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<document id="4EE3F4C10B153A583AE01F6232E680AF" ID-CLB-Dataset="39229" ID-DOI="10.11646/zootaxa.4059.1.9" ID-GBIF-Dataset="3ed35bba-4eb0-454e-970d-b0caabfc8c4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="238194" ID-ZooBank="9A0F1819-753E-471E-81F3-06A9719223D8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461361189615" checkinUser="plazi" docAuthor="Zwick, Peter" docDate="2015" docId="0E0C0B57FFFDFFD32BBC275BF819FE72" docLanguage="en" docName="zt04059p180.pdf" docOrigin="Zootaxa 4059 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Madenemura culminalis Paulian 1951" docType="treatment" docVersion="9" lastPageNumber="178" masterDocId="F235732FFFF5FFDA2B2B241FFD55FFDE" masterDocTitle="Notes on Madagascan Stoneflies (Plecoptera: Notonemouridae)" masterLastPageNumber="180" masterPageNumber="169" pageNumber="177" updateTime="1698661804956" updateUser="ExternalLinkService">
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<mods:title id="7B138A42E3063CCA1F08983D93CDE97D">Notes on Madagascan Stoneflies (Plecoptera: Notonemouridae)</mods:title>
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<mods:namePart id="72E976519FE81AB6FF11D38987D142C5">Zwick, Peter</mods:namePart>
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<mods:typeOfResource id="B1677ED87D4CDD93FD841C8A84A167D5">text</mods:typeOfResource>
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<mods:title id="F97230E48C225DFD255AC132A2B9BB38">Zootaxa</mods:title>
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<mods:date id="D763EF9483B9F5303D4D96941DBB3751">2015</mods:date>
<mods:detail id="FD9E8C22E8A66FBCE3362652E4A73C69" type="volume">
<mods:number id="7AE2C1970975889D7A2B3CDC037176BD">4059</mods:number>
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<mods:classification id="5DB98AD54DCF4D50B33675B66BD57B76">journal article</mods:classification>
<mods:identifier id="C7D30BF1968D54B3670A476730968C20" type="CLB-Dataset">39229</mods:identifier>
<mods:identifier id="04245E30CBD42AC03B181FAF5A3E13A9" type="DOI">10.11646/zootaxa.4059.1.9</mods:identifier>
<mods:identifier id="1F7B2476D08ABC009BD99228F2EFCC72" type="GBIF-Dataset">3ed35bba-4eb0-454e-970d-b0caabfc8c4b</mods:identifier>
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<treatment id="0E0C0B57FFFDFFD32BBC275BF819FE72" ID-DOI="http://doi.org/10.5281/zenodo.6100646" ID-GBIF-Taxon="119648562" ID-Zenodo-Dep="6100646" LSID="urn:lsid:plazi:treatment:0E0C0B57FFFDFFD32BBC275BF819FE72" httpUri="http://treatment.plazi.org/id/0E0C0B57FFFDFFD32BBC275BF819FE72" lastPageId="9" lastPageNumber="178" pageId="8" pageNumber="177">
<subSubSection id="CEBFE9CAFFFDFFD22BBC275BFF73FC1A" pageId="8" pageNumber="177" type="nomenclature">
<paragraph id="861ABA41FFFDFFD22BBC275BFF24FC80" blockId="8.[151,625,836,895]" box="[151,625,836,862]" pageId="8" pageNumber="177">
<heading id="DD520D2DFFFDFFD22BBC275BFF24FC80" bold="true" box="[151,625,836,862]" fontSize="11" level="1" pageId="8" pageNumber="177" reason="1">
<taxonomicName id="41A5C1C2FFFDFFD22BBC275BFF24FC80" ID-CoL="6QRN4" authority="Paulian, 1951" authorityName="Paulian" authorityYear="1951" box="[151,625,836,862]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="culminalis">
<emphasis id="B4D16653FFFDFFD22BBC275BFF24FC80" bold="true" box="[151,625,836,862]" pageId="8" pageNumber="177">
<emphasis id="B4D16653FFFDFFD22BBC275BFC95FC80" bold="true" box="[151,448,836,862]" italics="true" pageId="8" pageNumber="177">Madenemura culminalis</emphasis>
<bibRefCitation id="E234C7B0FFFDFFD22AEC275BFF24FC80" author="Paulian" box="[455,625,836,862]" pageId="8" pageNumber="177" refString="Paulian, R. (1951) Faune des eaux douces de Madagascar. Plecopteres et Megalopteres. Memoires de l'Institut Scientifique de Madagascar, (A), 6, 53 - 61." type="journal article" year="1951">Paulian, 1951</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="861ABA41FFFDFFD22BBC2779FCC8FCA1" blockId="8.[151,625,836,895]" box="[151,413,870,895]" pageId="8" pageNumber="177">
(
<figureCitation id="1E9EA6C4FFFDFFD22BB42779FDBBFCA1" box="[159,238,870,895]" captionStart="FIGURES 1 – 9" captionStartId="4.[151,264,1828,1851]" captionTargetBox="[182,1435,205,1808]" captionTargetId="figure@4.[159,1444,195,1821]" captionTargetPageId="4" captionText="FIGURES 1 – 9. Madenemura spp. 1, Madenemura sp., wings, after a photograph; 2, M. culminalis, male abdominal tip, lateral; 3 – 9, M. andringitrensis: 3, detached and opened ventral portion of abdominal segment 9, oblique dorsal view; 4, the detached paraproct body with the retractor muscles; 5, female sternite 8 plus 9, in situ; 6, female sternite 9, detached; 7, dorsal view of center of male T 10 with epiproct; 8, caudal part of baseplate and hook of epiproct in side view; 9, the isolated paraproct body, dorsal view. 3 and 4 not to scale, 5, 6 and 7, 8 to the same scale, respectively. bb, basal bar; bp, epiproct baseplate; ce, cercus; gp, gonopore; mu, muscles; pb, paraproct body; rm, retractor muscle; S, T, sternite and tergite, respectively, with number of corresponding segment; sd, sperm duct; sp, sperm pump; vl, ventral lobe." httpUri="https://zenodo.org/record/238195/files/figure.png" pageId="8" pageNumber="177">Figs. 2</figureCitation>
,
<figureCitation id="1E9EA6C4FFFDFFD22BD12778FC16FCA1" box="[250,323,871,895]" captionStart="FIGURES 10 – 17" captionStartId="5.[151,264,1816,1839]" captionTargetBox="[170,1433,206,1765]" captionTargetId="figure@5.[156,1441,193,1783]" captionTargetPageId="5" captionText="FIGURES 10 – 17, Madenemura spp., males. 10 – 12, M. descarpentriesi: 10, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 11, dorsoventral view of slide-mounted paraproct body; 12, apex of baseplate and epiproct hook, lateral view. 13, 14, M. culminalis: 13, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 14, dorsoventral view of slide-mounted paraproct body; 15, apex of baseplate and epiproct hook, lateral view. 16, 17, Madenemura sp., perhaps sp. 3 of Paulian (1959): 16, abdominal tip, lateral, semidiagrammatic, paraproct body shaded, arrow points at raised antecosta 10, inset below shows caudal edge of T 10 in dorsal view; 17, epiproct, lateral. 12 and 15 – 17 not to scale." httpUri="https://zenodo.org/record/238196/files/figure.png" pageId="8" pageNumber="177">13–17</figureCitation>
,
<figureCitation id="1E9EA6C4FFFDFFD22A652778FCC0FCA1" box="[334,405,871,895]" captionStart="FIGURES 18 – 24" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[184,1393,201,1924]" captionTargetId="figure@6.[175,1406,193,1932]" captionTargetPageId="6" captionText="FIGURES 18 – 24, Madenemura spp., female abdominal tips, in lateral (even numbers) and ventral (uneven numbers) view; after photographs, semidiagrammatic: 18, 19: M. andringitrensis. 20, 21: M. descarpentriesi. 22, 23: M. culminalis; 24: M. pauliani sp. n. (holotype). Same scale for all." httpUri="https://zenodo.org/record/238197/files/figure.png" pageId="8" pageNumber="177">22, 23</figureCitation>
)
</paragraph>
<paragraph id="861ABA41FFFDFFD22BBC27B0FF73FC1A" blockId="8.[151,550,942,964]" box="[151,550,942,964]" pageId="8" pageNumber="177">
=?
<taxonomicName id="41A5C1C2FFFDFFD22B9627B1FC11FC1A" box="[189,324,942,964]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B4D16653FFFDFFD22B9627B1FC11FC1A" box="[189,324,942,964]" italics="true" pageId="8" pageNumber="177">Madenemura</emphasis>
</taxonomicName>
sp. 3 of
<bibRefCitation id="E234C7B0FFFDFFD22AB727B1FF73FC1A" author="Paulian" box="[412,550,942,964]" pageId="8" pageNumber="177" refString="Paulian, R. (1959) Recherches sur les insectes d'importance biologique a Madagascar. XXXIII. Nouveaux Plecopteres malgaches. Memoires de l'Institut Scientifique de Madagascar, (E), 11, 9 - 16." type="journal article" year="1959">Paulian, 1959</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="CEBFE9CAFFFDFFD22BBC27EEF970F994" pageId="8" pageNumber="177" type="materials_examined">
<paragraph id="861ABA41FFFDFFD22BBC27EEFF4BFA00" blockId="8.[151,1437,1009,2006]" pageId="8" pageNumber="177">
<emphasis id="B4D16653FFFDFFD22BBC27EEFC25FBD4" bold="true" box="[151,368,1009,1034]" pageId="8" pageNumber="177">Material studied.</emphasis>
<collectingCountry id="FEB2FAD1FFFDFFD22AA827EEFF59FBD4" box="[387,524,1009,1034]" name="Madagascar" pageId="8" pageNumber="177">Madagascar</collectingCountry>
, Ranomafana National Pk., Mariavaratra stream, 2115.629'S, 4725.149'E; emergence trap, J.Benstead:
<date id="F21B9C81FFFDFFD22AF7200AFF09FBF3" box="[476,604,1045,1069]" pageId="8" pageNumber="177" value="1997-11-25">25.11.1997</date>
,
<specimenCount id="90A371C8FFFDFFD22943200AFFDBFBF3" box="[616,654,1045,1069]" pageId="8" pageNumber="177" type="generic">2♂</specimenCount>
;
<date id="F21B9C81FFFDFFD229B0200AFE49FBF3" box="[667,796,1045,1069]" pageId="8" pageNumber="177" value="1997-12-16">16.12.1997</date>
,
<specimenCount id="90A371C8FFFDFFD22802200AFE1BFBF3" box="[809,846,1045,1069]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
;
<date id="F21B9C81FFFDFFD22877200AFE89FBF3" box="[860,988,1045,1069]" pageId="8" pageNumber="177" value="1998-10-16">16.10.1998</date>
,
<specimenCount id="90A371C8FFFDFFD228C2200AF95AFBF3" box="[1001,1039,1045,1069]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
;
<date id="F21B9C81FFFDFFD22F37200AF9C9FBF3" box="[1052,1180,1045,1069]" pageId="8" pageNumber="177" value="1998-11-27">27.11.1998</date>
,
<specimenCount id="90A371C8FFFDFFD22F82200AF99BFBF3" box="[1193,1230,1045,1069]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
;
<specimenCount id="90A371C8FFFDFFD22FF7200AF857FBF3" box="[1244,1282,1045,1069]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
:
<collectingCountry id="FEB2FAD1FFFDFFD22E24200AF8CDFBF3" box="[1295,1432,1045,1069]" name="Madagascar" pageId="8" pageNumber="177">Madagascar</collectingCountry>
, Prov. Fianarantsoa,
<quantity id="415D17A4FFFDFFD22AAF2026FCE0FB8C" box="[388,437,1081,1106]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="177" unit="km" value="7.0">7km</quantity>
W Ranomanfana
<quantity id="415D17A4FFFDFFD229BC2025FF8FFB8C" box="[663,730,1082,1106]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="177" unit="m" value="900.0">900m</quantity>
, 14.–
<date id="F21B9C81FFFDFFD228312025FECAFB8C" box="[794,927,1082,1106]" pageId="8" pageNumber="177" value="1990-03-19">19.03.1990</date>
, W.E.Steiner \ Malaise trap across stream montane rain forest (
<collectionCode id="E0B42284FFFDFFD22AA62041FCB1FBA8" LSID="urn:lsid:biocol.org:col:34871" box="[397,484,1118,1142]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="177">USNM</collectionCode>
).
<specimenCount id="90A371C8FFFDFFD22AD72041FF74FBA8" box="[508,545,1118,1142]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
: Preparation 0 194,
<date id="F21B9C81FFFDFFD228212041FE38FBA8" box="[778,877,1118,1142]" pageId="8" pageNumber="177" value="1994-03-29">29/03/94</date>
Piège lumineux soir Bassin: Ambre 1 Riv. Pte Cascade Local: Camp Base WWF [
<collectionCode id="E0B42284FFFDFFD2296A209DFF2AFB47" box="[577,639,1154,1177]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="8" pageNumber="177">MZL</collectionCode>
, received on slide]. 1♀:
<collectingCountry id="FEB2FAD1FFFDFFD22894209DF9DEFB44" box="[959,1163,1154,1178]" name="Madagascar" pageId="8" pageNumber="177">MADAGASCAR</collectingCountry>
: Antsiranana R.N.I. de Marojejy
<quantity id="415D17A4FFFDFFD22A3E20BAFC3CFB60" box="[277,361,1189,1214]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="177" unit="km" value="10.5">10.5km</quantity>
NW Manantenina
<date id="F21B9C81FFFDFFD2295B20BAFFCBFB60" box="[624,670,1189,1214]" pageId="8" pageNumber="177">XI.6</date>
–12.1996
<quantity id="415D17A4FFFDFFD2280920BAFE22FB60" box="[802,887,1189,1214]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="8" pageNumber="177" unit="m" value="1625.0">1,625m</quantity>
Eric L. Quinter along tributary at head Andranomifototra R. 1426.4'S, 4944.5'E Malaise traps; 1 pharate ♂, 1♀. E1000-4 P1013
<collectingCountry id="FEB2FAD1FFFDFFD22F5420D6F853FB3F" box="[1151,1286,1225,1249]" name="Madagascar" pageId="8" pageNumber="177">Madagascar</collectingCountry>
Bas. Lokoho Riv. Andranomifotatra Loc. Marojejy Camp4 4944'48''E, 1426'85''S
<date id="F21B9C81FFFDFFD22F3F20F2F9C4FADB" box="[1044,1169,1261,1285]" pageId="8" pageNumber="177" value="1999-11-29">29.11.1999</date>
Doumenq, E.; 7♀:
<collectingCountry id="FEB2FAD1FFFDFFD22BBC210DFC36FAF4" box="[151,355,1298,1322]" name="Madagascar" pageId="8" pageNumber="177">MADAGASCAR</collectingCountry>
: Antsiranana R.N.I. de Marojejy
<quantity id="415D17A4FFFDFFD229C1210EFE6AFAF4" box="[746,831,1297,1322]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="177" unit="km" value="10.5">10.5km</quantity>
NW Manantenina
<date id="F21B9C81FFFDFFD22F0B210EF91AFAF4" box="[1056,1103,1297,1322]" pageId="8" pageNumber="177">XI.3</date>
–XI.19.1996
<quantity id="415D17A4FFFDFFD22FC0210EF815FAF4" box="[1259,1344,1297,1322]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="8" pageNumber="177" unit="m" value="1625.0">1,625m</quantity>
Eric L. Quinter 1426.2'S, 4944.5'E Malaise traps;
<specimenCount id="90A371C8FFFDFFD2295F212AFFC8FA93" box="[628,669,1333,1357]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
,1♀:
<collectingCountry id="FEB2FAD1FFFDFFD229FB212AFEC9FA93" box="[720,924,1333,1357]" name="Madagascar" pageId="8" pageNumber="177">MADAGASCAR</collectingCountry>
: Antsiranana R.N.I. de Marojejy
<quantity id="415D17A4FFFDFFD22E3A212AF833FA93" box="[1297,1382,1333,1357]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="177" unit="km" value="10.5">10.5km</quantity>
NW Manantenina
<date id="F21B9C81FFFDFFD22A132146FC32FAAC" box="[312,359,1369,1394]" pageId="8" pageNumber="177">XI.6</date>
–12.1996
<quantity id="415D17A4FFFDFFD22AC92146FF62FAAC" box="[482,567,1369,1394]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="8" pageNumber="177" unit="m" value="1625.0">1,625m</quantity>
Eric L. Quinter along tributary at head Andranomifototra R. 1426.4'S, 4944.5'E Malaise traps; 1♀:
<collectingCountry id="FEB2FAD1FFFDFFD22ACB2162FFF9FA4B" box="[480,684,1405,1429]" name="Madagascar" pageId="8" pageNumber="177">MADAGASCAR</collectingCountry>
: Antsiranana R.N.I. de Marojejy
<quantity id="415D17A4FFFDFFD22F012162F92AFA4B" box="[1066,1151,1405,1429]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="8" pageNumber="177" unit="km" value="10.5">10.5km</quantity>
NW Manantenina
<date id="F21B9C81FFFDFFD22E702162F8D8FA4B" box="[1371,1421,1405,1429]" pageId="8" pageNumber="177">XI.6</date>
– 12.1996
<quantity id="415D17A4FFFDFFD22BD721BEFC04FA64" box="[252,337,1441,1466]" metricMagnitude="3" metricUnit="m" metricValue="1.625" pageId="8" pageNumber="177" unit="m" value="1625.0">1,625m</quantity>
Eric L. Quinter along tributary at head Andranomifototra R. 1426.4'S, 4944.5'E Malaise traps handschr.:
<date id="F21B9C81FFFDFFD22A3B21D9FCCAFA00" box="[272,415,1478,1502]" pageId="8" pageNumber="177">XI.12–XI.19</date>
(all
<collectionCode id="E0B42284FFFDFFD22AF821D9FF46FA00" box="[467,531,1478,1502]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="8" pageNumber="177">MZL</collectionCode>
).
</paragraph>
<paragraph id="861ABA41FFFDFFD22BEC21F6FE8BF9F8" blockId="8.[151,1437,1009,2006]" pageId="8" pageNumber="177">
<taxonomicName id="41A5C1C2FFFDFFD22BEC21F6FC09F9DC" box="[199,348,1513,1538]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B4D16653FFFDFFD22BEC21F6FC09F9DC" box="[199,348,1513,1538]" italics="true" pageId="8" pageNumber="177">Madenemura</emphasis>
</taxonomicName>
sp. 3 of
<bibRefCitation id="E234C7B0FFFDFFD22AE321F6FF3CF9DC" author="Paulian" box="[456,617,1513,1538]" pageId="8" pageNumber="177" refString="Paulian, R. (1959) Recherches sur les insectes d'importance biologique a Madagascar. XXXIII. Nouveaux Plecopteres malgaches. Memoires de l'Institut Scientifique de Madagascar, (E), 11, 9 - 16." type="journal article" year="1959">Paulian, 1959</bibRefCitation>
:
<specimenCount id="90A371C8FFFDFFD2295D21F5FFC9F9DC" box="[630,668,1514,1538]" pageId="8" pageNumber="177" type="generic">1♂</specimenCount>
:
<collectingCountry id="FEB2FAD1FFFDFFD2298021F5FE27F9DC" box="[683,882,1514,1538]" name="Madagascar" pageId="8" pageNumber="177">MADAGASCAR</collectingCountry>
<quantity id="415D17A4FFFDFFD2285721F6FEEEF9DC" box="[892,955,1513,1538]" metricMagnitude="4" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="177" unit="km" value="19.0">19km</quantity>
NW Man. [probably Manantenina]
<quantity id="415D17A4FFFDFFD22E7621F6FD82F9F8" metricMagnitude="6" metricUnit="m" metricValue="1.99615" pageId="8" pageNumber="177" unit="m" value="1996150.0">1996, 150m</quantity>
RNI Marojejy
<date id="F21B9C81FFFDFFD22AA82211FCEFF9F8" box="[387,442,1550,1574]" pageId="8" pageNumber="177" value="1922-10">X.22</date>
<date id="F21B9C81FFFDFFD22AEA2211FF71F9F8" box="[449,548,1549,1574]" pageId="8" pageNumber="177" value="2014-11-19">
XI.19
<geoCoordinate id="E391DC86FFFDFFD229232212FF71F9F8" box="[520,548,1549,1574]" direction="south" orientation="latitude" pageId="8" pageNumber="177" precision="92" value="-14.1">14</geoCoordinate>
</date>
°6.0'S
<geoCoordinate id="E391DC86FFFDFFD229452212FF89F9F8" box="[622,732,1549,1574]" direction="east" orientation="longitude" pageId="8" pageNumber="177" precision="92" value="49.751667">49°45.1'E</geoCoordinate>
Malaise Traps (
<collectionCode id="E0B42284FFFDFFD228BA2211FE87F9F8" box="[913,978,1550,1574]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="8" pageNumber="177">MZL</collectionCode>
).
</paragraph>
<paragraph id="861ABA41FFFDFFD22BEC222EF970F994" blockId="8.[151,1437,1009,2006]" box="[199,1061,1585,1610]" pageId="8" pageNumber="177">
<emphasis id="B4D16653FFFDFFD22BEC222EFC73F994" bold="true" box="[199,294,1585,1610]" pageId="8" pageNumber="177">Habitus</emphasis>
. As for the genus; WL of males
<quantity id="415D17A4FFFDFFD229BF222EFE42F994" box="[660,791,1585,1610]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" metricValueMax="7.1" metricValueMin="6.5" pageId="8" pageNumber="177" unit="mm" value="6.8" valueMax="7.1" valueMin="6.5">6.5–7.1mm</quantity>
, of females
<quantity id="415D17A4FFFDFFD228B5222EF974F994" box="[926,1057,1585,1610]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.5" metricValueMin="6.3" pageId="8" pageNumber="177" unit="mm" value="6.9" valueMax="7.5" valueMin="6.3">6.3–7.5mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="CEBFE9CAFFFDFFD32BEC224AF819FE72" lastPageId="9" lastPageNumber="178" pageId="8" pageNumber="177" type="description">
<paragraph id="861ABA41FFFDFFD22BEC224AF8DFF8FC" blockId="8.[151,1437,1009,2006]" pageId="8" pageNumber="177">
<emphasis id="B4D16653FFFDFFD22BEC224AFC51F9B0" bold="true" box="[199,260,1621,1646]" pageId="8" pageNumber="177">Male</emphasis>
. The front of the epiproct baseplate projects deeply into T10, dividing it into hemitergites. The overall shape of the baseplate is trapezoidal, with sinuous sides and wide truncate caudal edge with distinct caudolateral angles. The epiproct is of similar width and has a wide peduncle. In dorsal view, the front of the epiproct is slightly convex, the rear face flat to shallowly concave (
<figureCitation id="1E9EA6C4FFFDFFD2298422DEFE57F904" box="[687,770,1729,1754]" captionStart="FIGURES 10 – 17" captionStartId="5.[151,264,1816,1839]" captionTargetBox="[170,1433,206,1765]" captionTargetId="figure@5.[156,1441,193,1783]" captionTargetPageId="5" captionText="FIGURES 10 – 17, Madenemura spp., males. 10 – 12, M. descarpentriesi: 10, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 11, dorsoventral view of slide-mounted paraproct body; 12, apex of baseplate and epiproct hook, lateral view. 13, 14, M. culminalis: 13, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 14, dorsoventral view of slide-mounted paraproct body; 15, apex of baseplate and epiproct hook, lateral view. 16, 17, Madenemura sp., perhaps sp. 3 of Paulian (1959): 16, abdominal tip, lateral, semidiagrammatic, paraproct body shaded, arrow points at raised antecosta 10, inset below shows caudal edge of T 10 in dorsal view; 17, epiproct, lateral. 12 and 15 – 17 not to scale." httpUri="https://zenodo.org/record/238196/files/figure.png" pageId="8" pageNumber="177">Fig. 13</figureCitation>
). The subterminal spinose area is transverse, in side view it appears as a hump. The epiproct is more deeply excavated than in
<taxonomicName id="41A5C1C2FFFDFFD2285122F9F919F920" box="[890,1100,1765,1790]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B4D16653FFFDFFD2285122F9F919F920" box="[890,1100,1765,1790]" italics="true" pageId="8" pageNumber="177">M. descarpentriesi</emphasis>
</taxonomicName>
(
<figureCitation id="1E9EA6C4FFFDFFD22F7022FAF9F8F920" box="[1115,1197,1765,1790]" captionStart="FIGURES 10 – 17" captionStartId="5.[151,264,1816,1839]" captionTargetBox="[170,1433,206,1765]" captionTargetId="figure@5.[156,1441,193,1783]" captionTargetPageId="5" captionText="FIGURES 10 – 17, Madenemura spp., males. 10 – 12, M. descarpentriesi: 10, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 11, dorsoventral view of slide-mounted paraproct body; 12, apex of baseplate and epiproct hook, lateral view. 13, 14, M. culminalis: 13, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 14, dorsoventral view of slide-mounted paraproct body; 15, apex of baseplate and epiproct hook, lateral view. 16, 17, Madenemura sp., perhaps sp. 3 of Paulian (1959): 16, abdominal tip, lateral, semidiagrammatic, paraproct body shaded, arrow points at raised antecosta 10, inset below shows caudal edge of T 10 in dorsal view; 17, epiproct, lateral. 12 and 15 – 17 not to scale." httpUri="https://zenodo.org/record/238196/files/figure.png" pageId="8" pageNumber="177">Fig. 15</figureCitation>
). The paraproct body (
<figureCitation id="1E9EA6C4FFFDFFD22BB42316FDA4F8FC" box="[159,241,1801,1826]" captionStart="FIGURES 10 – 17" captionStartId="5.[151,264,1816,1839]" captionTargetBox="[170,1433,206,1765]" captionTargetId="figure@5.[156,1441,193,1783]" captionTargetPageId="5" captionText="FIGURES 10 – 17, Madenemura spp., males. 10 – 12, M. descarpentriesi: 10, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 11, dorsoventral view of slide-mounted paraproct body; 12, apex of baseplate and epiproct hook, lateral view. 13, 14, M. culminalis: 13, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 14, dorsoventral view of slide-mounted paraproct body; 15, apex of baseplate and epiproct hook, lateral view. 16, 17, Madenemura sp., perhaps sp. 3 of Paulian (1959): 16, abdominal tip, lateral, semidiagrammatic, paraproct body shaded, arrow points at raised antecosta 10, inset below shows caudal edge of T 10 in dorsal view; 17, epiproct, lateral. 12 and 15 – 17 not to scale." httpUri="https://zenodo.org/record/238196/files/figure.png" pageId="8" pageNumber="177">Fig. 14</figureCitation>
) resembles
<taxonomicName id="41A5C1C2FFFDFFD22A5C2315FF1CF8FC" box="[375,585,1801,1826]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="177" phylum="Arthropoda" rank="species" species="descarpentriesi">
<emphasis id="B4D16653FFFDFFD22A5C2315FF1CF8FC" box="[375,585,1801,1826]" italics="true" pageId="8" pageNumber="177">M. descarpentriesi</emphasis>
</taxonomicName>
but is longer and more slender, each blade being 7.5 times as long a wide.
</paragraph>
<paragraph id="861ABA41FFFDFFD22BEC2332F86AF850" blockId="8.[151,1437,1009,2006]" pageId="8" pageNumber="177">
<emphasis id="B4D16653FFFDFFD22BEC2332FC70F898" bold="true" box="[199,293,1837,1862]" pageId="8" pageNumber="177">Female.</emphasis>
The long and narrow ovipositor is of uniform width over most of its length. It is a firm brown structure without macroscopic pilosity, except the two setae at the shallowly bilobed tip of S8. In ventral view the ovipositor extends beyond the abdominal tip for a distance that equals the basal width of S8 (
<figureCitation id="1E9EA6C4FFFDFFD22F82236AF866F850" box="[1193,1331,1909,1934]" captionStart="FIGURES 18 – 24" captionStartId="6.[151,264,1952,1975]" captionTargetBox="[184,1393,201,1924]" captionTargetId="figure@6.[175,1406,193,1932]" captionTargetPageId="6" captionText="FIGURES 18 – 24, Madenemura spp., female abdominal tips, in lateral (even numbers) and ventral (uneven numbers) view; after photographs, semidiagrammatic: 18, 19: M. andringitrensis. 20, 21: M. descarpentriesi. 22, 23: M. culminalis; 24: M. pauliani sp. n. (holotype). Same scale for all." httpUri="https://zenodo.org/record/238197/files/figure.png" pageId="8" pageNumber="177">Figs. 22, 23</figureCitation>
).
</paragraph>
<paragraph id="861ABA41FFFDFFD32BEC2386F95AFF0B" blockId="8.[151,1437,1009,2006]" lastBlockId="9.[151,1436,151,429]" lastPageId="9" lastPageNumber="178" pageId="8" pageNumber="177">
<emphasis id="B4D16653FFFDFFD22BEC2386FC53F86C" bold="true" box="[199,262,1945,1970]" pageId="8" pageNumber="177">Note.</emphasis>
This species is recognized by the ovipositor. The species was first described from northern
<collectingCountry id="FEB2FAD1FFFDFFD22E3E2386F8C9F86C" box="[1301,1436,1945,1970]" name="Madagascar" pageId="8" pageNumber="177">Madagascar</collectingCountry>
but is actually widespread on the island although not common. Paulian's (1959) illustrations of the male, especially the long paraproct body, agree with the present specimens. In two distant regions, sexes were found in the same area and occurred in small numbers in trap catches dominated by other species.
</paragraph>
<paragraph id="861ABA41FFFCFFD32BEC24FFF819FE72" blockId="9.[151,1436,151,429]" pageId="9" pageNumber="178">
In a single seemingly teneral male (WL
<quantity id="415D17A4FFFCFFD3299D24C0FE52FF26" box="[694,775,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="9" pageNumber="178" unit="mm" value="5.7">5.7mm</quantity>
) the divided antecosta of T10 forms a pair of slightly asymmetrical, slightly raised lobes (
<figureCitation id="1E9EA6C4FFFCFFD32907251BFFDFFEC3" box="[556,650,260,285]" captionStart="FIGURES 10 – 17" captionStartId="5.[151,264,1816,1839]" captionTargetBox="[170,1433,206,1765]" captionTargetId="figure@5.[156,1441,193,1783]" captionTargetPageId="5" captionText="FIGURES 10 – 17, Madenemura spp., males. 10 – 12, M. descarpentriesi: 10, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 11, dorsoventral view of slide-mounted paraproct body; 12, apex of baseplate and epiproct hook, lateral view. 13, 14, M. culminalis: 13, central part of T 10 with epiproct baseplate and epiproct, cerci diagrammatic; 14, dorsoventral view of slide-mounted paraproct body; 15, apex of baseplate and epiproct hook, lateral view. 16, 17, Madenemura sp., perhaps sp. 3 of Paulian (1959): 16, abdominal tip, lateral, semidiagrammatic, paraproct body shaded, arrow points at raised antecosta 10, inset below shows caudal edge of T 10 in dorsal view; 17, epiproct, lateral. 12 and 15 – 17 not to scale." httpUri="https://zenodo.org/record/238196/files/figure.png" pageId="9" pageNumber="178">Figs. 16</figureCitation>
). Insofar it corresponds to Male sp. 3 of
<bibRefCitation id="E234C7B0FFFCFFD32F78251BF9ACFEC3" author="Paulian" box="[1107,1273,260,285]" pageId="9" pageNumber="178" refString="Paulian, R. (1959) Recherches sur les insectes d'importance biologique a Madagascar. XXXIII. Nouveaux Plecopteres malgaches. Memoires de l'Institut Scientifique de Madagascar, (E), 11, 9 - 16." type="journal article" year="1959">Paulian (1959)</bibRefCitation>
which has not been illustrated. Paulian compared it to
<taxonomicName id="41A5C1C2FFFCFFD329732536FEF0FE9E" authority="Paulian, 1959" authorityName="Paulian" authorityYear="1959" box="[600,933,295,320]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="bezanozano">
<emphasis id="B4D16653FFFCFFD329732536FE57FE9E" box="[600,770,295,320]" italics="true" pageId="9" pageNumber="178">M. bezanozano</emphasis>
<bibRefCitation id="E234C7B0FFFCFFD328212538FEF0FE9E" author="Paulian" box="[778,933,295,320]" pageId="9" pageNumber="178" refString="Paulian, R. (1959) Recherches sur les insectes d'importance biologique a Madagascar. XXXIII. Nouveaux Plecopteres malgaches. Memoires de l'Institut Scientifique de Madagascar, (E), 11, 9 - 16." type="journal article" year="1959">Paulian, 1959</bibRefCitation>
</taxonomicName>
which I have not seen, also no male with an epiproct as wide and with prominent rear corners as shown for
<taxonomicName id="41A5C1C2FFFCFFD3284F2552F95BFEBB" box="[868,1038,332,357]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="bezanozano">
<emphasis id="B4D16653FFFCFFD3284F2552F95BFEBB" box="[868,1038,332,357]" italics="true" pageId="9" pageNumber="178">M. bezanozano</emphasis>
</taxonomicName>
. The paraproct body of the present male agrees with
<emphasis id="B4D16653FFFCFFD32A45256EFF18FE56" box="[366,589,367,392]" italics="true" pageId="9" pageNumber="178">
<taxonomicName id="41A5C1C2FFFCFFD32A45256EFF1DFE56" box="[366,584,367,392]" class="Insecta" family="Notonemouridae" genus="Madenemura" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="178" phylum="Arthropoda" rank="species" species="descarpentriesi">M. descarpentriesi</taxonomicName>
,
</emphasis>
the epiproct resembles it also but is more strongly bent forward. The crumpled cerci suggest an artifact. It is unclear if this male is malformed or if it represents a separate taxon.
</paragraph>
</subSubSection>
</treatment>
</document>
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