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<document id="9CE82CD87BFE67D06D1B3FBD56779B60" ID-DOI="10.5281/zenodo.5393630" ID-ISSN="1638-9387" ID-Zenodo-Dep="5393630" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630319175263" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="B36487D8FFD2D52BD2E2FDA3FB81FA9E" docLanguage="en" docName="Zoosystema.2007.29.3.417-456.pdf" docOrigin="Zoosystema 29 (3)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus costulifer Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="436" masterDocId="4F5DFFA0FFC1D538D20EFFF0FFA5FC3D" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group" masterLastPageNumber="456" masterPageNumber="417" pageNumber="436" updateTime="1699225420998" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="824DA83BB5E0F7004AE1565F60CF7220" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3BC616D6CD18E26B8C79B9D88013862A">
<mods:title id="EF0949F02B93FDD8E6A953DE4BB2A396">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 2: the javanicus-group</mods:title>
</mods:titleInfo>
<mods:name id="D6B15AC10726F294D9D9F13A6ABA8DE1" type="personal">
<mods:role id="AC5E7B0F98C6986C5D9F8FC5C3E316FF">
<mods:roleTerm id="6549276F5FED53B10EF8CC39BB23F7BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CC48B6ED89F1489DDA420F61FFB29679">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="227C254FFD7417D75C68CF5919C9C08D">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="1636BE36BA1FE2F080CE56DFDB70D9CE" type="email">sgol@orc.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="0DD400BE35588B1033F609DF14292D13" type="personal">
<mods:role id="FA8BAC871CB8772E5CEF0BCD6B54D7BB">
<mods:roleTerm id="A5736A236F50B293B1F3816E118B4175">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AF46EA602F15DF7666DF651D0D6330E9">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="49A6A8F104024CF4265A09BC444754F4">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="06D2B8640ACA7AC240DCF5CB045A8996" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="FB2A1E6EB11778427EE4B562C09736D1" type="personal">
<mods:role id="D1F050F1F1C5D7E78E129F0DE49FE6FB">
<mods:roleTerm id="650955557B91631D92F94CB321E6996A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DDD0DD094D22560203D6B71C5C34655B">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="63247B4C196CADC1928477B267A6B886">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="DF4AB99357CEFF599657B1921EADDBCC" type="email">collmill@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="7658612C3B79600518B78816F2AB6FD7" type="personal">
<mods:role id="DC33C9EFEDE823B761A8D656FE3042B6">
<mods:roleTerm id="D7535F2050899944A39E1063CC0536B3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3E5D4D839E577333031D4247A6FF202">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="18FDB9909AD05F4E71ACB51D9E67B5FB">Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
</mods:name>
<mods:typeOfResource id="27DE28A33880BA73BF843B9E4EB10BFE">text</mods:typeOfResource>
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<mods:title id="D1753E42C11D432CD5476565430D1BFC">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="D2317EDB5A9D6D3CF05EDA61EB5B9F7B">
<mods:date id="564D2AAB36EC882ED9BBDF970E7AE44C">2007</mods:date>
<mods:detail id="7A01F49728DA2F63F5D26C76B881E971" type="volume">
<mods:number id="9A4F978D9030EECC1369DC64A99F5B74">29</mods:number>
</mods:detail>
<mods:detail id="43492593F6576935D252C00D0C491DB3" type="issue">
<mods:number id="81BC5BAE868275CA318EDAA52212FDEB">3</mods:number>
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<mods:start id="1A34011CDEEBFDBFE88F6DE71C23BB77">417</mods:start>
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<mods:classification id="86A7DBF07B24FCA8B4BC922229E19080">journal article</mods:classification>
<mods:identifier id="09EDD45A3087518CCAFEE6AADE427B4E" type="DOI">10.5281/zenodo.5393630</mods:identifier>
<mods:identifier id="02E389D5851D78680AF0A18B1C9561A0" type="ISSN">1638-9387</mods:identifier>
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</mods:mods>
<treatment id="B36487D8FFD2D52BD2E2FDA3FB81FA9E" LSID="urn:lsid:plazi:treatment:B36487D8FFD2D52BD2E2FDA3FB81FA9E" httpUri="http://treatment.plazi.org/id/B36487D8FFD2D52BD2E2FDA3FB81FA9E" lastPageNumber="436" pageId="19" pageNumber="436">
<subSubSection id="73D76545FFD2D52BD2E2FDA3FDA9FE52" box="[236,524,595,623]" pageId="19" pageNumber="436" type="nomenclature">
<paragraph id="3B7236CEFFD2D52BD2E2FDA3FDA9FE52" blockId="19.[236,524,595,655]" box="[236,524,595,623]" pageId="19" pageNumber="436">
<heading id="603A81A2FFD2D52BD2E2FDA3FDA9FE52" box="[236,524,595,623]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="436" reason="2">
<taxonomicName id="FCCD4D4DFFD2D52BD2E2FDA3FE6EFE53" ID-CoL="3GGJN" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[236,459,595,622]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="436" phylum="Arthropoda" rank="species" species="costulifer" status="sp. nov.">
<emphasis id="09B9EADCFFD2D52BD2E2FDA3FE6EFE53" bold="true" box="[236,459,595,622]" italics="true" pageId="19" pageNumber="436">Glyphiulus costulifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="128A57A7FFD2D52BD3DFFDA5FDA9FE52" box="[465,524,597,623]" pageId="19" pageNumber="436" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD2D52BD337FD84FE1AFEB2" box="[313,447,628,655]" pageId="19" pageNumber="436" type="description">
<paragraph id="3B7236CEFFD2D52BD337FD84FE1AFEB2" blockId="19.[236,524,595,655]" box="[313,447,628,655]" pageId="19" pageNumber="436">
(
<figureCitation id="A3F62A4BFFD2D52BD34FFD84FE13FEB2" box="[321,438,628,655]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="20.[113,124,1657,1674]" captionStartId-1="21.[113,124,1635,1652]" captionStartId-2="22.[113,124,1149,1166]" captionTargetBox-0="[117,1206,216,1545]" captionTargetBox-1="[259,1063,215,1592]" captionTargetBox-2="[116,1207,225,1110]" captionTargetId-0="figure-120@20.[641,729,688,1087]" captionTargetId-1="figure-148@21.[629,890,854,1037]" captionTargetId-2="figure-24@22.[94,1229,215,1123]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." captionText-1="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." captionText-2="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5393660" figureDoi-1="http://doi.org/10.5281/zenodo.5393664" figureDoi-2="http://doi.org/10.5281/zenodo.5393666" httpUri-0="https://zenodo.org/record/5393660/files/figure.png" httpUri-1="https://zenodo.org/record/5393664/files/figure.png" httpUri-2="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">Figs 14-16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD2D52BD27FFD5DFDECFF74" pageId="19" pageNumber="436" type="materials_examined">
<paragraph id="3B7236CEFFD2D52BD27FFD5DFDECFF74" blockId="19.[113,647,685,841]" pageId="19" pageNumber="436">
<materialsCitation id="8BA53C93FFD2D52BD27FFD5DFD85FF29" collectingDate="2000-12-29" collectionCode="TYPE, MATERIAL, MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="Laos" county="Nong Kiaw" latitude="20.72" location="Tham Pha Kouang Cave" longLatPrecision="760" longitude="102.68" municipality="Muang Ngoy" pageId="19" pageNumber="436" specimenCode="GA 049" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang" typeStatus="holotype">
<collectionCode id="5DDCAE0BFFD2D52BD27FFD5DFEA3FEFE" box="[113,262,685,708]" pageId="19" pageNumber="436">TYPE MATERIAL</collectionCode>
. —
<emphasis id="09B9EADCFFD2D52BD338FD5DFECEFEF8" bold="true" box="[310,363,685,709]" pageId="19" pageNumber="436">
<collectingCountry id="43DA765EFFD2D52BD338FD5DFEC3FEF8" box="[310,358,685,709]" name="Laos" pageId="19" pageNumber="436">Laos</collectingCountry>
.
</emphasis>
<collectingRegion id="F909F82CFFD2D52BD37DFD5DFDE1FEF9" box="[371,580,685,709]" country="Laos" name="Louangphabang" pageId="19" pageNumber="436">Luang Prabang Prov.</collectingRegion>
,
<collectingCounty id="D2134E42FFD2D52BD040FD5DFF07FEE2" pageId="19" pageNumber="436">Nong Kiaw</collectingCounty>
(
<collectingMunicipality id="DB16ACB4FFD2D52BD2A3FD37FE8BFEE2" box="[173,302,711,735]" pageId="19" pageNumber="436">Muang Ngoy</collectingMunicipality>
),
<location id="3E126015FFD2D52BD332FD37FDBAFEE2" LSID="urn:lsid:plazi:treatment:B36487D8FFD2D52BD2E2FDA3FB81FA9E:3E126015FFD2D52BD332FD37FDBAFEE2" box="[316,543,711,736]" country="Laos" county="Nong Kiaw" latitude="20.72" longLatPrecision="760" longitude="102.68" municipality="Muang Ngoy" name="Tham Pha Kouang Cave" pageId="19" pageNumber="436" stateProvince="Louangphabang">Tham Pha Kouang Cave</location>
,
<geoCoordinate id="5EF95009FFD2D52BD029FD37FD27FEE2" box="[551,642,711,735]" degrees="102.68" direction="east" orientation="longitude" pageId="19" pageNumber="436" precision="555" value="102.68">102.68°E</geoCoordinate>
,
<geoCoordinate id="5EF95009FFD2D52BD27FFD12FF6DFEC7" box="[113,200,738,762]" degrees="20.72" direction="north" orientation="latitude" pageId="19" pageNumber="436" precision="555" value="20.72">20.72°N</geoCoordinate>
,
<date id="4F73100EFFD2D52BD2DDFD12FEF5FEC7" box="[211,336,738,762]" pageId="19" pageNumber="436" value="2000-12-29">
<collectingDate id="5F37E9E6FFD2D52BD2DDFD12FEF5FEC7" box="[211,336,738,762]" pageId="19" pageNumber="436" value="2000-12-29">29.XII.2000</collectingDate>
</date>
, leg.
<collectorName id="96385318FFD2D52BD38AFD12FE78FEC7" box="[388,477,738,762]" pageId="19" pageNumber="436">A. Bedos</collectorName>
&amp;
<collectorName id="96385318FFD2D52BD3F1FD12FD23FEC7" box="[511,646,738,762]" pageId="19" pageNumber="436">L. Deharveng</collectorName>
(LAO-099),
<typeStatus id="E476886CFFD2D52BD2E0FD0CFEE1FF29" box="[238,324,764,788]" pageId="19" pageNumber="436" type="holotype">holotype</typeStatus>
<specimenCount id="2DCBFD47FFD2D52BD345FCF1FEFAFF2E" box="[331,351,769,787]" pageId="19" pageNumber="436" type="male">♂</specimenCount>
(
<collectionCode id="5DDCAE0BFFD2D52BD362FD0CFE65FF29" box="[364,448,764,788]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="436">MNHN</collectionCode>
<specimenCode id="6B6B9EB5FFD2D52BD3C9FD0CFDBCFF29" box="[455,537,764,788]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="436">GA 049</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD2D52BD025FD0CFE09FF12" collectingDate="2000-12-29" collectionCode="MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="Laos" county="Nong Kiaw" isTailEnumeration="true" latitude="20.72" location="Tham Pha Kouang Cave" longLatPrecision="760" longitude="102.68" municipality="Muang Ngoy" pageId="19" pageNumber="436" specimenCode="GA 049" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Louangphabang" typeStatus="paratype">
<typeStatus id="E476886CFFD2D52BD025FD0CFD22FF29" box="[555,647,764,788]" pageId="19" pageNumber="436" type="paratype">paratypes</typeStatus>
<specimenCount id="2DCBFD47FFD2D52BD27FFCE7FF09FF13" box="[113,172,791,815]" pageId="19" pageNumber="436" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="2DCBFD47FFD2D52BD2B6FCE7FF48FF13" box="[184,237,791,815]" pageId="19" pageNumber="436" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="5DDCAE0BFFD2D52BD2F4FCE7FEEBFF12" box="[250,334,791,815]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="436">MNHN</collectionCode>
<specimenCode id="6B6B9EB5FFD2D52BD35DFCE7FE00FF12" box="[339,421,791,815]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="19" pageNumber="436">GA 049</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD2D52BD3B8FCE7FD82FF12" box="[438,551,791,815]" collectionCode="SEM" country="Laos" pageId="19" pageNumber="436" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2DCBFD47FFD2D52BD3B8FCE7FE79FF13" box="[438,476,791,815]" pageId="19" pageNumber="436" type="male">1 ♂</specimenCount>
(
<collectionCode id="5DDCAE0BFFD2D52BD3E7FCE7FD85FF12" box="[489,544,791,815]" pageId="19" pageNumber="436">SEM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD2D52BD03EFCE7FF70FF74" collectionCode="ZMUM" country="Laos" pageId="19" pageNumber="436" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2DCBFD47FFD2D52BD03EFCE7FDF2FF13" box="[560,599,791,815]" pageId="19" pageNumber="436" type="male">1 ♂</specimenCount>
,
<specimenCount id="2DCBFD47FFD2D52BD06DFCE7FD23FF13" box="[611,646,791,815]" pageId="19" pageNumber="436" type="female">1 ♀</specimenCount>
(
<collectionCode id="5DDCAE0BFFD2D52BD277FCC1FF6AFF74" box="[121,207,817,841]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="19" pageNumber="436">ZMUM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD2D52BD2EEFCC1FE39FF74" box="[224,412,817,841]" collectionCode="ZMUC" country="Laos" pageId="19" pageNumber="436" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="2DCBFD47FFD2D52BD2EEFCC1FEA2FF75" box="[224,263,817,841]" pageId="19" pageNumber="436" type="male">1 ♂</specimenCount>
,
<specimenCount id="2DCBFD47FFD2D52BD31DFCC1FE92FF75" box="[275,311,817,841]" pageId="19" pageNumber="436" type="female">1 ♀</specimenCount>
(
<collectionCode id="5DDCAE0BFFD2D52BD34BFCC1FE30FF74" box="[325,405,817,841]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="19" pageNumber="436">ZMUC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="8BA53C93FFD2D52BD3A9FCC1FDE1FF74" box="[423,580,817,841]" collectionCode="NMNHS" country="Laos" pageId="19" pageNumber="436" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="2DCBFD47FFD2D52BD3A9FCC1FE6EFF75" box="[423,459,817,841]" pageId="19" pageNumber="436" type="female">1 ♀</specimenCount>
(
<collectionCode id="5DDCAE0BFFD2D52BD3D7FCC1FD98FF74" box="[473,573,817,841]" pageId="19" pageNumber="436">NMNHS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD2D52BD27FFC91FF18FFAE" pageId="19" pageNumber="436" type="etymology">
<paragraph id="3B7236CEFFD2D52BD27FFC91FF18FFAE" blockId="19.[113,646,864,915]" pageId="19" pageNumber="436">ETYMOLOGY. — To emphasize the normally carinate collum.</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD2D52BD27FFC5BFDCCF811" pageId="19" pageNumber="436" type="diagnosis">
<paragraph id="3B7236CEFFD2D52BD27FFC5BFDCCF811" blockId="19.[113,647,939,1069]" pageId="19" pageNumber="436">DIAGNOSIS. — Differs from congeners by the nearly typical carinotaxy pattern of the collum, coupled with highly unusual traces of both ventrolateral crests, the very little modified male coxae 1, the long but bare flagellum of the posterior gonopods, etc. (see also key below).</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD2D52BD27FFBBFFCE4F97A" pageId="19" pageNumber="436" type="description">
<paragraph id="3B7236CEFFD2D52BD27FFBBFFEA7F85A" blockId="19.[113,647,1103,1700]" box="[113,258,1103,1129]" pageId="19" pageNumber="436">DESCRIPTION</paragraph>
<paragraph id="3B7236CEFFD2D52BD27FFB9EFDABF8F5" blockId="19.[113,647,1103,1700]" pageId="19" pageNumber="436">
<materialsCitation id="8BA53C93FFD2D52BD27FFB9EFDABF8F5" collectionCode="T" pageId="19" pageNumber="436" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="E476886CFFD2D52BD27FFB9EFF72F8B5" box="[113,215,1134,1160]" pageId="19" pageNumber="436" type="holotype">Holotype</typeStatus>
with 56p+1a+
<collectionCode id="5DDCAE0BFFD2D52BD37AFB9FFE26F8B4" box="[372,387,1135,1161]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="436">T</collectionCode>
, about
<quantity id="FC359B2BFFD2D52BD3DAFB9FFD87F8B4" box="[468,546,1135,1161]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="436" unit="mm" value="28.0">28 mm</quantity>
long and
<quantity id="FC359B2BFFD2D52BD27FFB7EFF64F894" box="[113,193,1166,1193]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="19" pageNumber="436" unit="mm" value="1.8">1.8 mm</quantity>
wide;
<typeStatus id="E476886CFFD2D52BD30FFB7FFEC5F894" box="[257,352,1167,1193]" pageId="19" pageNumber="436" type="paratype">paratypes</typeStatus>
(
<specimenCount id="2DCBFD47FFD2D52BD363FB64FE26F895" box="[365,387,1172,1192]" pageId="19" pageNumber="436" type="male">♂</specimenCount>
,
<specimenCount id="2DCBFD47FFD2D52BD383FB64FE05F895" box="[397,416,1172,1192]" pageId="19" pageNumber="436" type="female">♀</specimenCount>
) with 48-62p+3-1a+
<collectionCode id="5DDCAE0BFFD2D52BD07FFB7FFDDAF894" box="[625,639,1167,1193]" name="Tavera, Department of Geology and Geophysics" pageId="19" pageNumber="436">T</collectionCode>
, length
<quantity id="FC359B2BFFD2D52BD2B3FB5EFE91F8F5" box="[189,308,1198,1225]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" metricValueMax="3.5" metricValueMin="2.3" pageId="19" pageNumber="436" unit="mm" value="29.0" valueMax="35.0" valueMin="23.0">23-35 mm</quantity>
, width
<quantity id="FC359B2BFFD2D52BD38BFB5EFDAEF8F5" box="[389,523,1198,1225]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" metricValueMax="1.9" metricValueMin="1.5" pageId="19" pageNumber="436" unit="mm" value="1.7" valueMax="1.9" valueMin="1.5">1.5-1.9 mm</quantity>
.
</materialsCitation>
</paragraph>
<paragraph id="3B7236CEFFD2D52BD286FB3DFEE3F9BA" blockId="19.[113,647,1103,1700]" pageId="19" pageNumber="436">
Coloration light yellow-brown to red-brown, tergal crests often dark castaneous brown. Greyish to red-brown, subtriangular eye patches with 7-9 ocelli each. Antennae usual (
<figureCitation id="A3F62A4BFFD2D52BD3D9FADDFD99F97A" box="[471,572,1325,1351]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Figs 14A</figureCitation>
;
<figureCitation id="A3F62A4BFFD2D52BD047FADCFDDFF97A" box="[585,634,1324,1351]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">16A</figureCitation>
), gnathochilarium (mentum divided, n = 3) oligotrichous (
<figureCitation id="A3F62A4BFFD2D52BD2D6FA9CFE9CF9BA" box="[216,313,1388,1415]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">Fig. 16B</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD286FA7CFD90FA38" blockId="19.[113,647,1103,1700]" pageId="19" pageNumber="436">
Head width = segment 2 = 6 &lt;collum = 7(8)&gt; segment 3 = 5 &lt;7 &lt;8 &lt;9 &lt;10 &lt;11 = midbody segments; body gradually tapering toward telson. Postcollar constriction evident (
<figureCitation id="A3F62A4BFFD2D52BD3C8FA1BFD8DFA38" box="[454,552,1515,1541]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14B</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD286F9FAFBD7FD4C" blockId="19.[113,647,1103,1700]" lastBlockId="19.[677,1210,216,1351]" pageId="19" pageNumber="436">
Collum almost typically crested, only median crest (ma) unusually abbreviated; carinotaxy pattern typical of the
<taxonomicName id="FCCD4D4DFFD2D52BD2A1F9BAFEB0FA59" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[175,277,1610,1636]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="436" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="09B9EADCFFD2D52BD2A1F9BAFEB0FA59" box="[175,277,1610,1636]" italics="true" pageId="19" pageNumber="436">granulatus</emphasis>
</taxonomicName>
-group (I-VI+7a+pc+ma+pc+7a+VI- I) (
<figureCitation id="A3F62A4BFFD2D52BD29AF999FEBDFAB9" box="[148,280,1641,1668]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14A, B</figureCitation>
). Subsequent metaterga evidently crested as well (
<figureCitation id="A3F62A4BFFD2D52BD307F979FEEBFA9E" box="[265,334,1673,1699]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14</figureCitation>
A-E).Carinotaxy formula 2+I/ i+3/3+I/i+2, i.e. all dorsal crests subdivided (
<figureCitation id="A3F62A4BFFD2D52BD657FF28FD19FD2F" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14C, D</figureCitation>
), while both ventrolateral crests undivided, barely traceable as ribbons of micropunctate/shagreened texture (
<figureCitation id="A3F62A4BFFD2D52BD0F2FEC7FCFEFD6F" box="[764,859,311,338]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="19" pageNumber="436">Fig. 15B</figureCitation>
). Ozoporiferous tubercles round, only slightly wider than high (
<figureCitation id="A3F62A4BFFD2D52BD1E8FEA6FBC0FD4C" box="[998,1125,342,369]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="19" pageNumber="436">Fig. 15A, B</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD0B5FE87FC1FFF11" blockId="19.[677,1210,216,1351]" pageId="19" pageNumber="436">
Tegument delicately alveolate-areolate, dull throughout. Fine longitudinal striations in front of stricture between pro- and metazona ventrolaterally, laterally and dorsally, remaining surface of prozona very delicately shagreened. Segments round in crosssection (
<figureCitation id="A3F62A4BFFD2D52BD0F2FDE5FCFEFE12" box="[764,859,532,559]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="19" pageNumber="436">Fig. 15A</figureCitation>
). Epiproct with a distinct, humplike ridge dorsally in distal 3/4 extent (
<figureCitation id="A3F62A4BFFD2D52BD63DFDC4FB37FE73" box="[1075,1170,564,590]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14E</figureCitation>
), in ventral view rounded and subtriangular in caudal part (
<figureCitation id="A3F62A4BFFD2D52BD0D3FD84FC9FFEB3" box="[733,826,628,654]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14F</figureCitation>
). Paraprocts flattened ventrally, with only a shallow but evident submarginal depression beside caudal edge, each with a row of about a dozen setae at medial margin (
<figureCitation id="A3F62A4BFFD2D52BD191FD23FC5BFED0" box="[927,1022,723,749]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14F</figureCitation>
). Hypoproct sublanceolate, very delicately and broadly emarginate caudomedially (
<figureCitation id="A3F62A4BFFD2D52BD141FCE2FC0BFF11" box="[847,942,786,812]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14F</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD0B5FCC2FC33FFB1" blockId="19.[677,1210,216,1351]" pageId="19" pageNumber="436">
Ventral flaps behind gonopod opening on male segment 7 distinguishable as a low but clear transverse ridge (
<figureCitation id="A3F62A4BFFD2D52BD12BFC81FC2DFFB1" box="[805,904,881,908]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="19" pageNumber="436">Fig. 15C</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD0B5FC61FC90F836" blockId="19.[677,1210,216,1351]" pageId="19" pageNumber="436">
Legs long, those on midbody segments about as long as segment diameter (
<figureCitation id="A3F62A4BFFD2D52BD1F2FC40FBCDFFF6" box="[1020,1128,944,971]" captionStart="FIG" captionStartId="20.[113,124,1657,1674]" captionTargetBox="[117,1206,216,1545]" captionTargetId="figure-120@20.[641,729,688,1087]" captionTargetPageId="20" captionText="FIG. 14. — Glyphiulus costulifer n. sp., paratype ♂: A, anterior part of body, lateral view; B, same, dorsal view; C, midbody segments, lateral view; D, same, dorsal view; E, caudal body end, lateral view; F, same, ventral view. Scale bars: A, E, F, 0.2 mm; B-D, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5393660" httpUri="https://zenodo.org/record/5393660/files/figure.png" pageId="19" pageNumber="436">Fig. 14C</figureCitation>
). Claw with a short but evident accessory spine at base (
<figureCitation id="A3F62A4BFFD2D52BD0A2FC00FD5DF836" box="[684,760,1008,1035]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">Fig. 16</figureCitation>
C-F).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD0B5FBFFFC3DF894" blockId="19.[677,1210,216,1351]" pageId="19" pageNumber="436">
Male legs 1 highly characteristic (
<figureCitation id="A3F62A4BFFD2D52BD620FBFFFB37F817" box="[1070,1170,1039,1066]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">Fig. 16C</figureCitation>
) in showing only slightly modified coxae with two low, paramedian, contiguous cones; telopodites 5-segmented, normal. Male legs 2 and 3 modified as usual (
<figureCitation id="A3F62A4BFFD2D52BD109FB7EFC2EF894" box="[775,907,1166,1193]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">Fig. 16D, E</figureCitation>
).
</paragraph>
<paragraph id="3B7236CEFFD2D52BD0B5FB5EFCE4F97A" blockId="19.[677,1210,216,1351]" pageId="19" pageNumber="436">
Anterior gonopods (
<figureCitation id="A3F62A4BFFD2D52BD185FB5EFC4BF8F4" box="[907,1006,1198,1225]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="19" pageNumber="436">Figs 15D</figureCitation>
;
<figureCitation id="A3F62A4BFFD2D52BD1F9FB5EFB82F8F4" box="[1015,1063,1198,1225]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">16G</figureCitation>
) not notched, paramedian processes (d) high, telopodites evidently curved. Posterior gonopods (
<figureCitation id="A3F62A4BFFD2D52BD1E6FB1DFBCAF935" box="[1000,1135,1261,1288]" captionStart="FIG" captionStartId="21.[113,124,1635,1652]" captionTargetBox="[259,1063,215,1592]" captionTargetId="figure-148@21.[629,890,854,1037]" captionTargetPageId="21" captionText="FIG. 15. — Glyphiulus costulifer n. sp., paratype ♂: A, midbody segment section; B, midbody metazonite enlarged, lateral view; C, segment 7, frontolateral view;D, anterior gonopod,caudal view;E, posterior gonopod,caudal view; F, posterior gonopod,enlarged caudal view. Scale bars: A, C, 0.2 mm; B, D, E, 0.1 mm; F, 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5393664" httpUri="https://zenodo.org/record/5393664/files/figure.png" pageId="19" pageNumber="436">Figs 15E, F</figureCitation>
;
<figureCitation id="A3F62A4BFFD2D52BD672FB1DFB15F935" box="[1148,1200,1261,1288]" captionStart="FIG" captionStartId="22.[113,124,1149,1166]" captionTargetBox="[116,1207,225,1110]" captionTargetId="figure-24@22.[94,1229,215,1123]" captionTargetPageId="22" captionText="FIG. 16. — Glyphiulus costulifer n. sp., paratype ♂: A, antenna; B, gnathochilarium; C, leg 1, frontal view; D, leg 2; E, leg 3; F, leg 13; G, anterior gonopods, caudal view; H, posterior gonopods, caudal view. Abbreviations: d, digitiform process; f, flagellum. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5393666" httpUri="https://zenodo.org/record/5393666/files/figure.png" pageId="19" pageNumber="436">16H</figureCitation>
) highly compact, typical of the group, flagella (f) high and bare.
</paragraph>
</subSubSection>
<subSubSection id="73D76545FFD2D52BD0ABFA9DFB81FA9E" pageId="19" pageNumber="436" type="discussion">
<paragraph id="3B7236CEFFD2D52BD0ABFA9DFCACF9B8" blockId="19.[677,1210,1389,1700]" box="[677,777,1389,1415]" pageId="19" pageNumber="436">REMARKS</paragraph>
<paragraph id="3B7236CEFFD2D52BD0ABFA7CFB3CF9FB" blockId="19.[677,1210,1389,1700]" pageId="19" pageNumber="436">Due to the nearly unmodified male legs 1, this species is probably among the basalmost in the genus.</paragraph>
<paragraph id="3B7236CEFFD2D52BD0B5FA3CFB81FA9E" blockId="19.[677,1210,1389,1700]" pageId="19" pageNumber="436">
In the absence of any troglomorphic traits, this species can only be considered troglophilic at most. This cave is known to support another congener,
<taxonomicName id="FCCD4D4DFFD2D52BD0ABF9DAFCD7FA59" authority="Golovatch, Geoffroy, Mauries &amp; Van den Spiegel, 2007" authorityName="Golovatch, Geoffroy, Mauries &amp; Van den Spiegel" authorityYear="2007" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="19" pageNumber="436" phylum="Arthropoda" rank="species" species="bedosae">
<emphasis id="09B9EADCFFD2D52BD0ABF9DAFCB3FA79" box="[677,790,1578,1604]" italics="true" pageId="19" pageNumber="436">G. bedosae</emphasis>
<bibRefCitation id="5F5C4B3FFFD2D52BD111F9DAFCD7FA59" author="GOLOVATCH S. I. &amp; GEOFFROY J. - J. &amp; MAURIES J. - P. &amp; VAN DEN SPIEGEL D." pageId="19" pageNumber="436" pagination="7 - 49" refId="ref17405" refString="GOLOVATCH S. I., GEOFFROY J. - J., MAURIES J. - P. &amp; VAN DEN SPIEGEL D. 2007. - Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatusgroup. Zoosystema 29 (1): 7 - 49." type="journal article" year="2007">Golovatch, Geoffroy, Mauriès &amp; Van den Spiegel, 2007</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="5F5C4B3FFFD2D52BD1A1F9BAFB0BFA59" author="GOLOVATCH S. I. &amp; GEOFFROY J. - J. &amp; MAURIES J. - P. &amp; VAN DEN SPIEGEL D." box="[943,1198,1610,1636]" pageId="19" pageNumber="436" pagination="7 - 49" refId="ref17405" refString="GOLOVATCH S. I., GEOFFROY J. - J., MAURIES J. - P. &amp; VAN DEN SPIEGEL D. 2007. - Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatusgroup. Zoosystema 29 (1): 7 - 49." type="journal article" year="2007">
Golovatch
<emphasis id="09B9EADCFFD2D52BD623F9BBFBC0FA59" box="[1069,1125,1610,1636]" italics="true" pageId="19" pageNumber="436">et al.</emphasis>
2007
</bibRefCitation>
). Interestingly, both species share the same, rather conspicuous structure of the telson.
</paragraph>
</subSubSection>
</treatment>
</document>
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