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<document id="5DA6615FC00C43DEE4BB6F92459FE884" ID-DOI="10.1206/0003-0090-408.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5380942" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630291433617" checkinUser="carolina" docAuthor="Grimaldi, David A." docDate="2016" docId="CF1987FEE925ED1640C4FD14CA4E76B8" docLanguage="en" docName="B408.pdf" docOrigin="Bulletin of the American Museum of Natural History 2016 (408)" docSource="http://www.bioone.org/doi/10.1206/0003-0090-408.1.1" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Burmapsilocephala evocoa Grimaldi 2016, new species" docType="treatment" docVersion="4" lastPageNumber="79" masterDocId="3320FF86E968ED594067FFE6CB5D7470" masterDocTitle="Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi" masterLastPageNumber="132" masterPageNumber="1" pageNumber="77" updateTime="1699216921357" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6EDFA4A0E3B84E765385C895EEB88BC2">Diverse Orthorrhaphan Flies (Insecta: Diptera: Brachycera) In Amber From The Cretaceous Of Myanmar: Brachycera In Cretaceous Amber, Part Vii David A. Grimaldi</mods:title>
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<subSubSection id="0FAA6563E925ED1440C4FD14C900777B" box="[163,605,754,779]" pageId="77" pageNumber="77" type="nomenclature">
<paragraph id="470F36E8E925ED1440C4FD14C900777B" blockId="77.[163,605,754,779]" box="[163,605,754,779]" pageId="77" pageNumber="77">
<heading id="1C478184E925ED1440C4FD14C900777B" box="[163,605,754,779]" centered="true" fontSize="9" level="2" pageId="77" pageNumber="77" reason="2">
<emphasis id="75C4EAFAE925ED1440C4FD14CA88777B" bold="true" box="[163,469,754,779]" pageId="77" pageNumber="77">
<taxonomicName id="80B04D6BE925ED1440C4FD14CA93777A" ID-CoL="8B9QB" authority="Grimaldi, 2016" authorityName="Grimaldi" authorityYear="2016" box="[163,462,754,778]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" pageNumber="77" phylum="Arthropoda" rank="species" species="evocoa" status="sp. nov.">
<emphasis id="75C4EAFAE925ED1440C4FD14CA93777A" bold="true" box="[163,462,754,778]" italics="true" pageId="77" pageNumber="77">Burmapsilocephala evocoa</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="6EF75781E925ED1441BBFD15C900777B" box="[476,605,755,779]" pageId="77" pageNumber="77" rank="species">new species</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FAA6563E925ED14417DFCC2CAB8774C" box="[282,485,804,828]" pageId="77" pageNumber="77" type="description">
<paragraph id="470F36E8E925ED14417DFCC2CAB8774C" blockId="77.[282,485,804,828]" box="[282,485,804,828]" pageId="77" pageNumber="77">
<heading id="1C478184E925ED14417DFCC2CAB8774C" box="[282,485,804,828]" centered="true" fontSize="9" level="2" pageId="77" pageNumber="77" reason="2">
<figureCitation id="DF8B2A6DE925ED14417DFCC2CA9D774C" box="[282,448,804,828]" captionStart="FIG" captionStartId="78.[120,162,1562,1583]" captionTargetBox="[135,1224,236,1532]" captionTargetId="figure-9@78.[120,1224,224,1536]" captionTargetPageId="78" captionText="FIG. 26. Burmapsilocephala spp. (Apsilocephalidae) in Burmese amber. AC. Burmapsilocephala evocoa, new species. D. B. cockerelli Gaimari and Mostovski (ex: Gaimari and Mostovski, 2000). A. Head, oblique frontal view. B. Male terminalia, oblique lateral view. C. Wing of B. evocoa (portion are not fully observable). Arrows indicate diagnostic venational differences. D. Wing of B. cockerelli. Panels A, B to same scale." figureDoi="http://doi.org/10.5281/zenodo.5380997" httpUri="https://zenodo.org/record/5380997/files/figure.png" pageId="77" pageNumber="77">Figures 26AC</figureCitation>
,
<figureCitation id="DF8B2A6DE925ED1441ACFCC2CAB8774C" box="[459,485,804,828]" captionStart="FIG" captionStartId="118.[120,162,1709,1730]" captionTargetBox="[216,1127,227,1682]" captionTargetId="figure-30@118.[120,1224,227,1683]" captionTargetPageId="118" captionText="FIG. 58. Burmapsilocephala evocoa, new species (Apsilocephalidae), holotype AMNH Bu-SE2/4. A. Habitus, left lateral. B. Head and thorax, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5381063" httpUri="https://zenodo.org/record/5381063/files/figure.png" pageId="77" pageNumber="77">58</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0FAA6563E925ED1440FFFCB0CA7070E7" pageId="77" pageNumber="77" type="diagnosis">
<paragraph id="470F36E8E925ED1440FFFCB0CA7070E7" blockId="77.[120,649,854,1738]" pageId="77" pageNumber="77">
DIAGNOSIS: Distinguished from
<taxonomicName id="80B04D6BE925ED144261FCB1C9D5771E" box="[518,648,854,878]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" pageNumber="77" phylum="Arthropoda" rank="species" species="cockerelli">
<emphasis id="75C4EAFAE925ED144261FCB1C9D5771E" box="[518,648,854,878]" italics="true" pageId="77" pageNumber="77">B. cockerelli</emphasis>
</taxonomicName>
(fig. 26D) also in Burmese amber, by possession of 14 thoracic macrosetae (vs. 12) as follows: 1 pair of notopleural setae (vs. 2), 2 pairs supraalars (vs. 1), 2 pairs postalars (vs. 1); notopleural setae distinctly shorter than in
<taxonomicName id="80B04D6BE925ED144199FC1DC93D7063" box="[510,608,1019,1043]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="77" pageNumber="77" phylum="Arthropoda" rank="species" species="cockerelli">
<emphasis id="75C4EAFAE925ED144199FC1DC93D7063" box="[510,608,1019,1043]" italics="true" pageId="77" pageNumber="77">cockerelli</emphasis>
</taxonomicName>
; R
<subScript id="DB3434ADE925ED144218FBE1C9D57068" attach="left" box="[639,648,1031,1048]" fontSize="7" pageId="77" pageNumber="77">5</subScript>
ending at wing tip (vs. posterior to it), crossvein r-m at middle of cell d (vs. in basal third); A
<subScript id="DB3434ADE925ED144218FBAFC9D5702A" attach="left" box="[639,648,1097,1114]" fontSize="7" pageId="77" pageNumber="77">2</subScript>
and anal lobe more developed (possibly sexually dimorphic).
</paragraph>
</subSubSection>
<subSubSection id="0FAA6563E925ED1640FFFB47C9487634" lastPageId="79" lastPageNumber="79" pageId="77" pageNumber="77" type="description">
<paragraph id="470F36E8E925ED1640FFFB47C9487634" blockId="77.[120,649,854,1738]" lastBlockId="79.[120,650,225,712]" lastPageId="79" lastPageNumber="79" pageId="77" pageNumber="77">
DESCRIPTION: Based on male. A slender, delicate, gracile fly with long, slender legs and abdomen. Body length
<quantity id="80489B0DE925ED14412EFB05CAF3708A" box="[329,430,1251,1275]" metricMagnitude="-3" metricUnit="m" metricValue="4.38" pageId="77" pageNumber="77" unit="mm" value="4.38">4.38 mm</quantity>
; thorax length 1.00 mm; abdomen length
<quantity id="80489B0DE925ED144108FAE2CA8F716B" box="[367,466,1284,1308]" metricMagnitude="-3" metricUnit="m" metricValue="2.87" pageId="77" pageNumber="77" unit="mm" value="2.87">2.87 mm</quantity>
; wing length ca.
<quantity id="80489B0DE925ED14401FFAC3CB84714C" box="[120,217,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="77" pageNumber="77" unit="mm" value="2.9">2.90 mm</quantity>
; hind leg length
<quantity id="80489B0DE925ED1441EBFAC3CAB3714C" box="[396,494,1317,1341]" metricMagnitude="-3" metricUnit="m" metricValue="4.57" pageId="77" pageNumber="77" unit="mm" value="4.57">4.57 mm</quantity>
.
<emphasis id="75C4EAFAE925ED14419EFAC3C966714D" bold="true" box="[505,571,1317,1341]" pageId="77" pageNumber="77">Head:</emphasis>
Dorsal surface flattened, broad. Eyes completely bare (no interfacetal setulae), holoptic in male, occupying most of head dorsally, frontally, laterally; medial margins of eyes meeting from ocellar triangle to just above bases of antennae, small portion of frons exposed; anteromedian margins of eyes slightly emarginate around bases of antennae; facets on dorsal surfaces ca. 2× diameter of ventral and lateralmost facets. Ocelli on small, low tubercle on posterodorsal margin of head capsule, with several stiff setulae in triangle. Face, including clypeus, deeply recessed, clypeus not visible.
<emphasis id="75C4EAFAE925ED1442DFFF07C87C7489" bold="true" box="[696,801,225,249]" pageId="77" pageNumber="77">Antenna:</emphasis>
Basal three segments stout, approximately equal in length; scape short, cylindrical; pedicel scoop shaped (longer setulae ventrally). Basal flagellomere asymmetrically drop shaped, attached dorsally to ventral apex of scape. Terminal article of antenna aristalike; long, flagellate, with short pubescence, length of article approximately equal to length of head. Occiput behind posterior margin of eyes with fringe of long, stiff, very fine setae.
<emphasis id="75C4EAFAE925ED144312FDEDCF587653" bold="true" box="[885,1029,523,547]" pageId="77" pageNumber="77">Mouthparts:</emphasis>
Proboscis short, slightly flattened laterally, only labellum visible beyond oral margin; palp and number of pseudotracheae not visible.
<emphasis id="75C4EAFAE925ED1443C0FD89CF5D76F7" bold="true" box="[935,1024,623,647]" pageId="77" pageNumber="77">Thorax:</emphasis>
Virtually bare of setulae, with only very sparse setulae on scutum and coxae. Bristlelike macrosetae as follows: 1 pair scutellars (apical), 1 postpronotal; 1 notopleural; 2 supraalars, 2 postalars. Scutum slightly arched, dorsum of scutellum slightly below that of scutum, scutellum small, no proscutellum present. Tranverse suture short, S-shaped, fully lateral.
<emphasis id="75C4EAFAE925ED1442DFFC91C9A177FF" bold="true" box="[696,764,887,911]" pageId="77" pageNumber="77">Wing:</emphasis>
Slender, middle portion of wing obscured due to preservation and folding. Tip of Sc reaching nearly to middle of wing; tip of R
<subScript id="DB3434ADE925ED144400FC23CF2D77A6" attach="left" box="[1127,1136,965,982]" fontSize="7" pageId="77" pageNumber="77">1</subScript>
slightly beyond middle of wing. R
<subScript id="DB3434ADE925ED1443AAFC00C88B7787" attach="left" box="[973,982,998,1015]" fontSize="7" pageId="77" pageNumber="77">1</subScript>
and R
<subScript id="DB3434ADE925ED14447CFC00CF6B7787" attach="left" box="[1051,1078,998,1015]" fontSize="7" pageId="77" pageNumber="77">2+3</subScript>
very straight, nearly parallel; R
<subScript id="DB3434ADE925ED144308FBE1C82A7068" attach="right" box="[879,887,1031,1048]" fontSize="7" pageId="77" pageNumber="77">4</subScript>
-R
<subScript id="DB3434ADE925ED1443F6FBE1C8C77068" attach="left" box="[913,922,1031,1048]" fontSize="7" pageId="77" pageNumber="77">5</subScript>
forked, base of fork slightly distal to level of apex of cell d; R
<subScript id="DB3434ADE925ED144469FBCECF4A7049" attach="left" box="[1038,1047,1064,1081]" fontSize="7" pageId="77" pageNumber="77">4</subScript>
slightly sinuous, tip of R
<subScript id="DB3434ADE925ED14436EFBAFC84F702A" attach="left" box="[777,786,1097,1114]" fontSize="7" pageId="77" pageNumber="77">5</subScript>
ending at apex of wing. Crossvein rm at middle of cell d. Cell br significantly longer than cell bm. Cells m3 and cup present (CuA
<subScript id="DB3434ADE925ED1444E7FB6DCFD470EC" attach="left" box="[1152,1161,1163,1180]" fontSize="7" pageId="77" pageNumber="77">1</subScript>
+ A
<subScript id="DB3434ADE925ED1444D8FB6DCF9570EC" attach="left" box="[1215,1224,1163,1180]" fontSize="7" pageId="77" pageNumber="77">1</subScript>
meeting before wing apex). A
<subScript id="DB3434ADE925ED144395FB4AC8A670CD" attach="left" box="[1010,1019,1196,1213]" fontSize="7" pageId="77" pageNumber="77">2</subScript>
well developed but apically evanescent; anal lobe well developed, alula and upper calypter not developed.
<emphasis id="75C4EAFAE925ED1444E9FB05CF95708B" bold="true" box="[1166,1224,1251,1275]" pageId="77" pageNumber="77">Legs:</emphasis>
Metacoxa with well-developed knob on posterior surface. All tibiae with dorsal and ventral longitudinal rows of short, erect macrosetae (lengths of setae approximately equal to width of tibia), as follows: protibia with 2 dorsal, 2 ventral setae; mesotibia with 3 dorsal, 5 ventral; metatibia with 810 dorsal, 56 ventral setae. Apices of tibiae with circlet of approximately 4 small setae. Tibial spurs absent. Leg segment lengths: tibia&gt; femur&gt; tarsomere 1&gt; tarsomere 2&gt; 3&gt; 4 = 5. Tarsomeres 14 on all legs with circlets of minute setae. Pulvilli slender, slightly shorter than claw; empodium tiny, bristleform.
<emphasis id="75C4EAFAE925ED144317F976C8B872D8" bold="true" box="[880,997,1680,1704]" pageId="77" pageNumber="77">Abdomen:</emphasis>
Long, slender, nearly cylindrical. Eight tergites exposed; lateral margins of tergites and sternites meeting.
<emphasis id="75C4EAFAE927ED16426DFF07CB95756A" bold="true" pageId="79" pageNumber="79">Terminalia (male):</emphasis>
Projected posteriad, not flexed dorsad or ventrad. Epandrium apparently widely divided into pair of flat, setose lobes dorsally. Pair of small, setose lobes between epandrium and gonocoxae, probably the cerci. Gonocoxae setose, entirely free from epandrium and hypandrium, longer than gonostyli; gonostyli with only microtrichia (no setae), slightly pointed apically, without teeth; gonocoxae and gonostyli apparently articulating in oblique plane. Aedeagus not visible.
</paragraph>
</subSubSection>
<caption id="13CF6660E926ED17401FF9FCCF7372F8" ID-DOI="http://doi.org/10.5281/zenodo.5380997" ID-Zenodo-Dep="5380997" httpUri="https://zenodo.org/record/5380997/files/figure.png" pageId="78" pageNumber="78" startId="78.[120,162,1562,1583]" targetBox="[135,1224,236,1532]" targetPageId="78">
<paragraph id="470F36E8E926ED17401FF9FCCF7372F8" blockId="78.[120,1224,1560,1672]" pageId="78" pageNumber="78">
FIG. 26.
<taxonomicName id="80B04D6BE926ED1740B7F9FEC9DF7240" box="[208,642,1560,1584]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="75C4EAFAE926ED1740B7F9FECAD07240" box="[208,397,1560,1584]" italics="true" pageId="78" pageNumber="78">Burmapsilocephala</emphasis>
spp. (Apsilocephalidae)
</taxonomicName>
in Burmese amber.
<emphasis id="75C4EAFAE926ED174337F9FEC8DB7240" bold="true" box="[848,902,1560,1584]" pageId="78" pageNumber="78">AC.</emphasis>
<taxonomicName id="80B04D6BE926ED1743ECF9FECFCF7240" authority="Grimaldi, 2016" authorityName="Grimaldi" authorityYear="2016" box="[907,1170,1560,1584]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="species" species="evocoa" status="sp. nov.">
<emphasis id="75C4EAFAE926ED1743ECF9FECFCF7240" box="[907,1170,1560,1584]" italics="true" pageId="78" pageNumber="78">Burmapsilocephala evocoa</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="6EF75781E926ED1744F9F9FECB9E723D" pageId="78" pageNumber="78" rank="species">new species</taxonomicNameLabel>
.
<emphasis id="75C4EAFAE926ED1740ABF9D3CBB8723D" bold="true" box="[204,229,1589,1613]" pageId="78" pageNumber="78">D.</emphasis>
<taxonomicName id="80B04D6BE926ED17408BF9D0C905723D" authority="Gaimari and Mostovski" authorityName="Gaimari and Mostovski" box="[236,600,1589,1613]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="species" species="cockerelli">
<emphasis id="75C4EAFAE926ED17408BF9D0CA3C723D" box="[236,353,1589,1613]" italics="true" pageId="78" pageNumber="78">B. cockerelli</emphasis>
Gaimari and Mostovski
</taxonomicName>
(ex: Gaimari and
<bibRefCitation id="23214B19E926ED174373F9D3C8E3723D" author="Mostovski, M. B. &amp; X. Martinez-Delclos" box="[788,958,1589,1613]" pageId="78" pageNumber="78" pagination="137 - 148" refId="ref59314" refString="Mostovski, M. B., and X. Martinez-Delclos. 2000. New Nemestrinoidea (Diptera: Brachycera) from the Upper Jurassic-Lower Cretaceous of Eurasia, taxonomy and palaeobiology. Entomological Problems 31 (2): 137 - 148." type="journal article" year="2000">Mostovski, 2000</bibRefCitation>
).
<emphasis id="75C4EAFAE926ED1743B7F9D3C8B4723D" bold="true" box="[976,1001,1589,1613]" pageId="78" pageNumber="78">A.</emphasis>
Head, oblique frontal view.
<emphasis id="75C4EAFAE926ED1740D6F9B5CB9A721B" bold="true" box="[177,199,1619,1643]" pageId="78" pageNumber="78">B.</emphasis>
Male terminalia, oblique lateral view.
<emphasis id="75C4EAFAE926ED17422FF9B4C93D721A" bold="true" box="[584,608,1618,1642]" pageId="78" pageNumber="78">C.</emphasis>
Wing of
<taxonomicName id="80B04D6BE926ED1742DAF9B5C846721A" authorityName="Grimaldi" authorityYear="2016" box="[701,795,1619,1642]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="species" species="evocoa">
<emphasis id="75C4EAFAE926ED1742DAF9B5C846721A" box="[701,795,1619,1642]" italics="true" pageId="78" pageNumber="78">B. evocoa</emphasis>
</taxonomicName>
(portion are not fully observable). Arrows indicate diagnostic venational differences.
<emphasis id="75C4EAFAE926ED17424DF996C91E72F8" bold="true" box="[554,579,1648,1672]" pageId="78" pageNumber="78">D.</emphasis>
Wing of
<taxonomicName id="80B04D6BE926ED1742C3F997C84572F8" box="[676,792,1648,1672]" class="Insecta" family="Apsilocephalidae" genus="Burmapsilocephala" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="species" species="cockerelli">
<emphasis id="75C4EAFAE926ED1742C3F997C84572F8" box="[676,792,1648,1672]" italics="true" pageId="78" pageNumber="78">B. cockerelli</emphasis>
</taxonomicName>
. Panels A, B to same scale.
</paragraph>
</caption>
<subSubSection id="0FAA6563E927ED1640FFFDA8C9067615" box="[152,603,589,614]" pageId="79" pageNumber="79" type="materials_examined">
<paragraph id="470F36E8E927ED1640FFFDA8C9067615" blockId="79.[120,650,225,712]" box="[152,603,589,614]" pageId="79" pageNumber="79">
<materialsCitation id="F7D83CB5E927ED1640FFFDA8C9067615" box="[152,603,589,614]" collectionCode="TYPE, AMNH" pageId="79" pageNumber="79" specimenCount="1" typeStatus="holotype">
<collectionCode id="21A1AE2DE927ED1640FFFDA8CB927615" box="[152,207,590,613]" pageId="79" pageNumber="79">TYPE</collectionCode>
:
<typeStatus id="980B884AE927ED1640BBFDABCA1E7615" box="[220,323,589,613]" pageId="79" pageNumber="79" type="holotype">Holotype</typeStatus>
, male,
<collectionCode id="21A1AE2DE927ED1641F6FDA8CAB87616" box="[401,485,590,614]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="79" pageNumber="79">AMNH</collectionCode>
Bu-SE2/4.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0FAA6563E927ED1640FFFD89CA4E76B8" pageId="79" pageNumber="79" type="etymology">
<paragraph id="470F36E8E927ED1640FFFD89CA4E76B8" blockId="79.[120,650,225,712]" pageId="79" pageNumber="79">
ETYMOLOGY: Taken directly from the name of the
<typeStatus id="980B884AE927ED1640C5FD76CB8C76D8" box="[162,209,656,680]" pageId="79" pageNumber="79">type</typeStatus>
genus of
<taxonomicName id="80B04D6BE927ED16415BFD69CAF376D7" box="[316,430,655,679]" class="Insecta" family="Evocoidae" kingdom="Animalia" order="Diptera" pageId="79" pageNumber="79" phylum="Arthropoda" rank="family">Evocoidae</taxonomicName>
, to which this fossil seems related.
</paragraph>
</subSubSection>
</treatment>
</document>