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<document id="2F51AD0DBEEB42E2ACB465CD2087B1CD" ID-DOI="10.5281/zenodo.199162" ID-GBIF-Dataset="d399490b-bdf4-43c8-9314-829859fdce53" ID-ISSN="1175-5326" ID-Zenodo-Dep="199162" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460328808760" checkinUser="plazi" docAuthor="Jesionowska, Katarzyna" docDate="2010" docId="D8273543FFE6544BB6BBA668FEB3FC0A" docLanguage="en" docName="zt02672p049.pdf" docOrigin="Zootaxa 2672" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Penthalodes hawaiiensis Jesionowska, 2010, n. sp." docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="241E4D3BFFE55458B62CA45AFFCCFFE7" masterDocTitle="A morphological study of the genus Penthalodes (Acari, Prostigmata, Eupodoidea, Penthalodidae) with description of a new species" masterLastPageNumber="49" masterPageNumber="29" pageNumber="32" updateTime="1698258321494" updateUser="plazi">
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<mods:title id="834F27B539E3F290BC38C487F088880F">A morphological study of the genus Penthalodes (Acari, Prostigmata, Eupodoidea, Penthalodidae) with description of a new species</mods:title>
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<mods:date id="CBBE0F74D086282F11CDC4DE2513A2C7">2010</mods:date>
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<treatment id="D8273543FFE6544BB6BBA668FEB3FC0A" ID-DOI="http://doi.org/10.5281/zenodo.5631878" ID-GBIF-Taxon="119404222" ID-Zenodo-Dep="5631878" LSID="urn:lsid:plazi:treatment:D8273543FFE6544BB6BBA668FEB3FC0A" httpUri="http://treatment.plazi.org/id/D8273543FFE6544BB6BBA668FEB3FC0A" lastPageId="19" lastPageNumber="48" pageId="3" pageNumber="32">
<subSubSection id="1894D7DEFFE6545BB6BBA668FAD4FD56" pageId="3" pageNumber="32" type="nomenclature">
<paragraph id="50318455FFE6545BB6BBA668FDC5FDAB" blockId="3.[151,521,562,588]" box="[151,521,562,588]" pageId="3" pageNumber="32">
<heading id="0B793339FFE6545BB6BBA668FDC5FDAB" bold="true" box="[151,521,562,588]" fontSize="11" level="1" pageId="3" pageNumber="32" reason="1">
<emphasis id="62FA5847FFE6545BB6BBA668FDC5FDAB" bold="true" box="[151,521,562,588]" pageId="3" pageNumber="32">
<taxonomicName id="978EFFD6FFE6545BB6BBA668FE0CFDAB" box="[151,448,562,588]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="62FA5847FFE6545BB6BBA668FE0CFDAB" bold="true" box="[151,448,562,588]" italics="true" pageId="3" pageNumber="32">Penthalodes hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE6545BB7EBA668FDC5FDAB" box="[455,521,562,588]" pageId="3" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="50318455FFE6545BB6BBA620FC30FD76" blockId="3.[151,1304,634,689]" box="[151,1020,634,657]" pageId="3" pageNumber="32">
Synonym:
<taxonomicName id="978EFFD6FFE6545BB727A620FD41FD76" authority="Strandtmann &amp; Goff, 1978" authorityName="Strandtmann &amp; Goff" authorityYear="1978" box="[267,653,634,657]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE6545BB727A620FEA9FD76" box="[267,357,634,657]" italics="true" pageId="3" pageNumber="32">P. ovalis</emphasis>
<bibRefCitation id="341FF9A4FFE6545BB740A620FD41FD76" author="Strandtmann" box="[364,653,634,657]" pageId="3" pageNumber="32" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandtmann &amp; Goff, 1978</bibRefCitation>
</taxonomicName>
,
<emphasis id="62FA5847FFE6545BB4BBA620FCFFFD76" box="[663,819,634,657]" italics="true" pageId="3" pageNumber="32">Pacific Insects</emphasis>
19, 34, 121143.
</paragraph>
<paragraph id="50318455FFE6545BB6BBA6C0FAD4FD56" blockId="3.[151,1304,634,689]" box="[151,1304,666,689]" pageId="3" pageNumber="32">
<emphasis id="62FA5847FFE6545BB6BBA6C0FEE5FD56" box="[151,297,666,689]" italics="true" pageId="3" pageNumber="32">Locus typicus</emphasis>
: Hawaii, vicinity of Volcano-Hilo road, samples from grass, lichen, treefern forest, cane fields
</paragraph>
</subSubSection>
<subSubSection id="1894D7DEFFE6545BB6BBA685FD51FA58" pageId="3" pageNumber="32" type="diagnosis">
<paragraph id="50318455FFE6545BB6BBA685FBCCFC53" blockId="3.[151,1437,735,986]" pageId="3" pageNumber="32">
<emphasis id="62FA5847FFE6545BB6BBA685FED5FD1E" bold="true" box="[151,281,735,761]" pageId="3" pageNumber="32">Diagnosis.</emphasis>
Epirostrum trilobed; middle part of epirostrum large, triangular; lateral lobes weakly developed with slightly pointed, resembling triangles. Dorsal idiosomal setae, except setae
<emphasis id="62FA5847FFE6545BB267A75FFBAFFCF9" box="[1099,1123,773,798]" italics="true" pageId="3" pageNumber="32">ro</emphasis>
and
<emphasis id="62FA5847FFE6545BB2B2A75FFB77FCF9" box="[1182,1211,773,798]" italics="true" pageId="3" pageNumber="32">bo</emphasis>
, loosely feathered. Rhagidial organ I with two recumbent baculiform solenidia, in tandem, in confluent depressions and one tiny spatulate solenidion in separate pit inserted laterally antiaxially in relation to rhagidial organ I. Famulus absent. Rhagidial organ II with three recumbent baculiform solenidia, staggered, in confluent depressions. Solenidia of rhagidial organ II of unequal size, not overlapping each other.
</paragraph>
<paragraph id="50318455FFE6545BB6E9A79AFAE4FC3D" blockId="3.[151,1437,735,986]" box="[197,1320,960,986]" pageId="3" pageNumber="32">
For the original description and deposition of
<typeStatus id="8F353AF7FFE6545BB4F3A79AFCDDFC3D" box="[735,785,960,986]" pageId="3" pageNumber="32">type</typeStatus>
specimens see
<bibRefCitation id="341FF9A4FFE6545BB5EAA79AFAE8FC3D" author="Strandtmann" box="[966,1316,960,986]" pageId="3" pageNumber="32" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandtmann and Goff (1978)</bibRefCitation>
.
</paragraph>
<paragraph id="50318455FFE6545BB6BBA075FEDCFB88" blockId="3.[151,496,1071,1135]" pageId="3" pageNumber="32">
<emphasis id="62FA5847FFE6545BB6BBA075FE25FBAE" bold="true" box="[151,489,1071,1097]" pageId="3" pageNumber="32">
<taxonomicName id="978EFFD6FFE6545BB6BBA075FE6BFBAE" box="[151,423,1071,1097]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFE6545BB6BBA075FE6BFBAE" bold="true" box="[151,423,1071,1097]" italics="true" pageId="3" pageNumber="32">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE6545BB783A075FE25FBAE" box="[431,489,1071,1097]" pageId="3" pageNumber="32" rank="species">n. sp</taxonomicNameLabel>
</emphasis>
. (Figs 17)
</paragraph>
<paragraph id="50318455FFE6545BB6BBA0C5FD51FA58" blockId="3.[151,1437,1183,1996]" pageId="3" pageNumber="32">
<emphasis id="62FA5847FFE6545BB6BBA0C5FED6FB5E" bold="true" box="[151,282,1183,1209]" pageId="3" pageNumber="32">Diagnosis.</emphasis>
Epirostrum trilobed, lateral lobes broadly rounded; in lateral view, base of epirostrum sloping; without reticulation. Dorsal idiosomal setae, except setae
<emphasis id="62FA5847FFE6545BB565A09FFCADFB39" box="[841,865,1221,1246]" italics="true" pageId="3" pageNumber="32">ro</emphasis>
and
<emphasis id="62FA5847FFE6545BB5B2A09FFC77FB39" box="[926,955,1221,1246]" italics="true" pageId="3" pageNumber="32">bo</emphasis>
, tridactylate, with short stem. Rhagidial organ of tarsus I with two banana-shaped solenidia in tandem in a confluent depressions and one T-shaped solenidion in a separate pit inserted laterally to proximal solenidion. Small stellate famulus inserted laterally on mid-region of tarsus I, well separated from rhagidial organ. Rhagidial organ of tarsus II with three thin baculiform solenidia, in tandem, in a common depression with the distal part of each solenidion overlapping the preceding one. One, small spine-like solenidion (possibly a famulus) inserted in pit laterally to rhagidial organ II, at the level of proximal solenidion.
</paragraph>
</subSubSection>
<subSubSection id="1894D7DEFFE65451B6E9A190FE45F8EA" lastPageId="9" lastPageNumber="38" pageId="3" pageNumber="32" type="description">
<paragraph id="50318455FFE6545BB6E9A190FC08F9B0" blockId="3.[151,1437,1183,1996]" pageId="3" pageNumber="32">
<emphasis id="62FA5847FFE6545BB6E9A190FE91FA03" bold="true" box="[197,349,1482,1508]" pageId="3" pageNumber="32">Description.</emphasis>
Body with well developed ornamentation, moderately sclerotized. Idiosoma angular (
<figureCitation id="C8B598D0FFE6545BB360A190FA5EFA03" box="[1356,1426,1482,1508]" captionStart="FIGURE 3" captionStartId="6.[151,259,1787,1811]" captionTargetBox="[218,1372,205,1741]" captionTargetId="figure@6.[212,1376,194,1765]" captionTargetPageId="6" captionText="FIGURE 3. Penthalodes polonicus n. sp. Female. (A) chelicera, lateral view. (B) cheliceral digits in detail. (C) epirostrum, dorsal view. (D) epirostrum, lateral view. (E) Penthalodes sp., epirostrum, lateral view." httpUri="https://zenodo.org/record/199164/files/figure.png" pageId="3" pageNumber="32">Fig. 3</figureCitation>
) with ornamental costulae forming pentagons (i.e. reticulate pattern). Dorsum with two “V”-shaped furrows. Idiosomal length (excluding epirostrum) about 400 µm, width (at the level of setae
<emphasis id="62FA5847FFE6545BB2AFA24FFB5CF9C9" box="[1155,1168,1557,1582]" italics="true" pageId="3" pageNumber="32">c</emphasis>
2) about 250 µm. Legs shorter than idiosoma. Ratio of idiosomal length to leg I length: 1:15.
</paragraph>
<paragraph id="50318455FFE6545BB6E9A239FE49F82B" blockId="3.[151,1437,1183,1996]" pageId="3" pageNumber="32">
<emphasis id="62FA5847FFE6545BB6E9A239FE90F99B" box="[197,348,1635,1660]" italics="true" pageId="3" pageNumber="32">Aspidosoma</emphasis>
. (Figs 1, 2, 3C, D) Prodorsum forms a steep anterior wall of idiosoma. Ornamentation consists of large tear-shaped costulae (i.e. point-shaped with tiny spine) and minute costulae discernible as dots. Large costulae form pentagons, while minute costulae form parallel lines on the surface of each pentagon. Naso small, smooth, and spherical, with one hemisphere hidden in a cavity. Epirostrum trilobed with lateral lobes broadly rounded, distinctly shorter than medial lobe (
<figureCitation id="C8B598D0FFE6545BB5D9A2A2FB8DF8F5" box="[1013,1089,1784,1810]" captionStart="FIGURE 3" captionStartId="6.[151,259,1787,1811]" captionTargetBox="[218,1372,205,1741]" captionTargetId="figure@6.[212,1376,194,1765]" captionTargetPageId="6" captionText="FIGURE 3. Penthalodes polonicus n. sp. Female. (A) chelicera, lateral view. (B) cheliceral digits in detail. (C) epirostrum, dorsal view. (D) epirostrum, lateral view. (E) Penthalodes sp., epirostrum, lateral view." httpUri="https://zenodo.org/record/199164/files/figure.png" pageId="3" pageNumber="32">Figs 3</figureCitation>
CD). Four pairs of prodorsal setae:
<emphasis id="62FA5847FFE6545BB6CDA344FF35F8D0" box="[225,249,1822,1847]" italics="true" pageId="3" pageNumber="32">ro</emphasis>
,
<emphasis id="62FA5847FFE6545BB725A344FEEAF8D0" box="[265,294,1822,1847]" italics="true" pageId="3" pageNumber="32">bo</emphasis>
,
<emphasis id="62FA5847FFE6545BB719A344FE86F8D0" box="[309,330,1822,1847]" italics="true" pageId="3" pageNumber="32">le</emphasis>
,
<emphasis id="62FA5847FFE6545BB775A344FEB9F8D0" box="[345,373,1822,1847]" italics="true" pageId="3" pageNumber="32">xa</emphasis>
(about 6, 56, 37 and 25 long, respectively). Setae
<emphasis id="62FA5847FFE6545BB5E9A344FC11F8D0" box="[965,989,1822,1847]" italics="true" pageId="3" pageNumber="32">ro</emphasis>
tiny, nude, inserted on frontal surface of naso, adjacent in a common depression (possibly a result of slide-mounting). Setae
<emphasis id="62FA5847FFE6545BB2A0A319FB65F8BB" box="[1164,1193,1859,1884]" italics="true" pageId="3" pageNumber="32">bo</emphasis>
filiform, inserted on prodorsum in deep bothridia dorsally; setae
<emphasis id="62FA5847FFE6545BB4B0A332FD7DF866" box="[668,689,1896,1921]" italics="true" pageId="3" pageNumber="32">le</emphasis>
and
<emphasis id="62FA5847FFE6545BB4C6A332FCCAF866" box="[746,774,1896,1921]" italics="true" pageId="3" pageNumber="32">xa</emphasis>
situated frontally. Setae
<emphasis id="62FA5847FFE6545BB205A332FBF2F866" box="[1065,1086,1896,1921]" italics="true" pageId="3" pageNumber="32">le</emphasis>
and
<emphasis id="62FA5847FFE6545BB25BA332FB5EF866" box="[1143,1170,1896,1921]" italics="true" pageId="3" pageNumber="32">xa</emphasis>
tridactylate with short basal branches. V-shaped furrows start lateral to setae
<emphasis id="62FA5847FFE6545BB51BA3D4FC99F840" box="[823,853,1934,1959]" italics="true" pageId="3" pageNumber="32">bo</emphasis>
. Transverse furrow separating prodorsum from opisthosoma absent.
</paragraph>
<paragraph id="50318455FFE1545CB6BBA3D9FEE3F83A" blockId="4.[151,1436,1923,2014]" pageId="4" pageNumber="33">
<emphasis id="62FA5847FFE1545CB6BBA3D9FED5F87C" bold="true" box="[151,281,1923,1947]" pageId="4" pageNumber="33">FIGURE 1.</emphasis>
<taxonomicName id="978EFFD6FFE1545CB70CA3DEFDDEF87C" box="[288,530,1924,1947]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="4" pageNumber="33" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFE1545CB70CA3DEFDDEF87C" box="[288,530,1924,1947]" italics="true" pageId="4" pageNumber="33">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE1545CB435A3D9FD99F87C" box="[537,597,1923,1947]" pageId="4" pageNumber="33" rank="species">
<emphasis id="62FA5847FFE1545CB435A3D9FD99F87C" bold="true" box="[537,597,1923,1947]" pageId="4" pageNumber="33">n. sp.</emphasis>
</taxonomicNameLabel>
Female. (A) Dorsum; prodorsal setae:
<emphasis id="62FA5847FFE1545CB5D9A3DEFBB7F87C" box="[1013,1147,1924,1947]" italics="true" pageId="4" pageNumber="33">ro, le, bo, xa</emphasis>
; opisthosomal setae:
<emphasis id="62FA5847FFE1545CB34EA3DEFAA2F87C" box="[1378,1390,1924,1947]" italics="true" pageId="4" pageNumber="33">c</emphasis>
1,
<emphasis id="62FA5847FFE1545CB3AEA3DEFA42F87C" box="[1410,1422,1924,1947]" italics="true" pageId="4" pageNumber="33">c</emphasis>
2,
<emphasis id="62FA5847FFE1545CB6BBA3FDFEF0F859" box="[151,316,1959,1982]" italics="true" pageId="4" pageNumber="33">d, e, f, h, ps, ad</emphasis>
. Lyrifissures: ia, im, ip, ih. (B) Venter; coxal regions: cx 14; trochanter: tr 14; aggenital setae: ag 18; anal setae:
<emphasis id="62FA5847FFE1545CB721A39DFEE4F839" box="[269,296,1991,2014]" italics="true" pageId="4" pageNumber="33">an</emphasis>
.
</paragraph>
<caption id="04F1D4DDFFE0545DB6BBA7A6FC66FBF4" box="[151,938,1020,1044]" httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="5" pageNumber="34" targetBox="[161,1430,205,979]" targetPageId="5">
<paragraph id="50318455FFE0545DB6BBA7A6FC66FBF4" blockId="5.[151,938,1020,1044]" box="[151,938,1020,1044]" pageId="5" pageNumber="34">
<emphasis id="62FA5847FFE0545DB6BBA7A6FED5FBF3" bold="true" box="[151,281,1020,1044]" pageId="5" pageNumber="34">FIGURE 2.</emphasis>
<taxonomicName id="978EFFD6FFE0545DB70CA7A6FDDEFBF4" box="[288,530,1020,1043]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFE0545DB70CA7A6FDDEFBF4" box="[288,530,1020,1043]" italics="true" pageId="5" pageNumber="34">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE0545DB434A7A6FD98FBF3" box="[536,596,1020,1044]" pageId="5" pageNumber="34" rank="species">
<emphasis id="62FA5847FFE0545DB434A7A6FD98FBF3" bold="true" box="[536,596,1020,1044]" pageId="5" pageNumber="34">n. sp.</emphasis>
</taxonomicNameLabel>
Female. Idiosoma, lateral view.
</paragraph>
</caption>
<paragraph id="50318455FFE0545DB6E9A018FAF0F8C0" blockId="5.[151,1437,1090,1980]" pageId="5" pageNumber="34">
<emphasis id="62FA5847FFE0545DB6E9A018FEADFBBC" box="[197,353,1090,1115]" italics="true" pageId="5" pageNumber="34">Gnathosoma</emphasis>
(
<figureCitation id="C8B598D0FFE0545DB75FA018FE0CFBBB" box="[371,448,1090,1116]" captionStart="FIGURE 3" captionStartId="6.[151,259,1787,1811]" captionTargetBox="[218,1372,205,1741]" captionTargetId="figure@6.[212,1376,194,1765]" captionTargetPageId="6" captionText="FIGURE 3. Penthalodes polonicus n. sp. Female. (A) chelicera, lateral view. (B) cheliceral digits in detail. (C) epirostrum, dorsal view. (D) epirostrum, lateral view. (E) Penthalodes sp., epirostrum, lateral view." httpUri="https://zenodo.org/record/199164/files/figure.png" pageId="5" pageNumber="34">Figs 3</figureCitation>
AB, 4). Subcapitulum (
<figureCitation id="C8B598D0FFE0545DB4C8A018FCE3FBBB" box="[740,815,1090,1116]" captionStart="FIGURE 4" captionStartId="8.[151,255,1869,1893]" captionTargetBox="[202,1390,194,1846]" captionTargetId="figure@8.[188,1398,194,1847]" captionTargetPageId="8" captionText="FIGURE 4. Penthalodes polonicus n. sp. (A) Subcapitulum, ventral view; (B) Subcapitulum, dorsal view; LB-labrum, LL-lateral lips, LI-labium, H-hypostome, sc-internal sclerites, n, m-subcapitular setae, cxp-coxal region of pedipalp. (C) Pedipalp, lateral view; (D) Pedipalpal tarsus in detail, lateral view." httpUri="https://zenodo.org/record/199165/files/figure.png" pageId="5" pageNumber="34">Fig. 4</figureCitation>
AB) about 110 long, conical, with well developed ornamentation comprising spine- to point-shaped costulae on smooth integument. Lateral lips (LL) connected by a smooth membrane dorsally for approximately two thirds their total length. Ventrally, the free parts of lateral lips bear one pair of setae
<emphasis id="62FA5847FFE0545DB41FA0E8FD8EFB2C" box="[563,578,1202,1227]" italics="true" pageId="5" pageNumber="34">n</emphasis>
, inserted slightly distal to labial apex; and one pair of setae
<emphasis id="62FA5847FFE0545DB302A0E8FA8FFB2C" box="[1326,1347,1202,1227]" italics="true" pageId="5" pageNumber="34">m</emphasis>
laterobasally. Setae
<emphasis id="62FA5847FFE0545DB76EA08DFE9DFB17" box="[322,337,1239,1264]" italics="true" pageId="5" pageNumber="34">n</emphasis>
and
<emphasis id="62FA5847FFE0545DB7A1A08DFE6EFB17" box="[397,418,1239,1264]" italics="true" pageId="5" pageNumber="34">m</emphasis>
plumose with long outgrowths. On ventral side, lateral lips fused to hypostome (H), delimited by sclerites
<emphasis id="62FA5847FFE0545DB7BBA0A6FE7CFAF2" box="[407,432,1276,1301]" italics="true" pageId="5" pageNumber="34">sc</emphasis>
. Top of hypostome forms small labium (LI). Hypostome divided by inner sclerite into a basal rectangular and distal triangular part. Paraxial surfaces of free part of lateral lips, as well as the ventral part of labrum (LB), strongly sclerotized. Distal antiaxial parts of lateral lips smooth and plicate. Adoral setae difficult to discern. Free part of labrum not longer than those of lateral lips. Chelicera (
<figureCitation id="C8B598D0FFE0545DB289A136FB26FA61" box="[1189,1258,1388,1414]" captionStart="FIGURE 3" captionStartId="6.[151,259,1787,1811]" captionTargetBox="[218,1372,205,1741]" captionTargetId="figure@6.[212,1376,194,1765]" captionTargetPageId="6" captionText="FIGURE 3. Penthalodes polonicus n. sp. Female. (A) chelicera, lateral view. (B) cheliceral digits in detail. (C) epirostrum, dorsal view. (D) epirostrum, lateral view. (E) Penthalodes sp., epirostrum, lateral view." httpUri="https://zenodo.org/record/199164/files/figure.png" pageId="5" pageNumber="34">Fig. 3</figureCitation>
AB) about 115 long, elongated, ornamented with tiny spines. Cheliceral seta absent. Fixed digit hood-shaped, distally bifurcate covering most of movable digit. Movable digit strongly sclerotized, narrows distally into hooked claw. Pedipalps (
<figureCitation id="C8B598D0FFE0545DB771A186FE6FFA11" box="[349,419,1500,1526]" captionStart="FIGURE 4" captionStartId="8.[151,255,1869,1893]" captionTargetBox="[202,1390,194,1846]" captionTargetId="figure@8.[188,1398,194,1847]" captionTargetPageId="8" captionText="FIGURE 4. Penthalodes polonicus n. sp. (A) Subcapitulum, ventral view; (B) Subcapitulum, dorsal view; LB-labrum, LL-lateral lips, LI-labium, H-hypostome, sc-internal sclerites, n, m-subcapitular setae, cxp-coxal region of pedipalp. (C) Pedipalp, lateral view; (D) Pedipalpal tarsus in detail, lateral view." httpUri="https://zenodo.org/record/199165/files/figure.png" pageId="5" pageNumber="34">Fig. 4</figureCitation>
CD) four-segmented, Tr-FG-Tb-Ts; coxal regions well formed and fused partially with dorso-basal subcapitulum (
<figureCitation id="C8B598D0FFE0545DB7F1A258FDEBF9FB" box="[477,551,1538,1564]" captionStart="FIGURE 4" captionStartId="8.[151,255,1869,1893]" captionTargetBox="[202,1390,194,1846]" captionTargetId="figure@8.[188,1398,194,1847]" captionTargetPageId="8" captionText="FIGURE 4. Penthalodes polonicus n. sp. (A) Subcapitulum, ventral view; (B) Subcapitulum, dorsal view; LB-labrum, LL-lateral lips, LI-labium, H-hypostome, sc-internal sclerites, n, m-subcapitular setae, cxp-coxal region of pedipalp. (C) Pedipalp, lateral view; (D) Pedipalpal tarsus in detail, lateral view." httpUri="https://zenodo.org/record/199165/files/figure.png" pageId="5" pageNumber="34">Fig. 4</figureCitation>
B). Pedipalp about 166 long,
<collectingCountry id="2899C4C5FFE0545DB5A6A258FC6AF9FB" box="[906,934,1538,1564]" name="Chile" pageId="5" pageNumber="34">Tr</collectingCountry>
to Ts: 25-55-57-29. Number of setae and solenidia: 0-2-3-9(1). Setae on femorogenu (
<emphasis id="62FA5847FFE0545DB49EA27DFD0DF9A7" box="[690,705,1575,1600]" italics="true" pageId="5" pageNumber="34">d</emphasis>
1-15,
<emphasis id="62FA5847FFE0545DB52DA27DFCDCF9A7" box="[769,784,1575,1600]" italics="true" pageId="5" pageNumber="34">d</emphasis>
2-21 long) and tibia (
<emphasis id="62FA5847FFE0545DB226A27DFBD5F9A7" box="[1034,1049,1575,1600]" italics="true" pageId="5" pageNumber="34">d</emphasis>
1-18,
<emphasis id="62FA5847FFE0545DB275A27DFBA6F9A7" box="[1113,1130,1575,1600]" italics="true" pageId="5" pageNumber="34">lt</emphasis>
-11,
<emphasis id="62FA5847FFE0545DB28DA27DFB7CF9A7" box="[1185,1200,1575,1600]" italics="true" pageId="5" pageNumber="34">d</emphasis>
2-13 long) plumose. Seta
<emphasis id="62FA5847FFE0545DB6FEA20AFF2DF98E" box="[210,225,1616,1641]" italics="true" pageId="5" pageNumber="34">d</emphasis>
2 on FG with a very long terminal spine. Pedipalpal tarsus elongated, sharply tipped, linear in profile with nine setae and one recumbent solenidion in depression dorsally; seta
<emphasis id="62FA5847FFE0545DB23EA222FB9EF976" box="[1042,1106,1656,1681]" italics="true" pageId="5" pageNumber="34">acmg</emphasis>
(=
<emphasis id="62FA5847FFE0545DB259A222FB48F976" box="[1141,1156,1656,1681]" italics="true" pageId="5" pageNumber="34">d</emphasis>
; 10 long) thick, pilose, angled basally, the largest; seta
<emphasis id="62FA5847FFE0545DB420A2C7FDE2F951" box="[524,558,1693,1718]" italics="true" pageId="5" pageNumber="34">cm</emphasis>
' (11 long) positioned paraxially, almost smooth, spiralled; seta
<emphasis id="62FA5847FFE0545DB337A2C7FAFEF951" box="[1307,1330,1693,1718]" italics="true" pageId="5" pageNumber="34">ul</emphasis>
'' slightly thickened, shortly pilose; setae
<emphasis id="62FA5847FFE0545DB435A298FDFDF93C" box="[537,561,1730,1755]" italics="true" pageId="5" pageNumber="34">ul</emphasis>
',
<emphasis id="62FA5847FFE0545DB46BA298FD9AF93C" box="[583,598,1730,1755]" italics="true" pageId="5" pageNumber="34">u</emphasis>
'' and
<emphasis id="62FA5847FFE0545DB4B2A298FD61F93C" box="[670,685,1730,1755]" italics="true" pageId="5" pageNumber="34">u</emphasis>
' (5 long each) cylindrical, hollow, each terminating in strong sclerotized solid (black) carina; seta
<emphasis id="62FA5847FFE0545DB464A2B2FD9BF8E6" box="[584,599,1768,1793]" italics="true" pageId="5" pageNumber="34">a</emphasis>
(7 long), thickened, with distal spine; setae
<emphasis id="62FA5847FFE0545DB24FA2B2FBA2F8E6" box="[1123,1134,1768,1793]" italics="true" pageId="5" pageNumber="34">s</emphasis>
plumose,
<emphasis id="62FA5847FFE0545DB2C5A2B2FB33F8E6" box="[1257,1279,1768,1793]" italics="true" pageId="5" pageNumber="34">la</emphasis>
'' pilose, very short (3 and 4 long, respectively). Ventro-distal part of tibia terminating in distinct spine-like process.
</paragraph>
<paragraph id="50318455FFE0545DB6E9A368FA50F85B" blockId="5.[151,1437,1090,1980]" pageId="5" pageNumber="34">
<emphasis id="62FA5847FFE0545DB6E9A368FEAAF8AC" box="[197,358,1842,1867]" italics="true" pageId="5" pageNumber="34">Opisthosoma</emphasis>
(Figs 1, 2;
<tableCitation id="1D0CB1EEFFE0545DB7DBA368FD9FF8AB" box="[503,595,1842,1868]" captionStart="TABLE 1" captionStartId="7.[151,239,487,511]" captionTargetBox="[151,1391,538,1086]" captionTargetPageId="7" captionText="TABLE 1. Homology of opisthosomal setae in representatives of the family Penthalodidae." httpUri="http://table.plazi.org/id/04F1D4DDFFE2545FB6BBA5BDFBA4FE18" pageId="5" pageNumber="34" tableUuid="04F1D4DDFFE2545FB6BBA5BDFBA4FE18">Table 1</tableCitation>
). Opisthosoma with characteristic “V”-shaped furrows, lacking any transverse furrows (as “das” or disjugal furrow). Ornamentation same as prodorsum, i.e. tear-shaped costulae forming pentagons, each covered by minute point-shaped costulae forming parallel lines (Fig. 1). Ornamentation on opisthosoma venter more delicate, with costulae being visibly smaller than dorsal costulae.
</paragraph>
<caption id="04F1D4DDFFE3545EB6BBA2A1FB7BF8D5" httpUri="https://zenodo.org/record/199164/files/figure.png" pageId="6" pageNumber="35" targetBox="[218,1372,205,1741]" targetPageId="6">
<paragraph id="50318455FFE3545EB6BBA2A1FB7BF8D5" blockId="6.[151,1437,1787,1842]" pageId="6" pageNumber="35">
<emphasis id="62FA5847FFE3545EB6BBA2A1FEEDF8F4" bold="true" box="[151,289,1787,1811]" pageId="6" pageNumber="35">FIGURE 3.</emphasis>
<taxonomicName id="978EFFD6FFE3545EB705A2A1FDE0F8F5" box="[297,556,1787,1810]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFE3545EB705A2A1FDE0F8F5" box="[297,556,1787,1810]" italics="true" pageId="6" pageNumber="35">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE3545EB418A2A1FDB9F8F4" box="[564,629,1787,1811]" pageId="6" pageNumber="35" rank="species">
<emphasis id="62FA5847FFE3545EB418A2A1FDB9F8F4" bold="true" box="[564,629,1787,1811]" pageId="6" pageNumber="35">n. sp.</emphasis>
</taxonomicNameLabel>
Female. (A) chelicera, lateral view. (B) cheliceral digits in detail. (C) epirostrum, dorsal view. (D) epirostrum, lateral view. (E)
<taxonomicName id="978EFFD6FFE3545EB4D0A341FC4CF8D5" box="[764,896,1819,1842]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="6" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE3545EB4D0A341FC4CF8D5" box="[764,896,1819,1842]" italics="true" pageId="6" pageNumber="35">Penthalodes</emphasis>
</taxonomicName>
sp., epirostrum, lateral view.
</paragraph>
</caption>
<paragraph id="50318455FFE3545FB6BBA33AFCD9FE50" blockId="6.[151,1436,1888,2029]" lastBlockId="7.[151,1436,152,439]" lastPageId="7" lastPageNumber="36" pageId="6" pageNumber="35">
Opisthosoma composed of nine fused segments: C, D, E, F, H, PS, AD, AN, PE. Segment C with two pairs of setae
<emphasis id="62FA5847FFE3545EB6F4A3DCFF29F878" box="[216,229,1926,1951]" italics="true" pageId="6" pageNumber="35">c</emphasis>
1 and
<emphasis id="62FA5847FFE3545EB709A3DCFEFEF878" box="[293,306,1926,1951]" italics="true" pageId="6" pageNumber="35">c</emphasis>
2;
<emphasis id="62FA5847FFE3545EB765A3DCFE9AF878" box="[329,342,1926,1951]" italics="true" pageId="6" pageNumber="35">c</emphasis>
2 inserted slightly distad of
<emphasis id="62FA5847FFE3545EB4B8A3DCFD6DF878" box="[660,673,1926,1951]" italics="true" pageId="6" pageNumber="35">c</emphasis>
1, almost at the level of setae
<emphasis id="62FA5847FFE3545EB5DFA3DCFBDCF878" box="[1011,1040,1926,1951]" italics="true" pageId="6" pageNumber="35">bo</emphasis>
. Segments D to AN with one pair of setae each, i.e.,
<emphasis id="62FA5847FFE3545EB750A3F4FE47F820" box="[380,395,1966,1991]" italics="true" pageId="6" pageNumber="35">d</emphasis>
, e,
<emphasis id="62FA5847FFE3545EB796A3F4FE0EF820" box="[442,450,1966,1991]" italics="true" pageId="6" pageNumber="35">f</emphasis>
,
<emphasis id="62FA5847FFE3545EB7FFA3F4FE2EF820" box="[467,482,1966,1991]" italics="true" pageId="6" pageNumber="35">h</emphasis>
,
<emphasis id="62FA5847FFE3545EB7DEA3F4FDC1F820" box="[498,525,1966,1991]" italics="true" pageId="6" pageNumber="35">ps</emphasis>
,
<emphasis id="62FA5847FFE3545EB431A3F4FDF7F820" box="[541,571,1966,1991]" italics="true" pageId="6" pageNumber="35">ad</emphasis>
and
<emphasis id="62FA5847FFE3545EB455A3F4FD5BF820" box="[633,663,1966,1991]" italics="true" pageId="6" pageNumber="35">an</emphasis>
, respectively. Segments H to PE form the posterior wall of the opisthosoma which is steep, beginning at about the level of setae
<emphasis id="62FA5847FFE3545EB58FA389FC7EF80B" box="[931,946,2003,2028]" italics="true" pageId="6" pageNumber="35">h</emphasis>
(
<figureCitation id="C8B598D0FFE3545EB5EFA389FBC1F80A" box="[963,1037,2003,2029]" captionStart="FIGURE 2" captionStartId="5.[151,255,1020,1044]" captionTargetBox="[161,1430,205,979]" captionTargetId="figure@5.[151,1436,194,999]" captionTargetPageId="5" captionText="FIGURE 2. Penthalodes polonicus n. sp. Female. Idiosoma, lateral view." httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="6" pageNumber="35">Fig. 2</figureCitation>
). Distances between insertions of setae
<emphasis id="62FA5847FFE2545FB6F5A4C2FED6FF56" box="[217,282,152,177]" italics="true" pageId="7" pageNumber="36">ps-ad</emphasis>
,
<emphasis id="62FA5847FFE2545FB706A4C2FEA2FF56" box="[298,366,152,177]" italics="true" pageId="7" pageNumber="36">ad-an</emphasis>
are almost equal and rows of these setae arranged one under another; setae
<emphasis id="62FA5847FFE2545FB2C2A4C2FAC7FF56" box="[1262,1291,152,177]" italics="true" pageId="7" pageNumber="36">ad</emphasis>
well distant of anal cleft, hence the anal cleft is accompanied by only one pair of setae,
<emphasis id="62FA5847FFE2545FB26FA4E7FBADFF31" box="[1091,1121,189,214]" italics="true" pageId="7" pageNumber="36">an</emphasis>
. Setal pair
<emphasis id="62FA5847FFE2545FB2DDA4E7FB35FF31" box="[1265,1273,189,214]" italics="true" pageId="7" pageNumber="36">f</emphasis>
aligned with lyrifissures
<emphasis id="62FA5847FFE2545FB709A4B8FEF0FF1C" box="[293,316,226,251]" italics="true" pageId="7" pageNumber="36">ip</emphasis>
. Setae
<emphasis id="62FA5847FFE2545FB7BAA4B8FE69FF1C" box="[406,421,226,251]" italics="true" pageId="7" pageNumber="36">h</emphasis>
are inserted laterally and well posterior to setae
<emphasis id="62FA5847FFE2545FB5D8A4B8FC30FF1C" box="[1012,1020,226,251]" italics="true" pageId="7" pageNumber="36">f</emphasis>
and lyrifissures
<emphasis id="62FA5847FFE2545FB2EBA4B8FB12FF1C" box="[1223,1246,226,251]" italics="true" pageId="7" pageNumber="36">ip</emphasis>
. V-furrows end about at the level of setae
<emphasis id="62FA5847FFE2545FB7E3A552FE1BFEC6" box="[463,471,264,289]" italics="true" pageId="7" pageNumber="36">f</emphasis>
. Distances between insertions of opisthosomal setae distinctly longer than length of setae. Opisthosomal setae
<emphasis id="62FA5847FFE2545FB7D8A577FDCDFEA1" box="[500,513,301,326]" italics="true" pageId="7" pageNumber="36">c</emphasis>
to
<emphasis id="62FA5847FFE2545FB405A577FD8AFEA1" box="[553,582,301,326]" italics="true" pageId="7" pageNumber="36">ad</emphasis>
tridactylate with short stem, and prominent medial “dactyl”. With four pairs of lyrifissures (
<emphasis id="62FA5847FFE2545FB7A7A508FE6EFE8C" box="[395,418,338,363]" italics="true" pageId="7" pageNumber="36">ia</emphasis>
,
<emphasis id="62FA5847FFE2545FB79DA508FE02FE8C" box="[433,462,338,363]" italics="true" pageId="7" pageNumber="36">im</emphasis>
,
<emphasis id="62FA5847FFE2545FB7F1A508FE38FE8C" box="[477,500,338,363]" italics="true" pageId="7" pageNumber="36">ip</emphasis>
,
<emphasis id="62FA5847FFE2545FB42FA508FDD6FE8C" box="[515,538,338,363]" italics="true" pageId="7" pageNumber="36">ih</emphasis>
?), slit-like with strong sclerotized margins. Anal cleft very short, positioned terminally. Anal valves well developed, inserted on a small protuberance of segment PE. Setae
<emphasis id="62FA5847FFE2545FB327A522FAE4FE76" box="[1291,1320,376,401]" italics="true" pageId="7" pageNumber="36">an</emphasis>
plumose, close to anterior margin of PE, similar to genital setae.
</paragraph>
<caption id="04F1D4DDFFE2545FB6BBA5BDFBA4FE18" ID-Table-UUID="04F1D4DDFFE2545FB6BBA5BDFBA4FE18" box="[151,1128,487,511]" httpUri="http://table.plazi.org/id/04F1D4DDFFE2545FB6BBA5BDFBA4FE18" pageId="7" pageNumber="36" targetBox="[151,1391,538,1086]" targetIsTable="true" targetPageId="7">
<paragraph id="50318455FFE2545FB6BBA5BDFBA4FE18" blockId="7.[151,1128,487,511]" box="[151,1128,487,511]" pageId="7" pageNumber="36">
<emphasis id="62FA5847FFE2545FB6BBA5BDFEC4FE18" bold="true" box="[151,264,487,511]" pageId="7" pageNumber="36">TABLE 1.</emphasis>
Homology of opisthosomal setae in representatives of the family
<taxonomicName id="978EFFD6FFE2545FB5E5A5B2FBA8FE18" box="[969,1124,488,511]" class="Arachnida" family="Penthalodidae" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="36" phylum="Arthropoda" rank="family">Penthalodidae</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="50318455FFE2545FB6B3A640FAC3FBDD" pageId="7" pageNumber="36">
<table id="228E76F5FFE2ABA7B6BBA640FAA3FBD9" box="[151,1391,538,1086]" gridcols="6" gridrows="6" pageId="7" pageNumber="36">
<tr id="EEBE8617FFE2ABA7B6BBA640FAA3FD8D" box="[151,1391,538,618]" gridrow="0" pageId="7" pageNumber="36">
<th id="AD6FEF6BFFE2ABA7B6BBA640FE5AFD8D" box="[151,406,538,618]" gridcol="0" gridrow="0" pageId="7" pageNumber="36">Names of setae according to Strandtmann 1967, 1971</th>
<th id="AD6FEF6BFFE2ABA7B787A640FDBBFD8D" box="[427,631,538,618]" gridcol="1" gridrow="0" pageId="7" pageNumber="36">
<taxonomicName id="978EFFD6FFE2545FB787A640FDE8FDC8" box="[427,548,538,559]" class="Arachnida" family="Penthalodidae" genus="Stereotydeus" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE2545FB787A640FDE8FDC8" box="[427,548,538,559]" italics="true" pageId="7" pageNumber="36">Stereotydeus</emphasis>
</taxonomicName>
with new proposition for this taxon
</th>
<th id="AD6FEF6BFFE2ABA7B4B1A640FC9EFD8D" box="[669,850,538,618]" gridcol="2" gridrow="0" pageId="7" pageNumber="36">
<taxonomicName id="978EFFD6FFE2545FB4B1A640FC9EFDAB" authority="Jesionowska, 1989" authorityName="Jesionowska" authorityYear="1989" box="[669,850,538,588]" class="Arachnida" family="Penthalodidae" genus="Protopenthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE2545FB4B1A640FC8BFDC8" box="[669,839,538,559]" italics="true" pageId="7" pageNumber="36">Protopenthalodes</emphasis>
Jesionowska, 1989
</taxonomicName>
</th>
<th id="AD6FEF6BFFE2ABA7B55CA640FBE8FD8D" box="[880,1060,538,618]" gridcol="3" gridrow="0" pageId="7" pageNumber="36">
<taxonomicName id="978EFFD6FFE2545FB55CA640FC2BFDC8" box="[880,999,538,559]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE2545FB55CA640FC2BFDC8" box="[880,999,538,559]" italics="true" pageId="7" pageNumber="36">Penthalodes</emphasis>
</taxonomicName>
according to Baker 1987
</th>
<th id="AD6FEF6BFFE2ABA7B215A640FB7CFD8D" box="[1081,1200,538,618]" gridcol="4" gridrow="0" pageId="7" pageNumber="36">
<taxonomicName id="978EFFD6FFE2545FB215A640FB63FDC8" box="[1081,1199,538,559]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE2545FB215A640FB63FDC8" box="[1081,1199,538,559]" italics="true" pageId="7" pageNumber="36">Penthalodes</emphasis>
</taxonomicName>
according to Kaluz 2000
</th>
<th id="AD6FEF6BFFE2ABA7B2D5A640FAA3FD8D" box="[1273,1391,538,618]" gridcol="5" gridrow="0" pageId="7" pageNumber="36">
<taxonomicName id="978EFFD6FFE2545FB2D5A640FAA3FDC8" box="[1273,1391,538,559]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="7" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE2545FB2D5A640FAA3FDC8" box="[1273,1391,538,559]" italics="true" pageId="7" pageNumber="36">Penthalodes</emphasis>
</taxonomicName>
new proposition
</th>
</tr>
<tr id="EEBE8617FFE2ABA7B6BBA627FAA3FD27" box="[151,1391,637,704]" gridrow="1" pageId="7" pageNumber="36">
<th id="AD6FEF6BFFE2ABA7B6BBA627FE5AFD27" box="[151,406,637,704]" gridcol="0" gridrow="1" pageId="7" pageNumber="36">i.h. (internal humeral) e.h. (external humeral)</th>
<td id="AD6FEF6BFFE2ABA7B787A627FDBBFD27" box="[427,631,637,704]" gridcol="1" gridrow="1" pageId="7" pageNumber="36">present; (c1) present; (c2)</td>
<td id="AD6FEF6BFFE2ABA7B4B1A627FC9EFD27" box="[669,850,637,704]" gridcol="2" gridrow="1" pageId="7" pageNumber="36">present present</td>
<td id="AD6FEF6BFFE2ABA7B55CA627FBE8FD27" box="[880,1060,637,704]" gridcol="3" gridrow="1" pageId="7" pageNumber="36">c1 c2</td>
<td id="AD6FEF6BFFE2ABA7B215A627FB7CFD27" box="[1081,1200,637,704]" gridcol="4" gridrow="1" pageId="7" pageNumber="36">c1 sc</td>
<td id="AD6FEF6BFFE2ABA7B2D5A627FAA3FD27" box="[1273,1391,637,704]" gridcol="5" gridrow="1" pageId="7" pageNumber="36">c1 c2</td>
</tr>
<tr id="EEBE8617FFE2ABA7B6BBA688FAA3FCF2" box="[151,1391,722,789]" gridrow="2" pageId="7" pageNumber="36">
<th id="AD6FEF6BFFE2ABA7B6BBA688FE5AFCF2" box="[151,406,722,789]" gridcol="0" gridrow="2" pageId="7" pageNumber="36">d1 (first dorsal) d2 (second dorsal)</th>
<td id="AD6FEF6BFFE2ABA7B787A688FDBBFCF2" box="[427,631,722,789]" gridcol="1" gridrow="2" pageId="7" pageNumber="36">present; (d) present; (e)</td>
<td id="AD6FEF6BFFE2ABA7B4B1A688FC9EFCF2" box="[669,850,722,789]" gridcol="2" gridrow="2" pageId="7" pageNumber="36">absent present</td>
<td id="AD6FEF6BFFE2ABA7B55CA688FBE8FCF2" box="[880,1060,722,789]" gridcol="3" gridrow="2" pageId="7" pageNumber="36">d1 e1</td>
<td id="AD6FEF6BFFE2ABA7B215A688FB7CFCF2" box="[1081,1200,722,789]" gridcol="4" gridrow="2" pageId="7" pageNumber="36">d1 e1</td>
<td id="AD6FEF6BFFE2ABA7B2D5A688FAA3FCF2" box="[1273,1391,722,789]" gridcol="5" gridrow="2" pageId="7" pageNumber="36">- d</td>
</tr>
<tr id="EEBE8617FFE2ABA7B6BBA77DFAA3FCD8" box="[151,1391,807,831]" gridrow="3" pageId="7" pageNumber="36">
<th id="AD6FEF6BFFE2ABA7B6BBA77DFE5AFCD8" box="[151,406,807,831]" gridcol="0" gridrow="3" pageId="7" pageNumber="36">i.l. (internal lumbar)</th>
<td id="AD6FEF6BFFE2ABA7B787A77DFDBBFCD8" box="[427,631,807,831]" gridcol="1" gridrow="3" pageId="7" pageNumber="36">present; (f)</td>
<td id="AD6FEF6BFFE2ABA7B4B1A77DFC9EFCD8" box="[669,850,807,831]" gridcol="2" gridrow="3" pageId="7" pageNumber="36">present</td>
<td id="AD6FEF6BFFE2ABA7B55CA77DFBE8FCD8" box="[880,1060,807,831]" gridcol="3" gridrow="3" pageId="7" pageNumber="36">f1</td>
<td id="AD6FEF6BFFE2ABA7B215A77DFB7CFCD8" box="[1081,1200,807,831]" gridcol="4" gridrow="3" pageId="7" pageNumber="36">f1</td>
<td id="AD6FEF6BFFE2ABA7B2D5A77DFAA3FCD8" box="[1273,1391,807,831]" gridcol="5" gridrow="3" pageId="7" pageNumber="36">e</td>
</tr>
<tr id="EEBE8617FFE2ABA7B6BBA70BFAA3FC73" box="[151,1391,849,916]" gridrow="4" pageId="7" pageNumber="36">
<th id="AD6FEF6BFFE2ABA7B6BBA70BFE5AFC73" box="[151,406,849,916]" gridcol="0" gridrow="4" pageId="7" pageNumber="36">e.l. (external lumbar) i.s. (internal sacral)</th>
<td id="AD6FEF6BFFE2ABA7B787A70BFDBBFC73" box="[427,631,849,916]" gridcol="1" gridrow="4" pageId="7" pageNumber="36">present; (h) present; (ps1)</td>
<td id="AD6FEF6BFFE2ABA7B4B1A70BFC9EFC73" box="[669,850,849,916]" gridcol="2" gridrow="4" pageId="7" pageNumber="36">absent present</td>
<td id="AD6FEF6BFFE2ABA7B55CA70BFBE8FC73" box="[880,1060,849,916]" gridcol="3" gridrow="4" pageId="7" pageNumber="36">f2 absent</td>
<td id="AD6FEF6BFFE2ABA7B215A70BFB7CFC73" box="[1081,1200,849,916]" gridcol="4" gridrow="4" pageId="7" pageNumber="36">f2 h1</td>
<td id="AD6FEF6BFFE2ABA7B2D5A70BFAA3FC73" box="[1273,1391,849,916]" gridcol="5" gridrow="4" pageId="7" pageNumber="36">- f</td>
</tr>
<tr id="EEBE8617FFE2ABA7B6BBA7FCFAA3FBD9" box="[151,1391,934,1086]" gridrow="5" pageId="7" pageNumber="36">
<th id="AD6FEF6BFFE2ABA7B6BBA7FCFE5AFBD9" box="[151,406,934,1086]" gridcol="0" gridrow="5" pageId="7" pageNumber="36">e.s. (external sacral) a3 (anal seta No 3) or a1 a2 (anal seta No 2) a1 (anal seta No 1) or a3</th>
<td id="AD6FEF6BFFE2ABA7B787A7FCFDBBFBD9" box="[427,631,934,1086]" gridcol="1" gridrow="5" pageId="7" pageNumber="36">present; (ps2) present;(ad1) present; (ad2) present; (an)</td>
<td id="AD6FEF6BFFE2ABA7B4B1A7FCFC9EFBD9" box="[669,850,934,1086]" gridcol="2" gridrow="5" pageId="7" pageNumber="36">present present present present</td>
<td id="AD6FEF6BFFE2ABA7B55CA7FCFBE8FBD9" box="[880,1060,934,1086]" gridcol="3" gridrow="5" pageId="7" pageNumber="36">absent ps1 ps2 ps3</td>
<td id="AD6FEF6BFFE2ABA7B215A7FCFB7CFBD9" box="[1081,1200,934,1086]" gridcol="4" gridrow="5" pageId="7" pageNumber="36">absent ps ps absent</td>
<td id="AD6FEF6BFFE2ABA7B2D5A7FCFAA3FBD9" box="[1273,1391,934,1086]" gridcol="5" gridrow="5" pageId="7" pageNumber="36">h ps ad an</td>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFE2545FB6E9A0D8FD6CFA20" blockId="7.[151,1438,1154,2001]" pageId="7" pageNumber="36">
<emphasis id="62FA5847FFE2545FB6E9A0D8FEBDFB7C" box="[197,369,1154,1179]" italics="true" pageId="7" pageNumber="36">Genital region</emphasis>
(Figs 1B, 5;
<tableCitation id="1D0CB1EEFFE2545FB423A0D8FDA6FB7B" box="[527,618,1154,1180]" captionStart="TABLE 2" captionStartId="12.[151,239,601,625]" captionText="TABLE 2. Number of aggenital and genital setae in representatives of Penthalodes." pageId="7" pageNumber="36">Table 2</tableCitation>
). Costulae forming pentagons and irregular lines or points, distinctly smaller and not as homogenous as those on dorsum. Reticulation distinct in pregenital area, but punctate in lateral aggenital region. Genital valves strongly sclerotized, well separated from aggenital region, differently ornamented with irregular tiny spines. Between coxal regions IV and genital valves (pregenital area) eight aggenital setae form two alignments, the first with six and the second with two setae; another four pairs of aggenital setae are arranged lateral to the genital valves (i.e. total aggenital setae = 16). Nine pairs of plumose genital setae, inserted close to inner margin of genital valves except for the laterally situated fifth pair. Seven pairs of plumose eugenital setae, most probably eupathidial in form, inserted on well sclerotized protuberances. Two pairs of genital papillae.
</paragraph>
<paragraph id="50318455FFE2545FB6E9A188FAE3F966" blockId="7.[151,1438,1154,2001]" pageId="7" pageNumber="36">
<emphasis id="62FA5847FFE2545FB6E9A188FE8DFA0C" box="[197,321,1490,1515]" italics="true" pageId="7" pageNumber="36">Podosoma</emphasis>
(Figs 1B, 2, 6, 7). Coxal regions well developed, strongly sclerotized and ornamented; dorsolateral surface rugose, especially on coxal regions I and II. Ornament on podosoma delicately punctate; between coxal regions IV with delicate pentagons. Contiguous coxal regions I and II clearly separated from contiguous coxal regions III and IV; coxal formula: 2-1-2-1. Sternal region well developed, much more delicately ornamented than coxal regions; sternal formula: 2-0-2-2. Sternal and coxal setae plumose.
</paragraph>
<paragraph id="50318455FFE25451B6E9A2D7FCE0FB5F" blockId="7.[151,1438,1154,2001]" lastBlockId="9.[151,1436,1149,1212]" lastPageId="9" lastPageNumber="38" pageId="7" pageNumber="36">
<emphasis id="62FA5847FFE2545FB6E9A2D7FF30F941" box="[197,252,1677,1702]" italics="true" pageId="7" pageNumber="36">Legs</emphasis>
(
<figureCitation id="C8B598D0FFE2545FB720A2D7FE94F940" box="[268,344,1677,1703]" captionStart="FIGURE 6" captionStartId="10.[151,255,1938,1962]" captionTargetBox="[270,1329,202,1900]" captionTargetId="figure@10.[257,1329,162,1901]" captionTargetPageId="10" captionText="FIGURE 6. Penthalodes polonicus n. sp. (A) Leg I, lateral view. (B) Leg II, lateral view. (C) Leg III, lateral view. (D) Leg IV, tarsus and tibia-lateral view, femur and genu-dorsal view. Solenidia and famuli arrowed." httpUri="https://zenodo.org/record/199166/files/figure.png" pageId="7" pageNumber="36">Figs 6</figureCitation>
,
<figureCitation id="C8B598D0FFE2545FB748A2D7FEB9F940" box="[356,373,1677,1703]" captionStart="FIGURE 7" captionStartId="11.[151,256,1694,1718]" captionTargetBox="[312,1280,219,1660]" captionTargetId="figure@11.[293,1293,197,1673]" captionTargetPageId="11" captionText="FIGURE 7. Penthalodes polonicus n. sp. (A) Tarsus I, dorsal view. (B) Tarsus II, dorsal view. Solenidia and famuli arrowed." httpUri="https://zenodo.org/record/199167/files/figure.png" pageId="7" pageNumber="36">7</figureCitation>
). Legs shorter than idiosoma, leg I 348 long, Tr-Ts 30-110-50-58-100 long, leg II 265 (30- 75-40-50-70 long), leg III 275 (30-85-40-50-70 long), leg IV 325 (30-100-50-65-80 long). Chaetotaxy and solenidiotaxy (in parentheses) for legs I to IV:
<collectingCountry id="2899C4C5FFE2545FB494A28DFD1FF916" box="[696,723,1751,1777]" name="Chile" pageId="7" pageNumber="36">Tr</collectingCountry>
1-1-1-1, F 9-7-5-7, G 5(1+f)-5(1+f)-5(1)-4(1), Tb 6(2)-6(2)- 6(1)-6(1), Ts 23(2+1+f)-16(3+1)-15-15. Solenidia inserted dorsally, smooth, without striae. Rhagidial organ I with two banana-shaped solenidia (10 long each) in tandem in confluent depressions (
<figureCitation id="C8B598D0FFE2545FB288A378FB3DF8DB" box="[1188,1265,1826,1852]" captionStart="FIGURE 6" captionStartId="10.[151,255,1938,1962]" captionTargetBox="[270,1329,202,1900]" captionTargetId="figure@10.[257,1329,162,1901]" captionTargetPageId="10" captionText="FIGURE 6. Penthalodes polonicus n. sp. (A) Leg I, lateral view. (B) Leg II, lateral view. (C) Leg III, lateral view. (D) Leg IV, tarsus and tibia-lateral view, femur and genu-dorsal view. Solenidia and famuli arrowed." httpUri="https://zenodo.org/record/199166/files/figure.png" pageId="7" pageNumber="36">Figs 6</figureCitation>
A, 7A). Distal depression below proximal depression, shorter than solenidion. Laterally, at level of proximal solenidion, one T-shaped solenidion (6 long) inserted in pit which is considerably larger than solenidion. Small stellate famulus positioned more proximally, completely distant from rhagidial organ I. Rhagidial organ II with three thin, baculiform solenidia (11, 10 and 9 long, respectively), in tandem in common depression; proximal
<emphasis id="62FA5847FFEC5451B6BBA027FED1FB72" bold="true" box="[151,285,1149,1173]" pageId="9" pageNumber="38">FIGURE 5.</emphasis>
<taxonomicName id="978EFFD6FFEC5451B708A024FDD3FB72" box="[292,543,1150,1173]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="9" pageNumber="38" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFEC5451B708A024FDD3FB72" box="[292,543,1150,1173]" italics="true" pageId="9" pageNumber="38">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFEC5451B40AA027FDA9FB72" box="[550,613,1149,1173]" pageId="9" pageNumber="38" rank="species">
<emphasis id="62FA5847FFEC5451B40AA027FDA9FB72" bold="true" box="[550,613,1149,1173]" pageId="9" pageNumber="38">n. sp.</emphasis>
</taxonomicNameLabel>
Genital region. Genital valve with nine genital setae,
<emphasis id="62FA5847FFEC5451B292A024FB07FB72" box="[1214,1227,1150,1173]" italics="true" pageId="9" pageNumber="38">g</emphasis>
1-
<emphasis id="62FA5847FFEC5451B2F1A024FB26FB72" box="[1245,1258,1150,1173]" italics="true" pageId="9" pageNumber="38">g</emphasis>
9; seven pairs of eugenital setae,
<emphasis id="62FA5847FFEC5451B76EA0FBFEA4FB5F" box="[322,360,1185,1208]" italics="true" pageId="9" pageNumber="38">eug</emphasis>
1-
<emphasis id="62FA5847FFEC5451B754A0FBFE53FB5F" box="[376,415,1185,1208]" italics="true" pageId="9" pageNumber="38">eug</emphasis>
7; two pairs of genital papillae, pap1-2.
</paragraph>
<caption id="04F1D4DDFFED5450B6BBA317FC97F843" httpUri="https://zenodo.org/record/199165/files/figure.png" pageId="8" pageNumber="37" targetBox="[202,1390,194,1846]" targetPageId="8">
<paragraph id="50318455FFED5450B6BBA317FC97F843" blockId="8.[151,1436,1869,1956]" pageId="8" pageNumber="37">
<emphasis id="62FA5847FFED5450B6BBA317FED6F882" bold="true" box="[151,282,1869,1893]" pageId="8" pageNumber="37">FIGURE 4.</emphasis>
<taxonomicName id="978EFFD6FFED5450B70EA317FDD9F883" box="[290,533,1869,1892]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="8" pageNumber="37" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFED5450B70EA317FDD9F883" box="[290,533,1869,1892]" italics="true" pageId="8" pageNumber="37">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFED5450B431A317FD95F882" box="[541,601,1869,1893]" pageId="8" pageNumber="37" rank="species">
<emphasis id="62FA5847FFED5450B431A317FD95F882" bold="true" box="[541,601,1869,1893]" pageId="8" pageNumber="37">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Subcapitulum, ventral view; (B) Subcapitulum, dorsal view; LB-labrum, LL-lateral lips, LI-labium, H-hypostome, sc-internal sclerites, n,m-subcapitular setae, cxp-coxal region of pedipalp. (C) Pedipalp, lateral view; (D) Pedipalpal tarsus in detail, lateral view.
</paragraph>
</caption>
<paragraph id="50318455FFEC5451B6BBA0B3FE45F8EA" blockId="9.[151,1437,1257,2029]" pageId="9" pageNumber="38">
solenidion the thickest; solenidia adjacent and overlapping one another because distal part of depression is positioned below proximal part (like cascade) (
<figureCitation id="C8B598D0FFEC5451B497A154FCCAFACF" box="[699,774,1294,1320]" captionStart="FIGURE 6" captionStartId="10.[151,255,1938,1962]" captionTargetBox="[270,1329,202,1900]" captionTargetId="figure@10.[257,1329,162,1901]" captionTargetPageId="10" captionText="FIGURE 6. Penthalodes polonicus n. sp. (A) Leg I, lateral view. (B) Leg II, lateral view. (C) Leg III, lateral view. (D) Leg IV, tarsus and tibia-lateral view, femur and genu-dorsal view. Solenidia and famuli arrowed." httpUri="https://zenodo.org/record/199166/files/figure.png" pageId="9" pageNumber="38">Figs 6</figureCitation>
B, 7B). Spine-like tiny solenidion (or famulus; 2 long) in small pit lateral to proximal solenidion. Tibia I with two dorsodistal solenidia, proximal solenidion (6 long) erect and baculiform, distal solenidion (8 long) banana-shaped, recumbent in depression; distal solenidion thickest; distance between solenidia 9. Tibia II with two dorsodistal solenidia; proximal solenidion (5 long), rod-like, distal solenidion (10 long) thick, banana-shaped, in depression; distance between solenidia 6. Tibia III and IV each with one erect, baculiform dorsoproximal solenidion (6 and 7 long, respectively). Genua IIV each distally with large (especially large on G I, II and IV) banana-shaped solenidion (13, 13, 9 and 13 long, respectively); genu I and II each with tiny spiniform famulus (4 and 3 long, respectively; in own pore) located near distal margin of article; famulus on genu I with one lateral spine (bifurcate). Three pairs of setae expanded distally near apotele I-IV each. All leg setae shorter than leg article, plumose with long distal outgrowths (spicule), especially long on femur II setae. Ventral tarsal setae angled basally, i.e. foot-shaped. Leg integument strongly sclerotized with ornamentation composed of tiny spines and minute tubercles. Apotele IIV each with ambulacrum composed of two claws and pad-like, setuled empodium. Ambulacrum of leg IV the largest.
</paragraph>
</subSubSection>
<subSubSection id="1894D7DEFFEC5453B6E9A342FB7EF8BA" lastPageId="11" lastPageNumber="40" pageId="9" pageNumber="38" type="materials_examined">
<paragraph id="50318455FFEC5451B6E9A342FBEFF80A" blockId="9.[151,1437,1257,2029]" pageId="9" pageNumber="38">
<emphasis id="62FA5847FFEC5451B6E9A342FE09F8D5" bold="true" box="[197,453,1816,1842]" pageId="9" pageNumber="38">Material examined.</emphasis>
Female
<typeStatus id="8F353AF7FFEC5451B41CA343FD57F8D4" box="[560,667,1817,1843]" pageId="9" pageNumber="38" type="holotype">holotype</typeStatus>
(KJJ724-1), Zakopane. Tatrzański Park Narodowy (The Tatra Mountains National Park). Dolina Chochołowska (Chochołowska Valley). Stone under the peaks of &quot;Olejarnia&quot; in Niżna Brama Chochołowska; mosses, grass and soil.
<date id="2430A295FFEC5451B5C9A339FBB8F89A" box="[997,1140,1891,1917]" pageId="9" pageNumber="38" value="1979-09-20">20.09.1979</date>
. Leg. J. Rafalski. Three females, same data as
<typeStatus id="8F353AF7FFEC5451B7B1A3D3FDCBF844" box="[413,519,1929,1955]" pageId="9" pageNumber="38" type="holotype">holotype</typeStatus>
. Further material examined: two females (one
<typeStatus id="8F353AF7FFEC5451B200A3D3FB5DF844" box="[1068,1169,1929,1955]" pageId="9" pageNumber="38" type="paratype">paratype</typeStatus>
KJJ797-2), Zakopane. The Tatra Mountains National Park. &quot;Wąwóz Kraków&quot; (Krakow Ravine); mosses, grass, soil and detritus from stone cracks and cavities (limestone).
<date id="2430A295FFEC5451B4B8A389FCD0F80A" box="[660,796,2003,2029]" pageId="9" pageNumber="38" value="1983-10-10">10.10.1983</date>
. Leg. Z. Olszanowski.
</paragraph>
<caption id="04F1D4DDFFEF5452B6BBA3C8FB57F82E" httpUri="https://zenodo.org/record/199166/files/figure.png" pageId="10" pageNumber="39" targetBox="[270,1329,202,1900]" targetPageId="10">
<paragraph id="50318455FFEF5452B6BBA3C8FB57F82E" blockId="10.[151,1436,1938,1993]" pageId="10" pageNumber="39">
<emphasis id="62FA5847FFEF5452B6BBA3C8FED7F84D" bold="true" box="[151,283,1938,1962]" pageId="10" pageNumber="39">FIGURE 6.</emphasis>
<taxonomicName id="978EFFD6FFEF5452B70EA3C8FDDAF84E" box="[290,534,1938,1961]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="10" pageNumber="39" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFEF5452B70EA3C8FDDAF84E" box="[290,534,1938,1961]" italics="true" pageId="10" pageNumber="39">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFEF5452B431A3C8FD96F84D" box="[541,602,1938,1962]" pageId="10" pageNumber="39" rank="species">
<emphasis id="62FA5847FFEF5452B431A3C8FD96F84D" bold="true" box="[541,602,1938,1962]" pageId="10" pageNumber="39">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Leg I, lateral view. (B) Leg II, lateral view. (C) Leg III, lateral view. (D) Leg IV, tarsus and tibia-lateral view, femur and genu-dorsal view. Solenidia and famuli arrowed.
</paragraph>
</caption>
<caption id="04F1D4DDFFEE5453B6BBA2C4FF3AF932" httpUri="https://zenodo.org/record/199167/files/figure.png" pageId="11" pageNumber="40" targetBox="[312,1280,219,1660]" targetPageId="11">
<paragraph id="50318455FFEE5453B6BBA2C4FF3AF932" blockId="11.[151,1436,1694,1749]" pageId="11" pageNumber="40">
<emphasis id="62FA5847FFEE5453B6BBA2C4FED0F951" bold="true" box="[151,284,1694,1718]" pageId="11" pageNumber="40">FIGURE 7.</emphasis>
<taxonomicName id="978EFFD6FFEE5453B708A2C4FDD0F952" box="[292,540,1694,1717]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFEE5453B708A2C4FDD0F952" box="[292,540,1694,1717]" italics="true" pageId="11" pageNumber="40">Penthalodes polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFEE5453B408A2C4FDAEF951" box="[548,610,1694,1718]" pageId="11" pageNumber="40" rank="species">
<emphasis id="62FA5847FFEE5453B408A2C4FDAEF951" bold="true" box="[548,610,1694,1718]" pageId="11" pageNumber="40">n. sp.</emphasis>
</taxonomicNameLabel>
(A) Tarsus I, dorsal view. (B) Tarsus II, dorsal view. Solenidia and famuli arrowed.
</paragraph>
</caption>
<paragraph id="50318455FFEE5453B6E9A344FB7EF8BA" blockId="11.[151,1436,1822,2035]" pageId="11" pageNumber="40">The material is deposited in the author's collection. In future, specimens will be transferred to the Acarological Collection of the Faculty of Biology, Adam Mickiewicz University, Poznań.</paragraph>
</subSubSection>
<subSubSection id="1894D7DEFFEE5454B6E9A332FCD1FE75" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40" type="etymology">
<paragraph id="50318455FFEE5453B6E9A332FB90F864" blockId="11.[151,1436,1822,2035]" box="[197,1116,1896,1923]" pageId="11" pageNumber="40">
<emphasis id="62FA5847FFEE5453B6E9A332FE9DF865" bold="true" box="[197,337,1896,1922]" pageId="11" pageNumber="40">Etymology.</emphasis>
The species name refers to the Polish part of the Tatra Mountains.
</paragraph>
<paragraph id="50318455FFEE5454B6E9A3D4FCD1FE75" blockId="11.[151,1436,1822,2035]" lastBlockId="12.[151,1437,152,551]" lastPageId="12" lastPageNumber="41" pageId="11" pageNumber="40">
<emphasis id="62FA5847FFEE5453B6E9A3D4FE17F84F" bold="true" box="[197,475,1934,1960]" pageId="11" pageNumber="40">Differential diagnosis.</emphasis>
This new species, although also European, differs from
<taxonomicName id="978EFFD6FFEE5453B253A3D5FAB1F84F" authority="Kaluz" authorityName="Kaluz" box="[1151,1405,1934,1960]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="11" pageNumber="40" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEE5453B253A3D5FB28F84F" box="[1151,1252,1935,1960]" italics="true" pageId="11" pageNumber="40">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
in the following combination of characteristics: (1) setae
<emphasis id="62FA5847FFEE5453B534A3EEFCFCF82A" box="[792,816,1972,1997]" italics="true" pageId="11" pageNumber="40">ro</emphasis>
adjacent, i.e., inserted close to each other; (2) dorsal idiosomal setae tridactylate (except setae
<emphasis id="62FA5847FFEE5453B450A383FD58F815" box="[636,660,2009,2034]" italics="true" pageId="11" pageNumber="40">ro</emphasis>
and
<emphasis id="62FA5847FFEE5453B4E0A383FD25F815" box="[716,745,2009,2034]" italics="true" pageId="11" pageNumber="40">bo</emphasis>
); (3) epirostrum with weakly developed semi-round lateral lobes; (4) epirostral ornamentation composed of irregular tubercles, not pentagons or reticular pattern; (5) solenidia in rhagidial organ I banana-shaped, in tandem in confluent depressions; not lanceolate; (6) all solenidia in the rhagidial organ II in tandem, slightly overlapping each other, thin and baculiform, none lanceolate; (7) lateral solenidion on tarsus I T-shaped, not spiniform; (8) pedipalpal tarsus elongated, linear in profile; (9) stellate famulus on tarsus I well removed from rhagidial organ I; (10) number of aggenital setae 16, not 23; in pregenital region, eight setae in two alignments (six and two, respectively), not 13 (five and eight, respectively); (11) seven pairs of eugenital setae.
</paragraph>
</subSubSection>
<subSubSection id="1894D7DEFFE9544BB6E9A5C7FEB3FC0A" lastPageId="19" lastPageNumber="48" pageId="12" pageNumber="41" type="reference_group">
<paragraph id="50318455FFE95454B6E9A5C7FC8BFDC0" blockId="12.[151,1437,152,551]" pageId="12" pageNumber="41">
In
<taxonomicName id="978EFFD6FFE95454B6CAA5C7FD8CFE50" authority="Kaluz (2000)" authorityName="Kaluz" authorityYear="2000" box="[230,576,413,439]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE95454B6CAA5C7FE87FE51" box="[230,331,413,438]" italics="true" pageId="12" pageNumber="41">P. ovalis</emphasis>
sensu
<bibRefCitation id="341FF9A4FFE95454B7B2A5C7FD8CFE50" author="Kaluz" box="[414,576,413,439]" pageId="12" pageNumber="41" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
</taxonomicName>
, the solenidia on tarsi I and II are arranged in a specific way. Rhagidial organ I has two lanceolate solenidia arranged linearly in separate depressions, with a third spiniform solenidion placed laterally in a pit. Rhagidial organ II has a lanceolate proximal solenidion placed separately from two parallel distal solenidia in a common depression.
</paragraph>
<caption id="04F1D4DDFFE95454B6BBA603FBD9FD97" box="[151,1045,601,625]" pageId="12" pageNumber="41">
<paragraph id="50318455FFE95454B6BBA603FBD9FD97" blockId="12.[151,1408,601,825]" box="[151,1045,601,625]" pageId="12" pageNumber="41">
<emphasis id="62FA5847FFE95454B6BBA603FEC4FD96" bold="true" box="[151,264,601,625]" pageId="12" pageNumber="41">TABLE 2.</emphasis>
Number of aggenital and genital setae in representatives of
<taxonomicName id="978EFFD6FFE95454B5A7A600FBC2FD96" box="[907,1038,602,625]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE95454B5A7A600FBC2FD96" box="[907,1038,602,625]" italics="true" pageId="12" pageNumber="41">Penthalodes</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="50318455FFE95454B6B3A6D7FA4CFD43" blockId="12.[151,1408,601,825]" box="[159,1408,653,676]" pageId="12" pageNumber="41">Species Aggenital setae in pregenital Rest of Total sum of Genital setae</paragraph>
<paragraph id="50318455FFE95454B416A6F7FB11FD23" blockId="12.[151,1408,601,825]" box="[570,1245,685,708]" pageId="12" pageNumber="41">region aggenital setae aggenital setae</paragraph>
<paragraph id="50318455FFE95454B416A682FCE5FD08" blockId="12.[151,1408,601,825]" box="[570,809,728,751]" pageId="12" pageNumber="41">first second</paragraph>
<paragraph id="50318455FFE95454B416A6A2FC86FCE8" blockId="12.[151,1408,601,825]" box="[570,842,760,783]" pageId="12" pageNumber="41">alignment alignment</paragraph>
<paragraph id="50318455FFE95454B6B3A778FCA3F9D5" blockId="12.[151,1408,601,825]" lastBlockId="12.[151,1437,1410,1996]" pageId="12" pageNumber="41">
<emphasis id="62FA5847FFE95454B6B3A778FF39FCDE" box="[159,245,802,825]" italics="true" pageId="12" pageNumber="41">P. o v a l i s</emphasis>
sensu
<bibRefCitation id="341FF9A4FFE95454B76DA778FE70FCDE" author="Kaluz" box="[321,444,802,825]" pageId="12" pageNumber="41" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz 2000</bibRefCitation>
(
<figureCitation id="C8B598D0FFE95454B7E7A778FDC9FCDE" box="[459,517,802,825]" captionStart="FIGURE 2" captionStartId="5.[151,255,1020,1044]" captionTargetBox="[161,1430,205,979]" captionTargetId="figure@5.[151,1436,194,999]" captionTargetPageId="5" captionText="FIGURE 2. Penthalodes polonicus n. sp. Female. Idiosoma, lateral view." httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="12" pageNumber="41">Fig.2</figureCitation>
)
<date id="2430A295FFE95454B416A778FC6CFCDE" box="[570,928,802,825]" pageId="12" pageNumber="41" value="1910-08-05">5 8 10</date>
23 9+9
<emphasis id="62FA5847FFE95454B6BBA1D8FB08FA7B" bold="true" box="[151,1220,1410,1436]" pageId="12" pageNumber="41">
Morphological survey of hitherto described representatives of the
<taxonomicName id="978EFFD6FFE95454B231A1D8FB71FA7B" box="[1053,1213,1410,1436]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE95454B231A1D8FB71FA7B" bold="true" box="[1053,1213,1410,1436]" italics="true" pageId="12" pageNumber="41">Penthalodes</emphasis>
</taxonomicName>
.
</emphasis>
Eight species of
<taxonomicName id="978EFFD6FFE95454B6BBA1F2FEE4FA26" box="[151,296,1448,1473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE95454B6BBA1F2FEE4FA26" box="[151,296,1448,1473]" italics="true" pageId="12" pageNumber="41">Penthalodes</emphasis>
</taxonomicName>
are now identifiable. Four species,
<taxonomicName id="978EFFD6FFE95454B4E1A1F2FCBEFA26" box="[717,882,1448,1473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="alaskaensis" status="sp. nov.">
<emphasis id="62FA5847FFE95454B4E1A1F2FCBEFA26" box="[717,882,1448,1473]" italics="true" pageId="12" pageNumber="41">P. alaskaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE95454B556A1FDFC71FA26" box="[890,957,1447,1473]" pageId="12" pageNumber="41" rank="species">
<emphasis id="62FA5847FFE95454B556A1FDFC71FA26" bold="true" box="[890,957,1447,1473]" pageId="12" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
,
<taxonomicName id="978EFFD6FFE95454B5E0A1F2FBFEFA26" box="[972,1074,1448,1473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFE95454B5E0A1F2FBFEFA26" box="[972,1074,1448,1473]" italics="true" pageId="12" pageNumber="41">P. boneti</emphasis>
</taxonomicName>
,
<taxonomicName id="978EFFD6FFE95454B26EA1F2FB20FA26" box="[1090,1260,1448,1473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFE95454B26EA1F2FB20FA26" box="[1090,1260,1448,1473]" italics="true" pageId="12" pageNumber="41">P. oregonensis</emphasis>
</taxonomicName>
and
<emphasis id="62FA5847FFE95454B30AA1F2FA5CFA26" box="[1318,1424,1448,1473]" italics="true" pageId="12" pageNumber="41">P. t u r n e r i</emphasis>
, are known from North and Central
<collectingCountry id="2899C4C5FFE95454B46FA197FD66FA00" box="[579,682,1485,1511]" name="United States of America" pageId="12" pageNumber="41">America</collectingCountry>
(
<bibRefCitation id="341FF9A4FFE95454B497A197FC87FA00" author="Baker" box="[699,843,1485,1511]" pageId="12" pageNumber="41" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFE95454B497A197FCC8FA00" box="[699,772,1485,1511]" name="United States Pacific Island Wildlife Refuges" pageId="12" pageNumber="41">Baker</collectingCountry>
1946
</bibRefCitation>
;
<bibRefCitation id="341FF9A4FFE95454B575A197FBFEFA00" author="Strandmann" box="[857,1074,1485,1511]" pageId="12" pageNumber="41" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandmann 1971</bibRefCitation>
), two species,
<taxonomicName id="978EFFD6FFE95454B2C8A197FA86FA01" box="[1252,1354,1485,1510]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE95454B2C8A197FA86FA01" box="[1252,1354,1485,1510]" italics="true" pageId="12" pageNumber="41">P. ovalis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFE95454B3AAA197FEC5F9EB" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFE95454B3AAA197FEC5F9EB" italics="true" pageId="12" pageNumber="41">P. polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE95454B73CA1A8FE9EF9EB" box="[272,338,1522,1548]" pageId="12" pageNumber="41" rank="species">
<emphasis id="62FA5847FFE95454B73CA1A8FE9EF9EB" bold="true" box="[272,338,1522,1548]" pageId="12" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
are from Europe (
<bibRefCitation id="341FF9A4FFE95454B404A1A8FCAFF9EB" author="Kaluz" box="[552,867,1522,1548]" pageId="12" pageNumber="41" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz 2000; present paper</bibRefCitation>
),
<emphasis id="62FA5847FFE95454B55AA1A9FC32F9EB" box="[886,1022,1523,1548]" italics="true" pageId="12" pageNumber="41">P. c a r i n a t u s</emphasis>
is from
<collectingCountry id="2899C4C5FFE95454B248A1A8FB6BF9EB" box="[1124,1191,1522,1548]" name="Japan" pageId="12" pageNumber="41">Japan</collectingCountry>
(
<bibRefCitation id="341FF9A4FFE95454B29AA1A8FA8CF9EB" author="Shiba" box="[1206,1344,1522,1548]" pageId="12" pageNumber="41" refString="Shiba, M. (1978) On some eupodiform mites from Japan (Acarina: Prostigmata). Reports of research Matsuyama Shinonome Junior College, 19, 133 - 152." type="journal article" year="1978">Shiba 1978</bibRefCitation>
), and
<taxonomicName id="978EFFD6FFE95454B3AAA1A9FEEEF9D6" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="hawaiiensis" status="sp. nov.">
<emphasis id="62FA5847FFE95454B3AAA1A9FEEEF9D6" italics="true" pageId="12" pageNumber="41">P. hawaiiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE95454B704A24DFEA6F9D6" box="[296,362,1559,1585]" pageId="12" pageNumber="41" rank="species">
<emphasis id="62FA5847FFE95454B704A24DFEA6F9D6" bold="true" box="[296,362,1559,1585]" pageId="12" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
is from Hawaii (
<bibRefCitation id="341FF9A4FFE95454B41CA242FCAEF9D5" author="Strandmann" box="[560,866,1560,1586]" pageId="12" pageNumber="41" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandmann &amp; Goff 1978</bibRefCitation>
).
</paragraph>
<paragraph id="50318455FFE95454B6B3A717FED7FAB3" pageId="12" pageNumber="41">
<table id="228E76F5FFE9ABA7B6BBA717FA4CFAB3" box="[151,1408,845,1364]" gridcols="4" gridrows="9" pageId="12" pageNumber="41">
<tr id="EEBE8617FFE9ABA7B6BBA717FA4CFC83" box="[151,1408,845,868]" gridrow="0" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA717FD8BFC83" box="[151,583,845,868]" gridcol="0" gridrow="0" pageId="12" pageNumber="41">
<taxonomicName id="978EFFD6FFE95454B6B3A717FDDCFC83" authority="Strandtmann 1971" authorityName="Strandtmann" authorityYear="1971" box="[159,528,845,868]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFE95454B6B3A717FEF8FC83" box="[159,308,845,868]" italics="true" pageId="12" pageNumber="41">P. alaskaensis</emphasis>
(Strandtmann 1971)
</taxonomicName>
5
</th>
<th id="AD6FEF6BFFE9ABA7B4F3A717FD20FC83" box="[735,748,845,868]" gridcol="1" gridrow="0" pageId="12" pageNumber="41">4</th>
<th id="AD6FEF6BFFE9ABA7B5A9A717FC0FFC83" box="[901,963,845,868]" gridcol="2" gridrow="0" pageId="12" pageNumber="41">8 or 9</th>
<th id="AD6FEF6BFFE9ABA7B26CA717FA4CFC83" box="[1088,1408,845,868]" gridcol="3" gridrow="0" pageId="12" pageNumber="41">17 or 18 7+7</th>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA722FA4CFC68" box="[151,1408,888,911]" gridrow="1" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA722FD8BFC68" box="[151,583,888,911]" gridcol="0" gridrow="1" pageId="12" pageNumber="41">
<emphasis id="62FA5847FFE95454B6B3A722FF23FC68" box="[159,239,888,911]" italics="true" pageId="12" pageNumber="41">P.o v a l i s</emphasis>
sensu Baker 1990 2
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A722FD20FC68" box="[735,748,888,911]" gridcol="1" gridrow="1" pageId="12" pageNumber="41">4</td>
<td id="AD6FEF6BFFE9ABA7B5A9A722FC0FFC68" box="[901,963,888,911]" gridcol="2" gridrow="1" pageId="12" pageNumber="41">6</td>
<td id="AD6FEF6BFFE9ABA7B26CA722FA4CFC68" box="[1088,1408,888,911]" gridcol="3" gridrow="1" pageId="12" pageNumber="41">12 10+10</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA7F8FA4CFC5E" box="[151,1408,930,953]" gridrow="2" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA7F8FD8BFC5E" box="[151,583,930,953]" gridcol="0" gridrow="2" pageId="12" pageNumber="41">
<emphasis id="62FA5847FFE95454B6B3A7F8FED6FC5E" box="[159,282,930,953]" italics="true" pageId="12" pageNumber="41">P. c a r i n a t u s</emphasis>
Shiba 1978 4
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A7F8FD20FC5E" box="[735,748,930,953]" gridcol="1" gridrow="2" pageId="12" pageNumber="41">4</td>
<td id="AD6FEF6BFFE9ABA7B5A9A7F8FC0FFC5E" box="[901,963,930,953]" gridcol="2" gridrow="2" pageId="12" pageNumber="41">8</td>
<td id="AD6FEF6BFFE9ABA7B26CA7F8FA4CFC5E" box="[1088,1408,930,953]" gridcol="3" gridrow="2" pageId="12" pageNumber="41">16 8+8</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA797FA4CFC03" box="[151,1408,973,996]" gridrow="3" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA797FD8BFC03" box="[151,583,973,996]" gridcol="0" gridrow="3" pageId="12" pageNumber="41">
<emphasis id="62FA5847FFE95454B6B3A797FECCFC03" box="[159,256,973,996]" italics="true" pageId="12" pageNumber="41">P. t u r n e r i</emphasis>
Baker 1946?
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A797FD20FC03" box="[735,748,973,996]" gridcol="1" gridrow="3" pageId="12" pageNumber="41">?</td>
<td id="AD6FEF6BFFE9ABA7B5A9A797FC0FFC03" box="[901,963,973,996]" gridcol="2" gridrow="3" pageId="12" pageNumber="41">?</td>
<td id="AD6FEF6BFFE9ABA7B26CA797FA4CFC03" box="[1088,1408,973,996]" gridcol="3" gridrow="3" pageId="12" pageNumber="41">16 9+9</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA7A2FA4CFBE8" box="[151,1408,1016,1039]" gridrow="4" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA7A2FD8BFBE8" box="[151,583,1016,1039]" gridcol="0" gridrow="4" pageId="12" pageNumber="41">
<taxonomicName id="978EFFD6FFE95454B6B3A7A2FEB2FBE8" authority="Baker 1946" authorityName="Baker" authorityYear="1946" box="[159,382,1016,1039]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFE95454B6B3A7A2FF30FBE8" box="[159,252,1016,1039]" italics="true" pageId="12" pageNumber="41">P. boneti</emphasis>
Baker 1946
</taxonomicName>
4
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A7A2FD20FBE8" box="[735,748,1016,1039]" gridcol="1" gridrow="4" pageId="12" pageNumber="41">6</td>
<td id="AD6FEF6BFFE9ABA7B5A9A7A2FC0FFBE8" box="[901,963,1016,1039]" gridcol="2" gridrow="4" pageId="12" pageNumber="41">8</td>
<td id="AD6FEF6BFFE9ABA7B26CA7A2FA4CFBE8" box="[1088,1408,1016,1039]" gridcol="3" gridrow="4" pageId="12" pageNumber="41">18 9+9</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA078FA4CFBDD" box="[151,1408,1058,1082]" gridrow="5" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA078FD8BFBDD" box="[151,583,1058,1082]" gridcol="0" gridrow="5" pageId="12" pageNumber="41">
<taxonomicName id="978EFFD6FFE95454B6B3A078FEEDFBDE" box="[159,289,1058,1081]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="polonicus" status="sp. nov.">
<emphasis id="62FA5847FFE95454B6B3A078FEEDFBDE" box="[159,289,1058,1081]" italics="true" pageId="12" pageNumber="41">P. polonicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="79C9E53CFFE95454B70BA078FEAFFBDD" box="[295,355,1058,1082]" pageId="12" pageNumber="41" rank="species">
<emphasis id="62FA5847FFE95454B70BA078FEAFFBDD" bold="true" box="[295,355,1058,1082]" pageId="12" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
(present paper) 6
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A078FD20FBDD" box="[735,748,1058,1082]" gridcol="1" gridrow="5" pageId="12" pageNumber="41">2</td>
<td id="AD6FEF6BFFE9ABA7B5A9A078FC0FFBDD" box="[901,963,1058,1082]" gridcol="2" gridrow="5" pageId="12" pageNumber="41">8</td>
<td id="AD6FEF6BFFE9ABA7B26CA078FA4CFBDD" box="[1088,1408,1058,1082]" gridcol="3" gridrow="5" pageId="12" pageNumber="41">16 9+9</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA017FA4CFB63" box="[151,1408,1101,1156]" gridrow="6" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA017FD8BFB63" box="[151,583,1101,1156]" gridcol="0" gridrow="6" pageId="12" pageNumber="41">
<emphasis id="62FA5847FFE95454B6B3A017FEFFFB83" box="[159,307,1101,1124]" italics="true" pageId="12" pageNumber="41">P. h a w a i i e n s i s</emphasis>
(Strandtmann et Goff? 1978)
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A017FD20FB63" box="[735,748,1101,1156]" gridcol="1" gridrow="6" pageId="12" pageNumber="41">?</td>
<td id="AD6FEF6BFFE9ABA7B5A9A017FC0FFB63" box="[901,963,1101,1156]" gridcol="2" gridrow="6" pageId="12" pageNumber="41">?</td>
<td id="AD6FEF6BFFE9ABA7B26CA017FA4CFB63" box="[1088,1408,1101,1156]" gridcol="3" gridrow="6" pageId="12" pageNumber="41">? 8+8 or 9+9</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA0C2FA4CFB28" box="[151,1408,1176,1231]" gridrow="7" pageId="12" pageNumber="41">
<th id="AD6FEF6BFFE9ABA7B6BBA0C2FD8BFB28" box="[151,583,1176,1231]" gridcol="0" gridrow="7" pageId="12" pageNumber="41">
<taxonomicName id="978EFFD6FFE95454B6B3A0C2FDD9FB48" authority="Thor and Willmann" authorityName="Thor and Willmann" box="[159,533,1176,1199]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE95454B6B3A0C2FEF6FB48" box="[159,314,1176,1199]" italics="true" pageId="12" pageNumber="41">P. ovalis sensu</emphasis>
Thor and Willmann
</taxonomicName>
,? 1941
</th>
<td id="AD6FEF6BFFE9ABA7B4F3A0C2FD20FB28" box="[735,748,1176,1231]" gridcol="1" gridrow="7" pageId="12" pageNumber="41">?</td>
<td id="AD6FEF6BFFE9ABA7B5A9A0C2FC0FFB28" box="[901,963,1176,1231]" gridcol="2" gridrow="7" pageId="12" pageNumber="41">?</td>
<td id="AD6FEF6BFFE9ABA7B26CA0C2FA4CFB28" box="[1088,1408,1176,1231]" gridcol="3" gridrow="7" pageId="12" pageNumber="41">? 6+6</td>
</tr>
<tr id="EEBE8617FFE9ABA7B6BBA160FA4CFAB3" box="[151,1408,1338,1364]" gridrow="8" pageId="12" pageNumber="41" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="AD6FEF6BFFE9ABA7B6BBA160FD8BFAB3" box="[151,583,1338,1364]" gridcol="0" gridrow="8" pageId="12" pageNumber="41">Discussion</th>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFE95454B6E9A267FB55F90B" blockId="12.[151,1437,1410,1996]" pageId="12" pageNumber="41">
The descriptions by
<bibRefCitation id="341FF9A4FFE95454B793A267FDAFF9B0" author="Baker" box="[447,611,1597,1623]" pageId="12" pageNumber="41" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFE95454B793A267FDCBF9B0" box="[447,519,1597,1623]" name="United States Pacific Island Wildlife Refuges" pageId="12" pageNumber="41">Baker</collectingCountry>
(1946)
</bibRefCitation>
,
<bibRefCitation id="341FF9A4FFE95454B45DA267FC90F9B0" author="Strandmann" box="[625,860,1597,1623]" pageId="12" pageNumber="41" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandmann (1971)</bibRefCitation>
and
<bibRefCitation id="341FF9A4FFE95454B5B5A267FB2AF9B0" author="Strandmann" box="[921,1254,1597,1623]" pageId="12" pageNumber="41" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandmann &amp; Goff (1978)</bibRefCitation>
are all lacking detail to some degree, but allow their recognition as distinct species. Bakers (1946) description details the shape of epirostrum, an important feature for separating these species. Nevertheless, drawings of rhagidial organ I and II and detail of other leg solenidia are lacking. Some data, now considered important in descriptions, are also missing from
<bibRefCitation id="341FF9A4FFE95454B41BA288FCD6F90B" author="Strandmann" box="[567,794,1746,1772]" pageId="12" pageNumber="41" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandmann (1971)</bibRefCitation>
and
<bibRefCitation id="341FF9A4FFE95454B57FA288FB59F90B" author="Strandmann" box="[851,1173,1746,1772]" pageId="12" pageNumber="41" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandmann &amp; Goff (1978)</bibRefCitation>
.
</paragraph>
<paragraph id="50318455FFE95454B6E9A2A2FDD3F82B" blockId="12.[151,1437,1410,1996]" pageId="12" pageNumber="41">
Shiba's (1978) description of
<taxonomicName id="978EFFD6FFE95454B431A2A2FD65F8F6" box="[541,681,1784,1809]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFE95454B431A2A2FD65F8F6" box="[541,681,1784,1809]" italics="true" pageId="12" pageNumber="41">P. carinatus</emphasis>
</taxonomicName>
is also brief, but is supported by drawings where, among others, the shape and the arrangement of solenidia on the tarsi and tibiae of legs I and II, as well as the shape of the epirostrum, distinguish this species from all others. However,
<bibRefCitation id="341FF9A4FFE95454B58FA318FB89F8BB" author="Shiba" box="[931,1093,1858,1884]" pageId="12" pageNumber="41" refString="Shiba, M. (1978) On some eupodiform mites from Japan (Acarina: Prostigmata). Reports of research Matsuyama Shinonome Junior College, 19, 133 - 152." type="journal article" year="1978">Shiba (1978)</bibRefCitation>
thought the main character distinguishing it from similar species (
<taxonomicName id="978EFFD6FFE95454B470A332FD73F866" box="[604,703,1896,1921]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE95454B470A332FD73F866" box="[604,703,1896,1921]" italics="true" pageId="12" pageNumber="41">P. ovalis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFE95454B4D5A332FC93F866" box="[761,863,1896,1921]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFE95454B4D5A332FC93F866" box="[761,863,1896,1921]" italics="true" pageId="12" pageNumber="41">P. boneti</emphasis>
</taxonomicName>
) was the presence of eight pairs of genital setae. The redescription of
<taxonomicName id="978EFFD6FFE95454B7BFA3D7FE35F841" box="[403,505,1933,1958]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE95454B7BFA3D7FE35F841" box="[403,505,1933,1958]" italics="true" pageId="12" pageNumber="41">P. ovalis</emphasis>
</taxonomicName>
by
<bibRefCitation id="341FF9A4FFE95454B40BA3D7FD0AF840" author="Kaluz" box="[551,710,1933,1959]" pageId="12" pageNumber="41" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
presents good drawings for his putative
<taxonomicName id="978EFFD6FFE95454B29EA3D7FAD4F841" box="[1202,1304,1933,1958]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="12" pageNumber="41" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE95454B29EA3D7FAD4F841" box="[1202,1304,1933,1958]" italics="true" pageId="12" pageNumber="41">P. ovalis</emphasis>
</taxonomicName>
specimens collected in
<collectingCountry id="2899C4C5FFE95454B70AA3E8FE40F82B" box="[294,396,1970,1996]" name="Slovakia" pageId="12" pageNumber="41">Slovakia</collectingCountry>
and
<collectingCountry id="2899C4C5FFE95454B7E9A3E8FDD7F82B" box="[453,539,1970,1996]" name="Turkey" pageId="12" pageNumber="41">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="50318455FFE85455B6E9A4C2FBF7FE75" blockId="13.[151,1437,152,924]" pageId="13" pageNumber="42">
<bibRefCitation id="341FF9A4FFE85455B6E9A4C2FDDAFF55" author="Thor" box="[197,534,152,178]" pageId="13" pageNumber="42" refString="Thor, S. &amp; Willmann, C. (1941) Acarina, Prostigmata - Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. Das Tierreich, 71 A, 1 - 186." type="journal article" year="1941">Thor and Willmann (1941)</bibRefCitation>
promoted the use of the name
<taxonomicName id="978EFFD6FFE85455B58CA4C2FBC7FF56" box="[928,1035,152,177]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE85455B58CA4C2FBC7FF56" box="[928,1035,152,177]" italics="true" pageId="13" pageNumber="42">P. ovalis</emphasis>
</taxonomicName>
, and the genus
<taxonomicName id="978EFFD6FFE85455B2FFA4C2FAA1FF56" box="[1235,1389,152,177]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE85455B2FFA4C2FAA1FF56" box="[1235,1389,152,177]" italics="true" pageId="13" pageNumber="42">Penthalodes</emphasis>
</taxonomicName>
, by presenting a list of diagnostic characters for the genus: (1) epirostrum present; (2) reticulate ornament of the body composed of tetragons, pentagons or hexagons; (3) furrows in the form of the letter V on dorsum; (4) no epivertex (i.e. naso); instead of it, an eye-like organ with a pair of rudimentary hairs; (5) legs shorter than the body; and (6) setae of the body very small, plumose or branched. Due to Thor and Wilmanns (1941) work, most ecological works and check-lists use the name
<taxonomicName id="978EFFD6FFE85455B530A508FC48FE8C" box="[796,900,338,363]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE85455B530A508FC48FE8C" box="[796,900,338,363]" italics="true" pageId="13" pageNumber="42">P. ovalis</emphasis>
</taxonomicName>
when they probably mean
<taxonomicName id="978EFFD6FFE85455B2FFA508FAA5FE8C" box="[1235,1385,338,363]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE85455B2FFA508FAA5FE8C" box="[1235,1385,338,363]" italics="true" pageId="13" pageNumber="42">Penthalodes</emphasis>
</taxonomicName>
sp., because many do not make proper reference to the species-level characteristics.
</paragraph>
<paragraph id="50318455FFE85455B6E9A5C7FCC4FDE5" blockId="13.[151,1437,152,924]" pageId="13" pageNumber="42">
The following morphological review allows for a new understanding of the characteristic features of adults of the genus
<taxonomicName id="978EFFD6FFE85455B7A2A598FDE4FE3C" box="[398,552,450,475]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE85455B7A2A598FDE4FE3C" box="[398,552,450,475]" italics="true" pageId="13" pageNumber="42">Penthalodes</emphasis>
</taxonomicName>
, comparisons to other representatives of the Eupodoidea, and the new diagnoses and species already presented in this work.
</paragraph>
<paragraph id="50318455FFE85455B6E9A657FB7FFD70" blockId="13.[151,1437,152,924]" pageId="13" pageNumber="42">
<emphasis id="62FA5847FFE85455B6E9A657FEF7FDC0" bold="true" box="[197,315,525,551]" italics="true" pageId="13" pageNumber="42">Idiosoma</emphasis>
. The idiosoma has characteristic shape; angular in profile, while being slightly narrowed posteriorly in dorsal view. Its anterior and posterior part is steep (
<figureCitation id="C8B598D0FFE85455B5BCA668FC1BFDAB" box="[912,983,562,588]" captionStart="FIGURE 2" captionStartId="5.[151,255,1020,1044]" captionTargetBox="[161,1430,205,979]" captionTargetId="figure@5.[151,1436,194,999]" captionTargetPageId="5" captionText="FIGURE 2. Penthalodes polonicus n. sp. Female. Idiosoma, lateral view." httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="13" pageNumber="42">Fig. 2</figureCitation>
). When viewed in lactic acid, it can be flattened laterally. Other authors describe it as plump, ovoid or almost globose. On the dorsum, there are characteristic furrows arranged like the letter “V” or “
<collectingCountry id="2899C4C5FFE85455B522A627FCE4FD70" box="[782,808,637,663]" name="Italy" pageId="13" pageNumber="42">Y</collectingCountry>
”, depending on slide preparation.
</paragraph>
<paragraph id="50318455FFE85455B6E9A6F8FA5DFC7B" blockId="13.[151,1437,152,924]" pageId="13" pageNumber="42">
<emphasis id="62FA5847FFE85455B6E9A6F8FE83FD5B" bold="true" box="[197,335,674,700]" italics="true" pageId="13" pageNumber="42">Epirostrum</emphasis>
. This structure resembles a small roof over the gnathosoma. The epirostrum is positioned well below the naso and is either trilobed or monolobed (
<tableCitation id="1D0CB1EEFFE85455B4D4A692FC9EFD05" box="[760,850,712,738]" captionStart="TABLE 3" captionStartId="13.[151,239,972,996]" captionTargetBox="[151,1422,1024,1313]" captionTargetPageId="13" captionText="TABLE 3. Form of the epirostrum in identifiable species of Penthalodes; P. ovalis is sensu Kaluz (2000)." httpUri="http://table.plazi.org/id/04F1D4DDFFE85455B6BBA796FB32FC03" pageId="13" pageNumber="42" tableUuid="04F1D4DDFFE85455B6BBA796FB32FC03">Table 3</tableCitation>
). The central lobe is always obviously the largest. The epirostrum of
<taxonomicName id="978EFFD6FFE85455B75FA6B7FDC8FCE1" box="[371,516,749,774]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE85455B75FA6B7FDC8FCE1" box="[371,516,749,774]" italics="true" pageId="13" pageNumber="42">Penthalodes</emphasis>
</taxonomicName>
sp. presented by
<bibRefCitation id="341FF9A4FFE85455B4FEA6B7FCA1FCE0" author="Baker" box="[722,877,749,775]" pageId="13" pageNumber="42" refString="Baker, A. S. (1987) Caleupodes, a new genus of eupodoid mite (Acari: Acariformes) showing primary opisthosomal segmentation. Bulletin of the British Museum (Natural History) (Zoology Series), Miscellanea, 53 (2) 103 - 113. Baker, A. S. (1990) A survey of external morphology of mites of the superfamily Eupodoidea Banks, 1894 (Acari: Acariformes). Journal of Natural History, 24, 1227 - 1261." type="journal article" year="1987">
<collectingCountry id="2899C4C5FFE85455B4FEA6B7FCD4FCE0" box="[722,792,749,775]" name="United States Pacific Island Wildlife Refuges" pageId="13" pageNumber="42">Baker</collectingCountry>
(1987)
</bibRefCitation>
is most similar to
<emphasis id="62FA5847FFE85455B265A6B7FB78FCE1" box="[1097,1204,749,774]" italics="true" pageId="13" pageNumber="42">P. t u r n e r i</emphasis>
. Ornamentation on the epirostrum is usually reticulate in the basal part, and linear or punctate in the apical part, but does vary in some species. The profile of the epirostrum can differ, e.g. the dorsal line can be sloping from the base, as in
<taxonomicName id="978EFFD6FFE85455B3AAA762FEC5FC91" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFE85455B3AAA762FEC5FC91" italics="true" pageId="13" pageNumber="42">P. polonicus</emphasis>
</taxonomicName>
, or can initially form a small platform (like in other representatives of the genus collected in
<collectingCountry id="2899C4C5FFE85455B36FA707FA5BFC90" box="[1347,1431,861,887]" name="Poland" pageId="13" pageNumber="42">Poland</collectingCountry>
,
<figureCitation id="C8B598D0FFE85455B6BBA7D8FF16FC7B" box="[151,218,898,924]" captionStart="FIGURE 3" captionStartId="6.[151,259,1787,1811]" captionTargetBox="[218,1372,205,1741]" captionTargetId="figure@6.[212,1376,194,1765]" captionTargetPageId="6" captionText="FIGURE 3. Penthalodes polonicus n. sp. Female. (A) chelicera, lateral view. (B) cheliceral digits in detail. (C) epirostrum, dorsal view. (D) epirostrum, lateral view. (E) Penthalodes sp., epirostrum, lateral view." httpUri="https://zenodo.org/record/199164/files/figure.png" pageId="13" pageNumber="42">Fig. 3</figureCitation>
E). The epirostrum does not occur in the larva and tritonymph (
<bibRefCitation id="341FF9A4FFE85455B5EDA7D8FB50FC7B" author="Jesionowska" box="[961,1180,898,924]" pageId="13" pageNumber="42" refString="Jesionowska, K. (1996) Morphology of the juvenile stages of eupodoid mites (Eupodoidea), In: Mitchell R., Horn D. J., Needham G. R. &amp; Welbourn W. C. (Eds), Acarology IX- Proceedings. Vol. 1. Ohio Biological Survey, Ohio, USA, pp. 325 - 333." type="book" year="1996">Jesionowska 1996</bibRefCitation>
;
<bibRefCitation id="341FF9A4FFE85455B284A7D8FA48FC7B" author="Strandtmann" box="[1192,1412,898,924]" pageId="13" pageNumber="42" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann 1971</bibRefCitation>
).
</paragraph>
<caption id="04F1D4DDFFE85455B6BBA796FB32FC03" ID-Table-UUID="04F1D4DDFFE85455B6BBA796FB32FC03" box="[151,1278,972,996]" httpUri="http://table.plazi.org/id/04F1D4DDFFE85455B6BBA796FB32FC03" pageId="13" pageNumber="42" targetBox="[151,1422,1024,1313]" targetIsTable="true" targetPageId="13">
<paragraph id="50318455FFE85455B6BBA796FB32FC03" blockId="13.[151,1278,972,996]" box="[151,1278,972,996]" pageId="13" pageNumber="42">
<emphasis id="62FA5847FFE85455B6BBA796FEC4FC03" bold="true" box="[151,264,972,996]" pageId="13" pageNumber="42">TABLE 3.</emphasis>
Form of the epirostrum in identifiable species of
<emphasis id="62FA5847FFE85455B537A797FBC5FC03" box="[795,1033,973,996]" italics="true" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B537A797FC6DFC03" box="[795,929,973,996]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">Penthalodes</taxonomicName>
;
<taxonomicName id="978EFFD6FFE85455B582A797FBC5FC03" box="[942,1033,973,996]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="ovalis">P. ovalis</taxonomicName>
</emphasis>
is sensu Kaluz (2000).
</paragraph>
</caption>
<paragraph id="50318455FFE85455B6B3A05AFA4AFAC6" pageId="13" pageNumber="42">
<table id="228E76F5FFE8ABA7B6BBA05AFA42FAC6" box="[151,1422,1024,1313]" gridcols="5" gridrows="5" pageId="13" pageNumber="42">
<tr id="EEBE8617FFE8ABA7B6BBA05AFA42FBF0" box="[151,1422,1024,1047]" gridrow="0" pageId="13" pageNumber="42">
<th id="AD6FEF6BFFE8ABA7B6BBA05AFEECFBF0" box="[151,288,1024,1047]" gridcol="0" gridrow="0" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A05AFEECFBF0" box="[159,288,1024,1047]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="49" phylum="Arthropoda" rank="genus">Penthalodes</taxonomicName>
</th>
<th id="AD6FEF6BFFE8ABA7B779A05AFE5AFBF0" box="[341,406,1024,1047]" gridcol="1" gridrow="0" pageId="13" pageNumber="42">Lobes</th>
<th id="AD6FEF6BFFE8ABA7B799A05AFD7FFBF0" box="[437,691,1024,1047]" gridcol="2" gridrow="0" pageId="13" pageNumber="42">Central lobe</th>
<th id="AD6FEF6BFFE8ABA7B4CFA05AFB95FBF0" box="[739,1113,1024,1047]" gridcol="3" gridrow="0" pageId="13" pageNumber="42">Lateral lobes</th>
<th id="AD6FEF6BFFE8ABA7B259A05AFA42FBF0" box="[1141,1422,1024,1047]" gridcol="4" gridrow="0" pageId="13" pageNumber="42">Ornamentation</th>
</tr>
<tr id="EEBE8617FFE8ABA7B6BBA070FA42FBA6" box="[151,1422,1066,1089]" gridrow="1" pageId="13" pageNumber="42">
<th id="AD6FEF6BFFE8ABA7B6BBA070FEECFBA6" box="[151,288,1066,1089]" gridcol="0" gridrow="1" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A070FF13FBA6" box="[159,223,1066,1089]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFE85455B6B3A070FF13FBA6" box="[159,223,1066,1089]" italics="true" pageId="13" pageNumber="42">ovalis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFE8ABA7B779A070FE5AFBA6" box="[341,406,1066,1089]" gridcol="1" gridrow="1" pageId="13" pageNumber="42">3</td>
<td id="AD6FEF6BFFE8ABA7B799A070FD7FFBA6" box="[437,691,1066,1089]" gridcol="2" gridrow="1" pageId="13" pageNumber="42">Triangular</td>
<td id="AD6FEF6BFFE8ABA7B4CFA070FB95FBA6" box="[739,1113,1066,1089]" gridcol="3" gridrow="1" pageId="13" pageNumber="42">lingulate, directed obliquely</td>
<td id="AD6FEF6BFFE8ABA7B259A070FA42FBA6" box="[1141,1422,1066,1089]" gridcol="4" gridrow="1" pageId="13" pageNumber="42">Wholly reticulate</td>
</tr>
<tr id="EEBE8617FFE8ABA7B6BBA00FFA42FB6B" box="[151,1422,1109,1164]" gridrow="2" pageId="13" pageNumber="42">
<th id="AD6FEF6BFFE8ABA7B6BBA00FFEECFB6B" box="[151,288,1109,1164]" gridcol="0" gridrow="2" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A00FFED6FB8B" box="[159,282,1109,1132]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFE85455B6B3A00FFED6FB8B" box="[159,282,1109,1132]" italics="true" pageId="13" pageNumber="42">alaskaensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFE8ABA7B779A00FFE5AFB6B" box="[341,406,1109,1164]" gridcol="1" gridrow="2" pageId="13" pageNumber="42">3</td>
<td id="AD6FEF6BFFE8ABA7B799A00FFD7FFB6B" box="[437,691,1109,1164]" gridcol="2" gridrow="2" pageId="13" pageNumber="42">Pentagonal; broad base, short triangular tip</td>
<td id="AD6FEF6BFFE8ABA7B4CFA00FFB95FB6B" box="[739,1113,1109,1164]" gridcol="3" gridrow="2" pageId="13" pageNumber="42">sharp apices directed antiaxially</td>
<td id="AD6FEF6BFFE8ABA7B259A00FFA42FB6B" box="[1141,1422,1109,1164]" gridcol="4" gridrow="2" pageId="13" pageNumber="42">Wholly reticulate</td>
</tr>
<tr id="EEBE8617FFE8ABA7B6BBA0FAFA42FB30" box="[151,1422,1184,1239]" gridrow="3" pageId="13" pageNumber="42">
<th id="AD6FEF6BFFE8ABA7B6BBA0FAFEECFB30" box="[151,288,1184,1239]" gridcol="0" gridrow="3" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A0FAFED1FB50" box="[159,285,1184,1207]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFE85455B6B3A0FAFED1FB50" box="[159,285,1184,1207]" italics="true" pageId="13" pageNumber="42">hawaiiensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFE8ABA7B779A0FAFE5AFB30" box="[341,406,1184,1239]" gridcol="1" gridrow="3" pageId="13" pageNumber="42">3</td>
<td id="AD6FEF6BFFE8ABA7B799A0FAFD7FFB30" box="[437,691,1184,1239]" gridcol="2" gridrow="3" pageId="13" pageNumber="42">Triangular</td>
<td id="AD6FEF6BFFE8ABA7B4CFA0FAFB95FB30" box="[739,1113,1184,1239]" gridcol="3" gridrow="3" pageId="13" pageNumber="42">slightly pointed, weakly separated from central lobe</td>
<td id="AD6FEF6BFFE8ABA7B259A0FAFA42FB30" box="[1141,1422,1184,1239]" gridcol="4" gridrow="3" pageId="13" pageNumber="42">Wholly reticulate</td>
</tr>
<tr id="EEBE8617FFE8ABA7B6BBA0B0FA42FAC6" box="[151,1422,1258,1313]" gridrow="4" pageId="13" pageNumber="42">
<th id="AD6FEF6BFFE8ABA7B6BBA0B0FEECFAC6" box="[151,288,1258,1313]" gridcol="0" gridrow="4" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A0B0FECBFAE6" box="[159,263,1258,1281]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFE85455B6B3A0B0FECBFAE6" box="[159,263,1258,1281]" italics="true" pageId="13" pageNumber="42">polonicus</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFE8ABA7B779A0B0FE5AFAC6" box="[341,406,1258,1313]" gridcol="1" gridrow="4" pageId="13" pageNumber="42">3</td>
<td id="AD6FEF6BFFE8ABA7B799A0B0FD7FFAC6" box="[437,691,1258,1313]" gridcol="2" gridrow="4" pageId="13" pageNumber="42">Triangular</td>
<td id="AD6FEF6BFFE8ABA7B4CFA0B0FB95FAC6" box="[739,1113,1258,1313]" gridcol="3" gridrow="4" pageId="13" pageNumber="42">broadly rounded, weakly separated</td>
<td id="AD6FEF6BFFE8ABA7B259A0B0FA42FAC6" box="[1141,1422,1258,1313]" gridcol="4" gridrow="4" pageId="13" pageNumber="42">Wholly punctuate with large and minute costulae</td>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFE85455B6B3A16FFAE3FAAB" blockId="13.[159,1422,1333,1580]" box="[159,1327,1333,1356]" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A16FFEC8FAAB" box="[159,260,1333,1356]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFE85455B6B3A16FFEC8FAAB" box="[159,260,1333,1356]" italics="true" pageId="13" pageNumber="42">carinatus</emphasis>
</taxonomicName>
1 Triangular; edges slightly plicate Wholly reticulate
</paragraph>
<paragraph id="50318455FFE85455B6B3A13AFA42FA90" blockId="13.[159,1422,1333,1580]" box="[159,1422,1376,1399]" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A13AFF25FA90" box="[159,233,1376,1399]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="turneri">
<emphasis id="62FA5847FFE85455B6B3A13AFF25FA90" box="[159,233,1376,1399]" italics="true" pageId="13" pageNumber="42">turneri</emphasis>
</taxonomicName>
1 Triangular with rounded apex Basally reticulate, apically
</paragraph>
<paragraph id="50318455FFE85455B259A1DAFB7AFA70" blockId="13.[159,1422,1333,1580]" box="[1141,1206,1408,1431]" pageId="13" pageNumber="42">striate</paragraph>
<paragraph id="50318455FFE85455B6B3A1F0FA42FA26" blockId="13.[159,1422,1333,1580]" box="[159,1422,1450,1473]" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A1F0FF2EFA26" box="[159,226,1450,1473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFE85455B6B3A1F0FF2EFA26" box="[159,226,1450,1473]" italics="true" pageId="13" pageNumber="42">boneti</emphasis>
</taxonomicName>
1 Rectangular or trapezoidal transformed medially into ligulate Basally reticulate, apically
</paragraph>
<paragraph id="50318455FFE85455B799A190FB7AFA06" blockId="13.[159,1422,1333,1580]" box="[437,1206,1482,1505]" pageId="13" pageNumber="42">process, cusps slightly pointed striate</paragraph>
<paragraph id="50318455FFE85455B6B3A1AFFA41F9EB" blockId="13.[159,1422,1333,1580]" box="[159,1421,1525,1548]" pageId="13" pageNumber="42">
<taxonomicName id="978EFFD6FFE85455B6B3A1AFFED3F9EB" box="[159,287,1525,1548]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFE85455B6B3A1AFFED3F9EB" box="[159,287,1525,1548]" italics="true" pageId="13" pageNumber="42">oregonensis</emphasis>
</taxonomicName>
1 Large, undivided, lobed, with broadly rounded apical part Basally punctuate, distally
</paragraph>
<paragraph id="50318455FFE85455B259A24FFC1AF88B" blockId="13.[159,1422,1333,1580]" lastBlockId="13.[151,1437,1650,2015]" pageId="13" pageNumber="42">
with tiny hairs
<emphasis id="62FA5847FFE85455B6E9A228FEC8F96B" bold="true" box="[197,260,1650,1676]" italics="true" pageId="13" pageNumber="42">Naso</emphasis>
. The naso of
<taxonomicName id="978EFFD6FFE85455B785A229FDF0F96B" box="[425,572,1651,1676]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE85455B785A229FDF0F96B" box="[425,572,1651,1676]" italics="true" pageId="13" pageNumber="42">Penthalodes</emphasis>
</taxonomicName>
is unique. It is spherical, smooth and completely disconnected from the epirostrum below. Only the anterior hemisphere of the naso is visible in dorsal view, while the other hemisphere is hidden in a cavity of the prodorsum wall.
<bibRefCitation id="341FF9A4FFE85455B51DA2E7FB41F930" author="Strandtmann" box="[817,1165,1725,1751]" pageId="13" pageNumber="42" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandtmann and Goff (1978)</bibRefCitation>
observed that the naso is submerged in the anterior part of idiosoma. A detailed description of the appearance of the naso is presented by
<bibRefCitation id="341FF9A4FFE85455B690A352FE3AF8C5" author="Haupt" box="[188,502,1800,1826]" pageId="13" pageNumber="42" refString="Haupt, J. &amp; Coineau, Y. (2002) Morphologie comparee du naso des Penthalodidae (Acari, Actinedida, Penthalodidae). Acarologia, 42 (3), 247 - 256." type="journal article" year="2002">Haupt and Coineau (2002)</bibRefCitation>
, and its scanning electron micrograph is given by
<collectingCountry id="2899C4C5FFE85455B26EA352FB44F8C5" box="[1090,1160,1800,1826]" name="United States Pacific Island Wildlife Refuges" pageId="13" pageNumber="42">Baker</collectingCountry>
(1990). The naso bears a pair of minute, nude, unbranched setae
<emphasis id="62FA5847FFE85455B4B4A377FD7DF8A1" box="[664,689,1837,1862]" italics="true" pageId="13" pageNumber="42">ro</emphasis>
.
<bibRefCitation id="341FF9A4FFE85455B4EFA377FBDCF8A0" author="Thor" box="[707,1040,1837,1863]" pageId="13" pageNumber="42" refString="Thor, S. &amp; Willmann, C. (1941) Acarina, Prostigmata - Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. Das Tierreich, 71 A, 1 - 186." type="journal article" year="1941">Thor and Willmann (1941)</bibRefCitation>
considered the naso absent, but instead a an eye-like structure (organ) with a pair of rudimentary setae.
</paragraph>
<paragraph id="50318455FFE85456B6E9A32DFD08FEAD" blockId="13.[151,1437,1650,2015]" lastBlockId="14.[151,1437,152,2008]" lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42">
<emphasis id="62FA5847FFE85455B6E9A32DFBDEF876" bold="true" box="[197,1042,1911,1937]" italics="true" pageId="13" pageNumber="42">Idiosomal setae, except setae of the genital region and podosoma</emphasis>
(
<tableCitation id="1D0CB1EEFFE85455B20FA322FB4DF875" box="[1059,1153,1912,1938]" captionStart="TABLE 1" captionStartId="7.[151,239,487,511]" captionTargetBox="[151,1391,538,1086]" captionTargetPageId="7" captionText="TABLE 1. Homology of opisthosomal setae in representatives of the family Penthalodidae." httpUri="http://table.plazi.org/id/04F1D4DDFFE2545FB6BBA5BDFBA4FE18" pageId="13" pageNumber="42" tableUuid="04F1D4DDFFE2545FB6BBA5BDFBA4FE18">Table 1</tableCitation>
). The number and the arrangement of idiosomal setae are characteristic for
<taxonomicName id="978EFFD6FFE85455B539A3C7FC64F851" box="[789,936,1949,1974]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="13" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFE85455B539A3C7FC64F851" box="[789,936,1949,1974]" italics="true" pageId="13" pageNumber="42">Penthalodes</emphasis>
</taxonomicName>
. They lack setae
<emphasis id="62FA5847FFE85455B257A3C7FB46F851" box="[1147,1162,1949,1974]" italics="true" pageId="13" pageNumber="42">d</emphasis>
1 and
<emphasis id="62FA5847FFE85455B2E2A3C7FB3FF851" box="[1230,1267,1949,1974]" italics="true" pageId="13" pageNumber="42">e.l.</emphasis>
(
<bibRefCitation id="341FF9A4FFE85455B32EA3C7FF1AF838" author="Strandtmann" pageId="13" pageNumber="42" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann 1971</bibRefCitation>
), which are usually found in other Eupodoidea, making just 13 pairs of idiosomal setae, of which four pairs fall on the prodorsum and nine pairs on the opisthosoma. Because the idiosoma is angular (geometric solid body), some setae are located on the steep walls of its anterior and posterior parts (
<figureCitation id="C8B598D0FFEB5456B286A4E7FB3EFF30" box="[1194,1266,189,215]" captionStart="FIGURE 2" captionStartId="5.[151,255,1020,1044]" captionTargetBox="[161,1430,205,979]" captionTargetId="figure@5.[151,1436,194,999]" captionTargetPageId="5" captionText="FIGURE 2. Penthalodes polonicus n. sp. Female. Idiosoma, lateral view." httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="14" pageNumber="43">Fig. 2</figureCitation>
). Therefore, it is difficult to be confident about setal names of the dorsum and the venter because they can optically change their location after flattening of the idiosoma. For example, setae
<emphasis id="62FA5847FFEB5456B590A552FC05FEC6" box="[956,969,264,289]" italics="true" pageId="14" pageNumber="43">c</emphasis>
<date id="2430A295FFEB5456B5E6A542FBC3FEC5" box="[970,1039,264,294]" pageId="14" pageNumber="43">2 may</date>
be placed slightly anterior to
<emphasis id="62FA5847FFEB5456B352A552FA47FEC6" box="[1406,1419,264,289]" italics="true" pageId="14" pageNumber="43">c</emphasis>
1, giving the illusion that they are prodorsal setae.
</paragraph>
<paragraph id="50318455FFEB5456B6EAA50FFEB0FD44" blockId="14.[151,1437,152,2008]" pageId="14" pageNumber="43">
<bibRefCitation id="341FF9A4FFEB5456B6EAA50FFE7DFE88" author="Strandtmann" box="[198,433,341,367]" pageId="14" pageNumber="43" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann (1971)</bibRefCitation>
and
<bibRefCitation id="341FF9A4FFEB5456B7C6A50FFD4BFE88" author="Baker" box="[490,647,341,367]" pageId="14" pageNumber="43" refString="Baker, A. S. (1987) Caleupodes, a new genus of eupodoid mite (Acari: Acariformes) showing primary opisthosomal segmentation. Bulletin of the British Museum (Natural History) (Zoology Series), Miscellanea, 53 (2) 103 - 113. Baker, A. S. (1990) A survey of external morphology of mites of the superfamily Eupodoidea Banks, 1894 (Acari: Acariformes). Journal of Natural History, 24, 1227 - 1261." type="journal article" year="1987">
<collectingCountry id="2899C4C5FFEB5456B7C6A50FFDFCFE88" box="[490,560,341,367]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
(1987)
</bibRefCitation>
considered
<taxonomicName id="978EFFD6FFEB5456B538A50CFC69FE88" box="[788,933,342,367]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEB5456B538A50CFC69FE88" box="[788,933,342,367]" italics="true" pageId="14" pageNumber="43">Penthalodes</emphasis>
</taxonomicName>
with four pairs of prodorsal and nine pairs of opisthosomal setae, contrary to
<bibRefCitation id="341FF9A4FFEB5456B465A520FD22FE73" author="Kaluz" box="[585,750,378,404]" pageId="14" pageNumber="43" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
who proposed an unusual arrangement of five pairs of prodorsal and eight pairs of opisthosomal setae. The prodorsal setae are thought to be
<emphasis id="62FA5847FFEB5456B252A5FAFB5DFE5E" box="[1150,1169,416,441]" italics="true" pageId="14" pageNumber="43">iv</emphasis>
,
<emphasis id="62FA5847FFEB5456B28CA5FAFB76FE5E" box="[1184,1210,416,441]" italics="true" pageId="14" pageNumber="43">ev</emphasis>
1,
<emphasis id="62FA5847FFEB5456B2FCA5FAFB26FE5E" box="[1232,1258,416,441]" italics="true" pageId="14" pageNumber="43">ev</emphasis>
2,
<emphasis id="62FA5847FFEB5456B32CA5FAFADCFE5E" box="[1280,1296,416,441]" italics="true" pageId="14" pageNumber="43">T</emphasis>
and
<emphasis id="62FA5847FFEB5456B364A5FAFAACFE5E" box="[1352,1376,416,441]" italics="true" pageId="14" pageNumber="43">sc</emphasis>
. The extra prodorsal setae are obtained by considering the opisthosomal setae
<emphasis id="62FA5847FFEB5456B5C2A593FC37FE05" box="[1006,1019,457,482]" italics="true" pageId="14" pageNumber="43">c</emphasis>
2 as the scapular setae
<emphasis id="62FA5847FFEB5456B32CA593FAD4FE05" box="[1280,1304,457,482]" italics="true" pageId="14" pageNumber="43">sc</emphasis>
. Five pairs of prodorsal setae are atypical for Eupodoidea, which have just four pairs of fundamental prodorsal setae.
<bibRefCitation id="341FF9A4FFEB5456B6BBA64CFEFFFDD7" author="Baker" box="[151,307,534,560]" pageId="14" pageNumber="43" refString="Baker, A. S. (1987) Caleupodes, a new genus of eupodoid mite (Acari: Acariformes) showing primary opisthosomal segmentation. Bulletin of the British Museum (Natural History) (Zoology Series), Miscellanea, 53 (2) 103 - 113. Baker, A. S. (1990) A survey of external morphology of mites of the superfamily Eupodoidea Banks, 1894 (Acari: Acariformes). Journal of Natural History, 24, 1227 - 1261." type="journal article" year="1987">
<collectingCountry id="2899C4C5FFEB5456B6BBA64CFF11FDD7" box="[151,221,534,560]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
(1987)
</bibRefCitation>
distinguishes nine pairs of opisthosomal setae with total lack of setae
<emphasis id="62FA5847FFEB5456B246A64CFBB5FDC8" box="[1130,1145,534,559]" italics="true" pageId="14" pageNumber="43">h</emphasis>
, whereas Kaluz presents only one pair of these setae (instead of two pairs like in other Eupodoidea). They both recognize two pairs of setae
<emphasis id="62FA5847FFEB5456B6F6A63BFF2EFD9D" box="[218,226,609,634]" italics="true" pageId="14" pageNumber="43">f</emphasis>
, i.e.
<emphasis id="62FA5847FFEB5456B732A63BFEEAFD9D" box="[286,294,609,634]" italics="true" pageId="14" pageNumber="43">f</emphasis>
1 and
<emphasis id="62FA5847FFEB5456B746A63BFEBEFD9D" box="[362,370,609,634]" italics="true" pageId="14" pageNumber="43">f</emphasis>
2, and setae
<emphasis id="62FA5847FFEB5456B42CA63BFDD6FD9D" box="[512,538,609,634]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
, according to Kaluz- two pairs of setae
<emphasis id="62FA5847FFEB5456B22BA63BFBEDFD9D" box="[1031,1057,609,634]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
, and -according to Baker- three pairs of these setae.
</paragraph>
<paragraph id="50318455FFEB5456B6E9A6F4FBE1FC84" blockId="14.[151,1437,152,2008]" pageId="14" pageNumber="43">
Here, I follow
<bibRefCitation id="341FF9A4FFEB5456B7ABA6F4FD4CFD2F" author="Strandmann" box="[391,640,686,712]" pageId="14" pageNumber="43" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandmann (1971)</bibRefCitation>
by considering
<taxonomicName id="978EFFD6FFEB5456B57EA6F4FC3DFD20" box="[850,1009,686,711]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEB5456B57EA6F4FC3DFD20" box="[850,1009,686,711]" italics="true" pageId="14" pageNumber="43">Penthalodes</emphasis>
</taxonomicName>
to have four prodorsal, and nine opisthosomal pairs of setae. In my opinion there are two pairs of setae
<emphasis id="62FA5847FFEB5456B5F7A68EFC24FD0A" box="[987,1000,724,749]" italics="true" pageId="14" pageNumber="43">c</emphasis>
,
<emphasis id="62FA5847FFEB5456B5DBA68EFBC8FD0A" box="[1015,1028,724,749]" italics="true" pageId="14" pageNumber="43">c</emphasis>
1 and
<emphasis id="62FA5847FFEB5456B26BA68EFB98FD0A" box="[1095,1108,724,749]" italics="true" pageId="14" pageNumber="43">c</emphasis>
2, and only one pair of setae on segment from D to AN each, particularly only one pair of setae
<emphasis id="62FA5847FFEB5456B584A6A6FC7CFCF2" box="[936,944,764,789]" italics="true" pageId="14" pageNumber="43">f</emphasis>
and
<emphasis id="62FA5847FFEB5456B5C6A6A6FC35FCF2" box="[1002,1017,764,789]" italics="true" pageId="14" pageNumber="43">h</emphasis>
(differently than
<collectingCountry id="2899C4C5FFEB5456B2E9A6A6FAC7FCF1" box="[1221,1291,764,790]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
and Kaluz). Three pairs of setae
<emphasis id="62FA5847FFEB5456B7AAA77BFE6CFCDD" box="[390,416,801,826]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
(
<emphasis id="62FA5847FFEB5456B79DA77BFE07FCDD" box="[433,459,801,826]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
1,
<emphasis id="62FA5847FFEB5456B7CFA77BFE31FCDD" box="[483,509,801,826]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2,
<emphasis id="62FA5847FFEB5456B438A77BFDE2FCDD" box="[532,558,801,826]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
3 according to
<collectingCountry id="2899C4C5FFEB5456B4FBA77BFCEEFCDC" box="[727,802,801,827]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
), I recognize as
<emphasis id="62FA5847FFEB5456B5CDA77BFC37FCDD" box="[993,1019,801,826]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
,
<emphasis id="62FA5847FFEB5456B226A77BFBEBFCDD" box="[1034,1063,801,826]" italics="true" pageId="14" pageNumber="43">ad</emphasis>
, and
<emphasis id="62FA5847FFEB5456B244A77BFB49FCDD" box="[1128,1157,801,826]" italics="true" pageId="14" pageNumber="43">an</emphasis>
, respectively (
<tableCitation id="1D0CB1EEFFEB5456B31EA77BFA46FCDC" box="[1330,1418,801,827]" captionStart="TABLE 1" captionStartId="7.[151,239,487,511]" captionTargetBox="[151,1391,538,1086]" captionTargetPageId="7" captionText="TABLE 1. Homology of opisthosomal setae in representatives of the family Penthalodidae." httpUri="http://table.plazi.org/id/04F1D4DDFFE2545FB6BBA5BDFBA4FE18" pageId="14" pageNumber="43" tableUuid="04F1D4DDFFE2545FB6BBA5BDFBA4FE18">Table 1</tableCitation>
). With only one pair of setae accompanying the anal cleft (
<emphasis id="62FA5847FFEB5456B51AA713FC9FFC85" box="[822,851,841,866]" italics="true" pageId="14" pageNumber="43">an</emphasis>
), not three or two.
</paragraph>
<paragraph id="50318455FFEB5456B6E9A735FD43FB8B" blockId="14.[151,1437,152,2008]" pageId="14" pageNumber="43">
Setae
<emphasis id="62FA5847FFEB5456B720A735FED5FC6F" box="[268,281,879,904]" italics="true" pageId="14" pageNumber="43">c</emphasis>
1 and
<emphasis id="62FA5847FFEB5456B777A735FEA6FC6F" box="[347,362,879,904]" italics="true" pageId="14" pageNumber="43">d</emphasis>
are positioned between furrows “V” and lyrifissures
<emphasis id="62FA5847FFEB5456B5F2A735FC38FC6F" box="[990,1012,879,904]" italics="true" pageId="14" pageNumber="43">ia</emphasis>
externally. Setal pair
<emphasis id="62FA5847FFEB5456B2DBA735FAC8FC6F" box="[1271,1284,879,904]" italics="true" pageId="14" pageNumber="43">e</emphasis>
lies laterally in relation to the furrows. Setae
<emphasis id="62FA5847FFEB5456B422A7CDFDDAFC57" box="[526,534,919,944]" italics="true" pageId="14" pageNumber="43">f</emphasis>
are aligned with lyrifissures
<emphasis id="62FA5847FFEB5456B546A7CDFC4CFC57" box="[874,896,919,944]" italics="true" pageId="14" pageNumber="43">ip</emphasis>
, and are lateral in relation to the “V” furrows. Setae
<emphasis id="62FA5847FFEB5456B6CCA7E6FF23FC32" box="[224,239,956,981]" italics="true" pageId="14" pageNumber="43">h</emphasis>
form a separate row posterior to setae
<emphasis id="62FA5847FFEB5456B4E9A7E6FD01FC32" box="[709,717,956,981]" italics="true" pageId="14" pageNumber="43">f</emphasis>
and situated more laterally than the other setae because the opisthosoma changes into steep wall at this position. One pair each of setae
<emphasis id="62FA5847FFEB5456B235A7B8FBFFFC1C" box="[1049,1075,994,1019]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
,
<emphasis id="62FA5847FFEB5456B26EA7B8FB93FC1C" box="[1090,1119,994,1019]" italics="true" pageId="14" pageNumber="43">ad</emphasis>
and
<emphasis id="62FA5847FFEB5456B2B6A7B8FB7BFC1C" box="[1178,1207,994,1019]" italics="true" pageId="14" pageNumber="43">an</emphasis>
lie at this posterior wall of the opisthosoma one under another. Only setae
<emphasis id="62FA5847FFEB5456B534A05DFCF9FBC7" box="[792,821,1031,1056]" italics="true" pageId="14" pageNumber="43">an</emphasis>
are placed in close vicinity to the anal cleft (see also in larva,
<bibRefCitation id="341FF9A4FFEB5456B724A076FE3EFBA1" author="Jesionowska" box="[264,498,1068,1094]" pageId="14" pageNumber="43" refString="Jesionowska, K. (1996) Morphology of the juvenile stages of eupodoid mites (Eupodoidea), In: Mitchell R., Horn D. J., Needham G. R. &amp; Welbourn W. C. (Eds), Acarology IX- Proceedings. Vol. 1. Ohio Biological Survey, Ohio, USA, pp. 325 - 333." type="book" year="1996">Jesionowska 1996</bibRefCitation>
). This idiosomal chaetotaxy is also characteristic of
<taxonomicName id="978EFFD6FFEB5456B2A6A076FAABFBA2" box="[1162,1383,1068,1093]" class="Arachnida" family="Penthalodidae" genus="Protopenthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEB5456B2A6A076FAABFBA2" box="[1162,1383,1068,1093]" italics="true" pageId="14" pageNumber="43">Protopenthalodes</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFEB5456B6BBA008FE81FB8C" box="[151,333,1106,1131]" class="Arachnida" family="Eupodidae" genus="Hawaiieupodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEB5456B6BBA008FE81FB8C" box="[151,333,1106,1131]" italics="true" pageId="14" pageNumber="43">Hawaiieupodes</emphasis>
</taxonomicName>
(
<bibRefCitation id="341FF9A4FFEB5456B770A008FDFAFB8B" author="Jesionowska" box="[348,566,1106,1132]" pageId="14" pageNumber="43" refString="Jesionowska, K. (1989) New genus and new species of mite of the family Penthalodidae (Actinotrichida, Actinedida, Eupodoidea) from Poland. Acta zoologica cracoviensia, 32 (3), 57 - 67." type="journal article" year="1989">Jesionowska 1989</bibRefCitation>
,
<bibRefCitation id="341FF9A4FFEB5456B46FA008FD4EFB8B" author="Jesionowska" box="[579,642,1106,1132]" pageId="14" pageNumber="43" refString="Jesionowska, K. (2008) Redescription of Hawaiieupodes termophilus Strandtmann et Goff, 1978 (Acari: Prostigmata: Eupodoidea: Penthalodidae) from Hawaii, with a discussion of the systematic status of the taxon. Annales Zoologici (Warszawa), 58 (2), 337 - 346." type="journal article" year="2008">2008</bibRefCitation>
).
</paragraph>
<paragraph id="50318455FFEB5456B6E9A02DFE90F934" blockId="14.[151,1437,152,2008]" pageId="14" pageNumber="43">
On the scanning electron micrograph published by
<collectingCountry id="2899C4C5FFEB5456B509A02DFCA7FB76" box="[805,875,1143,1169]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
(Fig.19d in 1990), the arrangement of setae on the posterior part of idiosoma is visible, illustrating three pairs of setae,
<emphasis id="62FA5847FFEB5456B5D7A0C6FBDAFB52" box="[1019,1046,1180,1205]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
1,
<emphasis id="62FA5847FFEB5456B202A0C6FB84FB52" box="[1070,1096,1180,1205]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2 and
<emphasis id="62FA5847FFEB5456B2A0A0C6FB6BFB52" box="[1164,1191,1180,1205]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
3. Setae
<emphasis id="62FA5847FFEB5456B32BA0C6FAEDFB52" box="[1287,1313,1180,1205]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
1 are more removed from each other than setae
<emphasis id="62FA5847FFEB5456B46FA09FFD91FB39" box="[579,605,1221,1246]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2, while setae
<emphasis id="62FA5847FFEB5456B4D2A09FFCD4FB39" box="[766,792,1221,1246]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2 are more removed than setae
<emphasis id="62FA5847FFEB5456B2AEA09FFB50FB39" box="[1154,1180,1221,1246]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
3. This arrangement is usually different in other Eupodoidea. Moreover, distances between rows of these setae are considerable and seem to be equal, although setae
<emphasis id="62FA5847FFEB5456B430A148FDFAFACC" box="[540,566,1298,1323]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
1 are located at a slightly larger distance from setae
<emphasis id="62FA5847FFEB5456B2BBA148FB7DFACC" box="[1175,1201,1298,1323]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2 compared with the distance between setae
<emphasis id="62FA5847FFEB5456B782A161FE04FAB3" box="[430,456,1339,1364]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2 and
<emphasis id="62FA5847FFEB5456B420A161FDEAFAB3" box="[524,550,1339,1364]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
3. It is clearly visible that three pairs of setae are placed one under another because the opisthosoma forms a steep wall in this place. The very short anal cleft has anal valves and is located on a semispherical protuberance (also visible on the photo in
<collectingCountry id="2899C4C5FFEB5456B597A1D2FBCDFA45" box="[955,1025,1416,1442]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
1990). Earlier,
<bibRefCitation id="341FF9A4FFEB5456B29BA1D2FF29FA20" author="Thor" pageId="14" pageNumber="43" refString="Thor, S. &amp; Willmann, C. (1941) Acarina, Prostigmata - Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. Das Tierreich, 71 A, 1 - 186." type="journal article" year="1941">Thor and Willmann (1941)</bibRefCitation>
observed that the anal opening is located terminally and on a protuberance. This suggests the existence of a tiny peranal segment, PE. Proximally, just in front of it, there is a pair of setae
<emphasis id="62FA5847FFEB5456B24DA189FBB7FA0B" box="[1121,1147,1491,1516]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
3 according to
<collectingCountry id="2899C4C5FFEB5456B30FA188FAA7FA0B" box="[1315,1387,1490,1516]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
, but in my opinion it is a pair of setae
<emphasis id="62FA5847FFEB5456B409A1A1FD8EF9F3" box="[549,578,1531,1556]" italics="true" pageId="14" pageNumber="43">an</emphasis>
. When assuming that segments form rings (which is showed by location of setae
<emphasis id="62FA5847FFEB5456B6D1A27BFED4F9DD" box="[253,280,1569,1594]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
,
<emphasis id="62FA5847FFEB5456B705A27BFE8BF9DD" box="[297,327,1569,1594]" italics="true" pageId="14" pageNumber="43">ad</emphasis>
and
<emphasis id="62FA5847FFEB5456B7AFA27BFE6DF9DD" box="[387,417,1569,1594]" italics="true" pageId="14" pageNumber="43">an</emphasis>
), it is difficult to imagine one large segment PS with three pairs of setae one under another with a simultaneous lack of segment H and setae
<emphasis id="62FA5847FFEB5456B56AA21CFC99F9B8" box="[838,853,1606,1631]" italics="true" pageId="14" pageNumber="43">h</emphasis>
. The location of setae
<emphasis id="62FA5847FFEB5456B24AA21CFB4CF9B8" box="[1126,1152,1606,1631]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
1,
<emphasis id="62FA5847FFEB5456B2B5A21CFB7FF9B8" box="[1177,1203,1606,1631]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
2 and
<emphasis id="62FA5847FFEB5456B2DAA21CFADCF9B8" box="[1270,1296,1606,1631]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
3, i.e.
<emphasis id="62FA5847FFEB5456B378A21CFAA2F9B8" box="[1364,1390,1606,1631]" italics="true" pageId="14" pageNumber="43">ps</emphasis>
,
<emphasis id="62FA5847FFEB5456B352A21CFA57F9B8" box="[1406,1435,1606,1631]" italics="true" pageId="14" pageNumber="43">ad</emphasis>
and
<emphasis id="62FA5847FFEB5456B6E5A234FF2AF960" box="[201,230,1646,1671]" italics="true" pageId="14" pageNumber="43">an</emphasis>
respectively, shows in this case that most probably there are three separate segments, namely PS, AD and AN, each with one pair of setae, while the terminally located anus lies on the last segment, i.e. on PE (
<tableCitation id="1D0CB1EEFFEB5456B6B2A2E3FF34F934" box="[158,248,1721,1747]" captionStart="TABLE 1" captionStartId="7.[151,239,487,511]" captionTargetBox="[151,1391,538,1086]" captionTargetPageId="7" captionText="TABLE 1. Homology of opisthosomal setae in representatives of the family Penthalodidae." httpUri="http://table.plazi.org/id/04F1D4DDFFE2545FB6BBA5BDFBA4FE18" pageId="14" pageNumber="43" tableUuid="04F1D4DDFFE2545FB6BBA5BDFBA4FE18">Table 1</tableCitation>
;
<figureCitation id="C8B598D0FFEB5456B72BA2E3FE81F934" box="[263,333,1721,1747]" captionStart="FIGURE 2" captionStartId="5.[151,255,1020,1044]" captionTargetBox="[161,1430,205,979]" captionTargetId="figure@5.[151,1436,194,999]" captionTargetPageId="5" captionText="FIGURE 2. Penthalodes polonicus n. sp. Female. Idiosoma, lateral view." httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="14" pageNumber="43">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="50318455FFEB5457B6E9A284FE5BFDE5" blockId="14.[151,1437,152,2008]" lastBlockId="15.[151,1437,152,2010]" lastPageId="15" lastPageNumber="44" pageId="14" pageNumber="43">
The structure of idiosomal setae can differ, with prodorsal setae (except setae
<emphasis id="62FA5847FFEB5456B259A284FB41F910" box="[1141,1165,1758,1783]" italics="true" pageId="14" pageNumber="43">ro</emphasis>
and
<emphasis id="62FA5847FFEB5456B2E6A284FB24F910" box="[1226,1256,1758,1783]" italics="true" pageId="14" pageNumber="43">bo</emphasis>
) being usually much larger than opisthosomal setae (especially well visible on Fig.
<quantity id="977629B0FFEB5456B5F8A359FBCEF8FA" box="[980,1026,1795,1821]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="43" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="341FF9A4FFEB5456B227A359FB57F8FA" author="Kaluz" box="[1035,1179,1795,1821]" pageId="14" pageNumber="43" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz 2000</bibRefCitation>
).
<bibRefCitation id="341FF9A4FFEB5456B29CA359FF29F8A4" author="Thor" pageId="14" pageNumber="43" refString="Thor, S. &amp; Willmann, C. (1941) Acarina, Prostigmata - Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. Das Tierreich, 71 A, 1 - 186." type="journal article" year="1941">Thor and Willmann (1941)</bibRefCitation>
observed that setae in
<emphasis id="62FA5847FFEB5456B7D8A373FD98F8A5" box="[500,596,1833,1858]" italics="true" pageId="14" pageNumber="43">P. o v a l i s</emphasis>
are very small, plumose or branched. Setae in specimens described by
<bibRefCitation id="341FF9A4FFEB5456B6BBA314FEFDF88F" author="Kaluz" box="[151,305,1870,1896]" pageId="14" pageNumber="43" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
are branched with few outgrowths.
<bibRefCitation id="341FF9A4FFEB5456B4F8A314FCBDF88F" author="Baker" box="[724,881,1870,1896]" pageId="14" pageNumber="43" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFEB5456B4F8A314FCD6F88F" box="[724,794,1870,1896]" name="United States Pacific Island Wildlife Refuges" pageId="14" pageNumber="43">Baker</collectingCountry>
(1946)
</bibRefCitation>
, when comparing
<taxonomicName id="978EFFD6FFEB5456B26AA314FB60F880" box="[1094,1196,1870,1895]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFEB5456B26AA314FB60F880" box="[1094,1196,1870,1895]" italics="true" pageId="14" pageNumber="43">P. boneti</emphasis>
</taxonomicName>
with
<taxonomicName id="978EFFD6FFEB5456B2C1A314FA57F88F" authority="Koch" authorityName="Koch" box="[1261,1435,1870,1896]" class="Arachnida" family="Penthaleidae" genus="Penthaleus" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="62FA5847FFEB5456B2C1A314FA99F880" box="[1261,1365,1870,1895]" italics="true" pageId="14" pageNumber="43">P. ovatus</emphasis>
Koch
</taxonomicName>
(i.e.
<emphasis id="62FA5847FFEB5456B6E0A329FEE0F86B" box="[204,300,1907,1932]" italics="true" pageId="14" pageNumber="43">P. o v a l i s</emphasis>
), found that the form of setae of the dorsum differs. In
<taxonomicName id="978EFFD6FFEB5456B5EEA329FBE5F86B" box="[962,1065,1907,1932]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFEB5456B5EEA329FBE5F86B" box="[962,1065,1907,1932]" italics="true" pageId="14" pageNumber="43">P. boneti</emphasis>
</taxonomicName>
, branches are short, while in
<taxonomicName id="978EFFD6FFEB5456B3AAA329FF28F855" class="Arachnida" family="Penthaleidae" genus="Penthaleus" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="62FA5847FFEB5456B3AAA329FF28F855" italics="true" pageId="14" pageNumber="43">P. ovatus</emphasis>
</taxonomicName>
they are less numerous and can be as long as the main stem. This feature approximates
<taxonomicName id="978EFFD6FFEB5456B2C5A3C3FA9EF855" box="[1257,1362,1945,1970]" class="Arachnida" family="Penthaleidae" genus="Penthaleus" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="62FA5847FFEB5456B2C5A3C3FA9EF855" box="[1257,1362,1945,1970]" italics="true" pageId="14" pageNumber="43">P. ovatus</emphasis>
</taxonomicName>
to the newly described species, i.e.
<taxonomicName id="978EFFD6FFEB5456B7D8A3E4FD4AF830" box="[500,646,1982,2007]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="14" pageNumber="43" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFEB5456B7D8A3E4FD4AF830" box="[500,646,1982,2007]" italics="true" pageId="14" pageNumber="43">P. polonicus</emphasis>
</taxonomicName>
, where setae are tridactyl, although prodorsal setae
<emphasis id="62FA5847FFEB5456B2DBA3E4FADFF830" box="[1271,1299,1982,2007]" italics="true" pageId="14" pageNumber="43">xa</emphasis>
and
<emphasis id="62FA5847FFEB5456B362A3E4FAAFF830" box="[1358,1379,1982,2007]" italics="true" pageId="14" pageNumber="43">le</emphasis>
also have short outgrowths at the base of the stem. On the other hand, in
<emphasis id="62FA5847FFEA5457B580A4C2FBDAFF56" box="[940,1046,152,177]" italics="true" pageId="15" pageNumber="44">P. t u r n e r i</emphasis>
<bibRefCitation id="341FF9A4FFEA5457B20CA4C2FB77FF55" author="Baker" box="[1056,1211,152,178]" pageId="15" pageNumber="44" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFEA5457B20CA4C2FBAAFF55" box="[1056,1126,152,178]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
(1946)
</bibRefCitation>
distinguishes setae on the dorsum as “semi-plumose”, with three to five lateral branches, while the anterior setae are “six-rayed”. This resembles the shape of setae on the dorsum of
<taxonomicName id="978EFFD6FFEA5457B528A4B8FCA4FF1C" box="[772,872,226,251]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B528A4B8FCA4FF1C" box="[772,872,226,251]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
</taxonomicName>
described by
<bibRefCitation id="341FF9A4FFEA5457B23DA4B8FB7FFF1B" author="Kaluz" box="[1041,1203,226,252]" pageId="15" pageNumber="44" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
. In
<taxonomicName id="978EFFD6FFEA5457B2F3A4B8FABEFF1C" box="[1247,1394,226,251]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEA5457B2F3A4B8FABEFF1C" box="[1247,1394,226,251]" italics="true" pageId="15" pageNumber="44">Penthalodes</emphasis>
</taxonomicName>
sp. presented by
<bibRefCitation id="341FF9A4FFEA5457B718A552FE18FEC5" author="Baker" box="[308,468,264,290]" pageId="15" pageNumber="44" refString="Baker, A. S. (1987) Caleupodes, a new genus of eupodoid mite (Acari: Acariformes) showing primary opisthosomal segmentation. Bulletin of the British Museum (Natural History) (Zoology Series), Miscellanea, 53 (2) 103 - 113. Baker, A. S. (1990) A survey of external morphology of mites of the superfamily Eupodoidea Banks, 1894 (Acari: Acariformes). Journal of Natural History, 24, 1227 - 1261." type="journal article" year="1987">
<collectingCountry id="2899C4C5FFEA5457B718A552FEB6FEC5" box="[308,378,264,290]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
(1987)
</bibRefCitation>
, the setae are short and not loosely feathered, with a long apical spine, while in
<taxonomicName id="978EFFD6FFEA5457B3AAA552FF2FFEA1" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B3AAA552FF2FFEA1" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
</taxonomicName>
they are &quot;densely spinose terminating in long filament&quot; (
<collectingCountry id="2899C4C5FFEA5457B590A577FBC5FEA0" box="[956,1033,301,327]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
1990). In
<taxonomicName id="978EFFD6FFEA5457B2A3A577FA8BFEA1" box="[1167,1351,301,326]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFEA5457B2A3A577FA8BFEA1" box="[1167,1351,301,326]" italics="true" pageId="15" pageNumber="44">P. oregonensis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFEA5457B3A9A577FECAFE8C" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFEA5457B3A9A577FECAFE8C" italics="true" pageId="15" pageNumber="44">P. carinatus</emphasis>
</taxonomicName>
, the setae are pilose rather than weakly plumose. In
<taxonomicName id="978EFFD6FFEA5457B541A508FBDDFE8C" box="[877,1041,338,363]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFEA5457B541A508FBDDFE8C" box="[877,1041,338,363]" italics="true" pageId="15" pageNumber="44">P. alaskaensis</emphasis>
</taxonomicName>
, they are plumose, while those in
<taxonomicName id="978EFFD6FFEA5457B6BBA522FE8EFE76" box="[151,322,376,401]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFEA5457B6BBA522FE8EFE76" box="[151,322,376,401]" italics="true" pageId="15" pageNumber="44">P. hawaiiensis</emphasis>
</taxonomicName>
are loosely plumose, with a longer apical outgrowth and sometimes also with some slightly longer lateral outgrowth. Usually, the podosomal setae and those of the genital region and a pair of setae
<emphasis id="62FA5847FFEA5457B352A5C7FA57FE51" box="[1406,1435,413,438]" italics="true" pageId="15" pageNumber="44">an</emphasis>
differ from other idiosomal setae, being plumose-pilose, which is particularly well shown in
<emphasis id="62FA5847FFEA5457B2F7A598FAAAFE3C" box="[1243,1382,450,475]" italics="true" pageId="15" pageNumber="44">P. p o l o n i c u s</emphasis>
and
<emphasis id="62FA5847FFEA5457B6BBA5B2FF3AFDE6" box="[151,246,488,513]" italics="true" pageId="15" pageNumber="44">P. o v a l i s</emphasis>
sensu Kaluz.
</paragraph>
<paragraph id="50318455FFEA5457B6E9A657FE6CFD58" blockId="15.[151,1437,152,2010]" pageId="15" pageNumber="44">
<emphasis id="62FA5847FFEA5457B6E9A657FE9FFDC0" bold="true" box="[197,339,525,551]" italics="true" pageId="15" pageNumber="44">Lyrifissures</emphasis>
. In all species of
<taxonomicName id="978EFFD6FFEA5457B432A657FD63FDC1" box="[542,687,525,550]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEA5457B432A657FD63FDC1" box="[542,687,525,550]" italics="true" pageId="15" pageNumber="44">Penthalodes</emphasis>
</taxonomicName>
, lyrifissures are distinct and always present as four pairs (
<emphasis id="62FA5847FFEA5457B37EA657FAA4FDC1" box="[1362,1384,525,550]" italics="true" pageId="15" pageNumber="44">ia</emphasis>
,
<emphasis id="62FA5847FFEA5457B35BA657FA58FDC1" box="[1399,1428,525,550]" italics="true" pageId="15" pageNumber="44">im</emphasis>
,
<emphasis id="62FA5847FFEA5457B6BBA668FF62FDAC" box="[151,174,562,587]" italics="true" pageId="15" pageNumber="44">ip</emphasis>
,
<emphasis id="62FA5847FFEA5457B692A668FF19FDAC" box="[190,213,562,587]" italics="true" pageId="15" pageNumber="44">ih</emphasis>
?). Lyrifissures
<emphasis id="62FA5847FFEA5457B7BFA668FE66FDAC" box="[403,426,562,587]" italics="true" pageId="15" pageNumber="44">ia</emphasis>
,
<emphasis id="62FA5847FFEA5457B796A668FE14FDAC" box="[442,472,562,587]" italics="true" pageId="15" pageNumber="44">im</emphasis>
and
<emphasis id="62FA5847FFEA5457B438A668FDE0FDAC" box="[532,556,562,587]" italics="true" pageId="15" pageNumber="44">ip</emphasis>
are lateral to furrows V, with
<emphasis id="62FA5847FFEA5457B5B5A668FC7CFDAC" box="[921,944,562,587]" italics="true" pageId="15" pageNumber="44">ia</emphasis>
situated slightly posterior to setae
<emphasis id="62FA5847FFEA5457B375A668FAAAFDAC" box="[1369,1382,562,587]" italics="true" pageId="15" pageNumber="44">c</emphasis>
1,
<emphasis id="62FA5847FFEA5457B352A668FA50FDAC" box="[1406,1436,562,587]" italics="true" pageId="15" pageNumber="44">im</emphasis>
slightly anterior to setae
<emphasis id="62FA5847FFEA5457B794A601FE09FD93" box="[440,453,603,628]" italics="true" pageId="15" pageNumber="44">e</emphasis>
, and
<emphasis id="62FA5847FFEA5457B428A601FDD6FD93" box="[516,538,603,628]" italics="true" pageId="15" pageNumber="44">ip</emphasis>
at the same level as setae
<emphasis id="62FA5847FFEA5457B562A601FC9AFD93" box="[846,854,603,628]" italics="true" pageId="15" pageNumber="44">f</emphasis>
. The last pair,
<emphasis id="62FA5847FFEA5457B228A601FBD7FD93" box="[1028,1051,603,628]" italics="true" pageId="15" pageNumber="44">ih</emphasis>
, is positioned on the ventral side of opisthosoma anterior to setae
<emphasis id="62FA5847FFEA5457B404A6DAFD8AFD7E" box="[552,582,640,665]" italics="true" pageId="15" pageNumber="44">an</emphasis>
. The lyrifissures are slit-shaped with heavily sclerotized margins and arranged transversally.
</paragraph>
<paragraph id="50318455FFEA5457B6E9A690FF25FB98" blockId="15.[151,1437,152,2010]" pageId="15" pageNumber="44">
<emphasis id="62FA5847FFEA5457B6E9A690FEB8FD03" bold="true" box="[197,372,714,740]" italics="true" pageId="15" pageNumber="44">Genital region</emphasis>
(
<tableCitation id="1D0CB1EEFFEA5457B7A8A690FE2CFD03" box="[388,480,714,740]" captionStart="TABLE 2" captionStartId="12.[151,239,601,625]" captionText="TABLE 2. Number of aggenital and genital setae in representatives of Penthalodes." pageId="15" pageNumber="44">Table 2</tableCitation>
). The number of setae of the genital region in
<taxonomicName id="978EFFD6FFEA5457B224A691FB55FD03" box="[1032,1177,715,740]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEA5457B224A691FB55FD03" box="[1032,1177,715,740]" italics="true" pageId="15" pageNumber="44">Penthalodes</emphasis>
</taxonomicName>
is variable. However, the arrangement of aggenital setae on the region between the coxal regions IV and the anterior margin of the genital valves (i.e. the pregenital region) is characteristic for
<emphasis id="62FA5847FFEA5457B571A74CFC39FCC8" box="[861,1013,790,815]" italics="true" pageId="15" pageNumber="44">
<taxonomicName id="978EFFD6FFEA5457B571A74CFC3DFCC8" box="[861,1009,790,815]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="genus">Penthalodes</taxonomicName>
,
</emphasis>
being composed of two alignments. The other setae are arranged in a line around the genital valves. The number of setae in each alignment varies. For example, in
<taxonomicName id="978EFFD6FFEA5457B774A73AFD9DFC9D" authority="Kaluz" authorityName="Kaluz" box="[344,593,864,890]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B774A73AFE77FC9E" box="[344,443,864,889]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
(
<figureCitation id="C8B598D0FFEA5457B44DA73AFD6BFC9D" box="[609,679,864,890]" captionStart="FIGURE 2" captionStartId="5.[151,255,1020,1044]" captionTargetBox="[161,1430,205,979]" captionTargetId="figure@5.[151,1436,194,999]" captionTargetPageId="5" captionText="FIGURE 2. Penthalodes polonicus n. sp. Female. Idiosoma, lateral view." httpUri="https://zenodo.org/record/199163/files/figure.png" pageId="15" pageNumber="44">
Fig.
<quantity id="977629B0FFEA5457B4B4A73AFD6BFC9D" box="[664,679,864,890]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="15" pageNumber="44" unit="in" value="2.0">2</quantity>
</figureCitation>
in 2000), the two alignments of aggenital setae in the pregenital region comprises five setae in the outer row, and eight setae in the inner row; the remaining five pairs of setae are arranged around the genital valves (23 setae in total). In
<emphasis id="62FA5847FFEA5457B578A7F1FC26FC23" box="[852,1002,939,964]" italics="true" pageId="15" pageNumber="44">
<taxonomicName id="978EFFD6FFEA5457B578A7F1FC2AFC23" box="[852,998,939,964]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="polonicus">P. polonicus</taxonomicName>
,
</emphasis>
six setae form the outer row and two setae form the inner row; the remaining number of setae is four pairs (
<quantity id="977629B0FFEA5457B5E9A78AFBCCFC0D" box="[965,1024,976,1002]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="15" pageNumber="44" unit="in" value="16.0">16 in</quantity>
total; Fig. 1B). Other examples are presented in table 2. For
<taxonomicName id="978EFFD6FFEA5457B7F0A7ACFD86FBE8" box="[476,586,1014,1039]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B7F0A7ACFD86FBE8" box="[476,586,1014,1039]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
</taxonomicName>
specimens presented by
<bibRefCitation id="341FF9A4FFEA5457B5A3A7AFFBF1FBE8" author="Baker" box="[911,1085,1013,1039]" pageId="15" pageNumber="44" refString="Baker, A. S. (1987) Caleupodes, a new genus of eupodoid mite (Acari: Acariformes) showing primary opisthosomal segmentation. Bulletin of the British Museum (Natural History) (Zoology Series), Miscellanea, 53 (2) 103 - 113. Baker, A. S. (1990) A survey of external morphology of mites of the superfamily Eupodoidea Banks, 1894 (Acari: Acariformes). Journal of Natural History, 24, 1227 - 1261." type="journal article" year="1987">
<collectingCountry id="2899C4C5FFEA5457B5A3A7AFFC17FBE8" box="[911,987,1013,1039]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
(1987)
</bibRefCitation>
, as well as specimens of
<taxonomicName id="978EFFD6FFEA5457B3A9A7ACFEEEFBD3" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFEA5457B3A9A7ACFEEEFBD3" italics="true" pageId="15" pageNumber="44">P. hawaiiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFEA5457B775A041FDCDFBD3" box="[345,513,1051,1076]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFEA5457B775A041FDCDFBD3" box="[345,513,1051,1076]" italics="true" pageId="15" pageNumber="44">P. oregonensis</emphasis>
</taxonomicName>
, there is no information on the aggenital region. The number of aggenital setae in
<taxonomicName id="978EFFD6FFEA5457B69AA01AFE86FBBE" box="[182,330,1088,1113]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEA5457B69AA01AFE86FBBE" box="[182,330,1088,1113]" italics="true" pageId="15" pageNumber="44">Penthalodes</emphasis>
</taxonomicName>
representatives ranges from 12 to 23 setae and may fluctuate, in particular in the pregenital region.
</paragraph>
<paragraph id="50318455FFEA5457B6E9A0D0FEF4FAF3" blockId="15.[151,1437,152,2010]" pageId="15" pageNumber="44">
The genital setae are inserted on strongly sclerotized genital valves, arranged linearly, close to the inner margin, usually with one pair situated more laterally. The number of genital setae varies from 6 to 10 pairs between species (
<tableCitation id="1D0CB1EEFFEA5457B744A08FFE09FB08" box="[360,453,1237,1263]" captionStart="TABLE 2" captionStartId="12.[151,239,601,625]" captionText="TABLE 2. Number of aggenital and genital setae in representatives of Penthalodes." pageId="15" pageNumber="44">Table 2</tableCitation>
). In
<emphasis id="62FA5847FFEA5457B7D0A08CFD92FB08" box="[508,606,1238,1263]" italics="true" pageId="15" pageNumber="44">P. o v a l i s</emphasis>
sensu
<collectingCountry id="2899C4C5FFEA5457B498A08FFD37FB08" box="[692,763,1237,1263]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
(1990) the number of genital setae can be inferred from drawing only.
</paragraph>
<paragraph id="50318455FFEA5457B6E9A17AFF15FA63" blockId="15.[151,1437,152,2010]" pageId="15" pageNumber="44">
The number of eugenital setae has earlier been mentioned only by
<bibRefCitation id="341FF9A4FFEA5457B5F5A17AFB0AFADD" author="Strandtmann" box="[985,1222,1312,1338]" pageId="15" pageNumber="44" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandtmann (1978)</bibRefCitation>
and
<bibRefCitation id="341FF9A4FFEA5457B32CA17AFA50FADD" author="Kaluz" box="[1280,1436,1312,1338]" pageId="15" pageNumber="44" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
as six pairs in
<taxonomicName id="978EFFD6FFEA5457B710A11CFDF9FAB8" authority="Kaluz" authorityName="Kaluz" box="[316,565,1349,1375]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B710A11CFE52FAB8" box="[316,414,1350,1375]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
, contrary to
<taxonomicName id="978EFFD6FFEA5457B4EAA11CFCA1FAB8" box="[710,877,1350,1375]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFEA5457B4EAA11CFCA1FAB8" box="[710,877,1350,1375]" italics="true" pageId="15" pageNumber="44">P. hawaiiensis</emphasis>
</taxonomicName>
and
<emphasis id="62FA5847FFEA5457B588A11CFBE3FAB8" box="[932,1071,1350,1375]" italics="true" pageId="15" pageNumber="44">P. p o l o n i c u s</emphasis>
, where there are seven pairs of setae.
</paragraph>
<paragraph id="50318455FFEA5457B6E9A1CAFBF0F91D" blockId="15.[151,1437,152,2010]" pageId="15" pageNumber="44">
<emphasis id="62FA5847FFEA5457B6E9A1CAFD82FA4E" box="[197,590,1424,1450]" italics="true" pageId="15" pageNumber="44">
<emphasis id="62FA5847FFEA5457B6E9A1CAFD8AFA4D" bold="true" box="[197,582,1424,1450]" italics="true" pageId="15" pageNumber="44">Ornamentation of the idiosoma</emphasis>
.
</emphasis>
The integument of
<taxonomicName id="978EFFD6FFEA5457B517A1CAFC02FA4E" box="[827,974,1424,1449]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFEA5457B517A1CAFC02FA4E" box="[827,974,1424,1449]" italics="true" pageId="15" pageNumber="44">Penthalodes</emphasis>
</taxonomicName>
species is moderately sclerotized with characteristic reticulate ornamentation formed by tiny, tubercular costulae creating pentagons and hexagons. The ornamentation is best discerned dorsally where costulae are large and distinct. On the other hand, the pattern is much more delicate, smaller on lateral and ventral parts, and can even disappear in the sternal region where it may be delicately punctate (particularly in the location of sejugal furrow). Around the genital valves, i.e. in the aggenital region, the delicate reticulate pattern sometimes disappears and tiny costulae form an irregular punctation, as in
<taxonomicName id="978EFFD6FFEA5457B7FCA22AFD93F96E" box="[464,607,1648,1673]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFEA5457B7FCA22AFD93F96E" box="[464,607,1648,1673]" italics="true" pageId="15" pageNumber="44">P. polonicus</emphasis>
</taxonomicName>
, whereas in other representatives, e.g. in
<taxonomicName id="978EFFD6FFEA5457B261A22AFB3FF96E" box="[1101,1267,1648,1673]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFEA5457B261A22AFB3FF96E" box="[1101,1267,1648,1673]" italics="true" pageId="15" pageNumber="44">P. alaskaensis</emphasis>
</taxonomicName>
, the reticulate ornamentation can be preserved. The coxal regions are strongly sclerotized, with a punctate or reticulate pattern, or a combination of punctate and reticulate ornamentation. In
<taxonomicName id="978EFFD6FFEA5457B5F2A2E1FBA3F933" box="[990,1135,1723,1748]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFEA5457B5F2A2E1FBA3F933" box="[990,1135,1723,1748]" italics="true" pageId="15" pageNumber="44">P. polonicus</emphasis>
</taxonomicName>
, there are sigillae visible between the coxal regions II and III, indicating the region of the sejugal furrow.
</paragraph>
<paragraph id="50318455FFEA5448B6E9A35FFA50FF30" blockId="15.[151,1437,152,2010]" lastBlockId="16.[151,1437,152,551]" lastPageId="16" lastPageNumber="45" pageId="15" pageNumber="44">
The lateral margins of the pentagons are formed by tubercular costulae which can be tear-shaped (more or less elongated), like in
<taxonomicName id="978EFFD6FFEA5457B786A371FDF5F8A3" box="[426,569,1835,1860]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFEA5457B786A371FDF5F8A3" box="[426,569,1835,1860]" italics="true" pageId="15" pageNumber="44">P. polonicus</emphasis>
</taxonomicName>
(Fig. 1), or have a broad basis terminating in a hook-like process, like in some
<taxonomicName id="978EFFD6FFEA5457B6F1A30AFE26F88D" authority="Baker 1990" authorityName="Baker" authorityYear="1990" box="[221,490,1872,1898]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B6F1A30AFE8DF88E" box="[221,321,1872,1897]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
(
<collectingCountry id="2899C4C5FFEA5457B77EA30AFE56F88D" box="[338,410,1872,1898]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
1990)
</taxonomicName>
, or resemble a pyramid (
<bibRefCitation id="341FF9A4FFEA5457B53DA30AFC6EF88D" author="Kaluz" box="[785,930,1872,1898]" pageId="15" pageNumber="44" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz 2000</bibRefCitation>
). The surface area of the pentagons can be covered by tiny costulae in the form of dots forming parallel lines, like in
<emphasis id="62FA5847FFEA5457B230A32CFB67F868" box="[1052,1195,1910,1935]" italics="true" pageId="15" pageNumber="44">P. p o l o n i c u s</emphasis>
and
<taxonomicName id="978EFFD6FFEA5457B2C7A32CFEF9F853" authority="Baker (1990)" authorityName="Baker" authorityYear="1990" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B2C7A32CFA9DF868" box="[1259,1361,1910,1935]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
sensu
<collectingCountry id="2899C4C5FFEA5457B6BBA3C0FF11F853" box="[151,221,1946,1972]" name="United States Pacific Island Wildlife Refuges" pageId="15" pageNumber="44">Baker</collectingCountry>
(1990)
</taxonomicName>
, or can be visibly lined with single tiny costulae, like in
<emphasis id="62FA5847FFEA5457B5E5A3C1FB9DF853" box="[969,1105,1947,1972]" italics="true" pageId="15" pageNumber="44">P. c a r i n a t u s</emphasis>
, or smooth, like in
<taxonomicName id="978EFFD6FFEA5457B315A3C1FEF9F83D" authority="Kaluz." authorityName="Kaluz." class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFEA5457B315A3C1FA50F853" box="[1337,1436,1947,1972]" italics="true" pageId="15" pageNumber="44">P. ovalis</emphasis>
sensu Kaluz.
</taxonomicName>
<taxonomicName id="978EFFD6FFEA5457B711A39AFDBCF83E" box="[317,624,1984,2009]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="15" pageNumber="44" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFEA5457B711A39AFDBCF83E" box="[317,624,1984,2009]" italics="true" pageId="15" pageNumber="44">Penthalodes hawaiiensis</emphasis>
</taxonomicName>
has an ornamentation composed mainly of hexagons, with single spines arranged rather irregularly within their areas. In some representatives of
<taxonomicName id="978EFFD6FFF55448B2A7A4C2FAE9FF56" box="[1163,1317,152,177]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF55448B2A7A4C2FAE9FF56" box="[1163,1317,152,177]" italics="true" pageId="16" pageNumber="45">Penthalodes</emphasis>
</taxonomicName>
there are sometimes single, additional, large tubercular costulae on the pentagon surface. This may occur in all species.
</paragraph>
<paragraph id="50318455FFF55448B6E9A4B8FD1DFDC0" blockId="16.[151,1437,152,551]" pageId="16" pageNumber="45">
In
<taxonomicName id="978EFFD6FFF55448B6C9A4B8FE28FF1B" authority="Kaluz" authorityName="Kaluz" box="[229,484,226,252]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF55448B6C9A4B8FE85FF1C" box="[229,329,226,251]" italics="true" pageId="16" pageNumber="45">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
, the polygonal reticulate ornament occurs on the whole surface of the idiosoma, except the sternal region, and in some specimens, this polygonal pattern on the dorsum can be disturbed. In
<taxonomicName id="978EFFD6FFF55448B3AAA552FF2DFEA1" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFF55448B3AAA552FF2DFEA1" italics="true" pageId="16" pageNumber="45">P. boneti</emphasis>
</taxonomicName>
the pattern is similar: a reticulate pattern between the coxal regions, except the sternal region where it becomes tuberculate (punctate). According to
<bibRefCitation id="341FF9A4FFF55448B49FA508FC9DFE8B" author="Baker" box="[691,849,338,364]" pageId="16" pageNumber="45" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFF55448B49FA508FD35FE8B" box="[691,761,338,364]" name="United States Pacific Island Wildlife Refuges" pageId="16" pageNumber="45">Baker</collectingCountry>
(1946)
</bibRefCitation>
, it is similar to the ornamentation of
<taxonomicName id="978EFFD6FFF55448B2D2A508FAABFE8C" box="[1278,1383,338,363]" class="Arachnida" family="Penthaleidae" genus="Penthaleus" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="62FA5847FFF55448B2D2A508FAABFE8C" box="[1278,1383,338,363]" italics="true" pageId="16" pageNumber="45">P. ovatus</emphasis>
</taxonomicName>
(i.e.
<taxonomicName id="978EFFD6FFF55448B6BBA522FECFFE76" box="[151,259,376,401]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF55448B6BBA522FECFFE76" box="[151,259,376,401]" italics="true" pageId="16" pageNumber="45">P. ovalis</emphasis>
</taxonomicName>
). In
<taxonomicName id="978EFFD6FFF55448B76EA522FE15FE76" box="[322,473,376,401]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF55448B76EA522FE15FE76" box="[322,473,376,401]" italics="true" pageId="16" pageNumber="45">P. carinatus</emphasis>
</taxonomicName>
, the reticulate pattern of the whole idiosoma is composed of pentagons and hexagons. In
<emphasis id="62FA5847FFF55448B719A5C7FE6DFE51" box="[309,417,413,438]" italics="true" pageId="16" pageNumber="45">P. t u r n e r i</emphasis>
and
<taxonomicName id="978EFFD6FFF55448B7F3A5C7FD45FE51" box="[479,649,413,438]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFF55448B7F3A5C7FD45FE51" box="[479,649,413,438]" italics="true" pageId="16" pageNumber="45">P. oregonensis</emphasis>
</taxonomicName>
, it is composed of hexagons on the whole idiosoma (
<bibRefCitation id="341FF9A4FFF55448B32CA5C7FA5CFE50" author="Baker" box="[1280,1424,413,439]" pageId="16" pageNumber="45" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFF55448B32CA5C7FA84FE50" box="[1280,1352,413,439]" name="United States Pacific Island Wildlife Refuges" pageId="16" pageNumber="45">Baker</collectingCountry>
1946
</bibRefCitation>
). In
<emphasis id="62FA5847FFF55448B696A598FEABFE3C" box="[186,359,450,475]" italics="true" pageId="16" pageNumber="45">P. a l a s k a e n s i s</emphasis>
, the reticulate pattern occurs on the whole idiosoma but clear, regular pentagons and hexagons are most distinct on the dorsum and the ornament on the sternal region is reticulato-punctate (
<bibRefCitation id="341FF9A4FFF55448B6B2A657FEB0FDC0" author="Strandtmann" box="[158,380,525,551]" pageId="16" pageNumber="45" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann 1971</bibRefCitation>
). It is similar in
<taxonomicName id="978EFFD6FFF55448B417A657FD06FDC1" box="[571,714,525,550]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF55448B417A657FD06FDC1" box="[571,714,525,550]" italics="true" pageId="16" pageNumber="45">P. polonicus</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="04F1D4DDFFF55448B6BBA60DFDB9FD78" ID-Table-UUID="04F1D4DDFFF55448B6BBA60DFDB9FD78" httpUri="http://table.plazi.org/id/04F1D4DDFFF55448B6BBA60DFDB9FD78" pageId="16" pageNumber="45" targetBox="[151,1418,690,941]" targetIsTable="true" targetPageId="16">
<paragraph id="50318455FFF55448B6BBA60DFDB9FD78" blockId="16.[151,1149,599,671]" pageId="16" pageNumber="45">
<emphasis id="62FA5847FFF55448B6BBA60DFEC4FD88" bold="true" box="[151,264,599,623]" pageId="16" pageNumber="45">TABLE 4.</emphasis>
Solenidia of the rhagidial organ on tarsi I and II in representatives of
<taxonomicName id="978EFFD6FFF55448B5DEA602FBBAFD88" box="[1010,1142,600,623]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF55448B5DEA602FBBAFD88" box="[1010,1142,600,623]" italics="true" pageId="16" pageNumber="45">Penthalodes</emphasis>
</taxonomicName>
.
<taxonomicName id="978EFFD6FFF55448B6B3A6D0FE7BFD78" authority="Solenidia" authorityName="Solenidia" box="[159,439,650,671]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="genus">Penthalodes Solenidia</taxonomicName>
of rhagidial organ I
</paragraph>
</caption>
<paragraph id="50318455FFF55448B770A6E8FB97FC4B" pageId="16" pageNumber="45">
<table id="228E76F5FFF5ABA7B6BBA6E8FA46FC4A" box="[151,1418,690,941]" gridcols="4" gridrows="6" pageId="16" pageNumber="45">
<tr id="EEBE8617FFF5ABA7B6BBA6E8FA46FD20" box="[151,1418,690,711]" gridrow="0" pageId="16" pageNumber="45" rowspan-0="1">
<th id="AD6FEF6BFFF5ABA7B770A6E8FE66FD20" box="[348,426,690,711]" gridcol="1" gridrow="0" pageId="16" pageNumber="45">Number</th>
<th id="AD6FEF6BFFF5ABA7B7CEA6E8FC8EFD20" box="[482,834,690,711]" gridcol="2" gridrow="0" pageId="16" pageNumber="45">Shape</th>
<th id="AD6FEF6BFFF5ABA7B5BCA6E8FA46FD20" box="[912,1418,690,711]" gridcol="3" gridrow="0" pageId="16" pageNumber="45">Arrangement</th>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA680FA46FD08" box="[151,1418,730,751]" gridrow="1" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA680FEDEFD08" box="[151,274,730,751]" gridcol="0" gridrow="1" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A680FEC2FD08" box="[159,270,730,751]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF55448B6B3A680FEC2FD08" box="[159,270,730,751]" italics="true" pageId="16" pageNumber="45">alaskaensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A680FE66FD08" box="[348,426,730,751]" gridcol="1" gridrow="1" pageId="16" pageNumber="45">2</td>
<td id="AD6FEF6BFFF5ABA7B7CEA680FC8EFD08" box="[482,834,730,751]" gridcol="2" gridrow="1" pageId="16" pageNumber="45">baculiform; equal in size</td>
<td id="AD6FEF6BFFF5ABA7B5BCA680FA46FD08" box="[912,1418,730,751]" gridcol="3" gridrow="1" pageId="16" pageNumber="45">recumbent in tandem; in confluent depressions</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA758FA46FCF0" box="[151,1418,770,791]" gridrow="2" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA758FEDEFCF0" box="[151,274,770,791]" gridcol="0" gridrow="2" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A758FF17FCF0" box="[159,219,770,791]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFF55448B6B3A758FF17FCF0" box="[159,219,770,791]" italics="true" pageId="16" pageNumber="45">boneti</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A758FE66FCF0" box="[348,426,770,791]" gridcol="1" gridrow="2" pageId="16" pageNumber="45">2+1</td>
<td id="AD6FEF6BFFF5ABA7B7CEA758FC8EFCF0" box="[482,834,770,791]" gridcol="2" gridrow="2" pageId="16" pageNumber="45">rod-like</td>
<td id="AD6FEF6BFFF5ABA7B5BCA758FA46FCF0" box="[912,1418,770,791]" gridcol="3" gridrow="2" pageId="16" pageNumber="45">in line; third laterally</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA770FA46FCD8" box="[151,1418,810,831]" gridrow="3" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA770FEDEFCD8" box="[151,274,810,831]" gridcol="0" gridrow="3" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A770FF35FCD8" box="[159,249,810,831]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF55448B6B3A770FF35FCD8" box="[159,249,810,831]" italics="true" pageId="16" pageNumber="45">carinatus</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A770FE66FCD8" box="[348,426,810,831]" gridcol="1" gridrow="3" pageId="16" pageNumber="45">2</td>
<td id="AD6FEF6BFFF5ABA7B7CEA770FC8EFCD8" box="[482,834,810,831]" gridcol="2" gridrow="3" pageId="16" pageNumber="45">slightly T-shaped</td>
<td id="AD6FEF6BFFF5ABA7B5BCA770FA46FCD8" box="[912,1418,810,831]" gridcol="3" gridrow="3" pageId="16" pageNumber="45">in tandem, in separate depressions</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA708FA46FC63" box="[151,1418,850,900]" gridrow="4" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA708FEDEFC63" box="[151,274,850,900]" gridcol="0" gridrow="4" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A708FEDDFC80" box="[159,273,850,871]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF55448B6B3A708FEDDFC80" box="[159,273,850,871]" italics="true" pageId="16" pageNumber="45">hawaiiensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A708FE66FC63" box="[348,426,850,900]" gridcol="1" gridrow="4" pageId="16" pageNumber="45">2+1</td>
<td id="AD6FEF6BFFF5ABA7B7CEA708FC8EFC63" box="[482,834,850,900]" gridcol="2" gridrow="4" pageId="16" pageNumber="45">baculiform; third small and spatulate</td>
<td id="AD6FEF6BFFF5ABA7B5BCA708FA46FC63" box="[912,1418,850,900]" gridcol="3" gridrow="4" pageId="16" pageNumber="45">recumbent in tandem; in confluent depressions; third laterally</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA7CDFA46FC4A" box="[151,1418,919,941]" gridrow="5" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA7CDFEDEFC4A" box="[151,274,919,941]" gridcol="0" gridrow="5" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A7C2FEDEFC4A" box="[159,274,920,941]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFF55448B6B3A7C2FEDEFC4A" box="[159,274,920,941]" italics="true" pageId="16" pageNumber="45">oregonensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A7CDFE66FC4A" box="[348,426,919,941]" gridcol="1" gridrow="5" pageId="16" pageNumber="45">2+1</td>
<td id="AD6FEF6BFFF5ABA7B7CEA7CDFC8EFC4A" box="[482,834,919,941]" gridcol="2" gridrow="5" pageId="16" pageNumber="45">rod-like</td>
<td id="AD6FEF6BFFF5ABA7B5BCA7CDFA46FC4A" box="[912,1418,919,941]" gridcol="3" gridrow="5" pageId="16" pageNumber="45">in line; third laterally</td>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFF55448B6B3A79AFDCAFC15" blockId="16.[159,1342,959,1010]" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A79AFEDFFC32" box="[159,275,960,981]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF55448B6B3A79AFEDFFC32" box="[159,275,960,981]" italics="true" pageId="16" pageNumber="45">ovalis sensu</emphasis>
</taxonomicName>
2+1 lanceolate sharply tipped; third small rod- in line in separate depressions; third laterally
<bibRefCitation id="341FF9A4FFF55448B6B3A787FED3FC15" author="Kaluz" box="[159,287,989,1010]" pageId="16" pageNumber="45" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
like
</paragraph>
<paragraph id="50318455FFF55448B6B3A05FFEC4FB25" pageId="16" pageNumber="45">
<table id="228E76F5FFF5ABA7B6BBA05FFA5BFB25" box="[151,1431,1029,1218]" gridcols="4" gridrows="4" pageId="16" pageNumber="45">
<tr id="EEBE8617FFF5ABA7B6BBA05FFA5BFBD0" box="[151,1431,1029,1079]" gridrow="0" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA05FFED2FBD0" box="[151,286,1029,1079]" gridcol="0" gridrow="0" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A05FFED2FBD0" authority="sensu Baker (1990)" authorityName="Baker" authorityYear="1990" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF55448B6B3A05FFEDFFBFD" box="[159,275,1029,1050]" italics="true" pageId="16" pageNumber="45">ovalis sensu</emphasis>
Baker (1990)
</taxonomicName>
</th>
<th id="AD6FEF6BFFF5ABA7B770A05FFE4DFBD0" box="[348,385,1029,1079]" gridcol="1" gridrow="0" pageId="16" pageNumber="45">2+1</th>
<th id="AD6FEF6BFFF5ABA7B7CEA05FFC8DFBD0" box="[482,833,1029,1079]" gridcol="2" gridrow="0" pageId="16" pageNumber="45">baculiform, equal in size; third tiny papillate</th>
<th id="AD6FEF6BFFF5ABA7B5BCA05FFA5BFBD0" box="[912,1431,1029,1079]" gridcol="3" gridrow="0" pageId="16" pageNumber="45">semi-erect in line in short common depression; third laterally</th>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA010FA5BFBB8" box="[151,1431,1098,1119]" gridrow="1" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA010FED2FBB8" box="[151,286,1098,1119]" gridcol="0" gridrow="1" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A010FF30FBB8" box="[159,252,1098,1119]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF55448B6B3A010FF30FBB8" box="[159,252,1098,1119]" italics="true" pageId="16" pageNumber="45">polonicus</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A010FE4DFBB8" box="[348,385,1098,1119]" gridcol="1" gridrow="1" pageId="16" pageNumber="45">2+1</td>
<td id="AD6FEF6BFFF5ABA7B7CEA010FC8DFBB8" box="[482,833,1098,1119]" gridcol="2" gridrow="1" pageId="16" pageNumber="45">banana-shaped; third small T-shaped</td>
<td id="AD6FEF6BFFF5ABA7B5BCA010FA5BFBB8" box="[912,1431,1098,1119]" gridcol="3" gridrow="1" pageId="16" pageNumber="45">recumbent, in tandem in confluent depressions</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA028FA5BFB60" box="[151,1431,1138,1159]" gridrow="2" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA028FED2FB60" box="[151,286,1138,1159]" gridcol="0" gridrow="2" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A028FF2EFB60" box="[159,226,1138,1159]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="turneri">
<emphasis id="62FA5847FFF55448B6B3A028FF2EFB60" box="[159,226,1138,1159]" italics="true" pageId="16" pageNumber="45">turneri</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A028FE4DFB60" box="[348,385,1138,1159]" gridcol="1" gridrow="2" pageId="16" pageNumber="45">2+1</td>
<td id="AD6FEF6BFFF5ABA7B7CEA028FC8DFB60" box="[482,833,1138,1159]" gridcol="2" gridrow="2" pageId="16" pageNumber="45">rod-like</td>
<td id="AD6FEF6BFFF5ABA7B5BCA028FA5BFB60" box="[912,1431,1138,1159]" gridcol="3" gridrow="2" pageId="16" pageNumber="45">in line; third laterally</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA0F1FA5BFB25" box="[151,1431,1195,1218]" gridrow="3" pageId="16" pageNumber="45" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="AD6FEF6BFFF5ABA7B6BBA0F1FED2FB25" box="[151,286,1195,1218]" gridcol="0" gridrow="3" pageId="16" pageNumber="45">continued.</th>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFF55448B6B3A084FC7EFA12" blockId="16.[159,637,1246,1267]" lastBlockId="16.[159,1357,1475,1555]" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A084FE7BFB14" authority="Solenidia" authorityName="Solenidia" box="[159,439,1246,1267]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="genus">Penthalodes Solenidia</taxonomicName>
of rhagidial organ II
<taxonomicName id="978EFFD6FFF55448B6B3A199FEDDFA3F" box="[159,273,1475,1496]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF55448B6B3A199FEDDFA3F" box="[159,273,1475,1496]" italics="true" pageId="16" pageNumber="45">hawaiiensis</emphasis>
</taxonomicName>
3+1(f?) baculiform, unequal size: proximal longest and thickest; staggered in confluent depressions distal smallest and thinnest; fourth tiny spine-like
</paragraph>
<paragraph id="50318455FFF55448B770A15CFA5BFA57" pageId="16" pageNumber="45">
<table id="228E76F5FFF5ABA7B6BBA15CFA5BFA57" box="[151,1431,1286,1456]" gridcols="4" gridrows="4" pageId="16" pageNumber="45">
<tr id="EEBE8617FFF5ABA7B6BBA15CFA5BFAFC" box="[151,1431,1286,1307]" gridrow="0" pageId="16" pageNumber="45" rowspan-0="1">
<th id="AD6FEF6BFFF5ABA7B770A15CFE66FAFC" box="[348,426,1286,1307]" gridcol="1" gridrow="0" pageId="16" pageNumber="45">Number</th>
<th id="AD6FEF6BFFF5ABA7B7F4A15CFC0CFAFC" box="[472,960,1286,1307]" gridcol="2" gridrow="0" pageId="16" pageNumber="45">Shape</th>
<th id="AD6FEF6BFFF5ABA7B22EA15CFA5BFAFC" box="[1026,1431,1286,1307]" gridcol="3" gridrow="0" pageId="16" pageNumber="45">Arrangement</th>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA174FA5BFA87" box="[151,1431,1326,1376]" gridrow="1" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA174FEC2FA87" box="[151,270,1326,1376]" gridcol="0" gridrow="1" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A174FEC2FAA4" box="[159,270,1326,1347]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF55448B6B3A174FEC2FAA4" box="[159,270,1326,1347]" italics="true" pageId="16" pageNumber="45">alaskaensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A174FE66FA87" box="[348,426,1326,1376]" gridcol="1" gridrow="1" pageId="16" pageNumber="45">3</td>
<td id="AD6FEF6BFFF5ABA7B7F4A174FC0CFA87" box="[472,960,1326,1376]" gridcol="2" gridrow="1" pageId="16" pageNumber="45">baculiform; two proximal equal in size; third distal longest</td>
<td id="AD6FEF6BFFF5ABA7B22EA174FA5BFA87" box="[1026,1431,1326,1376]" gridcol="3" gridrow="1" pageId="16" pageNumber="45">recumbent in tandem; in confluent depressions</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA129FA5BFA6F" box="[151,1431,1395,1416]" gridrow="2" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA129FEC2FA6F" box="[151,270,1395,1416]" gridcol="0" gridrow="2" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A129FF17FA6F" box="[159,219,1395,1416]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFF55448B6B3A129FF17FA6F" box="[159,219,1395,1416]" italics="true" pageId="16" pageNumber="45">boneti</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A129FE66FA6F" box="[348,426,1395,1416]" gridcol="1" gridrow="2" pageId="16" pageNumber="45">3</td>
<td id="AD6FEF6BFFF5ABA7B7F4A129FC0CFA6F" box="[472,960,1395,1416]" gridcol="2" gridrow="2" pageId="16" pageNumber="45">rod-like</td>
<td id="AD6FEF6BFFF5ABA7B22EA129FA5BFA6F" box="[1026,1431,1395,1416]" gridcol="3" gridrow="2" pageId="16" pageNumber="45">in line</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA1C1FA5BFA57" box="[151,1431,1435,1456]" gridrow="3" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA1C1FEC2FA57" box="[151,270,1435,1456]" gridcol="0" gridrow="3" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A1C1FF35FA57" box="[159,249,1435,1456]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF55448B6B3A1C1FF35FA57" box="[159,249,1435,1456]" italics="true" pageId="16" pageNumber="45">carinatus</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF5ABA7B770A1C1FE66FA57" box="[348,426,1435,1456]" gridcol="1" gridrow="3" pageId="16" pageNumber="45">3</td>
<td id="AD6FEF6BFFF5ABA7B7F4A1C1FC0CFA57" box="[472,960,1435,1456]" gridcol="2" gridrow="3" pageId="16" pageNumber="45">slightly T-shaped</td>
<td id="AD6FEF6BFFF5ABA7B22EA1C1FA5BFA57" box="[1026,1431,1435,1456]" gridcol="3" gridrow="3" pageId="16" pageNumber="45">in separate depressions; two distal parallel</td>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFF55448B7F4A1A4FD8CF9F4" blockId="16.[159,1357,1475,1555]" box="[472,576,1534,1555]" pageId="16" pageNumber="45">posteriorly</paragraph>
<paragraph id="50318455FFF55448B6B3A27CFBC1F94F" pageId="16" pageNumber="45">
<table id="228E76F5FFF5ABA7B6BBA27CFA5BF922" box="[151,1431,1574,1733]" gridcols="3" gridrows="3" pageId="16" pageNumber="45">
<tr id="EEBE8617FFF5ABA7B6BBA27CFA5BF9DC" box="[151,1431,1574,1595]" gridrow="0" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA27CFEA4F9DC" box="[151,360,1574,1595]" gridcol="0" gridrow="0" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A27CFEDEF9DC" box="[159,274,1574,1595]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFF55448B6B3A27CFEDEF9DC" box="[159,274,1574,1595]" italics="true" pageId="16" pageNumber="45">oregonensis</emphasis>
</taxonomicName>
3
</th>
<th id="AD6FEF6BFFF5ABA7B7F4A27CFC43F9DC" box="[472,911,1574,1595]" gridcol="1" gridrow="0" pageId="16" pageNumber="45">rod-like</th>
<th id="AD6FEF6BFFF5ABA7B22EA27CFA5BF9DC" box="[1026,1431,1574,1595]" gridcol="2" gridrow="0" pageId="16" pageNumber="45">in line</th>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA214FA5BF967" box="[151,1431,1614,1664]" gridrow="1" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA214FEA4F967" box="[151,360,1614,1664]" gridcol="0" gridrow="1" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A214FEDFF984" box="[159,275,1614,1635]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF55448B6B3A214FEDFF984" box="[159,275,1614,1635]" italics="true" pageId="16" pageNumber="45">ovalis sensu</emphasis>
</taxonomicName>
3 Kaluz (2000)
</th>
<td id="AD6FEF6BFFF5ABA7B7F4A214FC43F967" box="[472,911,1614,1664]" gridcol="1" gridrow="1" pageId="16" pageNumber="45">proximal lanceolate sharply tipped; two distal baculiform</td>
<td id="AD6FEF6BFFF5ABA7B22EA214FA5BF967" box="[1026,1431,1614,1664]" gridcol="2" gridrow="1" pageId="16" pageNumber="45">proximal in own depression separately; two distal parallel in common depression</td>
</tr>
<tr id="EEBE8617FFF5ABA7B6BBA2C9FA5BF922" box="[151,1431,1683,1733]" gridrow="2" pageId="16" pageNumber="45">
<th id="AD6FEF6BFFF5ABA7B6BBA2C9FEA4F922" box="[151,360,1683,1733]" gridcol="0" gridrow="2" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A2C9FEDFF94F" authority="Baker" authorityName="Baker" authorityYear="1990" box="[159,275,1683,1704]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF55448B6B3A2C9FEDFF94F" box="[159,275,1683,1704]" italics="true" pageId="16" pageNumber="45">ovalis sensu</emphasis>
</taxonomicName>
? Baker (1990)
</th>
<td id="AD6FEF6BFFF5ABA7B7F4A2C9FC43F922" box="[472,911,1683,1733]" gridcol="1" gridrow="2" pageId="16" pageNumber="45">?</td>
<td id="AD6FEF6BFFF5ABA7B22EA2C9FA5BF922" box="[1026,1431,1683,1733]" gridcol="2" gridrow="2" pageId="16" pageNumber="45">?</td>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFF55448B6B3A283FB27F8EC" blockId="16.[159,1350,1752,1843]" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A283FF30F909" box="[159,252,1753,1774]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF55448B6B3A283FF30F909" box="[159,252,1753,1774]" italics="true" pageId="16" pageNumber="45">polonicus</emphasis>
</taxonomicName>
3+1(f?) thin baculiform, almost equal, proximal thickest; fourth in tandem in common depression, tiny spine-like laterally overlapping one another
</paragraph>
<paragraph id="50318455FFF55448B6B3A344FBF3F8D4" blockId="16.[159,1350,1752,1843]" box="[159,1087,1822,1843]" pageId="16" pageNumber="45">
<taxonomicName id="978EFFD6FFF55448B6B3A344FF2EF8D4" box="[159,226,1822,1843]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="16" pageNumber="45" phylum="Arthropoda" rank="species" species="turneri">
<emphasis id="62FA5847FFF55448B6B3A344FF2EF8D4" box="[159,226,1822,1843]" italics="true" pageId="16" pageNumber="45">turneri</emphasis>
</taxonomicName>
3 rod-like in line
</paragraph>
<paragraph id="50318455FFF55449B6E9A322FAB2FE75" blockId="16.[151,1437,1912,2013]" lastBlockId="17.[151,1437,152,402]" lastPageId="17" lastPageNumber="46" pageId="16" pageNumber="45">
<emphasis id="62FA5847FFF55448B6E9A322FD0CF875" bold="true" box="[197,704,1912,1938]" pageId="16" pageNumber="45">
<emphasis id="62FA5847FFF55448B6E9A322FE55F875" bold="true" box="[197,409,1912,1938]" italics="true" pageId="16" pageNumber="45">Rhagidial organs</emphasis>
,
<emphasis id="62FA5847FFF55448B785A322FD0CF875" bold="true" box="[425,704,1912,1938]" italics="true" pageId="16" pageNumber="45">solenidia and leg setae</emphasis>
</emphasis>
(
<tableCitation id="1D0CB1EEFFF55448B4FCA323FCF5F874" box="[720,825,1913,1939]" captionStart="TABLE 4" captionStartId="16.[151,239,599,623]" captionTargetBox="[151,1418,690,941]" captionTargetPageId="16" captionText="TABLE 4. Solenidia of the rhagidial organ on tarsi I and II in representatives of Penthalodes. Penthalodes Solenidia of rhagidial organ I" httpUri="http://table.plazi.org/id/04F1D4DDFFF55448B6BBA60DFDB9FD78" pageId="16" pageNumber="45" tableUuid="04F1D4DDFFF55448B6BBA60DFDB9FD78">Tables 4</tableCitation>
,
<tableCitation id="1D0CB1EEFFF55448B56AA323FC9BF874" box="[838,855,1913,1939]" captionStart="TABLE 5" captionStartId="17.[151,244,450,474]" captionText="TABLE 5. Solenidia on tibiae and genua in representatives of Penthalodes. No data available for P. boneti, P." pageId="16" pageNumber="45">5</tableCitation>
). The rhagidial organ on tarsus I consists of two dorsal solenidia and a small lateral solenidion. The rhagidial organ on tarsus II consists of three dorsal solenidia and a small, rod-like solenidion (or famulus?) lying laterally in a separate pit. Species differences result from solenidia shape, the manner of their arrangement as well as whether they lie in common or separate depressions. It seems that in species of
<taxonomicName id="978EFFD6FFF45449B4E2A4E7FC93FF31" box="[718,863,189,214]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF45449B4E2A4E7FC93FF31" box="[718,863,189,214]" italics="true" pageId="17" pageNumber="46">Penthalodes</emphasis>
</taxonomicName>
there are no famuli typical of other Eupodoidea. Most probably they are replaced by a tiny solenidion. In some species, however, there is a small, stellate famulus on tarsus I. This famulus is placed a small distance from the rhagidial organ in
<taxonomicName id="978EFFD6FFF45449B2C9A552FAB6FEC6" box="[1253,1402,264,289]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF45449B2C9A552FAB6FEC6" box="[1253,1402,264,289]" italics="true" pageId="17" pageNumber="46">P. carinatus</emphasis>
</taxonomicName>
or laterally and far behind it in
<emphasis id="62FA5847FFF45449B7D0A577FD43FEA1" box="[508,655,301,326]" italics="true" pageId="17" pageNumber="46">P. p o l o n i c u s</emphasis>
.
<bibRefCitation id="341FF9A4FFF45449B488A577FC5EFEA0" author="Strandmann" box="[676,914,301,327]" pageId="17" pageNumber="46" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandmann (1971)</bibRefCitation>
also reported a famulus in
<emphasis id="62FA5847FFF45449B2CBA577FA5DFEA1" box="[1255,1425,301,326]" italics="true" pageId="17" pageNumber="46">P. a l a s k a e n s i s</emphasis>
, accompanying the rhagidial organ. However, after repeated verification, he considered it is missing, particularly in specimens from Hawaii (
<bibRefCitation id="341FF9A4FFF45449B444A522FC6EFE75" author="Strandtmann" box="[616,930,376,402]" pageId="17" pageNumber="46" refString="Strandtmann, R. W. &amp; Goff, M. L. (1978) The Eupodoidea of Hawaii (Acarina: Prostigmata). Pacific Insects, 19 (3 - 4), 121 - 143." type="journal article" year="1978">Strandtmann &amp; Goff 1978</bibRefCitation>
). Famuli are not known in other species.
</paragraph>
<caption id="04F1D4DDFFF45449B6BBA598FA57FE3E" box="[151,1435,450,474]" pageId="17" pageNumber="46">
<paragraph id="50318455FFF45449B6BBA598FA57FE3E" blockId="17.[151,1435,450,1116]" box="[151,1435,450,474]" pageId="17" pageNumber="46">
<emphasis id="62FA5847FFF45449B6BBA598FEDEFE3D" bold="true" box="[151,274,450,474]" pageId="17" pageNumber="46">TABLE 5.</emphasis>
Solenidia on tibiae and genua in representatives of
<taxonomicName id="978EFFD6FFF45449B558A598FBCEFE3E" box="[884,1026,450,473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF45449B558A598FBCEFE3E" box="[884,1026,450,473]" italics="true" pageId="17" pageNumber="46">Penthalodes</emphasis>
</taxonomicName>
. No data available for
<taxonomicName id="978EFFD6FFF45449B33DA598FABBFE3E" box="[1297,1399,450,473]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFF45449B33DA598FABBFE3E" box="[1297,1399,450,473]" italics="true" pageId="17" pageNumber="46">P. boneti</emphasis>
</taxonomicName>
, P.
</paragraph>
</caption>
<paragraph id="50318455FFF45449B6BBA5B8FB2EFE1E" blockId="17.[151,1435,450,1116]" box="[151,1250,482,505]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6BBA5B8FEDBFE1E" box="[151,279,482,505]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="oregonensis">
<emphasis id="62FA5847FFF45449B6BBA5B8FEDBFE1E" box="[151,279,482,505]" italics="true" pageId="17" pageNumber="46">oregonensis</emphasis>
</taxonomicName>
and
<emphasis id="62FA5847FFF45449B766A5B8FE67FE1E" box="[330,427,482,505]" italics="true" pageId="17" pageNumber="46">P. t u r n e r i</emphasis>
. Abbreviations: DD = dorsodistal; DM = dorsomedial; DP = dorsoproximal.
</paragraph>
<paragraph id="50318455FFF45449B6B3A64FFB30FDCD" blockId="17.[151,1435,450,1116]" box="[159,1276,533,554]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A64FFEDFFDCD" box="[159,275,533,554]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="49" phylum="Arthropoda" rank="genus">Penthalodes</taxonomicName>
Tibia I Tibia II Tibia III Tibia IV
</paragraph>
<paragraph id="50318455FFF45449B6B3A667FC8CFD88" blockId="17.[151,1435,450,1116]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A667FEC2FDB5" box="[159,270,573,594]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF45449B6B3A667FEC2FDB5" box="[159,270,573,594]" italics="true" pageId="17" pageNumber="46">alaskaensis</emphasis>
</taxonomicName>
2 DD in line: one in pit and 2 DD in line: one in pit and 1 DM, erect 1 DM, erect longer one piliform longer one piliform
</paragraph>
<paragraph id="50318455FFF45449B6B3A6D8FCBFFD53" blockId="17.[151,1435,450,1116]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A6D8FF35FD70" box="[159,249,642,663]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF45449B6B3A6D8FF35FD70" box="[159,249,642,663]" italics="true" pageId="17" pageNumber="46">carinatus</emphasis>
</taxonomicName>
2 DD in line: one thicker 2 DD in line: one thicker 1 DM, erect 1 DM, erect baculiform (T) in pit and baculiform (T) in pit and
</paragraph>
<paragraph id="50318455FFF45449B778A6E7FCAFFD35" blockId="17.[151,1435,450,1116]" box="[340,867,701,722]" pageId="17" pageNumber="46">much longer one thin piliform slightly longer one thin</paragraph>
<paragraph id="50318455FFF45449B4A9A680FD18FD08" blockId="17.[151,1435,450,1116]" box="[645,724,730,751]" pageId="17" pageNumber="46">piliform</paragraph>
<paragraph id="50318455FFF45449B6B3A758FC53FCD3" blockId="17.[151,1435,450,1116]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A758FEDDFCF0" box="[159,273,770,791]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF45449B6B3A758FEDDFCF0" box="[159,273,770,791]" italics="true" pageId="17" pageNumber="46">hawaiiensis</emphasis>
</taxonomicName>
2 DD in line: one thicker rod- 2 DD in line: one thicker rod- 1 slender erect 1 slender erect like in pit and longer one thin like in pit and longer one thin
</paragraph>
<paragraph id="50318455FFF45449B778A767FD18FCB5" blockId="17.[151,1435,450,1116]" box="[340,724,829,850]" pageId="17" pageNumber="46">piliform piliform</paragraph>
<paragraph id="50318455FFF45449B6B3A73FFA47FC9D" blockId="17.[151,1435,450,1116]" box="[159,1419,869,890]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A73FFEDFFC9D" box="[159,275,869,890]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF45449B6B3A73FFEDFFC9D" box="[159,275,869,890]" italics="true" pageId="17" pageNumber="46">ovalis sensu</emphasis>
</taxonomicName>
2 DD in line: one longer rod- 2 DD in line: one slightly DP short baculiform DP slightly longer than
</paragraph>
<paragraph id="50318455FFF45449B6B3A7D8FAD9FC53" blockId="17.[151,1435,450,1116]" pageId="17" pageNumber="46">
<bibRefCitation id="341FF9A4FFF45449B6B3A7D8FED3FC70" author="Kaluz" box="[159,287,898,919]" pageId="17" pageNumber="46" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
like in pit and one thicker thicker and longer rod-like in that on Tb III sharply-tipped spiniform pit and one spiniform baculiform
</paragraph>
<paragraph id="50318455FFF45449B6B3A792FB7BFC3B" blockId="17.[151,1435,450,1116]" box="[159,1207,967,989]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A792FEDFFC3A" box="[159,275,968,989]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF45449B6B3A792FEDFFC3A" box="[159,275,968,989]" italics="true" pageId="17" pageNumber="46">ovalis sensu</emphasis>
</taxonomicName>
????
</paragraph>
<paragraph id="50318455FFF45449B6B3A7BFFED2FC1D" blockId="17.[151,1435,450,1116]" box="[159,286,997,1018]" pageId="17" pageNumber="46">
<collectingCountry id="2899C4C5FFF45449B6B3A7BFFF14FC1D" box="[159,216,997,1018]" name="United States Pacific Island Wildlife Refuges" pageId="17" pageNumber="46">Baker</collectingCountry>
(1990)
</paragraph>
<paragraph id="50318455FFF45449B6B3A057FA4CFBC5" blockId="17.[151,1435,450,1116]" box="[159,1408,1037,1058]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A057FF30FBC5" box="[159,252,1037,1058]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF45449B6B3A057FF30FBC5" box="[159,252,1037,1058]" italics="true" pageId="17" pageNumber="46">polonicus</emphasis>
</taxonomicName>
2 DD in line: one thicker 2 DD in line: one longer thick 1 DP erect baculiform 1 DP erect baculiform
</paragraph>
<paragraph id="50318455FFF45449B778A070FC51FBD8" blockId="17.[151,1435,450,1116]" box="[340,925,1066,1087]" pageId="17" pageNumber="46">banana-shaped in pit and banana-shaped in pit and one</paragraph>
<paragraph id="50318455FFF45449B778A01DFD1DFBBB" blockId="17.[151,1435,450,1116]" box="[340,721,1095,1116]" pageId="17" pageNumber="46">shorter one baculiform rod-like</paragraph>
<paragraph id="50318455FFF45449B6BBA0DAFEC4FB70" blockId="17.[151,264,1152,1175]" box="[151,264,1152,1175]" pageId="17" pageNumber="46">continued.</paragraph>
<paragraph id="50318455FFF45449B6B3A0E9FAE8FAB4" pageId="17" pageNumber="46">
<table id="228E76F5FFF4ABA7B6BBA0E9FA43FAB4" box="[151,1423,1203,1363]" gridcols="5" gridrows="3" pageId="17" pageNumber="46">
<tr id="EEBE8617FFF4ABA7B6BBA0E9FA43FB2F" box="[151,1423,1203,1224]" gridrow="0" pageId="17" pageNumber="46">
<th id="AD6FEF6BFFF4ABA7B6BBA0E9FEDFFB2F" box="[151,275,1203,1224]" gridcol="0" gridrow="0" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A0E9FEDFFB2F" box="[159,275,1203,1224]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="49" phylum="Arthropoda" rank="genus">Penthalodes</taxonomicName>
</th>
<th id="AD6FEF6BFFF4ABA7B778A0E9FD86FB2F" box="[340,586,1203,1224]" gridcol="1" gridrow="0" pageId="17" pageNumber="46">Genu I</th>
<th id="AD6FEF6BFFF4ABA7B451A0E9FCBFFB2F" box="[637,883,1203,1224]" gridcol="2" gridrow="0" pageId="17" pageNumber="46">Genu II</th>
<th id="AD6FEF6BFFF4ABA7B589A0E9FBA5FB2F" box="[933,1129,1203,1224]" gridcol="3" gridrow="0" pageId="17" pageNumber="46">Genu III</th>
<th id="AD6FEF6BFFF4ABA7B29DA0E9FA43FB2F" box="[1201,1423,1203,1224]" gridcol="4" gridrow="0" pageId="17" pageNumber="46">Genu IV</th>
</tr>
<tr id="EEBE8617FFF4ABA7B6BBA081FA43FACC" box="[151,1423,1243,1323]" gridrow="1" pageId="17" pageNumber="46">
<th id="AD6FEF6BFFF4ABA7B6BBA081FEDFFACC" box="[151,275,1243,1323]" gridcol="0" gridrow="1" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A081FEC2FB17" box="[159,270,1243,1264]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF45449B6B3A081FEC2FB17" box="[159,270,1243,1264]" italics="true" pageId="17" pageNumber="46">alaskaensis</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF4ABA7B778A081FD86FACC" box="[340,586,1243,1323]" gridcol="1" gridrow="1" pageId="17" pageNumber="46">1 DD, erect; plus 1 tiny famulus at dorsal anterior margin</td>
<td id="AD6FEF6BFFF4ABA7B451A081FCBFFACC" box="[637,883,1243,1323]" gridcol="2" gridrow="1" pageId="17" pageNumber="46">1 DD, erect; plus 1 tiny famulus at dorsal anterior margin</td>
<td id="AD6FEF6BFFF4ABA7B589A081FBA5FACC" box="[933,1129,1243,1323]" gridcol="3" gridrow="1" pageId="17" pageNumber="46">1 DD, erect piliform</td>
<td id="AD6FEF6BFFF4ABA7B29DA081FA43FACC" box="[1201,1423,1243,1323]" gridcol="4" gridrow="1" pageId="17" pageNumber="46">1 DD, erect piliform</td>
</tr>
<tr id="EEBE8617FFF4ABA7B6BBA164FA43FAB4" box="[151,1423,1342,1363]" gridrow="2" pageId="17" pageNumber="46">
<th id="AD6FEF6BFFF4ABA7B6BBA164FEDFFAB4" box="[151,275,1342,1363]" gridcol="0" gridrow="2" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A164FF35FAB4" box="[159,249,1342,1363]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF45449B6B3A164FF35FAB4" box="[159,249,1342,1363]" italics="true" pageId="17" pageNumber="46">carinatus</emphasis>
</taxonomicName>
</th>
<td id="AD6FEF6BFFF4ABA7B778A164FD86FAB4" box="[340,586,1342,1363]" gridcol="1" gridrow="2" pageId="17" pageNumber="46">1 DM, erect</td>
<td id="AD6FEF6BFFF4ABA7B451A164FCBFFAB4" box="[637,883,1342,1363]" gridcol="2" gridrow="2" pageId="17" pageNumber="46">1 DM, erect</td>
<td id="AD6FEF6BFFF4ABA7B589A164FBA5FAB4" box="[933,1129,1342,1363]" gridcol="3" gridrow="2" pageId="17" pageNumber="46">1 DM, erect</td>
<td id="AD6FEF6BFFF4ABA7B29DA164FA43FAB4" box="[1201,1423,1342,1363]" gridcol="4" gridrow="2" pageId="17" pageNumber="46">1 DM, erect</td>
</tr>
</table>
</paragraph>
<paragraph id="50318455FFF45449B6B3A13CFC82FA7F" blockId="17.[159,1423,1382,1728]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A13CFEDDFA9C" box="[159,273,1382,1403]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF45449B6B3A13CFEDDFA9C" box="[159,273,1382,1403]" italics="true" pageId="17" pageNumber="46">hawaiiensis</emphasis>
</taxonomicName>
1 DM, erect piliform; plus 1 1 DD, erect piliform; plus 1 1 slender erect 1 slender erect tiny famulus at dorsal tiny famulus at dorsal
</paragraph>
<paragraph id="50318455FFF45449B778A1FAFCDCFA52" blockId="17.[159,1423,1382,1728]" box="[340,784,1440,1461]" pageId="17" pageNumber="46">anterior margin anterior margin</paragraph>
<paragraph id="50318455FFF45449B6B3A193FA43FA3A" blockId="17.[159,1423,1382,1728]" box="[159,1423,1480,1502]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A193FEDFFA39" box="[159,275,1481,1502]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF45449B6B3A193FEDFFA39" box="[159,275,1481,1502]" italics="true" pageId="17" pageNumber="46">ovalis sensu</emphasis>
</taxonomicName>
1 DM, erect short spiniform 1 DM, erect short spiniform 1 DM erect baculiform 1 DD erect baculiform
</paragraph>
<paragraph id="50318455FFF45449B6B3A1BCFED3FA1C" blockId="17.[159,1423,1382,1728]" box="[159,287,1510,1531]" pageId="17" pageNumber="46">
<bibRefCitation id="341FF9A4FFF45449B6B3A1BCFED3FA1C" author="Kaluz" box="[159,287,1510,1531]" pageId="17" pageNumber="46" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
</paragraph>
<paragraph id="50318455FFF45449B6B3A254FB70F9C4" blockId="17.[159,1423,1382,1728]" box="[159,1212,1550,1571]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A254FEDFF9C4" box="[159,275,1550,1571]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="subSpecies" species="ovalis" subSpecies="sensu">
<emphasis id="62FA5847FFF45449B6B3A254FEDFF9C4" box="[159,275,1550,1571]" italics="true" pageId="17" pageNumber="46">ovalis sensu</emphasis>
</taxonomicName>
?? distal papilla (Fig.13f)?
</paragraph>
<paragraph id="50318455FFF45449B6B3A271FED2F9A7" blockId="17.[159,1423,1382,1728]" box="[159,286,1579,1600]" pageId="17" pageNumber="46">
<collectingCountry id="2899C4C5FFF45449B6B3A271FF14F9A7" box="[159,216,1579,1600]" name="United States Pacific Island Wildlife Refuges" pageId="17" pageNumber="46">Baker</collectingCountry>
(1990)
</paragraph>
<paragraph id="50318455FFF45449B6B3A209FA40F98F" blockId="17.[159,1423,1382,1728]" box="[159,1420,1619,1640]" pageId="17" pageNumber="46">
<taxonomicName id="978EFFD6FFF45449B6B3A209FF30F98F" box="[159,252,1619,1640]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF45449B6B3A209FF30F98F" box="[159,252,1619,1640]" italics="true" pageId="17" pageNumber="46">polonicus</emphasis>
</taxonomicName>
1 distal large banana-shaped; 1 distal large banana-shaped; 1 distal banana-shaped 1 distal banana-shaped
</paragraph>
<paragraph id="50318455FFF45449B778A22AFC89F962" blockId="17.[159,1423,1382,1728]" box="[340,837,1648,1669]" pageId="17" pageNumber="46">plus 1 tiny bifurcate famulus plus 1 tiny spiniform</paragraph>
<paragraph id="50318455FFF45449B778A2D4FCBFF944" blockId="17.[159,1423,1382,1728]" box="[340,883,1678,1699]" pageId="17" pageNumber="46">at dorsal anterior margin famulus at dorsal anterior</paragraph>
<paragraph id="50318455FFF45449B451A2F1FD0CF927" blockId="17.[159,1423,1382,1728]" box="[637,704,1707,1728]" pageId="17" pageNumber="46">margin</paragraph>
<paragraph id="50318455FFF45449B6E9A35CFBE5F83C" blockId="17.[151,1436,1798,2011]" pageId="17" pageNumber="46">
The lateral solenidion on tarsus I is always very small and situated in a separate depression. In
<taxonomicName id="978EFFD6FFF45449B31BA35CFEE5F8A2" authority="Kaluz" authorityName="Kaluz" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF45449B31BA35CFA50F8F8" box="[1335,1436,1798,1823]" italics="true" pageId="17" pageNumber="46">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
, it is rod-like (according to Kaluz, it is spiniform; see also Fig. 13c in
<collectingCountry id="2899C4C5FFF45449B244A371FB62F8A2" box="[1128,1198,1835,1861]" name="United States Pacific Island Wildlife Refuges" pageId="17" pageNumber="46">Baker</collectingCountry>
1990) and is placed behind the rhagidial organ. In
<emphasis id="62FA5847FFF45449B7D6A30BFD49F88D" box="[506,645,1873,1898]" italics="true" pageId="17" pageNumber="46">P. p o l o n i c u s</emphasis>
, on the other hand, it is relatively large, T-shaped and is situated at the level of the proximal solenidion.
<bibRefCitation id="341FF9A4FFF45449B47FA32CFD3EF877" author="Baker" box="[595,754,1910,1936]" pageId="17" pageNumber="46" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFF45449B47FA32CFD56F877" box="[595,666,1910,1936]" name="United States Pacific Island Wildlife Refuges" pageId="17" pageNumber="46">Baker</collectingCountry>
(1946)
</bibRefCitation>
mentions the lateral solenidion (as a sensory seta) of the rhagidial organ in all species described by him; it is rod-like as other solenidia. A similar situation occurs in
<taxonomicName id="978EFFD6FFF45449B3AAA3C1FEEDF83D" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF45449B3AAA3C1FEEDF83D" italics="true" pageId="17" pageNumber="46">P. hawaiiensis</emphasis>
</taxonomicName>
. In
<taxonomicName id="978EFFD6FFF45449B763A39BFDE1F83C" authority="Shiba" authorityName="Shiba" box="[335,557,1985,2011]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="17" pageNumber="46" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF45449B763A39BFE17F83D" box="[335,475,1985,2010]" italics="true" pageId="17" pageNumber="46">P. carinatus</emphasis>
, Shiba
</taxonomicName>
did not observe the solenidion of that
<typeStatus id="8F353AF7FFF45449B5C3A39BFBE9F83C" box="[1007,1061,1985,2011]" pageId="17" pageNumber="46">type</typeStatus>
.
</paragraph>
<paragraph id="50318455FFF7544AB6E9A4C2FA82FE8B" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="47">
The two solenidia of rhagidial organ I can be in tandem in confluent depressions, like in
<taxonomicName id="978EFFD6FFF7544AB2E4A4C2FAA7FF56" box="[1224,1387,152,177]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF7544AB2E4A4C2FAA7FF56" box="[1224,1387,152,177]" italics="true" pageId="18" pageNumber="47">P. alaskaensis</emphasis>
</taxonomicName>
and
<emphasis id="62FA5847FFF7544AB6BBA4E7FEE8FF31" box="[151,292,189,214]" italics="true" pageId="18" pageNumber="47">P. p o l o n i c u s</emphasis>
, or in separate depressions, like in
<taxonomicName id="978EFFD6FFF7544AB4E1A4E7FC97FF31" box="[717,859,189,214]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF7544AB4E1A4E7FC97FF31" box="[717,859,189,214]" italics="true" pageId="18" pageNumber="47">P. carinatus</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFF7544AB5BAA4E7FADFFF30" authority="Kaluz. Solenidia" authorityName="Kaluz. Solenidia" box="[918,1299,189,215]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF7544AB5BAA4E7FC36FF31" box="[918,1018,189,214]" italics="true" pageId="18" pageNumber="47">P. ovalis</emphasis>
sensu Kaluz. Solenidia
</taxonomicName>
are usually baculiform, except in
<taxonomicName id="978EFFD6FFF7544AB7B4A4B8FD5FFF1B" authority="Kaluz" authorityName="Kaluz" box="[408,659,226,252]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF7544AB7B4A4B8FE36FF1C" box="[408,506,226,251]" italics="true" pageId="18" pageNumber="47">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
, where they are lanceolate and sharply tipped; in
<taxonomicName id="978EFFD6FFF7544AB2FBA4B8FAAFFF1C" box="[1239,1379,226,251]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF7544AB2FBA4B8FAAFFF1C" box="[1239,1379,226,251]" italics="true" pageId="18" pageNumber="47">P. carinatus</emphasis>
</taxonomicName>
they resemble the letter “T”; and in
<emphasis id="62FA5847FFF7544AB42FA552FD42FEC6" box="[515,654,264,289]" italics="true" pageId="18" pageNumber="47">P. p o l o n i c u s</emphasis>
a banana. In
<taxonomicName id="978EFFD6FFF7544AB51CA552FC72FEC6" box="[816,958,264,289]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB51CA552FC72FEC6" box="[816,958,264,289]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
, the distal solenidion protrudes from the short depression; it is also characteristic that the distal depression is below the proximal one. The solenidia in
<emphasis id="62FA5847FFF7544AB6BBA508FF3AFE8C" box="[151,246,338,363]" italics="true" pageId="18" pageNumber="47">P. o v a l i s</emphasis>
sensu
<collectingCountry id="2899C4C5FFF7544AB765A508FE43FE8B" box="[329,399,338,364]" name="United States Pacific Island Wildlife Refuges" pageId="18" pageNumber="47">Baker</collectingCountry>
(1990) are semi-erect because the depression, in which they are inserted, is short.
</paragraph>
<paragraph id="50318455FFF7544AB6E9A522FB73FCCB" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="47">
The three solenidia of rhagidial organ II can be arranged one behind another in confluent depressions, like in
<taxonomicName id="978EFFD6FFF7544AB69AA5C7FE93FE51" box="[182,351,413,438]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF7544AB69AA5C7FE93FE51" box="[182,351,413,438]" italics="true" pageId="18" pageNumber="47">P. alaskaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="978EFFD6FFF7544AB743A5C7FE14FE51" box="[367,472,413,438]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="62FA5847FFF7544AB743A5C7FE14FE51" box="[367,472,413,438]" italics="true" pageId="18" pageNumber="47">P. boneti</emphasis>
</taxonomicName>
,
<emphasis id="62FA5847FFF7544AB7C4A5C7FD5CFE51" box="[488,656,413,438]" italics="true" pageId="18" pageNumber="47">P. h a w a i i e n s i s</emphasis>
and
<taxonomicName id="978EFFD6FFF7544AB4FCA5C7FCAEFE51" box="[720,866,413,438]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB4FCA5C7FCAEFE51" box="[720,866,413,438]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
. Otherwise, in
<emphasis id="62FA5847FFF7544AB231A5C7FBB3FE51" box="[1053,1151,413,438]" italics="true" pageId="18" pageNumber="47">P. o v a l i s</emphasis>
sensu Kaluz two distal solenidia are parallel and situated in a common depression.
<emphasis id="62FA5847FFF7544AB58CA598FBFBFE3C" box="[928,1079,450,475]" italics="true" pageId="18" pageNumber="47">P. c a r i n a t u s</emphasis>
is similar, but has separate depressions. In
<taxonomicName id="978EFFD6FFF7544AB762A5B2FE3BFDE6" box="[334,503,488,513]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF7544AB762A5B2FE3BFDE6" box="[334,503,488,513]" italics="true" pageId="18" pageNumber="47">P. hawaiiensis</emphasis>
</taxonomicName>
, the bases of the solenidia do not lie in one line but are staggered in tandem. In
<emphasis id="62FA5847FFF7544AB6BBA657FEEFFDC1" box="[151,291,525,550]" italics="true" pageId="18" pageNumber="47">P. p o l o n i c u s</emphasis>
, the solenidia are in one line but overlap each other because the depression is short resembling a cascade. In
<taxonomicName id="978EFFD6FFF7544AB706A668FDF4FDAB" authority="Kaluz" authorityName="Kaluz" box="[298,568,562,588]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF7544AB706A668FE59FDAC" box="[298,405,562,587]" italics="true" pageId="18" pageNumber="47">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
, two distal solenidia are baculiform, while the proximal solenidion is lanceolate. In
<taxonomicName id="978EFFD6FFF7544AB760A602FE28FD96" box="[332,484,600,625]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="carinatus">
<emphasis id="62FA5847FFF7544AB760A602FE28FD96" box="[332,484,600,625]" italics="true" pageId="18" pageNumber="47">P. carinatus</emphasis>
</taxonomicName>
, the solenidia slightly resembles the letter “T”. In
<taxonomicName id="978EFFD6FFF7544AB25CA602FAE9FD96" box="[1136,1317,600,625]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF7544AB25CA602FAE9FD96" box="[1136,1317,600,625]" italics="true" pageId="18" pageNumber="47">P. hawaiiensis</emphasis>
</taxonomicName>
, they are baculiform and of different length and thickness; the proximal solenidion is the thickest and longest, while the distal one is the thinnest and shortest. In
<taxonomicName id="978EFFD6FFF7544AB451A6F8FCC2FD5C" box="[637,782,674,699]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB451A6F8FCC2FD5C" box="[637,782,674,699]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
, differences between solenidia are slightly less visible; and laterally to rhagidial organ II lies a small, spiniform solenidion or famulus (?) in a pit, like in
<taxonomicName id="978EFFD6FFF7544AB3A9A692FEEEFCE1" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF7544AB3A9A692FEEEFCE1" italics="true" pageId="18" pageNumber="47">P. hawaiiensis</emphasis>
</taxonomicName>
. This structure resembles a solenidion because it is hollow (transparent) proximally, while being solid (non-transparent) distally like a famulus. In other species, this has not been observed.
</paragraph>
<paragraph id="50318455FFF7544AB6E9A762FC97FA40" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="47">
The occurrence of solenidia on all tibiae and genua, plus single famuli on genu I and II near the distal margin of the article, is characteristic for
<taxonomicName id="978EFFD6FFF7544AB456A707FCC7FC91" box="[634,779,861,886]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF7544AB456A707FCC7FC91" box="[634,779,861,886]" italics="true" pageId="18" pageNumber="47">Penthalodes</emphasis>
</taxonomicName>
. Famuli are not recorded in other descriptions, although
<bibRefCitation id="341FF9A4FFF7544AB6BBA7D8FE4EFC7B" author="Strandtmann" box="[151,386,898,924]" pageId="18" pageNumber="47" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann (1971)</bibRefCitation>
observed tiny sensory setae and
<collectingCountry id="2899C4C5FFF7544AB52BA7D8FC81FC7B" box="[775,845,898,924]" name="United States Pacific Island Wildlife Refuges" pageId="18" pageNumber="47">Baker</collectingCountry>
(1990) mentioned a characteristic papilla on genu III. On tibiae I and II, the distal solenidion is situated in a depression, while the proximal solenidion is erect. Particularly large disproportions in the size of solenidia on tibia II occur in
<taxonomicName id="978EFFD6FFF7544AB219A797FB06FC01" box="[1077,1226,973,998]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB219A797FB06FC01" box="[1077,1226,973,998]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
, where the distal solenidion is very thick, and at least by half as long as the tiny, rod-like proximal solenidion. In
<emphasis id="62FA5847FFF7544AB324A7A8FA5DFBEC" box="[1288,1425,1010,1035]" italics="true" pageId="18" pageNumber="47">P. c a r i n a t u s</emphasis>
, the proximal solenidia are erect, sharp-pointed and clearly longer than the distal solenidia, which are thicker, rather T-shaped, and pointed distally.
<taxonomicName id="978EFFD6FFF7544AB449A067FCD8FBB1" box="[613,788,1085,1110]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF7544AB449A067FCD8FBB1" box="[613,788,1085,1110]" italics="true" pageId="18" pageNumber="47">P. hawaiiensis</emphasis>
</taxonomicName>
is similar, where the proximal solenidia are thin and longer than the rod-like, slightly thicker distal solenidia. In
<emphasis id="62FA5847FFF7544AB5A2A039FC3AFB9B" box="[910,1014,1123,1148]" italics="true" pageId="18" pageNumber="47">P. o v a l i s</emphasis>
sensu Kaluz, on tibia I the distal solenidion is rod-like and slightly longer and thinner then the proximal solenidion; on tibia II, the proximal solenidion is short and piliform, while the distal solenidion is rod-like and slightly thicker and longer than the proximal solenidion. The bases of distal solenidia lie in a deep pit so that only the distal part is visible from the pit aperture. Solenidia on genua I and II in this species are very short and spiniform, contrary to
<taxonomicName id="978EFFD6FFF7544AB320A0A2FA50FAF6" box="[1292,1436,1272,1297]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB320A0A2FA50FAF6" box="[1292,1436,1272,1297]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
where they are large, thick and banana-shaped. Solenidia on genua III and IV in
<taxonomicName id="978EFFD6FFF7544AB278A147FA83FAD0" authority="Kaluz" authorityName="Kaluz" box="[1108,1359,1309,1335]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF7544AB278A147FB7BFAD1" box="[1108,1207,1309,1334]" italics="true" pageId="18" pageNumber="47">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
are of similar length and thickness, contrary to
<taxonomicName id="978EFFD6FFF7544AB452A119FCC3FABB" box="[638,783,1347,1372]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB452A119FCC3FABB" box="[638,783,1347,1372]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
where the solenidion on genu IV is thicker and longer than that on genu III. In both species, solenidia on tibiae III and IV are of subequal length and thickness. For other species, there are no detailed descriptions or drawings.
</paragraph>
<paragraph id="50318455FFF7544AB6E9A1E8FC27F960" blockId="18.[151,1437,152,2007]" pageId="18" pageNumber="47">
Leg setae are characteristic for
<taxonomicName id="978EFFD6FFF7544AB468A1E9FD15FA2B" box="[580,729,1459,1484]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF7544AB468A1E9FD15FA2B" box="[580,729,1459,1484]" italics="true" pageId="18" pageNumber="47">Penthalodes</emphasis>
</taxonomicName>
. In
<taxonomicName id="978EFFD6FFF7544AB526A1E9FC51FA2B" box="[778,925,1459,1484]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB526A1E9FC51FA2B" box="[778,925,1459,1484]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
, they are rather plumose and have a short shaft with triangular outgrowths, with the apical outgrowth being clearly the longest. At the base of each apotele are three pairs of thick setae, expanded distally, which are distinct in
<taxonomicName id="978EFFD6FFF7544AB20FA1A7FB7FF9F1" box="[1059,1203,1533,1558]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB20FA1A7FB7FF9F1" box="[1059,1203,1533,1558]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
and
<emphasis id="62FA5847FFF7544AB2C1A1A7FA5DF9F1" box="[1261,1425,1533,1558]" italics="true" pageId="18" pageNumber="47">P. h a w a i i e n s i s</emphasis>
. In other species there is no information, although
<bibRefCitation id="341FF9A4FFF7544AB4C2A278FC11F9DB" author="Strandtmann" box="[750,989,1570,1596]" pageId="18" pageNumber="47" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann (1971)</bibRefCitation>
noted the occurrence of four pairs of apical setae on all tarsi (i.e. those lying close the apotele) as well as the occurrence of a single dorsal seta in the proximal part of the article as a characteristic feature of
<taxonomicName id="978EFFD6FFF7544AB57FA237FC28F961" box="[851,996,1645,1670]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF7544AB57FA237FC28F961" box="[851,996,1645,1670]" italics="true" pageId="18" pageNumber="47">Penthalodes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="50318455FFF7544BB6E9A2C8FA90FE8B" blockId="18.[151,1437,152,2007]" lastBlockId="19.[151,1437,152,1005]" lastPageId="19" lastPageNumber="48" pageId="18" pageNumber="47">
<emphasis id="62FA5847FFF7544AB6E9A2C8FE90F94B" bold="true" box="[197,348,1682,1708]" italics="true" pageId="18" pageNumber="47">Aspidosoma</emphasis>
. The aspidosoma is not separated from the rest of the body because the transverse &quot;das&quot; furrow is absent. In other eupodoids it usually separates the prodorsum from the opisthodorsum. The usually characteristic prodorsal plate of eupodoids is also absent. The ornamentation of the prodorsum is the same as the opisthodorsum. On the sides of prodorsum, more or less behind setae
<emphasis id="62FA5847FFF7544AB5D4A359FBD8F8FB" box="[1016,1044,1795,1820]" italics="true" pageId="18" pageNumber="47">xa</emphasis>
, the V-shaped furrows begin. All
<taxonomicName id="978EFFD6FFF7544AB6BBA372FEE0F8A6" box="[151,300,1832,1857]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF7544AB6BBA372FEE0F8A6" box="[151,300,1832,1857]" italics="true" pageId="18" pageNumber="47">Penthalodes</emphasis>
</taxonomicName>
have a prodorsum with an anterior, steep wall of the aspidosoma, i.e. in fact of the idiosoma. Therefore, the naso lies frontally, more or less half-way up the anterior wall of the idiosoma. Setal pairs
<emphasis id="62FA5847FFF7544AB379A317FAA6F881" box="[1365,1386,1869,1894]" italics="true" pageId="18" pageNumber="47">le</emphasis>
and
<emphasis id="62FA5847FFF7544AB6BBA329FF78F86B" box="[151,180,1907,1932]" italics="true" pageId="18" pageNumber="47">xa</emphasis>
are also situated frontally. Setae
<emphasis id="62FA5847FFF7544AB46BA329FD91F86B" box="[583,605,1907,1932]" italics="true" pageId="18" pageNumber="47">le</emphasis>
are well removed from setae
<emphasis id="62FA5847FFF7544AB5EBA329FC29F86B" box="[967,997,1907,1932]" italics="true" pageId="18" pageNumber="47">bo</emphasis>
, while setae
<emphasis id="62FA5847FFF7544AB2A9A329FB6EF86B" box="[1157,1186,1907,1932]" italics="true" pageId="18" pageNumber="47">xa</emphasis>
lie laterally, slightly behind the line of setae
<emphasis id="62FA5847FFF7544AB781A3C2FE0EF856" box="[429,450,1944,1969]" italics="true" pageId="18" pageNumber="47">le</emphasis>
. Prodorsal setae
<emphasis id="62FA5847FFF7544AB4A7A3C2FD64F856" box="[651,680,1944,1969]" italics="true" pageId="18" pageNumber="47">bo</emphasis>
lie as far back as between furrows “V”, in deep bothridia, rather dorsally. In
<taxonomicName id="978EFFD6FFF7544AB709A3E7FE74F831" box="[293,440,1981,2006]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="18" pageNumber="47" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF7544AB709A3E7FE74F831" box="[293,440,1981,2006]" italics="true" pageId="18" pageNumber="47">P. polonicus</emphasis>
</taxonomicName>
, they are filiform (sensu
<bibRefCitation id="341FF9A4FFF7544AB4C4A3E7FC08F830" author="Jesionowska" box="[744,964,1981,2007]" pageId="18" pageNumber="47" refString="Jesionowska, K. (2002) Morfologia szczecin niektorych przedstawicieli roztoczy z rodziny Eupodidae (Actinotrichida, Actinedida, Eupodoidea) z Polski. [Setal morphology in some representatives of the family Eupodidae]. Zeszyty Naukowe Uniwersytetu Szczeci n skiego Nr 348. Acta Biologica, 9, 31 - 45 (In Polish)." type="journal article" year="2002">Jesionowska 2002</bibRefCitation>
) in their distal part, while the proximal part is covered with tiny hairs. It is similar in
<taxonomicName id="978EFFD6FFF6544BB483A4C2FC9BFF56" box="[687,855,152,177]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF6544BB483A4C2FC9BFF56" box="[687,855,152,177]" italics="true" pageId="19" pageNumber="48">P. hawaiiensis</emphasis>
</taxonomicName>
except they are weakly plumose in the distal part because they have more thickly placed outgrowths (pers. observations). The drawing from Strandtmann, however, seems to show they are filiform. In
<emphasis id="62FA5847FFF6544BB486A4B8FCC6FF1C" box="[682,778,226,251]" italics="true" pageId="19" pageNumber="48">P. o v a l i s</emphasis>
sensu Kaluz, they are spiculate distally. In
<emphasis id="62FA5847FFF6544BB325A4B8FA5DFF1C" box="[1289,1425,226,251]" italics="true" pageId="19" pageNumber="48">P. c a r i n a t u s</emphasis>
, they are apically pilose, like in the species described by
<bibRefCitation id="341FF9A4FFF6544BB511A552FC2CFEC5" author="Baker" box="[829,992,264,290]" pageId="19" pageNumber="48" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFF6544BB511A552FC48FEC5" box="[829,900,264,290]" name="United States Pacific Island Wildlife Refuges" pageId="19" pageNumber="48">Baker</collectingCountry>
(1946)
</bibRefCitation>
. In
<emphasis id="62FA5847FFF6544BB221A552FBECFEC6" box="[1037,1056,264,289]" italics="true" pageId="19" pageNumber="48">P.</emphasis>
sp. presented by
<bibRefCitation id="341FF9A4FFF6544BB2D9A552FA5BFEC5" author="Baker" box="[1269,1431,264,290]" pageId="19" pageNumber="48" refString="Baker, A. S. (1987) Caleupodes, a new genus of eupodoid mite (Acari: Acariformes) showing primary opisthosomal segmentation. Bulletin of the British Museum (Natural History) (Zoology Series), Miscellanea, 53 (2) 103 - 113. Baker, A. S. (1990) A survey of external morphology of mites of the superfamily Eupodoidea Banks, 1894 (Acari: Acariformes). Journal of Natural History, 24, 1227 - 1261." type="journal article" year="1987">
<collectingCountry id="2899C4C5FFF6544BB2D9A552FAF0FEC5" box="[1269,1340,264,290]" name="United States Pacific Island Wildlife Refuges" pageId="19" pageNumber="48">Baker</collectingCountry>
(1987)
</bibRefCitation>
, setae
<emphasis id="62FA5847FFF6544BB6F6A577FF34FEA1" box="[218,248,301,326]" italics="true" pageId="19" pageNumber="48">bo</emphasis>
are strikingly short and rather filiform on their whole length. Therefore, most probably, setae
<emphasis id="62FA5847FFF6544BB373A577FAB0FEA1" box="[1375,1404,301,326]" italics="true" pageId="19" pageNumber="48">bo</emphasis>
in representatives of the genus
<taxonomicName id="978EFFD6FFF6544BB7C8A508FDB9FE8C" box="[484,629,338,363]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF6544BB7C8A508FDB9FE8C" box="[484,629,338,363]" italics="true" pageId="19" pageNumber="48">Penthalodes</emphasis>
</taxonomicName>
are weakly plumose in their distal part (or spiculate, or pilose).
</paragraph>
<paragraph id="50318455FFF6544BB6E9A522FCA8FE3B" blockId="19.[151,1437,152,1005]" pageId="19" pageNumber="48">
<emphasis id="62FA5847FFF6544BB6E9A522FF32FE76" box="[197,254,376,401]" italics="true" pageId="19" pageNumber="48">Eyes</emphasis>
. Lentiform eyes have been observed in most representatives of
<taxonomicName id="978EFFD6FFF6544BB23CA522FB6AFE76" box="[1040,1190,376,401]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF6544BB23CA522FB6AFE76" box="[1040,1190,376,401]" italics="true" pageId="19" pageNumber="48">Penthalodes</emphasis>
</taxonomicName>
. On the photograph published by
<collectingCountry id="2899C4C5FFF6544BB713A5C7FE44FE50" box="[319,392,413,439]" name="United States Pacific Island Wildlife Refuges" pageId="19" pageNumber="48">Baker</collectingCountry>
(1990), fine striation of their surface is seen. The eyes are situated behind setae
<emphasis id="62FA5847FFF6544BB354A5C7FA59FE51" box="[1400,1429,413,438]" italics="true" pageId="19" pageNumber="48">xa</emphasis>
, laterally, more or less at the level of where furrows “V” start.
</paragraph>
<paragraph id="50318455FFF6544BB6E9A5BDFEB3FC0A" blockId="19.[151,1437,152,1005]" pageId="19" pageNumber="48">
<emphasis id="62FA5847FFF6544BB6E9A5BDFEACFDE6" bold="true" box="[197,352,487,513]" italics="true" pageId="19" pageNumber="48">Gnathosoma</emphasis>
. The lack of a cheliceral seta is most probably characteristic for
<taxonomicName id="978EFFD6FFF6544BB24EA5B2FB3FFDE6" box="[1122,1267,488,513]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="62FA5847FFF6544BB24EA5B2FB3FFDE6" box="[1122,1267,488,513]" italics="true" pageId="19" pageNumber="48">Penthalodes</emphasis>
</taxonomicName>
. The shape of the profile of the pedipalpal tarsus differs in some species. The tarsus can be elongated (
<taxonomicName id="978EFFD6FFF6544BB2E0A657FABAFDC1" box="[1228,1398,525,550]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="alaskaensis">
<emphasis id="62FA5847FFF6544BB2E0A657FABAFDC1" box="[1228,1398,525,550]" italics="true" pageId="19" pageNumber="48">P. alaskaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="978EFFD6FFF6544BB3AAA657FF2EFDAC" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF6544BB3AAA657FF2EFDAC" italics="true" pageId="19" pageNumber="48">P. ovalis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFF6544BB70CA668FE75FDAC" box="[288,441,562,587]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF6544BB70CA668FE75FDAC" box="[288,441,562,587]" italics="true" pageId="19" pageNumber="48">P. polonicus</emphasis>
</taxonomicName>
) or shorter, similar to an oval (
<emphasis id="62FA5847FFF6544BB564A668FC17FDAC" box="[840,987,562,587]" italics="true" pageId="19" pageNumber="48">P. c a r i n a t u s</emphasis>
). In
<emphasis id="62FA5847FFF6544BB231A668FABDFDAC" box="[1053,1393,562,587]" italics="true" pageId="19" pageNumber="48">P. a l a s k a e n s i s, P. c a r i n a t u s</emphasis>
,
<taxonomicName id="978EFFD6FFF6544BB3A9A668FEE7FD96" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF6544BB3A9A668FEE7FD96" italics="true" pageId="19" pageNumber="48">P. hawaiiensis</emphasis>
</taxonomicName>
,
<taxonomicName id="978EFFD6FFF6544BB710A602FE64FD96" box="[316,424,600,625]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF6544BB710A602FE64FD96" box="[316,424,600,625]" italics="true" pageId="19" pageNumber="48">P. ovalis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFF6544BB7CBA602FD4EFD96" box="[487,642,600,625]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF6544BB7CBA602FD4EFD96" box="[487,642,600,625]" italics="true" pageId="19" pageNumber="48">P. polonicus</emphasis>
</taxonomicName>
a small rod-like solenidion is recumbent in a depression. In
<taxonomicName id="978EFFD6FFF6544BB3A9A602FEEFFD71" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF6544BB3A9A602FEEFFD71" italics="true" pageId="19" pageNumber="48">P. hawaiiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="978EFFD6FFF6544BB772A627FE23FD71" box="[350,495,637,662]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF6544BB772A627FE23FD71" box="[350,495,637,662]" italics="true" pageId="19" pageNumber="48">P. polonicus</emphasis>
</taxonomicName>
, the tip of the three apical setae each on the pedipalpal tarsus is characteristic; they bear strongly sclerotized shuttles (carinas;
<figureCitation id="C8B598D0FFF6544BB4FFA6F8FCD6FD5B" box="[723,794,674,700]" captionStart="FIGURE 4" captionStartId="8.[151,255,1869,1893]" captionTargetBox="[202,1390,194,1846]" captionTargetId="figure@8.[188,1398,194,1847]" captionTargetPageId="8" captionText="FIGURE 4. Penthalodes polonicus n. sp. (A) Subcapitulum, ventral view; (B) Subcapitulum, dorsal view; LB-labrum, LL-lateral lips, LI-labium, H-hypostome, sc-internal sclerites, n, m-subcapitular setae, cxp-coxal region of pedipalp. (C) Pedipalp, lateral view; (D) Pedipalpal tarsus in detail, lateral view." httpUri="https://zenodo.org/record/199165/files/figure.png" pageId="19" pageNumber="48">Fig. 4</figureCitation>
CD). According to
<bibRefCitation id="341FF9A4FFF6544BB22DA6F8FB6FFD5B" author="Kaluz" box="[1025,1187,674,700]" pageId="19" pageNumber="48" refString="Kaluz, S. (2000) Redescription of Penthalodes ovalis (Acarina, Prostigmata, Penthalodidae) based on mites from Central Europe and Turkey. Biologia, Bratislava, 55 (5), 477 - 482." type="journal article" year="2000">Kaluz (2000)</bibRefCitation>
, two apical setae are solenidia, while
<bibRefCitation id="341FF9A4FFF6544BB74FA692FD95FD05" author="Strandtmann" box="[355,601,712,738]" pageId="19" pageNumber="48" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann (1971)</bibRefCitation>
and
<bibRefCitation id="341FF9A4FFF6544BB4BBA692FCF6FD05" author="Baker" box="[663,826,712,738]" pageId="19" pageNumber="48" refString="Baker, E. W. (1946) New species of North and Central American mites of the family Penthaleidae (Acarina). Journal of the Washington Academy of Sciences, 36 (12), 421 - 425." type="journal article" year="1946">
<collectingCountry id="2899C4C5FFF6544BB4BBA692FD2CFD05" box="[663,736,712,738]" name="United States Pacific Island Wildlife Refuges" pageId="19" pageNumber="48">Baker</collectingCountry>
(1946)
</bibRefCitation>
stated only that they are nude. For other species, sufficient descriptions and drawings are lacking. On the subcapitulum, a pair of setae
<emphasis id="62FA5847FFF6544BB252A6B7FB41FCE1" box="[1150,1165,749,774]" italics="true" pageId="19" pageNumber="48">n</emphasis>
(
<emphasis id="62FA5847FFF6544BB2B2A6B7FB09FCE1" box="[1182,1221,749,774]" italics="true" pageId="19" pageNumber="48">sbc</emphasis>
2) lies more or less at the level of the labium, while setae
<emphasis id="62FA5847FFF6544BB475A74CFDA2FCC8" box="[601,622,790,815]" italics="true" pageId="19" pageNumber="48">m</emphasis>
(
<emphasis id="62FA5847FFF6544BB4ACA74CFD6BFCC8" box="[640,679,790,815]" italics="true" pageId="19" pageNumber="48">sbc</emphasis>
1) lies laterally, antiaxially, at the base of the lateral lips fused in that part of subcapitulum with the hypostome. In
<taxonomicName id="978EFFD6FFF6544BB4C5A764FC5FFCB0" box="[745,915,830,855]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="hawaiiensis">
<emphasis id="62FA5847FFF6544BB4C5A764FC5FFCB0" box="[745,915,830,855]" italics="true" pageId="19" pageNumber="48">P. hawaiiensis</emphasis>
</taxonomicName>
, they are rather plumose, in
<taxonomicName id="978EFFD6FFF6544BB2C0A764FF2EFC9A" authority="Kaluz" authorityName="Kaluz" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="62FA5847FFF6544BB2C0A764FA9EFCB0" box="[1260,1362,830,855]" italics="true" pageId="19" pageNumber="48">P. ovalis</emphasis>
sensu Kaluz
</taxonomicName>
pilose, while in
<taxonomicName id="978EFFD6FFF6544BB794A739FD9FFC9B" box="[440,595,867,892]" class="Arachnida" family="Penthalodidae" genus="Penthalodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="19" pageNumber="48" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="62FA5847FFF6544BB794A739FD9FFC9B" box="[440,595,867,892]" italics="true" pageId="19" pageNumber="48">P. polonicus</emphasis>
</taxonomicName>
they are weakly plumose with rather long lateral outgrowths. The hypostome, which forms ventral part of subcapitulum, is composed of two parts, a basal rectangular and a distal triangular part. Usually, the inner sclerite is visible that separates these parts (
<figureCitation id="C8B598D0FFF6544BB281A7F7FB34FC20" box="[1197,1272,941,967]" captionStart="FIGURE 4" captionStartId="8.[151,255,1869,1893]" captionTargetBox="[202,1390,194,1846]" captionTargetId="figure@8.[188,1398,194,1847]" captionTargetPageId="8" captionText="FIGURE 4. Penthalodes polonicus n. sp. (A) Subcapitulum, ventral view; (B) Subcapitulum, dorsal view; LB-labrum, LL-lateral lips, LI-labium, H-hypostome, sc-internal sclerites, n, m-subcapitular setae, cxp-coxal region of pedipalp. (C) Pedipalp, lateral view; (D) Pedipalpal tarsus in detail, lateral view." httpUri="https://zenodo.org/record/199165/files/figure.png" pageId="19" pageNumber="48">Fig. 4</figureCitation>
A; Fig. 5B in
<bibRefCitation id="341FF9A4FFF6544BB6BBA789FEBEFC0A" author="Strandtmann" box="[151,370,979,1005]" pageId="19" pageNumber="48" refString="Strandtmann, R. W. (1971) The eupodid mites of Alaska. Pacific Insects, 13 (1), 75 - 118." type="journal article" year="1971">Strandtmann 1971</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>