174 lines
20 KiB
XML
174 lines
20 KiB
XML
<document id="9D604DCBBB619569F6958EACF11F4378" ID-DOI="10.1206/0003-0090-408.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5380942" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630291433617" checkinUser="carolina" docAuthor="Grimaldi, David A." docDate="2016" docId="CF1987FEE964ED5740A5FA80CAC77615" docLanguage="en" docName="B408.pdf" docOrigin="Bulletin of the American Museum of Natural History 2016 (408)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-408.1.1" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Cretasolva burmitica Grimaldi 2016, new species" docType="treatment" docVersion="3" lastPageNumber="14" masterDocId="3320FF86E968ED594067FFE6CB5D7470" masterDocTitle="Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi" masterLastPageNumber="132" masterPageNumber="1" pageNumber="12" updateTime="1699216921357" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="3FBB192996FBE0C1C66266AFD397DEE1">
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<mods:title id="A57D371FC9F25EB0078DB28983FF61CF">Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi</mods:title>
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<mods:name id="E192D9647B430BB4D51F46D9C8D19BBA" type="personal">
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<mods:roleTerm id="D07E6D6C0B55C35561946740A926C016">Author</mods:roleTerm>
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<mods:namePart id="421C234BB76D8679D549AD9623134E43">Grimaldi, David A.</mods:namePart>
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<mods:typeOfResource id="55E85A2872EAADC5ACD7A3C2022BB70F">text</mods:typeOfResource>
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<mods:title id="D9ECD5E1604F297E8D38C561D7EC3640">Bulletin of the American Museum of Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="B689E5023173360536BED918775973B2">
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<mods:date id="7CC0BCD9AD4E033F291EF5609D324217">2016</mods:date>
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<mods:number id="C128AAD7231F0720A5C8ADE0858CAA1D">2016-09-28</mods:number>
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<mods:detail id="7851BDC9AFF3BDFBB3866A4086314C6D" type="volume">
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<mods:number id="8F5B7C312295D6B065FE1B2DE42BE955">2016</mods:number>
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<mods:number id="56927F38793134BDF091D04688628733">408</mods:number>
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<mods:start id="72B874F1F45FA5E0AF352DBBA65838F9">1</mods:start>
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<mods:url id="995B1DCA394DF5E2E662BCC28DF9036F">http://www.bioone.org/doi/10.1206/0003-0090-408.1.1</mods:url>
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<mods:classification id="792151A20C150EA109DF3A17DA1FF4CB">journal article</mods:classification>
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<mods:identifier id="D0B37321993C463121D666CF0154FDAE" type="DOI">10.1206/0003-0090-408.1.1</mods:identifier>
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<mods:identifier id="7DFA09051A07DE65B717C1C3EDFCF339" type="ISSN">0003-0090</mods:identifier>
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<mods:identifier id="8455C3D20A8F98690008FD3D384EE089" type="Zenodo-Dep">5380942</mods:identifier>
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<treatment id="CF1987FEE964ED5740A5FA80CAC77615" LSID="urn:lsid:plazi:treatment:CF1987FEE964ED5740A5FA80CAC77615" httpUri="http://treatment.plazi.org/id/CF1987FEE964ED5740A5FA80CAC77615" lastPageId="14" lastPageNumber="14" pageId="12" pageNumber="12">
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<subSubSection id="0FAA6563E964ED5540A5FA80C960710F" box="[194,573,1382,1407]" pageId="12" pageNumber="12" type="nomenclature">
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<paragraph id="470F36E8E964ED5540A5FA80C960710F" blockId="12.[194,573,1382,1407]" box="[194,573,1382,1407]" pageId="12" pageNumber="12">
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<heading id="1C478184E964ED5540A5FA80C960710F" box="[194,573,1382,1407]" centered="true" fontSize="9" level="2" pageId="12" pageNumber="12" reason="2">
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<taxonomicName id="80B04D6BE964ED5540A5FA80CAF2710E" ID-CoL="8BLRW" authority="Grimaldi, 2016" authorityName="Grimaldi" authorityYear="2016" box="[194,431,1382,1406]" class="Insecta" family="Xylomyidae" genus="Cretasolva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="12" phylum="Arthropoda" rank="species" species="burmitica" status="sp. nov.">
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<emphasis id="75C4EAFAE964ED5540A5FA80CAF2710E" bold="true" box="[194,431,1382,1406]" italics="true" pageId="12" pageNumber="12">Cretasolva burmitica</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="6EF75781E964ED5541DAFA81C960710F" box="[445,573,1383,1407]" pageId="12" pageNumber="12" rank="species">new species</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FAA6563E964ED55415DFA7ECA9B71C1" box="[314,454,1432,1457]" pageId="12" pageNumber="12" type="description">
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<paragraph id="470F36E8E964ED55415DFA7ECA9B71C1" blockId="12.[314,454,1432,1457]" box="[314,454,1432,1457]" pageId="12" pageNumber="12">
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<heading id="1C478184E964ED55415DFA7ECA9B71C1" box="[314,454,1432,1457]" centered="true" fontSize="9" level="2" pageId="12" pageNumber="12" reason="2">
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<figureCitation id="DF8B2A6DE964ED55415DFA7ECAC271C1" box="[314,415,1432,1457]" captionStart="FIG" captionStartId="13.[120,160,1202,1223]" captionTargetBox="[222,1138,227,1176]" captionTargetId="figure-243@13.[120,1224,224,1176]" captionTargetPageId="13" captionText="FIG. 3. Cretasolva burmitica, new genus and species (Xylomyidae), AMNH Bu-SE05, holotype. A. Wing. B, C. Apices of meso- and metatibia, respectively, showing spurs. D. Head, lateral view, with detail of antennal apex." figureDoi="http://doi.org/10.5281/zenodo.5380948" httpUri="https://zenodo.org/record/5380948/files/figure.png" pageId="12" pageNumber="12">Figures 3</figureCitation>
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,
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<figureCitation id="DF8B2A6DE964ED5541CBFA7FCA9B71C1" box="[428,454,1433,1457]" captionStart="FIG" captionStartId="94.[120,162,1485,1506]" captionTargetBox="[120,1224,224,1458]" captionTargetId="figure-27@94.[120,1224,224,1459]" captionTargetPageId="94" captionText="FIG. 34. Cretosolva burmitica, new genus and species, holotype AMNH Bu-SE005 A. Right habitus. B. Right wing." figureDoi="http://doi.org/10.5281/zenodo.5381013" httpUri="https://zenodo.org/record/5381013/files/figure.png" pageId="12" pageNumber="12">34</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0FAA6563E964ED5540FFFA2CC9177193" box="[152,586,1482,1507]" pageId="12" pageNumber="12" type="diagnosis">
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<paragraph id="470F36E8E964ED5540FFFA2CC9177193" blockId="12.[120,648,1482,1738]" box="[152,586,1482,1507]" pageId="12" pageNumber="12">DIAGNOSIS: As for genus, by monotypy.</paragraph>
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</subSubSection>
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<subSubSection id="0FAA6563E964ED5440FFFA0DC8E67235" lastPageId="13" lastPageNumber="13" pageId="12" pageNumber="12" type="description">
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<paragraph id="470F36E8E964ED5440FFFA0DC8E67235" blockId="12.[120,648,1482,1738]" lastBlockId="13.[696,1225,1283,1737]" lastPageId="13" lastPageNumber="13" pageId="12" pageNumber="12">
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DESCRIPTION: Based on unique female. Slen- der bodied with slender wings, legs of moderate length. Body length (excluding antennae, including cerci)
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<quantity id="80489B0DE964ED55408EF9A8CA6A7216" box="[233,311,1614,1638]" metricMagnitude="0" metricUnit="m" metricValue="4.69" pageId="12" pageNumber="12" unit="m" value="4.69">4.69 m</quantity>
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; thorax length
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<quantity id="80489B0DE964ED554186F9A8C91B7216" box="[481,582,1614,1638]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="12" pageNumber="12" unit="mm" value="1.66">1.66 mm</quantity>
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; wing length
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<quantity id="80489B0DE964ED5540ABF989CA6A72F7" box="[204,311,1647,1671]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="12" pageNumber="12" unit="mm" value="3.6">3.60 mm</quantity>
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; abdomen length (including cerci)
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<quantity id="80489B0DE964ED5540DCF976CA4072D8" box="[187,285,1680,1704]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="12" unit="mm" value="2.5">2.50 mm</quantity>
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.
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<emphasis id="75C4EAFAE964ED554140F977CA3772D9" bold="true" box="[295,362,1681,1705]" pageId="12" pageNumber="12">Head:</emphasis>
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Slightly broader than thorax, occiput slightly concave. Frons relatively broad, width slightly greater than distance between outside margins of lateral ocelli. Face concave (clypeus not observable). Eye: Relatively large, occupying most of head surface in lateral, frontal, and dorsal views. Margins with frons parallel. Eye completely bare, facets not differentiated. Ocelli slightly raised above surface of frons.
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<emphasis id="75C4EAFAE964ED554299FE2FC8357591" bold="true" box="[766,872,457,481]" pageId="12" pageNumber="12">Antenna:</emphasis>
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Submoniliform, with 8 flagellomeres evenly tapered from broad basal one to narrow apical flagellomere, apical one with minute style. Flagellomere lengths: 8> 7> 6 = 1> 2–5. Scape and pedicel small; antennal sockets very close, situated in ventral half of frons.
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<emphasis id="75C4EAFAE964ED5542DFFD76C81676D8" bold="true" box="[696,843,656,680]" pageId="12" pageNumber="12">Mouthparts:</emphasis>
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Palps directed forward; 2-segmented, basal palpomere slender; apical one oval, nearly twice width of basal palpomere. Proboscis short, only labellum exposed; labellum large, fleshy, laterally flattened.
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<emphasis id="75C4EAFAE964ED554477FCF2CF35775C" bold="true" box="[1040,1128,788,812]" pageId="12" pageNumber="12">Thorax:</emphasis>
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Dorsum of scutum, scutellum, portion of anepisternum with dense, very fine, setigerous punctation, denser on scutellum; no macrosetae or long pilosity. Transverse suture short, mostly lateral (pleural), with faint impression dorsolaterally. Scutellum narrow in dorsal view, longer than wide, without spines; small, short but distinct proscutellum present. Pleural sutures not easily observed in unique specimen; pleura large oblique (vs. vertical).
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<emphasis id="75C4EAFAE964ED5543CEFBB8C8B37006" bold="true" box="[937,1006,1118,1142]" pageId="12" pageNumber="12">Wing:</emphasis>
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Long, slender, W/L 0.30, veins well developed, most are well sclerotized, lightest ones are R
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<subScript id="DB3434ADE964ED5543A5FB4AC88070CD" attach="left" box="[962,989,1196,1213]" fontSize="7" pageId="12" pageNumber="12">2+3</subScript>
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and base of M (latter faint, not sclerotized). Minute setulae on R veins (except R
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<subScript id="DB3434ADE964ED554341FB08C81F708F" attach="left" box="[806,834,1262,1279]" fontSize="7" pageId="12" pageNumber="12">2+3</subScript>
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) and M veins (except for stem), none on Cu veins. Vein C ends at apex of M
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<subScript id="DB3434ADE964ED5544F5FAE9CFC67150" attach="right" box="[1170,1179,1295,1312]" fontSize="7" pageId="12" pageNumber="12">1</subScript>
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. No pterostigma. Sc complete, ca. 0.5× wing length; R
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<subScript id="DB3434ADE964ED5542AEFAB7C98F7112" attach="left" box="[713,722,1361,1378]" fontSize="7" pageId="12" pageNumber="12">1</subScript>
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0.55× wing length, close and parallel to Sc; Rs and R
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<subScript id="DB3434ADE964ED55429DFA94C84B71F3" attach="left" box="[762,790,1394,1411]" fontSize="7" pageId="12" pageNumber="12">2+3</subScript>
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short, parallel to R
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<subScript id="DB3434ADE964ED554388FA94C8A571F3" attach="left" box="[1007,1016,1394,1411]" fontSize="7" pageId="12" pageNumber="12">1</subScript>
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; Rs without thickened node or weak area; fork of R
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<subScript id="DB3434ADE964ED55444CFA72CF6971D5" attach="left" box="[1067,1076,1428,1445]" fontSize="7" pageId="12" pageNumber="12">4</subScript>
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-R
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<subScript id="DB3434ADE964ED554429FA75CF0A71D4" attach="left" box="[1102,1111,1427,1444]" fontSize="7" pageId="12" pageNumber="12">5</subScript>
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asymmetrical, R
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<subScript id="DB3434ADE964ED554360FA53C84D71B6" attach="left" box="[775,784,1461,1478]" fontSize="7" pageId="12" pageNumber="12">4</subScript>
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0.66× length of R
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<subScript id="DB3434ADE964ED5543B0FA52C8BD71B5" attach="left" box="[983,992,1460,1477]" fontSize="7" pageId="12" pageNumber="12">5</subScript>
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. Crossvein r-m close to base of cell d, M veins connected to apex of cell d; M
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<subScript id="DB3434ADE964ED554372FA11C8437278" attach="right" box="[789,798,1527,1544]" fontSize="7" pageId="12" pageNumber="12">1</subScript>
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, M
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<subScript id="DB3434ADE964ED554326FA11C8177278" attach="left" box="[833,842,1527,1544]" fontSize="7" pageId="12" pageNumber="12">2</subScript>
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parallel; cell m long, slender, nearly reaching to wing margin (M
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<subScript id="DB3434ADE964ED554399F9FECF5A7259" attach="left" box="[1022,1031,1560,1577]" fontSize="7" pageId="12" pageNumber="12">3</subScript>
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+CuA
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<subScript id="DB3434ADE964ED55442DF9FECF0E7259" attach="left" box="[1098,1107,1560,1577]" fontSize="7" pageId="12" pageNumber="12">1</subScript>
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spur very short); crossvein m-cu present. Cell d long and narrow, W/L 0.3; cells br and bm of equal length. Vein A
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<subScript id="DB3434ADE964ED554364F99DC85172FC" attach="left" box="[771,780,1659,1676]" fontSize="7" pageId="12" pageNumber="12">1</subScript>
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very long, apex reaching to midlength of wing, cell cup closed. Vein A
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<subScript id="DB3434ADE964ED554462F97ACF5372DD" attach="left" box="[1029,1038,1692,1709]" fontSize="7" pageId="12" pageNumber="12">2</subScript>
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short; anal lobe and alula present but small.
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<emphasis id="75C4EAFAE964ED55446DF954CF1B72BA" bold="true" box="[1034,1094,1714,1738]" pageId="12" pageNumber="12">Legs:</emphasis>
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Robust, of moderate length. Coxae robust, meso- and metacoxae meeting, distant from procoxa. Relative lengths of leg segments: Pro- and mesofemur> tibiae> basitarsomere> combined lengths tarsomeres 2–5. Metatibia> femur> basitarsomere> combined lengths tarsomeres 2–5. Tibial spurs 0-2-2; spurs long and thin, lengths ca. 1.5× apical width of tibiae. No tubercles on ventral surface of hind femur. Pretarsus with well-developed simple claws; pulvilli small, ca 0.5× length of claw; empodium pulvilliform, short and broad.
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<emphasis id="75C4EAFAE965ED54401FF989CBAF72F7" bold="true" box="[120,242,1647,1671]" pageId="13" pageNumber="13">Abdomen:</emphasis>
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Only slightly longer than thorax, thickness and width tapered to narrow apex. Seven tergites visible, basal 2 tergites fully sclero- tized (no median membranous areas); tergite 1 narrow in dorsal view, width of its posterior margin ca. 0.5× that of tergite 2; tergite 2 large, nearly twice the length of tergite 3; broadest tergites t2, t3. Cerci (female) long, slender, 2-segmented, length approximately equal to length of segments 6+7, basal cercomere slightly longer than apical one; t10 small, lying between bases. Tergites 2–4 with shallow transverse sulci (possibly on t1 and t5 but not fully visible).
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</paragraph>
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</subSubSection>
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<caption id="13CF6660E965ED54401FFB54CF997095" ID-DOI="http://doi.org/10.5281/zenodo.5380948" ID-Zenodo-Dep="5380948" httpUri="https://zenodo.org/record/5380948/files/figure.png" pageId="13" pageNumber="13" startId="13.[120,160,1202,1223]" targetBox="[222,1138,227,1176]" targetPageId="13">
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<paragraph id="470F36E8E965ED54401FFB54CF997095" blockId="13.[120,1225,1200,1254]" pageId="13" pageNumber="13">
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FIG. 3.
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<taxonomicName id="80B04D6BE965ED5440A5FB56CAD370B8" authority="Grimaldi, 2016" authorityName="Grimaldi" authorityYear="2016" box="[194,398,1200,1224]" class="Insecta" family="Xylomyidae" genus="Cretasolva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="species" species="burmitica" status="new genus and species">
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<emphasis id="75C4EAFAE965ED5440A5FB56CAD370B8" box="[194,398,1200,1224]" italics="true" pageId="13" pageNumber="13">Cretasolva burmitica</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="6EF75781E965ED5441FCFB57C92370B8" box="[411,638,1200,1225]" pageId="13" pageNumber="13" rank="species">new genus and species</taxonomicNameLabel>
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(
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<taxonomicName id="80B04D6BE965ED5442ECFB56C85A70B8" box="[651,775,1200,1224]" class="Insecta" family="Xylomyidae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="13" phylum="Arthropoda" rank="family">Xylomyidae</taxonomicName>
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), AMNH Bu-SE05, holotype.
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<emphasis id="75C4EAFAE965ED544454FB56CF1670B8" bold="true" box="[1075,1099,1200,1224]" pageId="13" pageNumber="13">A.</emphasis>
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Wing.
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<emphasis id="75C4EAFAE965ED5444F2FB56CF9470B8" bold="true" box="[1173,1225,1200,1224]" pageId="13" pageNumber="13">B, C.</emphasis>
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Apices of meso- and metatibia, respectively, showing spurs.
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<emphasis id="75C4EAFAE965ED5442AAFB28C9BB7096" bold="true" box="[717,742,1230,1254]" pageId="13" pageNumber="13">D.</emphasis>
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Head, lateral view, with detail of antennal apex.
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</paragraph>
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</caption>
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<subSubSection id="0FAA6563E965ED5442BFF9A8CFF87216" box="[728,1189,1614,1638]" pageId="13" pageNumber="13" type="materials_examined">
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<paragraph id="470F36E8E965ED5442BFF9A8CFF87216" blockId="13.[696,1225,1283,1737]" box="[728,1189,1614,1638]" pageId="13" pageNumber="13">
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<materialsCitation id="F7D83CB5E965ED5442BFF9A8CFF87216" box="[728,1189,1614,1638]" collectionCode="TYPE, AMNH" pageId="13" pageNumber="13" specimenCount="1" typeStatus="holotype">
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<collectionCode id="21A1AE2DE965ED5442BFF9A8C84F7215" box="[728,786,1614,1637]" pageId="13" pageNumber="13">TYPE</collectionCode>
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:
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<typeStatus id="980B884AE965ED54437BF9A8C8DE7216" box="[796,899,1614,1638]" pageId="13" pageNumber="13" type="holotype">Holotype</typeStatus>
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, female,
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<collectionCode id="21A1AE2DE965ED544382F9A8CF647216" box="[997,1081,1614,1638]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="13" pageNumber="13">AMNH</collectionCode>
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Bu-SE05.
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</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FAA6563E965ED5442BFF989C87872B9" pageId="13" pageNumber="13" type="etymology">
|
||
<paragraph id="470F36E8E965ED5442BFF989C87872B9" blockId="13.[696,1225,1283,1737]" pageId="13" pageNumber="13">
|
||
ETYMOLOGY: Derived from English
|
||
<emphasis id="75C4EAFAE965ED54440AF989CF9572F7" box="[1133,1224,1647,1671]" italics="true" pageId="13" pageNumber="13">burmite,</emphasis>
|
||
another name for Burmese amber, and Latinized.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FAA6563E966ED5740FFFF04CAC77615" pageId="14" pageNumber="14" type="discussion">
|
||
<paragraph id="470F36E8E966ED5740FFFF04CAC77615" blockId="14.[120,650,226,613]" pageId="14" pageNumber="14">
|
||
COMMENTS: Specimen is in a small, irregular piece 4 × 5 ×
|
||
<quantity id="80489B0DE966ED574171FEE5CA06756A" box="[278,347,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="14" unit="mm" value="6.0">6 mm</quantity>
|
||
, with the sides closely surrounding the specimen, providing most views. The fly is complete, but it lies on a weathered internal flow; body of the fly (but not the wings) has a thin, reddish covering over the exocuticle, which, along with some fine fractures, obscures some details. This reddish layer indicates initial stages of pyritization. The section with the fly was trimmed from a cabochen that also contains two small moths,
|
||
<specimenCount id="51B6FD61E966ED57415FFDCBCAC07634" box="[312,413,556,580]" pageId="14" pageNumber="14" type="adult">one adult</specimenCount>
|
||
<taxonomicName id="80B04D6BE966ED5741C3FDCACA597615" authority="(Auchenorrhyncha)" baseAuthorityName="Auchenorrhyncha" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="14" phylum="Arthropoda" rank="family">Cicadellidae (Auchenorrhyncha)</taxonomicName>
|
||
, and a spider.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |