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<document id="3F51AD39CC0C9FCA04EC333457F056EC" ID="10.11646/zootaxa.4173.6.2" ID-CLB-Dataset="38065" ID-DOI="10.11646/zootaxa.4173.6.2" ID-GBIF-Dataset="f9f36ad7-2af7-4b6f-bc33-37418b11470b" ID-ISSN="1175-5326" ID-Zenodo-Dep="254767" ID-ZooBank="AEBD61A6-37B5-4413-A9B1-7895D3EE53F4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476092631919" checkinUser="plazi" docAuthor="Moeseneder, Christian H. &amp; Hutchinson, Paul M." docDate="2016" docId="F805B047FFF1FFBBFF538BE9FCE7CFC8" docLanguage="en" docName="zootaxa.4173.6.2.pdf" docOrigin="Zootaxa 4173 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Navigator Moeseneder &amp; Hutchinson, new genus" docType="treatment" docUuid="E71AE6FD-A65B-434A-94A1-33104F9EE945" docUuidSource="ZooBank" docVersion="10" lastPageNumber="536" masterDocId="043CC83FFFF2FFBDFFC48B7EFFD5CB0C" masterDocTitle="Navigator, a new endemic genus of Cetoniinae (Coleoptera: Scarabaeidae) from Australia, with descriptions of two new species and behavioural studies" masterLastPageNumber="556" masterPageNumber="530" pageNumber="533" updateTime="1698429497132" updateUser="ExternalLinkService">
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<mods:title id="288E9E4F28FD7C70847C8B18498C544B">Navigator, a new endemic genus of Cetoniinae (Coleoptera: Scarabaeidae) from Australia, with descriptions of two new species and behavioural studies</mods:title>
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<mods:namePart id="F7237DD1B2CBA2B047AE1FC0FE1139CA">Moeseneder, Christian H.</mods:namePart>
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<mods:namePart id="D79953A2BD1647948497512F52EF6C80">Hutchinson, Paul M.</mods:namePart>
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<mods:title id="8C7B3408DCB6B92DE39B0075F438BD95">Zootaxa</mods:title>
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<mods:date id="A514E8D7FC98BF5D8DBFDF2043C2C1E7">2016</mods:date>
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<mods:number id="93285977E9402329341F85451D6A8FA0">4173</mods:number>
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<mods:identifier id="1936F9A3EA4184C77E24EDB9BBE90A1C" type="CLB-Dataset">38065</mods:identifier>
<mods:identifier id="C6C45C3D4FBEED0CB961566E0D2736E1" type="DOI">10.11646/zootaxa.4173.6.2</mods:identifier>
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<treatment id="F805B047FFF1FFBBFF538BE9FCE7CFC8" ID-DOI="http://doi.org/10.5281/zenodo.6071748" ID-GBIF-Taxon="124997311" ID-Zenodo-Dep="6071748" LSID="urn:lsid:zoobank.org:act:E71AE6FD-A65B-434A-94A1-33104F9EE945" httpUri="http://treatment.plazi.org/id/F805B047FFF1FFBBFF538BE9FCE7CFC8" lastPageId="6" lastPageNumber="536" pageId="3" pageNumber="533">
<subSubSection id="38B652DAFFF1FFBEFF538BE9FE72CBDF" pageId="3" pageNumber="533" type="nomenclature">
<paragraph id="70130151FFF1FFBEFF538BE9FD20CBBE" blockId="3.[151,757,151,211]" box="[151,757,151,178]" pageId="3" pageNumber="533">
<heading id="2B5BB63DFFF1FFBEFF538BE9FD20CBBE" bold="true" box="[151,757,151,178]" fontSize="11" level="1" pageId="3" pageNumber="533" reason="1">
<emphasis id="42D8DD43FFF1FFBEFF538BE9FD20CBBE" bold="true" box="[151,757,151,178]" pageId="3" pageNumber="533">
<taxonomicName id="B7AC7AD2FFF1FFBEFF538BE9FDBDCBBE" ID-CoL="5XXJ" authority="Moeseneder &amp; Hutchinson" authorityName="Moeseneder &amp; Hutchinson" box="[151,616,151,178]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF1FFBEFF538BE9FEDACBBD" bold="true" box="[151,271,151,177]" italics="true" pageId="3" pageNumber="533">Navigator</emphasis>
Moeseneder &amp; Hutchinson
</taxonomicName>
,
<taxonomicNameLabel id="59EB6038FFF1FFBEFDB08BE6FD20CBBE" box="[628,757,152,178]" pageId="3" pageNumber="533" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="70130151FFF1FFBEFF538BC4FE72CBDF" blockId="3.[151,757,151,211]" box="[151,423,186,211]" pageId="3" pageNumber="533">
(
<figureCitation id="E8971DD4FFF1FFBEFF5B8BC4FEC2CBDF" box="[159,279,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="4.[151,250,1487,1509]" captionStartId-1="5.[151,250,1971,1993]" captionStartId-10="17.[151,250,1745,1767]" captionStartId-2="7.[151,250,1930,1952]" captionStartId-3="9.[151,250,1293,1315]" captionStartId-4="10.[151,250,931,953]" captionStartId-5="13.[151,250,867,889]" captionStartId-6="13.[151,250,1786,1808]" captionStartId-7="14.[151,250,853,875]" captionStartId-8="16.[151,250,1576,1598]" captionStartId-9="17.[151,250,1077,1099]" captionTargetBox-0="[179,1419,218,1444]" captionTargetBox-1="[313,1284,231,1925]" captionTargetBox-10="[163,1430,1168,1708]" captionTargetBox-2="[160,1429,1303,1899]" captionTargetBox-3="[209,1378,559,1267]" captionTargetBox-4="[166,1418,198,901]" captionTargetBox-5="[160,1429,198,829]" captionTargetBox-6="[159,1427,953,1762]" captionTargetBox-7="[158,1423,200,825]" captionTargetBox-8="[158,1429,847,1550]" captionTargetId-0="figure@4.[150,1436,193,1465]" captionTargetId-1="figure@5.[248,1336,193,1949]" captionTargetId-10="figure@17.[151,1436,1149,1723]" captionTargetId-2="figure@7.[151,1436,1297,1909]" captionTargetId-3="figure@9.[201,1386,553,1272]" captionTargetId-4="figure@10.[155,1432,193,910]" captionTargetId-5="figure@13.[151,1435,193,846]" captionTargetId-6="figure@13.[151,1436,939,1765]" captionTargetId-7="figure@14.[150,1436,193,831]" captionTargetId-8="figure@16.[151,1435,841,1555]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="17" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="13" captionTargetPageId-6="13" captionTargetPageId-7="14" captionTargetPageId-8="16" captionText-0="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." captionText-1="FIGURE 2. Navigator male aedeagi. A— Navigator fossor. B— Navigator pixii. C— Navigator ruficornis. Note: male N. interior are unknown." captionText-10="FIGURE 11. Navigator pixii allotype, female." captionText-2="FIGURE 3. Navigator fossor lectotype, female." captionText-3="FIGURE 4. Navigator fossor male." captionText-4="FIGURE 5. Navigator fossor female." captionText-5="FIGURE 6. Navigator ruficornis lectotype, male. Image © MNHN, A. Mantilleri." captionText-6="FIGURE 7. Navigator ruficornis male." captionText-7="FIGURE 8. Navigator ruficornis female." captionText-8="FIGURE 9. Navigator pixii holotype, male." captionText-9="FIGURE 10. Navigator pixii male, colour form. Legs and antennae digitally removed." httpUri-0="https://zenodo.org/record/254768/files/figure.png" httpUri-1="https://zenodo.org/record/254769/files/figure.png" httpUri-10="https://zenodo.org/record/254777/files/figure.png" httpUri-2="https://zenodo.org/record/254770/files/figure.png" httpUri-3="https://zenodo.org/record/254771/files/figure.png" httpUri-4="https://zenodo.org/record/254772/files/figure.png" httpUri-5="https://zenodo.org/record/254773/files/figure.png" httpUri-6="https://zenodo.org/record/254774/files/figure.png" httpUri-7="https://zenodo.org/record/254775/files/figure.png" httpUri-8="https://zenodo.org/record/254776/files/figure.png" pageId="3" pageNumber="533">Figs. 1–11</figureCitation>
,
<figureCitation id="E8971DD4FFF1FFBEFEE78BC5FE94CBDF" box="[291,321,187,211]" captionStart="FIGURE 13" captionStartId="20.[151,250,766,788]" captionTargetBox="[162,1432,208,733]" captionTargetId="figure@20.[151,1436,193,744]" captionTargetPageId="20" captionText="FIGURE 13. Navigator interior holotype, female." httpUri="https://zenodo.org/record/254779/files/figure.png" pageId="3" pageNumber="533">13</figureCitation>
,
<figureCitation id="E8971DD4FFF1FFBEFE898BC5FEBDCBDF" box="[333,360,187,211]" captionStart="FIGURE 14" captionStartId="22.[151,250,1888,1910]" captionTargetBox="[252,1343,212,1839]" captionTargetId="figure@22.[232,1366,193,1867]" captionTargetPageId="22" captionText="FIGURE 14. Navigator habitat. A – D— Navigator fossor (all images near or 10 km west of Menzies, Western Australia): A— Acacia leaf litter and branch thicket. B— Acacia with scattered Grevillea. C—male on leaf litter under Acacia. D—larva in decomposing leaf substrate. E – F— Navigator pixii (images at Boggomoss site BM 03, Queesland): E—collecting locations of adults and larvae. F—accumulations of dead leaves, predominantly from Acacia harpophylla, on the ground under which larvae were found." httpUri="https://zenodo.org/record/254780/files/figure.png" pageId="3" pageNumber="533">14</figureCitation>
C–D)
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF1FFBEFF538A7CFA8ACA6F" pageId="3" pageNumber="533" type="materials_examined">
<paragraph id="70130151FFF1FFBEFF538A7CFC54CA17" blockId="3.[151,1437,258,2011]" box="[151,897,258,283]" pageId="3" pageNumber="533">
<emphasis id="42D8DD43FFF1FFBEFF538A7CFEFECA17" bold="true" box="[151,299,258,283]" pageId="3" pageNumber="533">Type species</emphasis>
.
<taxonomicName id="B7AC7AD2FFF1FFBEFEFD8A7CFD76CA17" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" box="[313,675,258,283]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF1FFBEFEFD8A7CFDFCCA17" box="[313,553,258,283]" italics="true" pageId="3" pageNumber="533">Pseudoclithria fossor</emphasis>
<bibRefCitation id="143D7CA0FFF1FFBEFDF48A7DFD76CA17" author="Lea" box="[560,675,259,283]" pageId="3" pageNumber="533" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
</taxonomicName>
, hereby designated.
</paragraph>
<paragraph id="70130151FFF1FFBEFF038A58FA8ACA6F" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="533">
<emphasis id="42D8DD43FFF1FFBEFF038A58FE58CA33" bold="true" box="[199,397,294,319]" pageId="3" pageNumber="533">Included species</emphasis>
.
<taxonomicName id="B7AC7AD2FFF1FFBEFE5A8A58FCC7CA33" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" box="[414,786,294,319]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF1FFBEFE5A8A58FD44CA33" box="[414,657,294,319]" italics="true" pageId="3" pageNumber="533">Pseudoclithria fossor</emphasis>
<bibRefCitation id="143D7CA0FFF1FFBEFD588A59FCC7CA33" author="Lea" box="[668,786,295,319]" pageId="3" pageNumber="533" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B7AC7AD2FFF1FFBEFCE48A58FAC9CA33" authority="Westwood, 1874" authorityName="Westwood" authorityYear="1874" box="[800,1308,294,319]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF1FFBEFCE48A58FBE8CA33" box="[800,1085,294,319]" italics="true" pageId="3" pageNumber="533">Pseudoclithria ruficornis</emphasis>
(
<bibRefCitation id="143D7CA0FFF1FFBEFB8B8A58FAC1CA33" author="Westwood" box="[1103,1300,294,319]" pageId="3" pageNumber="533" refString="Westwood, J. O. (1874) Descriptions of some new species of exotic Cetoniidae. Transactions of the Entomological Society of London, 4, 477." type="journal article" year="1874">Westwood, 1874</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="B7AC7AD2FFF1FFBEFAEE8A59FE28CA6F" authority="Moeseneder &amp; Hutchinson" authorityName="Moeseneder &amp; Hutchinson" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="pixii" status="sp. nov.">
<emphasis id="42D8DD43FFF1FFBEFAEE8A59FF1CCA6F" italics="true" pageId="3" pageNumber="533">Navigator pixii</emphasis>
Moeseneder &amp; Hutchinson
</taxonomicName>
<taxonomicNameLabel id="59EB6038FFF1FFBEFDC08A34FD5BCA6F" box="[516,654,330,355]" pageId="3" pageNumber="533" rank="species">
<emphasis id="42D8DD43FFF1FFBEFDC08A34FD5BCA6F" bold="true" box="[516,654,330,355]" pageId="3" pageNumber="533">new species</emphasis>
</taxonomicNameLabel>
, and
<taxonomicName id="B7AC7AD2FFF1FFBEFD0F8A32FB1BCA6F" authority="Moeseneder &amp; Hutchinson" authorityName="Moeseneder &amp; Hutchinson" box="[715,1230,330,355]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="interior" status="sp. nov.">
<emphasis id="42D8DD43FFF1FFBEFD0F8A32FC4CCA6F" box="[715,921,332,355]" italics="true" pageId="3" pageNumber="533">Navigator interior</emphasis>
Moeseneder &amp; Hutchinson
</taxonomicName>
<taxonomicNameLabel id="59EB6038FFF1FFBEFB118A34FA8ACA6F" box="[1237,1375,330,355]" pageId="3" pageNumber="533" rank="species">
<emphasis id="42D8DD43FFF1FFBEFB118A34FA8ACA6F" bold="true" box="[1237,1375,330,355]" pageId="3" pageNumber="533">new species</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF1FFBEFF038A10FDABCF5A" pageId="3" pageNumber="533" type="description">
<paragraph id="70130151FFF1FFBEFF038A10FDABCF5A" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="533">
<emphasis id="42D8DD43FFF1FFBEFF038A10FE84CA8B" bold="true" box="[199,337,366,391]" pageId="3" pageNumber="533">Description</emphasis>
. Size
<quantity id="B754ACB4FFF1FFBEFE638A10FD92CA8A" box="[423,583,366,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.89" metricValueMin="0.96" pageId="3" pageNumber="533" unit="mm" value="14.25" valueMax="18.9" valueMin="9.6">9.6–18.9 mm</quantity>
. Form elongate ovate or ovate. Colour black or brown with metallic reflections.
<emphasis id="42D8DD43FFF1FFBEFEDE8AECFE8DCAA7" bold="true" box="[282,344,402,427]" italics="true" pageId="3" pageNumber="533">Head</emphasis>
. Clypeus subquadrate to elongate; lateral margins divergent (
<figureCitation id="E8971DD4FFF1FFBEFBC78AECFB94CAA7" box="[1027,1089,402,427]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
A, detail 1), widest preapically (
<figureCitation id="E8971DD4FFF1FFBEFF5B8AC8FF08CAC3" box="[159,221,438,463]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
A, detail 2;
<figureCitation id="E8971DD4FFF1FFBEFEA18AC8FE76CAC3" box="[357,419,438,463]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
C, detail 4); clypeolateral ridge present; anterior margin shallow arcuate to linear (
<figureCitation id="E8971DD4FFF1FFBEFAFB8AC8FAA8CAC3" box="[1343,1405,438,463]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
A, detail 3); margins gradually inclined to moderate elevations (
<figureCitation id="E8971DD4FFF1FFBEFCA38AA4FC7FCAFF" box="[871,938,474,499]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
A, detail 4); disc coarsely punctate. Frons bearing low, distinct medial elevation; coarsely punctate. Antenna club enlarged in male, longer than length of antennomeres 2–7.
<emphasis id="42D8DD43FFF1FFBEFEBB895CFE01C937" bold="true" box="[383,468,546,571]" italics="true" pageId="3" pageNumber="533">Thorax</emphasis>
. Pronotum weakly transverse, trapezoidal (
<figureCitation id="E8971DD4FFF1FFBEFC1B895CFBF0C937" box="[991,1061,546,571]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
C); basomedian margin weakly concave-linear (
<figureCitation id="E8971DD4FFF1FFBEFE948938FE41C953" box="[336,404,582,607]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
C, detail 1); lateral margins with broad medial angles (
<figureCitation id="E8971DD4FFF1FFBEFBE18938FBBCC953" box="[1061,1129,582,607]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
C, detail 2); disc coarsely punctate (
<figureCitation id="E8971DD4FFF1FFBEFEC28914FE93C98F" box="[262,326,618,643]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
C, detail 3). Scutellum base broad, elongate, finely or coarsely punctate. Elytron posthumeral arch weakly sinuate-linear (
<figureCitation id="E8971DD4FFF1FFBEFE5E89F0FE0EC9AB" box="[410,475,654,679]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
D, detail 1), slightly exposing metacoxa (
<figureCitation id="E8971DD4FFF1FFBEFC7389F0FC2CC9AB" box="[951,1017,654,679]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
D, detail 2), basal half parallel sided (
<figureCitation id="E8971DD4FFF1FFBEFF5B89CCFF35C9C7" box="[159,224,690,715]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
D, detail 3), apical half broadly arcuate to apex (
<figureCitation id="E8971DD4FFF1FFBEFCD789CCFC81C9C7" box="[787,852,690,715]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
D, detail 4); mesepimeron slightly protruding (
<figureCitation id="E8971DD4FFF1FFBEFAAB89CCFF77C9E3" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
D, detail 5); humeral and apical umbones indistinctly raised (
<figureCitation id="E8971DD4FFF1FFBEFC9C89A8FC4DC9E3" box="[856,920,726,751]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
D, details 6, 7); sutural costa distinct, bearing two or more distinct narrow costae; disc coarsely punctate, mostly coalesced, eroding costal margins. Mesometasternal process undeveloped, present as low rounded declivous process between mesocoxae (
<figureCitation id="E8971DD4FFF1FFBEFA868860FA51C83B" box="[1346,1412,798,823]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
F, detail 1).
<emphasis id="42D8DD43FFF1FFBEFEC8883DFE97C856" bold="true" box="[268,322,835,858]" italics="true" pageId="3" pageNumber="533">Legs</emphasis>
. Protibia bidentate or tridentate, denticles elongate; protarsomeres elongate, simple. Mesotibia unidentate. Mesotarsomeres elongate, simple. Metafemur elongate, anterior margin arcuate. Metatibia elongate, broad or narrow, unidentate (
<figureCitation id="E8971DD4FFF1FFBEFE2E88F4FDE2C8AF" box="[490,567,906,931]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Figs. 1</figureCitation>
H–J); internal spur spatulate in female (
<figureCitation id="E8971DD4FFF1FFBEFBC888F4FB9AC8AF" box="[1036,1103,906,931]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
H, detail 1). Metatarsomeres elongate, simple.
<emphasis id="42D8DD43FFF1FFBEFEA488D0FE1BC8CB" bold="true" box="[352,462,942,967]" italics="true" pageId="3" pageNumber="533">Abdomen</emphasis>
. Sternites distinctly convex and distended in female (
<figureCitation id="E8971DD4FFF1FFBEFBF188D0FBA3C8CB" box="[1077,1142,942,967]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
E, detail 1). Metasternum sparsely punctate medially, denser rugulose laterally. Pygidium concentrically rugose, female in normal plane (
<figureCitation id="E8971DD4FFF1FFBEFAAB88ACFF77CF03" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
E, detail 2) to elevated (
<figureCitation id="E8971DD4FFF1FFBEFE748888FE3ACF03" box="[432,495,1014,1039]" captionStart="FIGURE 1" captionStartId="4.[151,250,1487,1509]" captionTargetBox="[179,1419,218,1444]" captionTargetId="figure@4.[150,1436,193,1465]" captionTargetPageId="4" captionText="FIGURE 1. Navigator characters. A—characters of the head (example N. pixii male): A 1 — margins laterally divergent, A 2 — head widest preapically, A 3 — shallow arcuate to linear anterior margin, A 4 — gradually inclined margins, A 5 — shallow preclypeus, A 6 — cross-section, A 7 — lateral declivity absent. B—characters of the head (example N. ruficornis female): B 1 — lateral declivity present, B 2 — deep preclypeus, B 3 — cross-section, B 4 — lateral declivity present. C—characters of the pronotum (example N. pixii male): C 1 — concave linear basolateral margin, C 2 — broad median angle of lateral margins, C 3 — coarsely punctate disc. D—characters of the elytra (example N. pixii male, except D 9): D 1 — weakly sinuate-linear posthumeral arch, D 2 — slightly exposed metacoxa, D 3 — elytral basal half parallel sided, D 4 — apical half of elytra broadly arcuate to apex, D 5 — slightly protruding mesepimeron, D 6 – 7 — indistinctly raised apical umbone, D 8 — visible pygidium D 9 — pygidium not well visible (example N. ruficornis). E—characters of the female abdomen: E 1 — distinctly convex and distended sternites, E 2 — pygidium in normal plane (example N. ruficornis), E 3 — pygidium in elevated plane (example N. pixii). F—character of the abdomen: F 1 — undeveloped mesometasternal process. G—metacoxa posterolateral angle: G 1 — angulate (examples N. pixii and N. ruficornis), G 2 — broadly arcuate (example N. fossor), G 3 — spinose (example N. interior). H – K: characters of the metatibia: H— Navigator fossor female: H 1 — spatulate internal spur, H 2 — unidentate, H 3 — trispinose apex. I— Navigator ruficornis female: I 1 — large, median denticle, I 2 — trispinose apex. J— Navigator interior female: J 1 — small, median denticle. K— Navigator pixii male: K 1 — unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis." httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="3" pageNumber="533">Fig. 1</figureCitation>
E, detail 3). Genitalia (
<figureCitation id="E8971DD4FFF1FFBEFD348888FCE0CF03" box="[752,821,1014,1039]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[313,1284,231,1925]" captionTargetId="figure@5.[248,1336,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Navigator male aedeagi. A— Navigator fossor. B— Navigator pixii. C— Navigator ruficornis. Note: male N. interior are unknown." httpUri="https://zenodo.org/record/254769/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
). Phallobase 1.5 x length of parameres and widest near base of parameres. Parameres narrowing from base; margin sinuate or parallel, widening near apex; dorsal cleft basally acute-obtuse then attenuate to apex.
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF1FFBEFF038F1CFE43CFEB" pageId="3" pageNumber="533" type="diagnosis">
<paragraph id="70130151FFF1FFBEFF038F1CFE43CFEB" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="533">
<emphasis id="42D8DD43FFF1FFBEFF038F1CFE04CF77" bold="true" box="[199,465,1122,1147]" pageId="3" pageNumber="533">Differential diagnosis</emphasis>
. The following combination of characters distinguishes
<taxonomicName id="B7AC7AD2FFF1FFBEFB528F1AFADDCF77" box="[1174,1288,1124,1147]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF1FFBEFB528F1AFADDCF77" box="[1174,1288,1124,1147]" italics="true" pageId="3" pageNumber="533">Navigator</emphasis>
</taxonomicName>
from other
<taxonomicName id="B7AC7AD2FFF1FFBEFF538FF8FEFBCF93" box="[151,302,1158,1183]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="tribe" tribe="Schizorhinini">Schizorhinini</taxonomicName>
genera. Size
<quantity id="B754ACB4FFF1FFBEFE0A8FF8FDBDCF92" box="[462,616,1158,1183]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.89" metricValueMin="0.96" pageId="3" pageNumber="533" unit="mm" value="14.25" valueMax="18.9" valueMin="9.6">9.6–18.9 mm</quantity>
, colour with metallic reflections, clypeal anterior margin nearly linear, dorsal surfaces coarsely punctate, elytral costae distinct and narrow, male antennal club enlarged, female metatibial internal spur spatulate.
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF1FFBEFF038F8AFCE6CC0E" pageId="3" pageNumber="533" type="reference_group">
<paragraph id="70130151FFF1FFBEFF038F8AFF1ACD27" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="533">
<taxonomicName id="B7AC7AD2FFF1FFBEFF038F8AFE5FCE07" ID-CoL="45X23" box="[199,394,1267,1291]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="fossor" status="comb. nov.">
<emphasis id="42D8DD43FFF1FFBEFF038F8AFE5FCE07" box="[199,394,1267,1291]" italics="true" pageId="3" pageNumber="533">Navigator fossor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="59EB6038FFF1FFBEFE538F8CFDBFCE07" box="[407,618,1266,1291]" pageId="3" pageNumber="533">
<emphasis id="42D8DD43FFF1FFBEFE538F8CFDBFCE07" bold="true" box="[407,618,1266,1291]" pageId="3" pageNumber="533">new combination</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="B7AC7AD2FFF1FFBEFD6A8F8AFC96CE07" box="[686,835,1267,1291]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="ruficornis" status="comb. nov.">
<emphasis id="42D8DD43FFF1FFBEFD6A8F8AFC96CE07" box="[686,835,1267,1291]" italics="true" pageId="3" pageNumber="533">N. ruficornis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="59EB6038FFF1FFBEFC958F8CFBF6CE07" box="[849,1059,1266,1291]" pageId="3" pageNumber="533">
<emphasis id="42D8DD43FFF1FFBEFC958F8CFBF6CE07" bold="true" box="[849,1059,1266,1291]" pageId="3" pageNumber="533">new combination</emphasis>
</taxonomicNameLabel>
are transferred from the genus
<taxonomicName id="B7AC7AD2FFF1FFBEFF538E68FEE9CE23" box="[151,316,1302,1327]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFF538E68FEE9CE23" box="[151,316,1302,1327]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
. Here we distinguish
<taxonomicName id="B7AC7AD2FFF1FFBEFDF18E69FD72CE22" ID-CoL="5XXJ" box="[565,679,1303,1326]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF1FFBEFDF18E69FD72CE22" box="[565,679,1303,1326]" italics="true" pageId="3" pageNumber="533">Navigator</emphasis>
</taxonomicName>
from the two remaining
<taxonomicName id="B7AC7AD2FFF1FFBEFC078E68FBBDCE23" box="[963,1128,1302,1327]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFC078E68FBBDCE23" box="[963,1128,1302,1327]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
groups, which are the
<typeStatus id="AF17BFF3FFF1FFBEFAA88E69FA49CE23" box="[1388,1436,1303,1327]" pageId="3" pageNumber="533">type</typeStatus>
species
<taxonomicName id="B7AC7AD2FFF1FFBEFF368E42FDEDCE5F" authority="Macleay, 1871" authorityName="Macleay" authorityYear="1871" box="[242,568,1338,1363]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFF368E42FEA6CE5F" box="[242,371,1338,1363]" italics="true" pageId="3" pageNumber="533">P. hirticeps</emphasis>
(Macleay, 1871)
</taxonomicName>
and all others, which we refer to as
<taxonomicName id="B7AC7AD2FFF1FFBEFC2B8E44FB41CE5F" box="[1007,1172,1338,1363]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFC2B8E44FB41CE5F" box="[1007,1172,1338,1363]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. Specimens of
<taxonomicName id="B7AC7AD2FFF1FFBEFF538E21FEDCCE7A" ID-CoL="5XXJ" box="[151,265,1375,1398]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF1FFBEFF538E21FEDCCE7A" box="[151,265,1375,1398]" italics="true" pageId="3" pageNumber="533">Navigator</emphasis>
</taxonomicName>
are structurally, and based on our observations also behaviourally, similar to
<taxonomicName id="B7AC7AD2FFF1FFBEFBAD8E20FE51CE97" authority="Hutchinson &amp; Moeseneder, 2013" authorityName="Hutchinson &amp; Moeseneder" authorityYear="2013" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFBAD8E20FAC1CE7B" box="[1129,1300,1374,1399]" italics="true" pageId="3" pageNumber="533">Grandaustralis</emphasis>
<bibRefCitation id="143D7CA0FFF1FFBEFADF8E20FE51CE97" author="Hutchinson" pageId="3" pageNumber="533" refString="Hutchinson, P. &amp; Moeseneder, C. H. (2013 a) Chondropyga insignicosta, a new species of Cetoniinae (Coleoptera: Scarabaeidae) from Queensland, Australia. Zootaxa, 3710 (3), 281 - 288. http: // dx. doi. org / 10.11646 / zootaxa. 3710.3.6" type="journal article" year="2013">Hutchinson &amp; Moeseneder, 2013</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B7AC7AD2FFF1FFBEFE4B8EFCFC55CE97" authority="Moeseneder &amp; Hutchinson, 2012" authorityName="Moeseneder &amp; Hutchinson" authorityYear="2012" box="[399,896,1410,1435]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFE4B8EFCFDD7CE97" box="[399,514,1410,1435]" italics="true" pageId="3" pageNumber="533">Octocollis</emphasis>
Moeseneder &amp; Hutchinson, 2012
</taxonomicName>
, and to a lesser degree
<taxonomicName id="B7AC7AD2FFF1FFBEFB478EFCFF06CEB3" authority="Thomson, 1880" authorityName="Thomson" authorityYear="1880" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFB478EFCFAF1CE97" box="[1155,1316,1410,1435]" italics="true" pageId="3" pageNumber="533">Tapinoschema</emphasis>
Thomson, 1880
</taxonomicName>
. The males of these genera possess greatly enlarged antennal clubs. Both sexes of these genera have a coarsely punctate head, pronotum, and elytra; clypeus with anterior margin sinuate to linear; mesometasternal process undeveloped (except moderately developed in
<taxonomicName id="B7AC7AD2FFF1FFBEFCCE8E90FC7ECD0B" box="[778,939,1518,1543]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFCCE8E90FC7ECD0B" box="[778,939,1518,1543]" italics="true" pageId="3" pageNumber="533">Tapinoschema</emphasis>
</taxonomicName>
); and weakly sinuate to linear posthumeral arch.
</paragraph>
<paragraph id="70130151FFF1FFBEFF038D48FCE6CC0E" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="533">
Differences to
<taxonomicName id="B7AC7AD2FFF1FFBEFEAB8D48FD08CD43" authority="Colour" authorityName="Colour" box="[367,733,1590,1615]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFEAB8D48FD53CD43" box="[367,646,1590,1615]" italics="true" pageId="3" pageNumber="533">Pseudoclithria hirticeps:</emphasis>
Colour
</taxonomicName>
with metallic reflections; matt in
<taxonomicName id="B7AC7AD2FFF1FFBEFB9A8D49FB08CD43" box="[1118,1245,1590,1615]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFB9A8D49FB08CD43" box="[1118,1245,1590,1615]" italics="true" pageId="3" pageNumber="533">P. hirticeps</emphasis>
</taxonomicName>
. Elytra with two or more distinct, narrow costae; in
<taxonomicName id="B7AC7AD2FFF1FFBEFDDE8D22FD42CD7F" box="[538,663,1626,1651]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFDDE8D22FD42CD7F" box="[538,663,1626,1651]" italics="true" pageId="3" pageNumber="533">P. hirticeps</emphasis>
</taxonomicName>
with single broad costa. Elytral posthumeral arch sinuate to linear; in
<taxonomicName id="B7AC7AD2FFF1FFBEFF538D01FEC1CD9B" box="[151,276,1662,1687]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFF538D01FEC1CD9B" box="[151,276,1662,1687]" italics="true" pageId="3" pageNumber="533">P. hirticeps</emphasis>
</taxonomicName>
sinuate, exposing abdomen. Mesometasternal process undeveloped; bud-shaped in
<taxonomicName id="B7AC7AD2FFF1FFBEFB7A8D01FAEECD9B" box="[1214,1339,1662,1687]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFB7A8D01FAEECD9B" box="[1214,1339,1662,1687]" italics="true" pageId="3" pageNumber="533">P. hirticeps</emphasis>
</taxonomicName>
. Female pygidium gradually convergent to apex; in
<taxonomicName id="B7AC7AD2FFF1FFBEFD5D8DDAFCC9CDB7" box="[665,796,1698,1723]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF1FFBEFD5D8DDAFCC9CDB7" box="[665,796,1698,1723]" italics="true" pageId="3" pageNumber="533">P. hirticeps</emphasis>
</taxonomicName>
highly transverse with distinctly obtuse apex. Female metatibia internal spur spatulate; in
<emphasis id="42D8DD43FFF1FFBEFDF58DB9FD79CDD2" box="[561,684,1734,1759]" italics="true" pageId="3" pageNumber="533">P. h i r t i c ep s</emphasis>
elongate, parallel. Male parameres (
<figureCitation id="E8971DD4FFF1FFBEFB968DB8FB42CDD3" box="[1106,1175,1734,1759]" captionStart="FIGURE 2" captionStartId="5.[151,250,1971,1993]" captionTargetBox="[313,1284,231,1925]" captionTargetId="figure@5.[248,1336,193,1949]" captionTargetPageId="5" captionText="FIGURE 2. Navigator male aedeagi. A— Navigator fossor. B— Navigator pixii. C— Navigator ruficornis. Note: male N. interior are unknown." httpUri="https://zenodo.org/record/254769/files/figure.png" pageId="3" pageNumber="533">Fig. 2</figureCitation>
) narrowing from base, widening to apex; in
<emphasis id="42D8DD43FFF1FFBEFE478D92FE28CC0F" box="[387,509,1770,1795]" italics="true" pageId="3" pageNumber="533">P. h i r t i ce p s</emphasis>
elongate attenuate to apex.
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF1FFBBFF038C70FC8DC8A9" lastPageId="6" lastPageNumber="536" pageId="3" pageNumber="533" type="description">
<paragraph id="70130151FFF1FFBEFF038C70FDF6CCD7" blockId="3.[151,1437,258,2011]" pageId="3" pageNumber="533">
Differences to
<taxonomicName id="B7AC7AD2FFF1FFBEFEB98C70FDF7CC2B" box="[381,546,1806,1831]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFEB98C70FDF7CC2B" box="[381,546,1806,1831]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species: colour with metallic reflections; matt in
<taxonomicName id="B7AC7AD2FFF1FFBEFB548C70FAE0CC2B" box="[1168,1333,1806,1831]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFB548C70FAE0CC2B" box="[1168,1333,1806,1831]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. Pronotum coarsely punctate; finely punctate in
<taxonomicName id="B7AC7AD2FFF1FFBEFD0E8C4CFCBACC47" box="[714,879,1842,1867]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFD0E8C4CFCBACC47" box="[714,879,1842,1867]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. Posthumeral arch sinuate to linear; in
<taxonomicName id="B7AC7AD2FFF1FFBEFF538C28FEE9CC63" box="[151,316,1878,1903]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFF538C28FEE9CC63" box="[151,316,1878,1903]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. sinuate, often exposing abdomen. Male antennal club greatly enlarged; in
<taxonomicName id="B7AC7AD2FFF1FFBEFB338C28FA49CC63" box="[1271,1436,1878,1903]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFB338C28FA49CC63" box="[1271,1436,1878,1903]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. not distinctly enlarged. Elytra with two distinct, narrow costae; in
<taxonomicName id="B7AC7AD2FFF1FFBEFC1A8C04FB56CC9F" box="[990,1155,1914,1939]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFC1A8C04FB56CC9F" box="[990,1155,1914,1939]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. with single low, wide costa. Legs elongate; in
<taxonomicName id="B7AC7AD2FFF1FFBEFE378CE0FD4DCCBB" box="[499,664,1950,1975]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFE378CE0FD4DCCBB" box="[499,664,1950,1975]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species. legs stout. Female metatibial internal spur spatulate; nonspatulate in
<taxonomicName id="B7AC7AD2FFF1FFBEFEDA8CBCFE16CCD7" box="[286,451,1986,2011]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="533" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF1FFBEFEDA8CBCFE16CCD7" box="[286,451,1986,2011]" italics="true" pageId="3" pageNumber="533">Pseudoclithria</emphasis>
</taxonomicName>
species.
</paragraph>
<caption id="24D351D9FFF6FFB9FF538EB1FD84CCBD" httpUri="https://zenodo.org/record/254768/files/figure.png" pageId="4" pageNumber="534" targetBox="[179,1419,218,1444]" targetPageId="4">
<paragraph id="70130151FFF6FFB9FF538EB1FD84CCBD" blockId="4.[151,1437,1487,1969]" pageId="4" pageNumber="534">
<emphasis id="42D8DD43FFF6FFB9FF538EB1FED8CEE8" bold="true" box="[151,269,1487,1509]" pageId="4" pageNumber="534">FIGURE 1</emphasis>
.
<taxonomicName id="B7AC7AD2FFF6FFB9FEDE8EAEFE54CEE9" box="[282,385,1488,1509]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF6FFB9FEDE8EAEFE54CEE9" box="[282,385,1488,1509]" italics="true" pageId="4" pageNumber="534">Navigator</emphasis>
</taxonomicName>
characters. A—characters of the head (example
<taxonomicName id="B7AC7AD2FFF6FFB9FCB78EAEFC6BCEE9" box="[883,958,1488,1509]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF6FFB9FCB78EAEFC6BCEE9" box="[883,958,1488,1509]" italics="true" pageId="4" pageNumber="534">N. pixii</emphasis>
</taxonomicName>
male): A1—margins laterally divergent, A2— head widest preapically, A3—shallow arcuate to linear anterior margin, A4—gradually inclined margins, A5—shallow preclypeus, A6—cross-section, A7—lateral declivity absent. B—characters of the head (example
<taxonomicName id="B7AC7AD2FFF6FFB9FB448D73FAD7CD2E" box="[1152,1282,1548,1570]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF6FFB9FB448D73FAD7CD2E" box="[1152,1282,1548,1570]" italics="true" pageId="4" pageNumber="534">N. ruficornis</emphasis>
</taxonomicName>
female): B1— lateral declivity present, B2—deep preclypeus, B3—cross-section, B4—lateral declivity present. C—characters of the pronotum (example
<taxonomicName id="B7AC7AD2FFF6FFB9FEA28D34FE64CD53" box="[358,433,1610,1631]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF6FFB9FEA28D34FE64CD53" box="[358,433,1610,1631]" italics="true" pageId="4" pageNumber="534">N. pixii</emphasis>
</taxonomicName>
male): C1—concave linear basolateral margin, C2—broad median angle of lateral margins, C3— coarsely punctate disc. D—characters of the elytra (example
<taxonomicName id="B7AC7AD2FFF6FFB9FD3A8D17FC9DCD72" box="[766,840,1641,1662]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF6FFB9FD3A8D17FC9DCD72" box="[766,840,1641,1662]" italics="true" pageId="4" pageNumber="534">N. pixii</emphasis>
</taxonomicName>
male, except D9): D1—weakly sinuate-linear posthumeral arch, D2—slightly exposed metacoxa, D3—elytral basal half parallel sided, D4—apical half of elytra broadly arcuate to apex, D5—slightly protruding mesepimeron, D6–7—indistinctly raised apical umbone, D8—visible pygidium D9—pygidium not well visible (example
<taxonomicName id="B7AC7AD2FFF6FFB9FE458DBBFDD0CDD6" box="[385,517,1732,1754]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF6FFB9FE458DBBFDD0CDD6" box="[385,517,1732,1754]" italics="true" pageId="4" pageNumber="534">N. ruficornis</emphasis>
</taxonomicName>
). E—characters of the female abdomen: E1—distinctly convex and distended sternites, E2—pygidium in normal plane (example
<taxonomicName id="B7AC7AD2FFF6FFB9FD808D9AFD12CDF5" box="[580,711,1763,1785]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF6FFB9FD808D9AFD12CDF5" box="[580,711,1763,1785]" italics="true" pageId="4" pageNumber="534">N. ruficornis</emphasis>
</taxonomicName>
), E3—pygidium in elevated plane (example
<taxonomicName id="B7AC7AD2FFF6FFB9FB5C8D9AFB31CDF5" box="[1176,1252,1764,1785]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF6FFB9FB5C8D9AFB31CDF5" box="[1176,1252,1764,1785]" italics="true" pageId="4" pageNumber="534">N. pixii</emphasis>
</taxonomicName>
). F—character of the abdomen: F1—undeveloped mesometasternal process. G—metacoxa posterolateral angle: G1—angulate (examples
<taxonomicName id="B7AC7AD2FFF6FFB9FA968C7CFA49CC1B" box="[1362,1436,1794,1815]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF6FFB9FA968C7CFA49CC1B" box="[1362,1436,1794,1815]" italics="true" pageId="4" pageNumber="534">N. pixii</emphasis>
</taxonomicName>
and
<taxonomicName id="B7AC7AD2FFF6FFB9FF028C5FFE99CC3A" box="[198,332,1824,1846]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF6FFB9FF028C5FFE99CC3A" box="[198,332,1824,1846]" italics="true" pageId="4" pageNumber="534">N. ruficornis</emphasis>
</taxonomicName>
), G2—broadly arcuate (example
<taxonomicName id="B7AC7AD2FFF6FFB9FD758C5FFCC4CC3A" box="[689,785,1824,1846]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF6FFB9FD758C5FFCC4CC3A" box="[689,785,1824,1846]" italics="true" pageId="4" pageNumber="534">N. fossor</emphasis>
</taxonomicName>
), G3—spinose (example
<taxonomicName id="B7AC7AD2FFF6FFB9FBE58C5FFB45CC3A" box="[1057,1168,1825,1846]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="interior">
<emphasis id="42D8DD43FFF6FFB9FBE58C5FFB45CC3A" box="[1057,1168,1825,1846]" italics="true" pageId="4" pageNumber="534">N. interior</emphasis>
</taxonomicName>
). H–K: characters of the metatibia: H—
<taxonomicName id="B7AC7AD2FFF6FFB9FEF78C3EFE36CC59" box="[307,483,1855,1877]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF6FFB9FEF78C3EFE36CC59" box="[307,483,1855,1877]" italics="true" pageId="4" pageNumber="534">Navigator fossor</emphasis>
</taxonomicName>
female: H1—spatulate internal spur, H2—unidentate, H3—trispinose apex. I—
<taxonomicName id="B7AC7AD2FFF6FFB9FAF18C3EFF2ECC7F" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF6FFB9FAF18C3EFF2ECC7F" italics="true" pageId="4" pageNumber="534">Navigator ruficornis</emphasis>
</taxonomicName>
female: I1—large, median denticle, I2—trispinose apex. J—
<taxonomicName id="B7AC7AD2FFF6FFB9FCAE8C21FBF0CC78" box="[874,1061,1887,1908]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="interior">
<emphasis id="42D8DD43FFF6FFB9FCAE8C21FBF0CC78" box="[874,1061,1887,1908]" italics="true" pageId="4" pageNumber="534">Navigator interior</emphasis>
</taxonomicName>
female: J1—small, median denticle. K—
<taxonomicName id="B7AC7AD2FFF6FFB9FF068C03FEB4CC9E" box="[194,353,1917,1938]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="534" phylum="Arthropoda" rank="species" species="pixii">
<emphasis id="42D8DD43FFF6FFB9FF068C03FEB4CC9E" box="[194,353,1917,1938]" italics="true" pageId="4" pageNumber="534">Navigator pixii</emphasis>
</taxonomicName>
male: K1—unispinose apex. Note: cross-sections in A and B are from clypeal anterolateral angle to clypeolateral margin, not in transverse axis.
</paragraph>
</caption>
<caption id="24D351D9FFF7FFB8FF538CCDFEA0CCE4" httpUri="https://zenodo.org/record/254769/files/figure.png" pageId="5" pageNumber="535" targetBox="[313,1284,231,1925]" targetPageId="5">
<paragraph id="70130151FFF7FFB8FF538CCDFEA0CCE4" blockId="5.[151,1437,1971,2024]" pageId="5" pageNumber="535">
<emphasis id="42D8DD43FFF7FFB8FF538CCDFEC4CCC4" bold="true" box="[151,273,1971,1993]" pageId="5" pageNumber="535">FIGURE 2</emphasis>
.
<taxonomicName id="B7AC7AD2FFF7FFB8FEE68CCAFE5CCCC5" box="[290,393,1972,1993]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF7FFB8FEE68CCAFE5CCCC5" box="[290,393,1972,1993]" italics="true" pageId="5" pageNumber="535">Navigator</emphasis>
</taxonomicName>
male aedeagi. A—
<taxonomicName id="B7AC7AD2FFF7FFB8FD9E8CCAFCDDCCC5" box="[602,776,1971,1993]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="535" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF7FFB8FD9E8CCAFCDDCCC5" box="[602,776,1971,1993]" italics="true" pageId="5" pageNumber="535">Navigator fossor</emphasis>
</taxonomicName>
. B—
<emphasis id="42D8DD43FFF7FFB8FC808CCAFC3FCCC5" box="[836,1002,1972,1993]" italics="true" pageId="5" pageNumber="535">
<taxonomicName id="B7AC7AD2FFF7FFB8FC808CCAFC33CCC5" box="[836,998,1972,1993]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="535" phylum="Arthropoda" rank="species" species="pixii">Navigator pixii</taxonomicName>
.
</emphasis>
C—
<emphasis id="42D8DD43FFF7FFB8FBDA8CCAFB2ECCC5" box="[1054,1275,1971,1993]" italics="true" pageId="5" pageNumber="535">
<taxonomicName id="B7AC7AD2FFF7FFB8FBDA8CCAFB22CCC5" box="[1054,1271,1971,1993]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="535" phylum="Arthropoda" rank="species" species="ruficornis">Navigator ruficornis</taxonomicName>
.
</emphasis>
Note: male
<taxonomicName id="B7AC7AD2FFF7FFB8FA418CCAFF31CCE4" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="535" phylum="Arthropoda" rank="species" species="interior">
<emphasis id="42D8DD43FFF7FFB8FA418CCAFF31CCE4" italics="true" pageId="5" pageNumber="535">N. interior</emphasis>
</taxonomicName>
are unknown.
</paragraph>
</caption>
<paragraph id="70130151FFF4FFBBFF038BE9FE35CA4C" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="536">
Differences to
<taxonomicName id="B7AC7AD2FFF4FFBBFEAB8BE9FDF7CBBC" authority="Size" authorityName="Size" box="[367,546,151,176]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEAB8BE9FE37CBBC" box="[367,482,151,176]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
: Size
</taxonomicName>
<quantity id="B754ACB4FFF4FFBBFDEE8BE9FD11CBA3" box="[554,708,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.89" metricValueMin="0.96" pageId="6" pageNumber="536" unit="mm" value="14.25" valueMax="18.9" valueMin="9.6">9.6–18.9 mm</quantity>
;
<quantity id="B754ACB4FFF4FFBBFD148BE9FC9FCBA3" box="[720,842,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="6" pageNumber="536" unit="mm" value="17.5" valueMax="19.0" valueMin="16.0">16–19 mm</quantity>
in
<taxonomicName id="B7AC7AD2FFF4FFBBFCB48BE9FC36CBBC" box="[880,995,151,176]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFCB48BE9FC36CBBC" box="[880,995,151,176]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
. Colour with metallic reflections; matt in
<taxonomicName id="B7AC7AD2FFF4FFBBFF738BC2FEFFCBD9" box="[183,298,188,213]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFF738BC2FEFFCBD9" box="[183,298,188,213]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
. Clypeus widest preapically; parallel in
<taxonomicName id="B7AC7AD2FFF4FFBBFCC78BC2FCA3CBD9" box="[771,886,188,213]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFCC78BC2FCA3CBD9" box="[771,886,188,213]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
. Pronotum widest basally; in
<taxonomicName id="B7AC7AD2FFF4FFBBFB118BC2FA9DCBD9" box="[1237,1352,188,213]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFB118BC2FA9DCBD9" box="[1237,1352,188,213]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
widest medially. Elytral costae distinct, narrow; indistinct in
<taxonomicName id="B7AC7AD2FFF4FFBBFCE18BA1FC4DCBF4" box="[805,920,223,248]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFCE18BA1FC4DCBF4" box="[805,920,223,248]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
. Female metatibial internal spur spatulate; unknown in female
<taxonomicName id="B7AC7AD2FFF4FFBBFEB88A7AFE3ACA11" box="[380,495,260,285]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEB88A7AFE3ACA11" box="[380,495,260,285]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
. Parameres narrowing from base, broadening preapically; in
<taxonomicName id="B7AC7AD2FFF4FFBBFB698A7AFAF5CA11" box="[1197,1312,260,285]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFB698A7AFAF5CA11" box="[1197,1312,260,285]" italics="true" pageId="6" pageNumber="536">Octocollis</emphasis>
</taxonomicName>
parameres broad with apical appendage.
</paragraph>
<paragraph id="70130151FFF4FFBBFF038A32FC53CAC3" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="536">
Differences to
<taxonomicName id="B7AC7AD2FFF4FFBBFEB18A32FDB1CA69" authority="Size" authorityName="Size" box="[373,612,332,357]" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEB18A32FDF5CA69" box="[373,544,332,357]" italics="true" pageId="6" pageNumber="536">Grandaustralis</emphasis>
: Size
</taxonomicName>
<quantity id="B754ACB4FFF4FFBBFDAB8A32FCDECA68" box="[623,779,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.89" metricValueMin="0.96" pageId="6" pageNumber="536" unit="mm" value="14.25" valueMax="18.9" valueMin="9.6">9.6–18.9 mm</quantity>
;
<quantity id="B754ACB4FFF4FFBBFCDF8A32FC17CA68" box="[795,962,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.55" metricValueMin="1.85" pageId="6" pageNumber="536" unit="mm" value="22.0" valueMax="25.5" valueMin="18.5">18.5–25.5 mm</quantity>
in
<taxonomicName id="B7AC7AD2FFF4FFBBFC2A8A32FB4CCA69" box="[1006,1177,332,357]" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFC2A8A32FB4CCA69" box="[1006,1177,332,357]" italics="true" pageId="6" pageNumber="536">Grandaustralis</emphasis>
</taxonomicName>
. Colour with metallic reflections; matt in
<taxonomicName id="B7AC7AD2FFF4FFBBFEBD8A11FDF1CA84" box="[377,548,367,392]" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEBD8A11FDF1CA84" box="[377,548,367,392]" italics="true" pageId="6" pageNumber="536">Grandaustralis</emphasis>
</taxonomicName>
. Elytra with two distinct narrow costae; indistinct in
<taxonomicName id="B7AC7AD2FFF4FFBBFB4A8A11FAECCA84" box="[1166,1337,367,392]" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFB4A8A11FAECCA84" box="[1166,1337,367,392]" italics="true" pageId="6" pageNumber="536">Grandaustralis</emphasis>
</taxonomicName>
. Female metatibial internal spur spatulate; parallel in
<taxonomicName id="B7AC7AD2FFF4FFBBFD008AEAFCBACAA1" box="[708,879,404,429]" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFD008AEAFCBACAA1" box="[708,879,404,429]" italics="true" pageId="6" pageNumber="536">Grandaustralis</emphasis>
</taxonomicName>
. Parameres narrowing from base, broadening preapically; in
<taxonomicName id="B7AC7AD2FFF4FFBBFEFB8AC9FE3FCADC" box="[319,490,439,464]" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEFB8AC9FE3FCADC" box="[319,490,439,464]" italics="true" pageId="6" pageNumber="536">Grandaustralis</emphasis>
</taxonomicName>
narrow at base, equal width to apex.
</paragraph>
<paragraph id="70130151FFF4FFBBFF038AA2FC34C9AB" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="536">
Differences to
<taxonomicName id="B7AC7AD2FFF4FFBBFEBD8AA2FD8BCAF9" authority="Size" authorityName="Size" box="[377,606,476,501]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEBD8AA2FDCFCAF9" box="[377,538,476,501]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
: Size
</taxonomicName>
<quantity id="B754ACB4FFF4FFBBFDA88AA2FCDFCAF8" box="[620,778,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.425" metricValueMax="1.89" metricValueMin="0.96" pageId="6" pageNumber="536" unit="mm" value="14.25" valueMax="18.9" valueMin="9.6">9.6–18.9 mm</quantity>
; 20.3–34.0 mm in
<taxonomicName id="B7AC7AD2FFF4FFBBFC378AA2FB41CAF9" box="[1011,1172,476,501]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFC378AA2FB41CAF9" box="[1011,1172,476,501]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
. Colour with metallic reflections; glossy in
<emphasis id="42D8DD43FFF4FFBBFE518A81FDEBC914" box="[405,574,511,536]" italics="true" pageId="6" pageNumber="536">
<taxonomicName id="B7AC7AD2FFF4FFBBFE518A81FDEFC914" box="[405,570,511,536]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">Tapinoschema</taxonomicName>
.
</emphasis>
Pronotal shape weakly transverse; in
<taxonomicName id="B7AC7AD2FFF4FFBBFBC78A81FB71C914" box="[1027,1188,511,536]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFBC78A81FB71C914" box="[1027,1188,511,536]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
distinctly transverse. Mesepimeron slightly exposed; in
<taxonomicName id="B7AC7AD2FFF4FFBBFDF6895AFD06C931" box="[562,723,548,573]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFDF6895AFD06C931" box="[562,723,548,573]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
distinctly exposed. Elytra with two distinct, narrow costae; indistinct in
<taxonomicName id="B7AC7AD2FFF4FFBBFEE68939FE16C96C" box="[290,451,583,608]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEE68939FE16C96C" box="[290,451,583,608]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
. Mesometasternal process undeveloped; in
<taxonomicName id="B7AC7AD2FFF4FFBBFC6F8939FB99C96C" box="[939,1100,583,608]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFC6F8939FB99C96C" box="[939,1100,583,608]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
parallel or attenuate elongate. Female metatibia internal spur spatulate; parallel in
<taxonomicName id="B7AC7AD2FFF4FFBBFD2B8912FC45C989" box="[751,912,620,645]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFD2B8912FC45C989" box="[751,912,620,645]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
. Parameres narrowing from base, broadening preapically; in
<taxonomicName id="B7AC7AD2FFF4FFBBFEFB89F1FE35C9A4" box="[319,480,655,680]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFEFB89F1FE35C9A4" box="[319,480,655,680]" italics="true" pageId="6" pageNumber="536">Tapinoschema</emphasis>
</taxonomicName>
narrow at base, broadening to widest at apex.
</paragraph>
<paragraph id="70130151FFF4FFBBFF0389CAFC8DC8A9" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="536">
Species examined:
<taxonomicName id="B7AC7AD2FFF4FFBBFE6D89CAFD6EC9C1" box="[425,699,692,717]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="hirticeps">
<emphasis id="42D8DD43FFF4FFBBFE6D89CAFD6EC9C1" box="[425,699,692,717]" italics="true" pageId="6" pageNumber="536">Pseudoclithria hirticeps</emphasis>
</taxonomicName>
male,
<taxonomicName id="B7AC7AD2FFF4FFBBFCD489CBFB8AC9C0" authority="Macleay, 1871" authorityName="Macleay" authorityYear="1871" box="[784,1119,692,717]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="mastersii">
<emphasis id="42D8DD43FFF4FFBBFCD489CBFC4DC9C0" box="[784,920,693,716]" italics="true" pageId="6" pageNumber="536">P. mastersii</emphasis>
(Macleay, 1871)
</taxonomicName>
female,
<taxonomicName id="B7AC7AD2FFF4FFBBFB0E89CBFF0DC9FC" authority="Janson, 1874" authorityName="Janson" authorityYear="1874" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="maura">
<emphasis id="42D8DD43FFF4FFBBFB0E89CBFAE6C9C0" box="[1226,1331,693,716]" italics="true" pageId="6" pageNumber="536">P. maura</emphasis>
(Janson, 1874)
</taxonomicName>
male and female,
<taxonomicName id="B7AC7AD2FFF4FFBBFE6D89A7FD68C9FC" authority="Janson, 1889" authorityName="Janson" authorityYear="1889" box="[425,701,727,752]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="adusta">
<emphasis id="42D8DD43FFF4FFBBFE6D89A7FDC5C9FC" box="[425,528,727,752]" italics="true" pageId="6" pageNumber="536">P. adusta</emphasis>
(Janson, 1889)
</taxonomicName>
male,
<taxonomicName id="B7AC7AD2FFF4FFBBFCCE89A7FC3AC9FC" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" box="[778,1007,727,752]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="dejecta">
<emphasis id="42D8DD43FFF4FFBBFCCE89A7FCA2C9FC" box="[778,887,727,752]" italics="true" pageId="6" pageNumber="536">P. dejecta</emphasis>
<bibRefCitation id="143D7CA0FFF4FFBBFCBB89A6FC3AC9FC" author="Lea" box="[895,1007,728,752]" pageId="6" pageNumber="536" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
</taxonomicName>
male,
<taxonomicName id="B7AC7AD2FFF4FFBBFBF889A7FA88C9FC" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" box="[1084,1373,727,752]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="erythroptera">
<emphasis id="42D8DD43FFF4FFBBFBF889A7FB33C9FC" box="[1084,1254,727,752]" italics="true" pageId="6" pageNumber="536">P. erythroptera</emphasis>
<bibRefCitation id="143D7CA0FFF4FFBBFB2989A6FA88C9FC" author="Lea" box="[1261,1373,728,752]" pageId="6" pageNumber="536" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
</taxonomicName>
male (from MNV image),
<taxonomicName id="B7AC7AD2FFF4FFBBFE428983FD40C818" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" box="[390,661,764,789]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="anchoralis">
<emphasis id="42D8DD43FFF4FFBBFE428983FDCEC819" box="[390,539,764,789]" italics="true" pageId="6" pageNumber="536">P. anchoralis</emphasis>
<bibRefCitation id="143D7CA0FFF4FFBBFDE08983FD40C818" author="Lea" box="[548,661,764,788]" pageId="6" pageNumber="536" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
</taxonomicName>
male and female,
<taxonomicName id="B7AC7AD2FFF4FFBBFCAE8983FB47C818" authority="Schaum, 1848" authorityName="Schaum" authorityYear="1848" box="[874,1170,764,789]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="42D8DD43FFF4FFBBFCAE8983FC00C818" box="[874,981,765,788]" italics="true" pageId="6" pageNumber="536">P. rugosa</emphasis>
(Schaum, 1848)
</taxonomicName>
male,
<taxonomicName id="B7AC7AD2FFF4FFBBFB248983FF1AC834" authority="Lea, 1914" authorityName="Lea" authorityYear="1914" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="kershawi">
<emphasis id="42D8DD43FFF4FFBBFB248983FAB7C819" box="[1248,1378,764,789]" italics="true" pageId="6" pageNumber="536">P. kershawi</emphasis>
<bibRefCitation id="143D7CA0FFF4FFBBFAAF8983FF1AC834" author="Lea" pageId="6" pageNumber="536" refString="Lea, A. M. (1914) Notes on Australian Cetonides, with a list of species and descriptions of some new ones. Transactions of the Royal Society of South Australia, 38, 132 - 218." type="journal article" year="1914">Lea, 1914</bibRefCitation>
</taxonomicName>
male (from MNV image), seven undescribed
<taxonomicName id="B7AC7AD2FFF4FFBBFD248861FC50C834" box="[736,901,799,824]" class="Insecta" family="Cetoniidae" genus="Pseudoclithria" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="42D8DD43FFF4FFBBFD248861FC50C834" box="[736,901,799,824]" italics="true" pageId="6" pageNumber="536">Pseudoclithria</emphasis>
</taxonomicName>
species (C.H.M. &amp; P.M.H., unpublished data),
<taxonomicName id="B7AC7AD2FFF4FFBBFF53883AFD85C850" authority="Janson, 1874" authorityName="Janson" authorityYear="1874" box="[151,592,836,861]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="digglesii">
<emphasis id="42D8DD43FFF4FFBBFF53883AFE75C851" box="[151,416,836,861]" italics="true" pageId="6" pageNumber="536">Tapinoschema digglesii</emphasis>
(Janson, 1874)
</taxonomicName>
male and female,
<taxonomicName id="B7AC7AD2FFF4FFBBFCE0883BFB90C851" authority="Macleay, 1863" authorityName="Macleay" authorityYear="1863" box="[804,1093,836,861]" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="impar">
<emphasis id="42D8DD43FFF4FFBBFCE0883BFC57C850" box="[804,898,837,860]" italics="true" pageId="6" pageNumber="536">T. impar</emphasis>
(
<bibRefCitation id="143D7CA0FFF4FFBBFC50883AFBE8C851" author="Macleay" box="[916,1085,836,861]" pageId="6" pageNumber="536" refString="Macleay, W. J. (1863) Descriptions of twenty new species of Australian Coleoptera, belonging to the families Cicindelidae and Cetoniidae. Transactions of the Royal Society of New South Wales, 1, 9 - 21." type="journal article" year="1863">Macleay, 1863</bibRefCitation>
)
</taxonomicName>
male and female,
<taxonomicName id="B7AC7AD2FFF4FFBBFADE883BFE96C88C" authority="Janson, 1874" authorityName="Janson" authorityYear="1874" class="Insecta" family="Cetoniidae" genus="Tapinoschema" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="lacunosa">
<emphasis id="42D8DD43FFF4FFBBFADE883BFA49C851" box="[1306,1436,836,861]" italics="true" pageId="6" pageNumber="536">T. lacunosa</emphasis>
(Janson, 1874)
</taxonomicName>
male and female,
<taxonomicName id="B7AC7AD2FFF4FFBBFDE98819FB42C88C" authority="Moeseneder &amp; Hutchinson, 2012" authorityName="Moeseneder &amp; Hutchinson" authorityYear="2012" box="[557,1175,871,896]" class="Insecta" family="Cetoniidae" genus="Octocollis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="42D8DD43FFF4FFBBFDE98819FCD5C88C" box="[557,768,871,896]" italics="true" pageId="6" pageNumber="536">Octocollis setosus</emphasis>
Moeseneder &amp; Hutchinson, 2012
</taxonomicName>
male,
<taxonomicName id="B7AC7AD2FFF4FFBBFB358819FD45C8A8" authority="Hutchinson &amp; Moeseneder, 2013" authorityName="Hutchinson &amp; Moeseneder" authorityYear="2013" class="Insecta" family="Cetoniidae" genus="Grandaustralis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="boomerang">
<emphasis id="42D8DD43FFF4FFBBFB358819FEC2C8A9" italics="true" pageId="6" pageNumber="536">Grandaustralis boomerang</emphasis>
<bibRefCitation id="143D7CA0FFF4FFBBFED988F2FD45C8A8" author="Hutchinson" box="[285,656,908,933]" pageId="6" pageNumber="536" refString="Hutchinson, P. &amp; Moeseneder, C. H. (2013 a) Chondropyga insignicosta, a new species of Cetoniinae (Coleoptera: Scarabaeidae) from Queensland, Australia. Zootaxa, 3710 (3), 281 - 288. http: // dx. doi. org / 10.11646 / zootaxa. 3710.3.6" type="journal article" year="2013">Hutchinson &amp; Moeseneder, 2013</bibRefCitation>
</taxonomicName>
male and female.
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF4FFBBFF0388D1FD28CF54" pageId="6" pageNumber="536" type="etymology">
<paragraph id="70130151FFF4FFBBFF0388D1FD28CF54" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="536">
<emphasis id="42D8DD43FFF4FFBBFF0388D1FE93C8C4" bold="true" box="[199,326,943,968]" pageId="6" pageNumber="536">Etymology</emphasis>
.
<taxonomicName id="B7AC7AD2FFF4FFBBFE9288CFFE1DC8C4" box="[342,456,945,968]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF4FFBBFE9288CFFE1DC8C4" box="[342,456,945,968]" italics="true" pageId="6" pageNumber="536">Navigator</emphasis>
</taxonomicName>
males, in the species where males are known, have enlarged antennae that may have developed to such a size to assist them in locating females. Hence we chose a name for the genus that is descriptive of their searching behaviour. We assume that males, using their enlarged antennae, follow pheromones released by the females and navigate to the source.
<taxonomicName id="B7AC7AD2FFF4FFBBFD8B8F63FD14CF38" box="[591,705,1053,1076]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF4FFBBFD8B8F63FD14CF38" box="[591,705,1053,1076]" italics="true" pageId="6" pageNumber="536">Navigator</emphasis>
</taxonomicName>
is the Latin noun for an entity that directs itself or a vessel over a planned course. The gender of
<taxonomicName id="B7AC7AD2FFF4FFBBFE368F3FFDB1CF54" box="[498,612,1089,1112]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF4FFBBFE368F3FFDB1CF54" box="[498,612,1089,1112]" italics="true" pageId="6" pageNumber="536">Navigator</emphasis>
</taxonomicName>
is masculine.
</paragraph>
</subSubSection>
<subSubSection id="38B652DAFFF4FFBBFF038F1AFCE7CFC8" pageId="6" pageNumber="536" type="discussion">
<paragraph id="70130151FFF4FFBBFF038F1AFCE7CFC8" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="536">
<emphasis id="42D8DD43FFF4FFBBFF038F1AFEE1CF71" bold="true" box="[199,308,1124,1149]" pageId="6" pageNumber="536">Remarks</emphasis>
. We assign
<taxonomicName id="B7AC7AD2FFF4FFBBFE788F1BFDFBCF70" box="[444,558,1125,1148]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="530" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="42D8DD43FFF4FFBBFE788F1BFDFBCF70" box="[444,558,1125,1148]" italics="true" pageId="6" pageNumber="536">Navigator</emphasis>
</taxonomicName>
to the tribe
<taxonomicName id="B7AC7AD2FFF4FFBBFD728F1AFC98CF71" box="[694,845,1124,1149]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="tribe" tribe="Schizorhinini">Schizorhinini</taxonomicName>
and, within it, to the subtribe Schizorhinina. In very old specimens of
<taxonomicName id="B7AC7AD2FFF4FFBBFE9F8FF7FE15CFAC" box="[347,448,1160,1184]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="42D8DD43FFF4FFBBFE9F8FF7FE15CFAC" box="[347,448,1160,1184]" italics="true" pageId="6" pageNumber="536">N. fossor</emphasis>
</taxonomicName>
and
<taxonomicName id="B7AC7AD2FFF4FFBBFE328FF7FD50CFAC" box="[502,645,1160,1184]" class="Insecta" family="Scarabaeidae" genus="Navigator" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="536" phylum="Arthropoda" rank="species" species="ruficornis">
<emphasis id="42D8DD43FFF4FFBBFE328FF7FD50CFAC" box="[502,645,1160,1184]" italics="true" pageId="6" pageNumber="536">N. ruficornis</emphasis>
</taxonomicName>
there is a loss of metallic reflections on all surfaces and only the black or brown base colours are retained (compare
<figureCitation id="E8971DD4FFF4FFBBFD558FD2FD08CFC9" box="[657,733,1196,1221]" captionStart="FIGURE 6" captionStartId="13.[151,250,867,889]" captionTargetBox="[160,1429,198,829]" captionTargetId="figure@13.[151,1435,193,846]" captionTargetPageId="13" captionText="FIGURE 6. Navigator ruficornis lectotype, male. Image © MNHN, A. Mantilleri." httpUri="https://zenodo.org/record/254773/files/figure.png" pageId="6" pageNumber="536">Figs. 6</figureCitation>
and
<figureCitation id="E8971DD4FFF4FFBBFCD08FD2FCF1CFC8" box="[788,804,1196,1220]" captionStart="FIGURE 9" captionStartId="16.[151,250,1576,1598]" captionTargetBox="[158,1429,847,1550]" captionTargetId="figure@16.[151,1435,841,1555]" captionTargetPageId="16" captionText="FIGURE 9. Navigator pixii holotype, male." httpUri="https://zenodo.org/record/254776/files/figure.png" pageId="6" pageNumber="536">9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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