444 lines
62 KiB
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444 lines
62 KiB
XML
<document ID-DOI="10.11646/zootaxa.5239.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7621749" ID-ZooBank="E446D1F2-B922-45A9-9F6D-01406154594E" approvalRequired="9" approvalRequired_for_textStreams="7" approvalRequired_for_treatments="2" checkinTime="1675872344917" checkinUser="plazi" docAuthor="Shinohara, Akihiko" docDate="2023" docId="C36200221219421A9FC7F9D8FE9495ED" docLanguage="en" docName="zootaxa.5239.1.1.pdf" docOrigin="Zootaxa 5239 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5239.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tremex okinawensis Togashi 1997" docType="treatment" docVersion="2" lastPageNumber="32" masterDocId="3F5B785A120242059F50FFEFFFA9966C" masterDocTitle="The woodwasp genus Tremex (Hymenoptera, Siricidae) of Japan" masterLastPageNumber="40" masterPageNumber="1" pageNumber="28" updateTime="1675873128288" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>The woodwasp genus Tremex (Hymenoptera, Siricidae) of Japan</mods:title>
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<taxonomicName authority="Togashi, 1997" authorityName="Togashi" authorityYear="1997" box="[151,576,1590,1617]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="okinawensis">
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<emphasis bold="true" box="[151,576,1590,1617]" pageId="27" pageNumber="28">
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<emphasis bold="true" box="[151,398,1590,1617]" italics="true" pageId="27" pageNumber="28">Tremex okinawensis</emphasis>
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<bibRefCitation author="Togashi, I." box="[405,576,1590,1616]" pageId="27" pageNumber="28" pagination="1 - 6" refId="ref29266" refString="Togashi, I. (1997) Two new species of Tremex Jurine (Hymenoptera: Siricidae) from Japan. Species Diversity, 2, 1 - 6. https: // doi. org / 10.12782 / specdiv. 2.1" type="journal article" year="1997">Togashi, 1997</bibRefCitation>
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</emphasis>
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<paragraph blockId="27.[151,960,1590,1653]" box="[151,960,1626,1653]" pageId="27" pageNumber="28">
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(
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<figureCitation box="[159,283,1627,1653]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="27" pageNumber="28">Figs 3K, L</figureCitation>
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,
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<figureCitation box="[295,332,1627,1653]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="27" pageNumber="28">4H</figureCitation>
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,
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<figureCitation box="[343,415,1626,1653]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="27" pageNumber="28">5G, N</figureCitation>
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,
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<figureCitation box="[426,456,1627,1653]" captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="27" pageNumber="28">6F</figureCitation>
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,
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<figureCitation box="[467,479,1627,1653]" captionStart="FIGURE 8" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[159,1428,568,1903]" captionTargetId="figure-186@15.[151,1435,561,1909]" captionTargetPageId="15" captionText="FIGURE 8. Wing, female.—A, T. nakanei, holotype of T. kaedei; B, T. nakanei, Tokyo; C, T. nakanei, Suzuka, Mie; D, T. nakanei, Narukawa-keikoku, Ehime; E, F, T.contractus, Okayama-shi, Okayama; G, T. okinawensis, holotype; H, T.okinawensis, Ginama, Okinawa; I, T. okinawensis, holotype of T. kurokivorus (left wings, ventral view); J, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621768" httpUri="https://zenodo.org/record/7621768/files/figure.png" pageId="27" pageNumber="28">8</figureCitation>
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G-J, 9F, 10K, L, 11F, 12H, 15;
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<tableCitation box="[829,951,1627,1653]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStart-3="TABLE 4" captionStartId-0="6.[151,239,980,1004]" captionStartId-1="7.[151,239,469,493]" captionStartId-2="7.[151,239,1276,1300]" captionStartId-3="20.[150,238,639,665]" captionText-0="TABLE 1. Variation of structural characters, female, 1." captionText-1="TABLE 2. Variation of structural characters, female, 2." captionText-2="TABLE 3. Variation of structural characters, male." captionText-3="TABLE 4. Number of specimens collected in each 10–11 days of April to November." pageId="27" pageNumber="28">Tables 1–4</tableCitation>
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)
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</paragraph>
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</subSubSection>
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<taxonomicName authority="Togashi, 1997: 2" authorityName="Togashi" authorityPageNumber="2" authorityYear="1997" box="[151,541,1695,1719]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="okinawensis">
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<emphasis box="[151,357,1695,1719]" italics="true" pageId="27" pageNumber="28">Tremex okinawensis</emphasis>
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<bibRefCitation author="Togashi, I." box="[365,541,1695,1719]" pageId="27" pageNumber="28" pagination="1 - 6" refId="ref29266" refString="Togashi, I. (1997) Two new species of Tremex Jurine (Hymenoptera: Siricidae) from Japan. Species Diversity, 2, 1 - 6. https: // doi. org / 10.12782 / specdiv. 2.1" type="journal article" year="1997">Togashi, 1997: 2</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Yafuso, M." box="[554,747,1695,1719]" page="328" pageId="27" pageNumber="28" year="2002">
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<bibRefCitation author="Yafuso, M." box="[554,747,1695,1719]" pageId="27" pageNumber="28" pagination="328" refId="ref29657" refString="Yafuso, M. (2002) Siricidae. In: Azuma, S., Yafuso, M., Kinjo, M., Hayashi, M., Kohama, T., Sasaki, T., Kimura, M. & Kawamura, M. (Eds.), Check List of the Insects of the Ryukyu Islands, Second Edition. The Biological Society of Okinawa, Nishihara, pp. 328. [in Japanese]" type="book chapter" year="2002">Yafuso, 2002: 328</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Togashi, I." box="[759,961,1695,1719]" page="939" pageId="27" pageNumber="28" year="2006">
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<bibRefCitation author="Togashi, I." box="[759,961,1695,1719]" pageId="27" pageNumber="28" pagination="938 - 944" refId="ref29340" refString="Togashi, I. (2006) Two new species of Tremex (Hymenoptera: Siricidae) reared from Castanopsis and Symplocos in Japan. Proceedings of the Entomological Society of Washington, 108, 938 - 944." type="journal article" year="2006">Togashi, 2006: 939</bibRefCitation>
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;
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<treatmentCitation author="Taeger, A. & Blank, S. M. & Liston, A. D." box="[973,1226,1695,1720]" page="108" pageId="27" pageNumber="28" year="2010">
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<bibRefCitation author="Taeger, A. & Blank, S. M. & Liston, A. D." box="[973,1226,1695,1720]" pageId="27" pageNumber="28" pagination="1 - 1064" refId="ref28562" refString="Taeger, A., Blank, S. M. & Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
|
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Taeger
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<emphasis box="[1050,1097,1695,1719]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
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., 2010: 108
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</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Kawano, T." box="[1238,1432,1695,1719]" page="19" pageId="27" pageNumber="28" year="2014">
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<bibRefCitation author="Kawano, T." box="[1238,1432,1695,1719]" pageId="27" pageNumber="28" pagination="19 - 20" refId="ref25072" refString="Kawano, T. (2014) The first record of Tremex okinawensis Togashi since the original description (Siricidae: Tremecinae). Tsunekibachi, Fukui, 24, 19 - 20. [in Japanese]" type="journal article" year="2014">Kawano, 2014: 19</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Naito, T." box="[199,360,1726,1750]" page="20" pageId="27" pageNumber="28" year="2019">
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<bibRefCitation author="Naito, T." box="[199,360,1726,1750]" pageId="27" pageNumber="28" pagination="18 - 20" refId="ref26445" refString="Naito, T. (2019) Family Siricidae. In: Editorial Committee of Catalogue of the Insects of Japan (Ed.), Catalogue of the Insects of Japan, Volume 9 Hymenoptera. Part 1. Symphyta. Entomological Society of Japan, Kyoto, pp. 18 - 20. [in Japanese]" type="book chapter" year="2019">Naito, 2019: 20</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Naito, T." box="[371,545,1726,1750]" page="467" pageId="27" pageNumber="28" year="2020">
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<bibRefCitation author="Naito, T." box="[371,545,1726,1750]" pageId="27" pageNumber="28" pagination="462 - 468" refId="ref26504" refString="Naito, T. (2020) Family Siricidae. In: Naito, T., Shinohara, A., Hara, H. & Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 179 - 182 + 462 - 468. [in Japanese]" type="book chapter" year="2020">Naito, 2020: 467</bibRefCitation>
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</treatmentCitation>
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.
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</paragraph>
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<paragraph blockId="27.[151,1437,1695,1781]" box="[151,1336,1756,1781]" pageId="27" pageNumber="28">
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<treatmentCitationGroup box="[151,1336,1756,1781]" pageId="27" pageNumber="28">
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<treatmentCitation author="Togashi, I." box="[151,555,1756,1781]" page="941" pageId="27" pageNumber="28" year="2006">
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<taxonomicName authority="Togashi, 2006: 941" authorityName="Togashi" authorityPageNumber="941" authorityYear="2006" box="[151,555,1756,1781]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="kurokivorus">
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<emphasis box="[151,351,1756,1780]" italics="true" pageId="27" pageNumber="28">Tremex kurokivorus</emphasis>
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<bibRefCitation author="Togashi, I." box="[357,555,1756,1781]" pageId="27" pageNumber="28" pagination="938 - 944" refId="ref29340" refString="Togashi, I. (2006) Two new species of Tremex (Hymenoptera: Siricidae) reared from Castanopsis and Symplocos in Japan. Proceedings of the Entomological Society of Washington, 108, 938 - 944." type="journal article" year="2006">Togashi, 2006: 941</bibRefCitation>
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;
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<treatmentCitation author="Taeger, A. & Blank, S. M. & Liston, A. D." box="[565,808,1756,1781]" page="108" pageId="27" pageNumber="28" year="2010">
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<bibRefCitation author="Taeger, A. & Blank, S. M. & Liston, A. D." box="[565,808,1756,1781]" pageId="27" pageNumber="28" pagination="1 - 1064" refId="ref28562" refString="Taeger, A., Blank, S. M. & Liston, A. D. (2010) World catalog of Symphyta (Hymenoptera). Zootaxa, 2580 (1), 1 - 1064. https: // doi. org / 10.11646 / zootaxa. 2580.1.1" type="journal article" year="2010">
|
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Taeger
|
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<emphasis box="[639,684,1756,1780]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
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., 2010: 108
|
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</bibRefCitation>
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</treatmentCitation>
|
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;
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<treatmentCitation author="Naito, T." box="[823,984,1756,1781]" page="20" pageId="27" pageNumber="28" year="2019">
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<bibRefCitation author="Naito, T." box="[823,984,1756,1781]" pageId="27" pageNumber="28" pagination="18 - 20" refId="ref26445" refString="Naito, T. (2019) Family Siricidae. In: Editorial Committee of Catalogue of the Insects of Japan (Ed.), Catalogue of the Insects of Japan, Volume 9 Hymenoptera. Part 1. Symphyta. Entomological Society of Japan, Kyoto, pp. 18 - 20. [in Japanese]" type="book chapter" year="2019">Naito, 2019: 20</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="Naito, T." box="[995,1169,1756,1781]" page="467" pageId="27" pageNumber="28" year="2020">
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<bibRefCitation author="Naito, T." box="[995,1169,1756,1781]" pageId="27" pageNumber="28" pagination="462 - 468" refId="ref26504" refString="Naito, T. (2020) Family Siricidae. In: Naito, T., Shinohara, A., Hara, H. & Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 179 - 182 + 462 - 468. [in Japanese]" type="book chapter" year="2020">Naito, 2020: 467</bibRefCitation>
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</treatmentCitation>
|
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.
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<emphasis bold="true" box="[1179,1330,1757,1781]" pageId="27" pageNumber="28">New synonym</emphasis>
|
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="27.[151,1437,1826,2033]" lastBlockId="28.[151,1437,151,501]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
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<emphasis bold="true" box="[151,420,1826,1853]" pageId="27" pageNumber="28">Diagnostic characters.</emphasis>
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<emphasis box="[429,511,1827,1853]" italics="true" pageId="27" pageNumber="28">Female</emphasis>
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. Length without ovipositor
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<quantity box="[828,952,1826,1853]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="27" pageNumber="28" unit="mm" value="22.0" valueMax="25.0" valueMin="19.0">19–25 mm</quantity>
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. Head pale brown, with clypeus and malar space black. Antenna wholly pale brown (
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<figureCitation box="[611,698,1862,1889]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="27" pageNumber="28">Fig. 5G</figureCitation>
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). Thorax black, with pronotum, propleuron, parascutal lobe, part of or entire mesoscutellum, part of metanotum pale brown; pronotum (
|
||
<figureCitation box="[896,1013,1899,1925]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="27" pageNumber="28">Fig. 3K, L</figureCitation>
|
||
) without blackish longitudinal median line; mesoscutum and mesopleuron sometimes marked with pale brown (
|
||
<figureCitation box="[946,1033,1935,1961]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="27" pageNumber="28">Fig. 3K</figureCitation>
|
||
). Fore and mid legs pale brown, with part of coxa and sometimes parts of trochanter and femur blackish to black; hind leg black, with tibia and tarsus dark brown (basal halves of tibia and tarsomere 1 creamy white). Forewing with distinct infuscation around cells R, 1R1, 2Rs, 3M, 3Cu and usually 2R1; cell 3R1 not infuscated basally and along anterior margin (
|
||
<figureCitation box="[1183,1308,151,177]" captionStart="FIGURE 8" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[159,1428,568,1903]" captionTargetId="figure-186@15.[151,1435,561,1909]" captionTargetPageId="15" captionText="FIGURE 8. Wing, female.—A, T. nakanei, holotype of T. kaedei; B, T. nakanei, Tokyo; C, T. nakanei, Suzuka, Mie; D, T. nakanei, Narukawa-keikoku, Ehime; E, F, T.contractus, Okayama-shi, Okayama; G, T. okinawensis, holotype; H, T.okinawensis, Ginama, Okinawa; I, T. okinawensis, holotype of T. kurokivorus (left wings, ventral view); J, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621768" httpUri="https://zenodo.org/record/7621768/files/figure.png" pageId="28" pageNumber="29">Fig. 8 G–J</figureCitation>
|
||
). Abdomen black, with following pale brown: Usually anterior margin of tergum 1, most of tergum 2 (except for posterior margin), lateral spot on terga 3–6, narrow anterior margin of tergum 7, anterior 1/3–1/2 of tergum 8 (except for anterior lateral corner), large mark on tergum 9, covering anterior half of precornal basin and large lateral surface, and cornus. Pale brown on head, thorax and tergum 9 often with dark orange tint. Antenna with 14–16 antennomeres. Pronotum (
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="28" pageNumber="29">Fig. 3K, L</figureCitation>
|
||
) with MPL 0.75–0.85 (average 0.81) times as long as OOCL. Hind tibia 0.99–1.05 (average 1.01) times as long as hind tarsomere 1; dorsal margin of hind tibia straight or very shallowly concave (
|
||
<figureCitation box="[1104,1190,367,393]" captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="28" pageNumber="29">Fig. 6F</figureCitation>
|
||
). Forewing 1.36–1.43 (average 1.38) times as long as ovipositor sheath. Abdominal tergum 8 1.00–1.27 (average 1.10) times as long as terga 5–7 combined; precornal basin short, 0.57–0.77 (average 0.63) times as long as wide, widest in posterior part (
|
||
<figureCitation captionStart="FIGURE 11" captionStartId="18.[151,250,1565,1589]" captionTargetBox="[175,1383,189,1534]" captionTargetId="figure-20@18.[151,1435,181,1542]" captionTargetPageId="18" captionText="FIGURE 11. Posterior part of abdomen, female.—A, T. bicinctus, holotype; B, T. longicollis, Okazaki, Aichi; C, T. katayamai, holotype; D, T. fuscicornis, Inchon, Korea; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, holotype." figureDoi="http://doi.org/10.5281/zenodo.7621776" httpUri="https://zenodo.org/record/7621776/files/figure.png" pageId="28" pageNumber="29">Fig. 11F</figureCitation>
|
||
); ovipositor sheath 2.69–2.75 (average 2.72) times as long as apical sheath.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7621781" ID-Zenodo-Dep="7621781" httpUri="https://zenodo.org/record/7621781/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,260,1784,1810]" targetBox="[171,1413,523,1760]" targetPageId="28">
|
||
<paragraph blockId="28.[151,1436,1784,1847]" pageId="28" pageNumber="29">
|
||
<emphasis bold="true" box="[151,303,1784,1810]" pageId="28" pageNumber="29">FIGURE 13.</emphasis>
|
||
Distribution of
|
||
<taxonomicName box="[484,601,1784,1810]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="apicalis">
|
||
<emphasis box="[484,601,1784,1810]" italics="true" pageId="28" pageNumber="29">T. apicalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName box="[615,744,1784,1810]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bicinctus">
|
||
<emphasis box="[615,744,1784,1810]" italics="true" pageId="28" pageNumber="29">T. bicinctus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Maa" authorityYear="1949" box="[800,946,1785,1810]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="contractus">
|
||
<emphasis box="[800,946,1785,1810]" italics="true" pageId="28" pageNumber="29">T. contractus</emphasis>
|
||
</taxonomicName>
|
||
in Japan based on the specimens examined (see Appendix for the collection records and the main text for the data of the type material of
|
||
<taxonomicName box="[1198,1326,1820,1846]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="bicinctus">
|
||
<emphasis box="[1198,1326,1820,1846]" italics="true" pageId="28" pageNumber="29">T. bicinctus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="29" pageNumber="30" type="description">
|
||
<paragraph blockId="29.[151,1437,151,609]" pageId="29" pageNumber="30">
|
||
<emphasis box="[199,255,151,177]" italics="true" pageId="29" pageNumber="30">Male</emphasis>
|
||
. Length
|
||
<quantity box="[353,474,151,177]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="29" pageNumber="30" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">11–16 mm</quantity>
|
||
. Head, antenna, thorax and legs black except as follows: terminal antennomere partly or entirely pale brown (
|
||
<figureCitation box="[383,472,186,213]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="29" pageNumber="30">Fig. 5N</figureCitation>
|
||
). Fore and mid femora (except for basal parts), tibiae and tarsi pale reddish brown and extreme base of hind tibia pale brown. Wings dark brown, basal and midapical parts more hyaline (
|
||
<figureCitation box="[1228,1311,223,249]" captionStart="FIGURE 9" captionStartId="16.[151,250,1266,1290]" captionTargetBox="[158,1429,276,1239]" captionTargetId="figure-20@16.[151,1435,271,1243]" captionTargetPageId="16" captionText="FIGURE 9. Wing, male.—A, T. apicalis, Tsukuba, Ibaraki; B, T. longicollis, Muroto, Kochi; C, T. fuscicornis, Chitose, Hokkaido; D, T. nakanei, Noda, Chiba; E, T. contractus, Tatsuno, Hyogo; F, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621770" httpUri="https://zenodo.org/record/7621770/files/figure.png" pageId="29" pageNumber="30">Fig. 9F</figureCitation>
|
||
). Abdomen (
|
||
<figureCitation box="[159,260,259,285]" captionStart="FIGURE 12" captionStartId="19.[151,250,1268,1292]" captionTargetBox="[159,1428,186,1239]" captionTargetId="figure-15@19.[151,1435,181,1244]" captionTargetPageId="19" captionText="FIGURE 12. Abdomen, male.—A, F, T. nakanei, Noda, Chiba; B, G, T. contractus, Tatsuno, Hyogo; C, T. apicalis, Tsukuba, Ibaraki; D, T. fuscicornis, Chitose, Hokkaido; E, T. longicollis, Muroto, Kochi; H, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621778" httpUri="https://zenodo.org/record/7621778/files/figure.png" pageId="29" pageNumber="30">Fig. 12H</figureCitation>
|
||
) pale brown, with following black: terga 1 and 2, basal part (extending posteriorly along median line) of terga 3–8, entire sternum 2, anterior and posterior margins and median line of sterna 3–7, most of sternum 8 (except for large lateral brownish spots) and sternum 9 (except for large median and posterior part). Antenna with 13 or 14 antennomeres. Pronotum (
|
||
<figureCitation box="[446,535,367,393]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="29" pageNumber="30">Fig. 4H</figureCitation>
|
||
) with MPL 0.81–0.84 (average 0.82) times as long as OOCL. Hind tibia short, 1.18–1.25 (average 1.22) times as long as hind tarsomere 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="29" pageNumber="30" type="materials_examined">
|
||
<paragraph blockId="29.[151,1437,151,609]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[199,484,438,465]" pageId="29" pageNumber="30">Type material examined.</emphasis>
|
||
<materialsCitation collectingDate="1971-05-03" collectionCode="NSMT" location="Benoki" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="holotype">
|
||
<specimenCount box="[487,507,439,465]" pageId="29" pageNumber="30" type="female">♀</specimenCount>
|
||
(
|
||
<typeStatus box="[518,608,439,465]" pageId="29" pageNumber="30">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Togashi" authorityYear="1997" box="[636,783,439,465]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis box="[636,783,439,465]" italics="true" pageId="29" pageNumber="30">T.okinawensis</emphasis>
|
||
</taxonomicName>
|
||
),“
|
||
<location LSID="urn:lsid:plazi:treatment:C36200221219421A9FC7F9D8FE9495ED:4E14E7EF121F42189C7AFE58FCD197BD" box="[810,888,439,465]" name="Benoki" pageId="29" pageNumber="30" stateProvince="Okinawa">Benoki</location>
|
||
,
|
||
<collectingRegion box="[895,988,439,465]" country="Japan" name="Okinawa" pageId="29" pageNumber="30">Okinawa</collectingRegion>
|
||
Is.,
|
||
<date box="[1025,1125,439,465]" pageId="29" pageNumber="30" value="1971-05-03">
|
||
<collectingDate box="[1025,1125,439,465]" pageId="29" pageNumber="30" value="1971-05-03">3.V.1971</collectingDate>
|
||
</date>
|
||
, K.
|
||
<collectingRegion box="[1161,1237,439,465]" country="Japan" name="Miyagi" pageId="29" pageNumber="30">Miyagi</collectingRegion>
|
||
”“
|
||
<typeStatus box="[1259,1357,439,465]" pageId="29" pageNumber="30">Holotype</typeStatus>
|
||
<taxonomicName authority="Togashi" authorityName="Togashi" authorityYear="1997" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis italics="true" pageId="29" pageNumber="30">Tremex okinawensis</emphasis>
|
||
Togashi
|
||
</taxonomicName>
|
||
” (
|
||
<collectionCode box="[395,473,475,501]" country="Japan" name="National Science Museum (Natural History)" pageId="29" pageNumber="30">NSMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectingDate="2003-10-12" collectingDateMax="2004-08-30" collectingDateMin="2003-10-12" collectionCode="NSMT" collectorName="H. Makihara" county="Oyama" location="Nakaharakuroki" municipality="Okinoerabu-jima" pageId="29" pageNumber="30" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa" typeStatus="holotype">
|
||
<specimenCount box="[493,513,475,501]" pageId="29" pageNumber="30" type="female">♀</specimenCount>
|
||
(
|
||
<typeStatus box="[530,620,475,501]" pageId="29" pageNumber="30">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Togashi" authorityYear="2006" box="[662,810,475,501]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kurokivorus">
|
||
<emphasis box="[662,810,475,501]" italics="true" pageId="29" pageNumber="30">T. kurokivorus</emphasis>
|
||
</taxonomicName>
|
||
), “
|
||
<collectingCounty box="[844,923,476,503]" pageId="29" pageNumber="30">Ôyama</collectingCounty>
|
||
,
|
||
<collectingMunicipality box="[937,1114,475,501]" pageId="29" pageNumber="30">Okinoerabu-jima</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:C36200221219421A9FC7F9D8FE9495ED:4E14E7EF121F42189B38FE34FAA49799" box="[1128,1293,475,501]" county="Oyama" municipality="Okinoerabu-jima" name="Nakaharakuroki" pageId="29" pageNumber="30" stateProvince="Okinawa">Nakaharakuroki</location>
|
||
[=
|
||
<taxonomicName class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="nakaharae">
|
||
<emphasis italics="true" pageId="29" pageNumber="30">Symplocos nakaharae</emphasis>
|
||
</taxonomicName>
|
||
],
|
||
<date box="[281,396,511,537]" pageId="29" pageNumber="30" value="2003-10-12">
|
||
<collectingDate box="[281,396,511,537]" pageId="29" pageNumber="30" value="2003-10-12">2003.10.12</collectingDate>
|
||
</date>
|
||
coll. wood,
|
||
<date box="[520,623,511,537]" pageId="29" pageNumber="30" value="2004-08-30">
|
||
<collectingDate box="[520,623,511,537]" pageId="29" pageNumber="30" value="2004-08-30">2004.8.30</collectingDate>
|
||
</date>
|
||
em.,
|
||
<collectorName box="[678,810,511,537]" pageId="29" pageNumber="30">H. Makihara</collectorName>
|
||
” “
|
||
<typeStatus box="[833,931,511,537]" pageId="29" pageNumber="30">Holotype</typeStatus>
|
||
<taxonomicName authority="Shinohara, 2023" authorityName="Shinohara" authorityYear="2023" box="[936,1139,511,537]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kurokivorus" status="sp. nov.">
|
||
<emphasis box="[936,1139,511,537]" italics="true" pageId="29" pageNumber="30">Tremex kurokivorus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1143,1223,511,537]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
|
||
” (
|
||
<collectionCode box="[1245,1324,511,537]" country="Japan" name="National Science Museum (Natural History)" pageId="29" pageNumber="30">NSMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="29.[151,1437,151,609]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[199,515,546,572]" pageId="29" pageNumber="30">Other material examined.</emphasis>
|
||
<materialsCitation county="Amami-oshima Island" location="See Appendix" municipality="Island" pageId="29" pageNumber="30" specimenCount="6" specimenCount-female="6" stateProvince="Okinawa">
|
||
<specimenCount box="[526,563,548,575]" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
|
||
4³ (3³ from
|
||
<collectingCounty box="[726,979,547,573]" pageId="29" pageNumber="30">Amami-ôshima Island</collectingCounty>
|
||
and
|
||
<specimenCount box="[1042,1084,548,575]" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
|
||
1³ from
|
||
<collectingRegion box="[1188,1289,547,573]" country="Japan" name="Okinawa" pageId="29" pageNumber="30">Okinawa</collectingRegion>
|
||
<collectingMunicipality box="[1300,1373,547,573]" pageId="29" pageNumber="30">Island</collectingMunicipality>
|
||
).
|
||
<location LSID="urn:lsid:plazi:treatment:C36200221219421A9FC7F9D8FE9495ED:4E14E7EF121F42189A24FDCCFEAC940D" county="Amami-oshima Island" municipality="Island" name="See Appendix" pageId="29" pageNumber="30" stateProvince="Okinawa">See Appendix</location>
|
||
for collection data
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7621785" ID-Zenodo-Dep="7621785" httpUri="https://zenodo.org/record/7621785/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1904,1928]" targetBox="[170,1415,636,1877]" targetPageId="29">
|
||
<paragraph blockId="29.[151,1436,1904,1964]" pageId="29" pageNumber="30">
|
||
<emphasis bold="true" box="[151,288,1904,1928]" pageId="29" pageNumber="30">FIGURE 14.</emphasis>
|
||
Distribution of
|
||
<taxonomicName authorityName=": Miyatake" authorityYear="1950" box="[451,583,1904,1928]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="longicollis">
|
||
<emphasis box="[451,583,1904,1928]" italics="true" pageId="29" pageNumber="30">T. longicollis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName=": Miyatake" authorityYear="1950" box="[633,769,1904,1928]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="fuscicornis">
|
||
<emphasis box="[633,769,1904,1928]" italics="true" pageId="29" pageNumber="30">T. fuscicornis</emphasis>
|
||
</taxonomicName>
|
||
in Japan based on the specimens examined (see Appendix for the collection records).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="30" pageNumber="31" type="distribution">
|
||
<paragraph blockId="30.[151,1436,150,285]" pageId="30" pageNumber="31">
|
||
<emphasis bold="true" box="[199,345,150,176]" pageId="30" pageNumber="31">Distribution</emphasis>
|
||
(
|
||
<figureCitation box="[369,460,150,177]" captionStart="FIGURE 15" captionStartId="30.[151,250,1941,1965]" captionTargetBox="[154,1434,310,1899]" captionTargetId="figure-50@30.[151,1436,309,1899]" captionTargetPageId="30" captionText="FIGURE 15. Distribution of T. nakanei, T. katayamai and T. okinawensis based on the specimens examined (see Appendix for the collection records and the main text for the data of the type material)." figureDoi="http://doi.org/10.5281/zenodo.7621787" httpUri="https://zenodo.org/record/7621787/files/figure.png" pageId="30" pageNumber="31">Fig. 15</figureCitation>
|
||
).
|
||
<collectingCountry box="[488,556,151,177]" name="Japan" pageId="30" pageNumber="31">Japan</collectingCountry>
|
||
: Nansei Islands (Amami-ôshima (new record), Okinoerabu and
|
||
<collectingRegion box="[1335,1436,151,177]" country="Japan" name="Okinawa" pageId="30" pageNumber="31">Okinawa</collectingRegion>
|
||
Islands).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="30" pageNumber="31" type="biology_ecology">
|
||
<paragraph blockId="30.[151,1436,150,285]" pageId="30" pageNumber="31">
|
||
<emphasis bold="true" box="[199,324,223,249]" pageId="30" pageNumber="31">Host plant</emphasis>
|
||
.
|
||
<taxonomicName box="[339,505,223,249]" class="Magnoliopsida" family="Symplocaceae" kingdom="Plantae" order="Ericales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="family">Symplocaceae</taxonomicName>
|
||
:
|
||
<taxonomicName authority="(Hayata) Masam." authorityName="Masam." baseAuthorityName="Hayata" box="[515,964,223,249]" class="Magnoliopsida" family="Symplocaceae" genus="Symplocos" kingdom="Plantae" order="Ericales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="nakaharae">
|
||
<emphasis box="[515,762,223,249]" italics="true" pageId="30" pageNumber="31">Symplocos nakaharae</emphasis>
|
||
(Hayata) Masam.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Togashi, I." box="[978,1134,223,249]" pageId="30" pageNumber="31" pagination="938 - 944" refId="ref29340" refString="Togashi, I. (2006) Two new species of Tremex (Hymenoptera: Siricidae) reared from Castanopsis and Symplocos in Japan. Proceedings of the Entomological Society of Washington, 108, 938 - 944." type="journal article" year="2006">Togashi 2006</bibRefCitation>
|
||
, and the Japanese name on the label).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7621787" ID-Zenodo-Dep="7621787" httpUri="https://zenodo.org/record/7621787/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1941,1965]" targetBox="[154,1434,310,1899]" targetPageId="30">
|
||
<paragraph blockId="30.[151,1437,1941,2001]" pageId="30" pageNumber="31">
|
||
<emphasis bold="true" box="[151,288,1941,1965]" pageId="30" pageNumber="31">FIGURE 15.</emphasis>
|
||
Distribution of
|
||
<taxonomicName authorityName="Takeuchi" authorityYear="1955" box="[451,556,1941,1965]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nakanei">
|
||
<emphasis box="[451,556,1941,1965]" italics="true" pageId="30" pageNumber="31">T. nakanei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Shinohara & Kurihara" authorityYear="2023" box="[569,700,1941,1965]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="katayamai">
|
||
<emphasis box="[569,700,1941,1965]" italics="true" pageId="30" pageNumber="31">T. katayamai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Togashi" authorityYear="1997" box="[749,898,1941,1965]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis box="[749,898,1941,1965]" italics="true" pageId="30" pageNumber="31">T. okinawensis</emphasis>
|
||
</taxonomicName>
|
||
based on the specimens examined (see Appendix for the collection records and the main text for the data of the type material).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph blockId="31.[151,1437,151,897]" pageId="31" pageNumber="32">
|
||
<emphasis bold="true" box="[199,308,151,177]" pageId="31" pageNumber="32">Remarks</emphasis>
|
||
. This species has been found only in northern part of the Nansei Islands and the male is described here for the first time. All the specimens were collected in late April to early May. The female is characterized by the pale brown antenna (
|
||
<figureCitation box="[387,476,222,249]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="31" pageNumber="32">Fig. 5G</figureCitation>
|
||
), the long pronotum (
|
||
<figureCitation box="[720,842,223,249]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="31" pageNumber="32">Fig. 3K, L</figureCitation>
|
||
), the hind tibia about as long as the hind tarsomere 1 and with the basal half creamy white and the apical half dark brown (
|
||
<figureCitation box="[916,1000,259,285]" captionStart="FIGURE 6" captionStartId="13.[151,250,1061,1085]" captionTargetBox="[155,1432,184,1034]" captionTargetId="figure-15@13.[151,1435,181,1037]" captionTargetPageId="13" captionText="FIGURE 6. Hind leg, female.—A, T. longicollis, Tsushima, Nagasaki; B, T. katayamai, holotype; C, T. fuscicornis, Inchon, Korea; D, T. nakanei, holotype of T. kaedei; E, T. contractus, Okayama-shi, Okayama; F, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621763" httpUri="https://zenodo.org/record/7621763/files/figure.png" pageId="31" pageNumber="32">Fig. 6F</figureCitation>
|
||
), the forewing with distinct infuscation around the cells R, 1R1, 2Rs, 3M, 3Cu and usually 2R1, the cell 3R1 not infuscated basally and along anterior margin (
|
||
<figureCitation box="[249,367,331,357]" captionStart="FIGURE 8" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[159,1428,568,1903]" captionTargetId="figure-186@15.[151,1435,561,1909]" captionTargetPageId="15" captionText="FIGURE 8. Wing, female.—A, T. nakanei, holotype of T. kaedei; B, T. nakanei, Tokyo; C, T. nakanei, Suzuka, Mie; D, T. nakanei, Narukawa-keikoku, Ehime; E, F, T.contractus, Okayama-shi, Okayama; G, T. okinawensis, holotype; H, T.okinawensis, Ginama, Okinawa; I, T. okinawensis, holotype of T. kurokivorus (left wings, ventral view); J, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621768" httpUri="https://zenodo.org/record/7621768/files/figure.png" pageId="31" pageNumber="32">Fig. 8G–J</figureCitation>
|
||
), the abdominal tergum 8 pale brown with posterior 1/2–3/4 black, the abdominal tergum 9 usually with black areas along the posterior margin laterally and the short precornal basin (
|
||
<figureCitation box="[1175,1311,367,393]" captionStart="FIGURE 10" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[180,1408,190,1875]" captionTargetId="figure-15@17.[173,1414,181,1884]" captionTargetPageId="17" captionText="FIGURE 10. Posterior part of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, F, T. fuscicornis Inchon, Korea; G, T. fuscicornis, Akigase, Saitama; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621772" httpUri="https://zenodo.org/record/7621772/files/figure.png" pageId="31" pageNumber="32">Fig. 10K, L</figureCitation>
|
||
). The male is characterized by the entirely black head (
|
||
<figureCitation box="[644,735,403,429]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="31" pageNumber="32">Fig. 4H</figureCitation>
|
||
), the antenna black with the narrow apex pale brown, with 13 or 14 antennomeres (
|
||
<figureCitation box="[389,479,438,465]" captionStart="FIGURE 5" captionStartId="12.[151,250,1787,1811]" captionTargetBox="[155,1432,565,1761]" captionTargetId="figure-186@12.[151,1435,561,1764]" captionTargetPageId="12" captionText="FIGURE 5. Antenna, female (A–G, I) and male (H, J–N).—A. T. apicalis, Minamichitose, Hokkaido; B, T. bicinctus, holotype; C, T. longicollis, Toyota, Aichi; D, T. fuscicornis, Inchon, Korea; E, T. nakanei, Ôginosen, Tottori; F, T. contractus, Okayamashi, Okayama; G, T. okinawensis, holotype; H. T. apicalis, Tsukuba, Ibaraki; I, T. katayamai, holotype; J, T. longicollis, Muroto, Kochi; K, T. fuscicornis, Shikotsu-ko, Hokkaido; L, T. nakanei, Noda, Chiba; M, T. contractus, Tatsuno, Hyogo; N, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621761" httpUri="https://zenodo.org/record/7621761/files/figure.png" pageId="31" pageNumber="32">Fig. 5N</figureCitation>
|
||
), the long pronotum (
|
||
<figureCitation box="[725,815,439,465]" captionStart="FIGURE 4" captionStartId="11.[151,250,1227,1251]" captionTargetBox="[158,1429,188,1197]" captionTargetId="figure-15@11.[151,1435,181,1203]" captionTargetPageId="11" captionText="FIGURE 4. Head, thorax and base of abdomen, male.—A, B, T. apicalis, Minamichitose, Hokkaido; C, D, T. fuscicornis, Shikotsu-ko, Hokkaido; E, T. longicollis, Muroto, Kochi; F, T. nakanei, Noda, Chiba; G, T. contractus, Okayama-shi, Okayama; H, T. okinawensis, Amami-oshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621759" httpUri="https://zenodo.org/record/7621759/files/figure.png" pageId="31" pageNumber="32">Fig. 4H</figureCitation>
|
||
), the short hind tibia (
|
||
<figureCitation box="[1065,1170,439,465]" captionStart="FIGURE 12" captionStartId="19.[151,250,1268,1292]" captionTargetBox="[159,1428,186,1239]" captionTargetId="figure-15@19.[151,1435,181,1244]" captionTargetPageId="19" captionText="FIGURE 12. Abdomen, male.—A, F, T. nakanei, Noda, Chiba; B, G, T. contractus, Tatsuno, Hyogo; C, T. apicalis, Tsukuba, Ibaraki; D, T. fuscicornis, Chitose, Hokkaido; E, T. longicollis, Muroto, Kochi; H, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621778" httpUri="https://zenodo.org/record/7621778/files/figure.png" pageId="31" pageNumber="32">Fig. 12H</figureCitation>
|
||
), the dark brown wings with the basal and midapical parts more hyaline (
|
||
<figureCitation box="[701,784,475,501]" captionStart="FIGURE 9" captionStartId="16.[151,250,1266,1290]" captionTargetBox="[158,1429,276,1239]" captionTargetId="figure-20@16.[151,1435,271,1243]" captionTargetPageId="16" captionText="FIGURE 9. Wing, male.—A, T. apicalis, Tsukuba, Ibaraki; B, T. longicollis, Muroto, Kochi; C, T. fuscicornis, Chitose, Hokkaido; D, T. nakanei, Noda, Chiba; E, T. contractus, Tatsuno, Hyogo; F, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621770" httpUri="https://zenodo.org/record/7621770/files/figure.png" pageId="31" pageNumber="32">Fig. 9F</figureCitation>
|
||
) and the black abdomen with pale brown marks (tergum 8 largely pale brown) (
|
||
<figureCitation box="[385,490,511,537]" captionStart="FIGURE 12" captionStartId="19.[151,250,1268,1292]" captionTargetBox="[159,1428,186,1239]" captionTargetId="figure-15@19.[151,1435,181,1244]" captionTargetPageId="19" captionText="FIGURE 12. Abdomen, male.—A, F, T. nakanei, Noda, Chiba; B, G, T. contractus, Tatsuno, Hyogo; C, T. apicalis, Tsukuba, Ibaraki; D, T. fuscicornis, Chitose, Hokkaido; E, T. longicollis, Muroto, Kochi; H, T. okinawensis, Amami-ôshima, Kagoshima." figureDoi="http://doi.org/10.5281/zenodo.7621778" httpUri="https://zenodo.org/record/7621778/files/figure.png" pageId="31" pageNumber="32">Fig. 12H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="31.[151,1437,151,897]" pageId="31" pageNumber="32">
|
||
<bibRefCitation author="Togashi, I." box="[199,369,547,573]" pageId="31" pageNumber="32" pagination="938 - 944" refId="ref29340" refString="Togashi, I. (2006) Two new species of Tremex (Hymenoptera: Siricidae) reared from Castanopsis and Symplocos in Japan. Proceedings of the Entomological Society of Washington, 108, 938 - 944." type="journal article" year="2006">Togashi (2006)</bibRefCitation>
|
||
distinguished
|
||
<taxonomicName authorityName="Shinohara" authorityYear="2023" box="[533,693,547,573]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kurokivorus">
|
||
<emphasis box="[533,693,547,573]" italics="true" pageId="31" pageNumber="32">T. kurokivorus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName authorityName="Togashi" authorityYear="1997" box="[762,926,547,573]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis box="[762,926,547,573]" italics="true" pageId="31" pageNumber="32">T. okinawensis</emphasis>
|
||
</taxonomicName>
|
||
by the differences in the color patterns of the mesoscutum, the fore and mid coxae, and the terga 8 and 9 of the abdomen, the shape of the antennal pedicel, the relative lengths of the flagellomeres 1 and 2, and the form of the precornal basin. These differences were actually between the female
|
||
<typeStatus box="[377,486,655,681]" pageId="31" pageNumber="32">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Shinohara" authorityYear="2023" box="[525,686,655,681]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kurokivorus">
|
||
<emphasis box="[525,686,655,681]" italics="true" pageId="31" pageNumber="32">T. kurokivorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Togashi" authorityYear="1997" box="[741,906,655,681]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis box="[741,906,655,681]" italics="true" pageId="31" pageNumber="32">T. okinawensis</emphasis>
|
||
</taxonomicName>
|
||
, because Togashi examined only the
|
||
<typeStatus box="[1327,1436,655,681]" pageId="31" pageNumber="32">holotypes</typeStatus>
|
||
of the two species. I have examined
|
||
<specimenCount box="[566,694,691,717]" pageId="31" pageNumber="32" type="female">five female</specimenCount>
|
||
specimens, including the
|
||
<typeStatus box="[990,1099,691,717]" pageId="31" pageNumber="32">holotypes</typeStatus>
|
||
of both species. Of these, the
|
||
<typeStatus box="[151,249,727,753]" pageId="31" pageNumber="32">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Togashi" authorityYear="1997" box="[293,460,727,753]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis box="[293,460,727,753]" italics="true" pageId="31" pageNumber="32">T. okinawensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[478,574,727,753]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="31" pageNumber="32">Figs 3K</figureCitation>
|
||
,
|
||
<figureCitation box="[590,626,727,753]" captionStart="FIGURE 8" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[159,1428,568,1903]" captionTargetId="figure-186@15.[151,1435,561,1909]" captionTargetPageId="15" captionText="FIGURE 8. Wing, female.—A, T. nakanei, holotype of T. kaedei; B, T. nakanei, Tokyo; C, T. nakanei, Suzuka, Mie; D, T. nakanei, Narukawa-keikoku, Ehime; E, F, T.contractus, Okayama-shi, Okayama; G, T. okinawensis, holotype; H, T.okinawensis, Ginama, Okinawa; I, T. okinawensis, holotype of T. kurokivorus (left wings, ventral view); J, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621768" httpUri="https://zenodo.org/record/7621768/files/figure.png" pageId="31" pageNumber="32">8G</figureCitation>
|
||
,
|
||
<figureCitation box="[641,691,727,753]" captionStart="FIGURE 10" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[180,1408,190,1875]" captionTargetId="figure-15@17.[173,1414,181,1884]" captionTargetPageId="17" captionText="FIGURE 10. Posterior part of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, F, T. fuscicornis Inchon, Korea; G, T. fuscicornis, Akigase, Saitama; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621772" httpUri="https://zenodo.org/record/7621772/files/figure.png" pageId="31" pageNumber="32">10K</figureCitation>
|
||
) is the palest specimen having the mesoscutum, mesoscutellum, mesopleuron and the fore and mid coxae largely pale brown, whereas these areas are mostly or entirely black in other specimens, including the
|
||
<typeStatus box="[504,602,799,825]" pageId="31" pageNumber="32">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authorityName="Shinohara" authorityYear="2023" box="[641,802,799,825]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kurokivorus">
|
||
<emphasis box="[641,802,799,825]" italics="true" pageId="31" pageNumber="32">T. kurokivorus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[818,909,799,825]" captionStart="FIGURE 3" captionStartId="10.[151,250,1870,1894]" captionTargetBox="[157,1430,190,1838]" captionTargetId="figure-20@10.[151,1435,181,1846]" captionTargetPageId="10" captionText="FIGURE 3. Head, thorax and base of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, T. fuscicornis, Schwedt, Germany, F, G, T. fuscicornis, Inchon, Korea; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621757" httpUri="https://zenodo.org/record/7621757/files/figure.png" pageId="31" pageNumber="32">Figs 3L</figureCitation>
|
||
,
|
||
<figureCitation box="[921,947,799,825]" captionStart="FIGURE 8" captionStartId="15.[151,250,1932,1956]" captionTargetBox="[159,1428,568,1903]" captionTargetId="figure-186@15.[151,1435,561,1909]" captionTargetPageId="15" captionText="FIGURE 8. Wing, female.—A, T. nakanei, holotype of T. kaedei; B, T. nakanei, Tokyo; C, T. nakanei, Suzuka, Mie; D, T. nakanei, Narukawa-keikoku, Ehime; E, F, T.contractus, Okayama-shi, Okayama; G, T. okinawensis, holotype; H, T.okinawensis, Ginama, Okinawa; I, T. okinawensis, holotype of T. kurokivorus (left wings, ventral view); J, T. okinawensis, Aha, Okinawa." figureDoi="http://doi.org/10.5281/zenodo.7621768" httpUri="https://zenodo.org/record/7621768/files/figure.png" pageId="31" pageNumber="32">8I</figureCitation>
|
||
,
|
||
<figureCitation box="[960,1009,799,825]" captionStart="FIGURE 10" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[180,1408,190,1875]" captionTargetId="figure-15@17.[173,1414,181,1884]" captionTargetPageId="17" captionText="FIGURE 10. Posterior part of abdomen, female.—A, T. apicalis, Tsukuba, Ibaraki; B, T. bicinctus, holotype; C, T. longicollis, Okazaki, Aichi; D, T. katayamai, holotype; E, F, T. fuscicornis Inchon, Korea; G, T. fuscicornis, Akigase, Saitama; H, T. nakanei, Ai, Shimane; I, T. nakanei, holotype of T. kaedei; J, T. contractus, Okayama-shi, Okayama; K, T. okinawensis, holotype; L, T. okinawensis, holotype of T. kurokivorus." figureDoi="http://doi.org/10.5281/zenodo.7621772" httpUri="https://zenodo.org/record/7621772/files/figure.png" pageId="31" pageNumber="32">10L</figureCitation>
|
||
). Otherwise, the
|
||
<specimenCount box="[1201,1370,799,825]" pageId="31" pageNumber="32" type="generic">five specimens</specimenCount>
|
||
show no significant differences and here I propose to treat
|
||
<taxonomicName authority="Togashi, 2006" authorityName="Togashi" authorityYear="2006" box="[743,1071,835,861]" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kurokivorus">
|
||
<emphasis box="[743,903,835,861]" italics="true" pageId="31" pageNumber="32">T. kurokivorus</emphasis>
|
||
<bibRefCitation author="Togashi, I." box="[910,1071,835,861]" pageId="31" pageNumber="32" pagination="938 - 944" refId="ref29340" refString="Togashi, I. (2006) Two new species of Tremex (Hymenoptera: Siricidae) reared from Castanopsis and Symplocos in Japan. Proceedings of the Entomological Society of Washington, 108, 938 - 944." type="journal article" year="2006">Togashi, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, as a synonym of
|
||
<taxonomicName authority="Togashi, 1997" authorityName="Togashi" authorityYear="1997" class="Insecta" family="Siricidae" genus="Tremex" kingdom="Animalia" order="Hymenoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="okinawensis">
|
||
<emphasis box="[1272,1436,835,861]" italics="true" pageId="31" pageNumber="32">T. okinawensis</emphasis>
|
||
<bibRefCitation author="Togashi, I." box="[151,312,871,897]" pageId="31" pageNumber="32" pagination="1 - 6" refId="ref29266" refString="Togashi, I. (1997) Two new species of Tremex Jurine (Hymenoptera: Siricidae) from Japan. Species Diversity, 2, 1 - 6. https: // doi. org / 10.12782 / specdiv. 2.1" type="journal article" year="1997">Togashi, 1997</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |