 Male</specimenCount>
metatibia internal margin with longitudinal setose groove, apex unispinose (
<figureCitation id="E8971DD4FFF4FFBBFC398DD5FBE1CDCD" box="[1021,1076,1707,1729]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
K, detail 1).
<collectingCounty id="997279DDFFF4FFBBFB778DD2FAC7CDCD" box="[1203,1298,1708,1729]" pageId="6" pageNumber="536">Metacoxa</collectingCounty>
posterolateral angle angulate (
<figureCitation id="E8971DD4FFF4FFBBFEBD8DB6FE7BCDD2" box="[377,430,1736,1758]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
G, detail 1).
<collectingMunicipality id="90779B2BFFF4FFBBFDE38DB6FD8DCDD2" box="[551,600,1736,1758]" pageId="6" pageNumber="536">Male</collectingMunicipality>
size
<quantity id="B754ACB4FFF4FFBBFD4E8DB6FCDECDD2" box="[650,779,1736,1758]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.17" metricValueMin="0.96" pageId="6" pageNumber="536" unit="mm" value="10.649999999999999" valueMax="11.7" valueMin="9.6">9.6–11.7 mm</quantity>
.
<location id="7573578AFFF4FFBBFCD08DB6FC8ECDD2" LSID="urn:lsid:plazi:treatment:F805B047FFF4FFBBFF538E66FA49CCC5:7573578AFFF4FFBBFCD08DB6FC8ECDD2" box="[788,859,1736,1758]" county="Metacoxa" municipality="Male" name="Female" pageId="6" pageNumber="536" stateProvince="Queensland">Female</location>
<quantity id="B754ACB4FFF4FFBBFCA48DB6FC09CDD2" box="[864,988,1736,1758]" metricMagnitude="-2" metricUnit="m" metricValue="1.325" metricValueMax="1.4" metricValueMin="1.25" pageId="6" pageNumber="536" unit="mm" value="13.25" valueMax="14.0" valueMin="12.5">12.5–14 mm</quantity>
, ovoid.
<location id="7573578AFFF4FFBBFBEC8DB7FB89CDD2" LSID="urn:lsid:plazi:treatment:F805B047FFF4FFBBFF538E66FA49CCC5:7573578AFFF4FFBBFBEC8DB7FB89CDD2" box="[1064,1116,1737,1758]" county="Metacoxa" municipality="Male" name="From" pageId="6" pageNumber="536" stateProvince="Queensland">From</location>
southeastern
<collectingRegion id="B268CFB3FFF4FFBBFB248DB6FA84CDD2" box="[1248,1361,1736,1758]" country="Australia" name="Queensland" pageId="6" pageNumber="536">Queensland</collectingRegion>
..........................................................................
<taxonomicName id="B7AC7AD2FFF4FFBBFBD58D98FA49CDF7" ID-CoL="45X25" authority="Moeseneder &amp; Hutchinson, 2016" authorityName="Moeseneder &amp; Hutchinson" authorityYear="2016" box="[1041,1436,1765,1787]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF4FFBBFBD58D98FB8DCDF7" box="[1041,1112,1766,1787]" italics="true" pageId="6" pageNumber="536">N. pixii</emphasis>
Moeseneder &amp; Hutchinson, 2016
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</paragraph>
<paragraph id="70130151FFF4FFBBFF538C7AFA49CC5F" blockId="6.[151,1436,1589,1993]" pageId="6" pageNumber="536">
<keyLead id="CB5DA264FFF4FFBBFF538C7AFA49CC5F" pageId="6" pageNumber="536">
– Male metatibia internal margin not grooved, apex trispinose (
<figureCitation id="E8971DD4FFF4FFBBFCEA8C7DFCB1CC15" box="[814,868,1795,1817]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
H, detail 3). Metacoxa posterolateral angle broadly arcuate (
<figureCitation id="E8971DD4FFF4FFBBFF2C8C5EFEC8CC3A" box="[232,285,1824,1846]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
G, detail 2). Male size
<quantity id="B754ACB4FFF4FFBBFE3D8C5EFD52CC3A" box="[505,647,1824,1846]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.6199999999999999" metricValueMin="1.48" pageId="6" pageNumber="536" unit="mm" value="15.5" valueMax="16.2" valueMin="14.8">14.8–16.2 mm</quantity>
. Female
<quantity id="B754ACB4FFF4FFBBFD188C5EFCBFCC3A" box="[732,874,1824,1846]" metricMagnitude="-2" metricUnit="m" metricValue="1.785" metricValueMax="1.89" metricValueMin="1.68" pageId="6" pageNumber="536" unit="mm" value="17.85" valueMax="18.9" valueMin="16.8">16.8–18.9 mm</quantity>
, elongate ovoid.
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From
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<collectingRegion id="B268CFB3FFF4FFBBFB838C5FFB40CC3A" box="[1095,1173,1825,1846]" country="American Samoa" name="Western" pageId="6" pageNumber="536">Western</collectingRegion>
Australia
</collectingRegion>
</collectingRegion>
.................................................................................................
<taxonomicName id="B7AC7AD2FFF4FFBBFB0B8C40FA49CC5F" ID-CoL="45X23" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" box="[1231,1436,1853,1875]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF4FFBBFB0B8C40FAF3CC5F" box="[1231,1318,1853,1875]" italics="true" pageId="6" pageNumber="536">N. fossor</emphasis>
(
<bibRefCitation id="143D7CA0FFF4FFBBFAF78C43FA40CC5F" author="Lea" box="[1331,1429,1853,1875]" pageId="6" pageNumber="536" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
)
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</paragraph>
</keyStep>
<keyStep id="CB5819F4FFF4FFBBFF538C25FA49CCC5" pageId="6" pageNumber="536">
<paragraph id="70130151FFF4FFBBFF538C25FA49CC82" blockId="6.[151,1436,1589,1993]" pageId="6" pageNumber="536">
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3 Metatibia with large, median denticle (
<figureCitation id="E8971DD4FFF4FFBBFD908C25FD5ECC7D" box="[596,651,1883,1905]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">
Fig.
<date id="04122791FFF4FFBBFD458C25FD5ECC7D" box="[641,651,1883,1905]" pageId="6" pageNumber="536">1</date>
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I, detail 1). Metacoxa posterolateral angle angulate (
<figureCitation id="E8971DD4FFF4FFBBFB458C25FB63CC7D" box="[1153,1206,1883,1905]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
G, detail 1).....................................................................................
<taxonomicName id="B7AC7AD2FFF4FFBBFBAE8C07FA49CC82" ID-CoL="45X26" authority="Westwood, 1874" authorityName="Westwood" authorityYear="1874" box="[1130,1436,1912,1934]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF4FFBBFBAE8C07FB31CC82" box="[1130,1252,1912,1934]" italics="true" pageId="6" pageNumber="536">N. ruficornis</emphasis>
(
<bibRefCitation id="143D7CA0FFF4FFBBFB368C06FA40CC82" author="Westwood" box="[1266,1429,1912,1934]" pageId="6" pageNumber="536" refString="Westwood, J. O. (1874) Descriptions of some new species of exotic Cetoniidae. Transactions of the Entomological Society of London, 4, 477." type="journal article" year="1874">Westwood, 1874</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="70130151FFF4FFBBFF538CE8FA49CCC5" blockId="6.[151,1436,1589,1993]" pageId="6" pageNumber="536">
<keyLead id="CB5DA264FFF4FFBBFF538CE8FA49CCC5" pageId="6" pageNumber="536">
– Metatibia with small, median denticle (
<figureCitation id="E8971DD4FFF4FFBBFD9D8CEBFD5ACCA7" box="[601,655,1941,1963]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
J, detail 1). Metacoxa posterolateral angle spinose (
<figureCitation id="E8971DD4FFF4FFBBFBB98CEBFB67CCA7" box="[1149,1202,1941,1963]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="6" pageNumber="536">Fig. 1</figureCitation>
G, detail 3)...........................................................................
<taxonomicName id="B7AC7AD2FFF4FFBBFC368CCAFA49CCC5" ID-CoL="45X24" authority="Moeseneder &amp; Hutchinson, 2016" authorityName="Moeseneder &amp; Hutchinson" authorityYear="2016" box="[1010,1436,1971,1993]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="interior">
<emphasis id="42D8DD43FFF4FFBBFC368CCAFB82CCC5" box="[1010,1111,1972,1993]" italics="true" pageId="6" pageNumber="536">N. interior</emphasis>
Moeseneder &amp; Hutchinson, 2016
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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