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<document id="E1C6F439CEE89CD0F76704A0FA7F2F58" ID-CLB-Dataset="27325" ID-DOI="10.11646/zootaxa.4585.2.4" ID-GBIF-Dataset="e1aa7494-63bf-4abd-9117-47f93fafd6bc" ID-ISSN="1175-5326" ID-Zenodo-Dep="2637340" ID-ZooBank="BF11D9BF-8C33-496F-AD6F-D5AC2A10020D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555050002028" checkinUser="plazi" docAuthor="Polhemus, Dan A., Molano, Fredy, Morales, Irina, Moreira, Felipe Ferraz Figueiredo &amp; Floriano, Carla Fernanda Burguez" docDate="2019" docId="8C1D87B02854E962FF295EE2FE454BAC" docLanguage="en" docName="zootaxa.4585.2.4.pdf" docOrigin="Zootaxa 4585 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Altavelia Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano 2019, gen. n." docType="treatment" docVersion="15" lastPageNumber="300" masterDocId="7024FFC82855E967FFBE5B25FF974F0C" masterDocTitle="Altavelia, a new genus of Neotropical Veliinae (Heteroptera: Gerromorpha: Veliidae), with a key to species and descriptions of four new species from Colombia" masterLastPageNumber="314" masterPageNumber="295" pageNumber="296" updateTime="1698719726503" updateUser="ExternalLinkService">
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<mods:title id="67CDBCC23696703BCB5AB99FAEF9DD68">Altavelia, a new genus of Neotropical Veliinae (Heteroptera: Gerromorpha: Veliidae), with a key to species and descriptions of four new species from Colombia</mods:title>
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<mods:namePart id="6378E49B81252410484C603AFEB4F4D0">Moreira, Felipe Ferraz Figueiredo</mods:namePart>
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<taxonomicName id="C3B44D252854E966FF295EE2FD834AEE" authority="Polhemus &amp; Moreira" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[151,532,1479,1506]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="296" phylum="Arthropoda" rank="genus" status="gen. nov.">
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Polhemus &amp; Moreira
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, 
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<figureCitation id="9C8F2A232854E966FF295ECFFE9A490F" box="[151,269,1514,1539]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="2.[151,250,1963,1985]" captionStartId-1="3.[151,250,1960,1982]" captionStartId-10="17.[151,250,1569,1591]" captionStartId-2="4.[151,250,1755,1777]" captionStartId-3="6.[151,250,1103,1125]" captionStartId-4="8.[151,250,1052,1074]" captionStartId-5="9.[151,250,1788,1810]" captionStartId-6="10.[151,250,1629,1651]" captionStartId-7="14.[151,250,1961,1983]" captionStartId-8="15.[151,250,1880,1902]" captionStartId-9="16.[151,250,1514,1536]" captionTargetBox-0="[156,1421,488,1876]" captionTargetBox-1="[155,1432,223,1902]" captionTargetBox-10="[151,1436,185,1551]" captionTargetBox-2="[177,1422,887,1728]" captionTargetBox-3="[159,1429,193,1074]" captionTargetBox-4="[160,1427,435,1031]" captionTargetBox-5="[149,1436,337,1668]" captionTargetBox-6="[158,1429,1021,1607]" captionTargetBox-7="[164,1428,193,1946]" captionTargetBox-8="[150,1436,193,1858]" captionTargetBox-9="[160,1426,903,1489]" captionTargetId-0="figure@2.[151,1435,488,1939]" captionTargetId-1="figure@3.[155,1432,223,1906]" captionTargetId-10="figure@17.[151,1436,185,1551]" captionTargetId-2="figure@4.[151,1436,877,1734]" captionTargetId-3="figure@6.[156,1430,193,1081]" captionTargetId-4="figure@8.[160,1427,435,1031]" captionTargetId-5="figure@9.[149,1437,337,1767]" captionTargetId-6="figure@10.[157,1430,1021,1607]" captionTargetId-7="figure@14.[164,1428,193,1947]" captionTargetId-8="figure@15.[150,1436,193,1859]" captionTargetId-9="figure@16.[160,1426,903,1492]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-10="17" captionTargetPageId-2="4" captionTargetPageId-3="6" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="14" captionTargetPageId-8="15" captionTargetPageId-9="16" captionText-0="FIGURE 1. A. Altavelia altoandina, male holotype, dorsal view. B. Altavelia altoandina, female paratype, dorsal view. C. Altavelia antioquia, male paratype, dorsal view. D. Altavelia antioquia, female paratype, dorsal view." captionText-1="FIGURE 2. A. Altavelia montana, male paratype, dorsal view. B. Altavelia montana, female paratype, dorsal view. C. Altavelia quindiana, female holotype, dorsal view." captionText-10="FIGURE 11. A Altavelia willei, male, dorsal view. B. Altavelia willei, male, dorsal view. C. Altavelia willei, male, apex of abdomen, lateral view. D. Altavelia willei, male, dorsal view, apex of abdomen (Figs. 11A, C and D, specimen from Peru, Dept. Ancash, as listed in D. Polhemus 2014)." captionText-2="FIGURE 3. A. Altavelia montana, thorax and base of abdomen, ventral view; white arrows show position of central tubercles. B. Altavelia flavomarginata, female, apex of thorax and abdomen, dorsal view." captionText-3="FIGURE 4. A. Altavelia altoandina, male proctiger, dorsal view. B. Altavelia antioquia, male proctiger, dorsal view. C. Altavelia columbiensis, male proctiger, dorsal view. D. Altavelia flavomarginata, male proctiger, dorsal view. E. Altavelia montana, male proctiger, dorsal view. F. Altavelia altoandina, paramere. G. Altavelia antioquia, paramere. H. Altavelia columbiensis, paramere. I. Altavelia daza, paramere. J. Altavelia flavomarginata, paramere. K. Altavelia montana, paramere. L. Altavelia inveruglas, male proctiger, dorsal view. M. Altavelia inveruglas, paramere. N. Altavelia osborniana, male proctiger, dorsal view. O. Altavelia osborniana, paramere." captionText-4="FIGURE 5. Distribution of Altavelia altoandina, A. antioquia, A. montana, and A. quindiana." captionText-5="FIGURE 6. A. Altavelia boliviana, male, dorsal view. B. Altavelia boliviana, female, apex of abdomen, left posterolateral view. C. Altavelia columbiensis, male, dorsal view. D. Altavelia columbiensis, male, lateral view." captionText-6="FIGURE 7. Distribution of Altavelia boliviana, A. columbiensis, and A. daza." captionText-7="FIGURE 8. A. Altavelia daza, male, dorsal view. B. Altavelia daza, male, proctiger and parameres, dorsal view. C. Altavelia daza, male, lateral view." captionText-8="FIGURE 9. A. Altavelia amoena (junior synonym of A. flavomarginata), male, dorsal view. B. Altavelia flavomarginata, male, dorsal view. C. Altavelia inveruglas, male, dorsal view. D. Altavelia osborniana, female, dorsal view" captionText-9="FIGURE 10. Distribution of Altavelia flavomarginata, A. inveruglas, A. osborniana, and A. willei." httpUri-0="https://zenodo.org/record/2637342/files/figure.png" httpUri-1="https://zenodo.org/record/2637344/files/figure.png" httpUri-10="https://zenodo.org/record/2637362/files/figure.png" httpUri-2="https://zenodo.org/record/2637346/files/figure.png" httpUri-3="https://zenodo.org/record/2637348/files/figure.png" httpUri-4="https://zenodo.org/record/2637350/files/figure.png" httpUri-5="https://zenodo.org/record/2637352/files/figure.png" httpUri-6="https://zenodo.org/record/2637354/files/figure.png" httpUri-7="https://zenodo.org/record/2637356/files/figure.png" httpUri-8="https://zenodo.org/record/2637358/files/figure.png" httpUri-9="https://zenodo.org/record/2637360/files/figure.png" pageId="1" pageNumber="296">Figs. 1–11</figureCitation>
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<emphasis id="36C0EAB42854E966FF295D17FEB64947" bold="true" box="[151,289,1586,1611]" pageId="1" pageNumber="296">Description</emphasis>
. Large-sized veliids, body elongate. Length 
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, usually greater than 
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; body length-towidth ratios ranging from 2.90 to 4.44.
</paragraph>
<paragraph id="040B36A62854E966FF795D5FFEF949BB" blockId="1.[151,1437,1586,2007]" pageId="1" pageNumber="296">Color. Ground color brown to black, in some species marked contrastingly with yellow or orange; silvery pubescence absent.</paragraph>
<paragraph id="040B36A62854E966FF795DE7FCFC48BF" blockId="1.[151,1437,1586,2007]" pageId="1" pageNumber="296">Structural characters. Head with eyes globose, exerted, separated by approximately 1.5× eye width, slightly removed from anterior pronotal margin, ocular setae present. Head declivant anteriorly, not recessed into pronotum, with three pairs of facial trichobothria; gular region short, often not visible. Labium long, extending onto metasternum, article I not attaining posterior margin of bucculae, articles I and IV subequal in length, article II very short, one-third length of article I, article III about 4× length of article I. Antennae with article I longest, II and III progressively shorter, article IV subequal to or slightly shorter than article II; setae on antennae short, greatest length less than 2× width of antennal article on which they arise.</paragraph>
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Pronotum of wingless and micropterous forms slightly bilobate (
<figureCitation id="9C8F2A232854E966FC225C9BFC7348DB" box="[924,996,1982,2007]" captionStart="FIGURE 1" captionStartId="2.[151,250,1963,1985]" captionTargetBox="[156,1421,488,1876]" captionTargetId="figure@2.[151,1435,488,1939]" captionTargetPageId="2" captionText="FIGURE 1. A. Altavelia altoandina, male holotype, dorsal view. B. Altavelia altoandina, female paratype, dorsal view. C. Altavelia antioquia, male paratype, dorsal view. D. Altavelia antioquia, female paratype, dorsal view." httpUri="https://zenodo.org/record/2637342/files/figure.png" pageId="1" pageNumber="296">Figs. 1</figureCitation>
A–D, 2A–C, 6A, 6C, 8A, 9A–C, 11A– B), with lateral margins of anterior lobe swollen; collar distinct, delineated by line of foveae; calli flat, unmodified; anterior lobe of pronotum bearing a few coarse foveae, numerous obscure foveae present on posterior lobe; humeri slightly expanded, lateral margins gently bowed outward; mesonotum exposed laterally. Pronotum of macropters longer and wider, central section domed; posterior margin broadly rounded, bearing numerous small foveae; extending caudad over wing bases and scutellum, humeri raised, prominent (
<figureCitation id="9C8F2A232857E965FB895A02FB0F4E4C" box="[1079,1176,295,320]" captionStart="FIGURE 9" captionStartId="15.[151,250,1880,1902]" captionTargetBox="[150,1436,193,1858]" captionTargetId="figure@15.[150,1436,193,1859]" captionTargetPageId="15" captionText="FIGURE 9. A. Altavelia amoena (junior synonym of A. flavomarginata), male, dorsal view. B. Altavelia flavomarginata, male, dorsal view. C. Altavelia inveruglas, male, dorsal view. D. Altavelia osborniana, female, dorsal view" httpUri="https://zenodo.org/record/2637358/files/figure.png" pageId="2" pageNumber="297">Fig. 9D</figureCitation>
). Forewing uniformly blackish, with four closed cells, without pale markings. Meso- and metasterna each with a pair of median tubercles, these tubercles meeting at suture, forming a cup-like cavity between (
<figureCitation id="9C8F2A232857E965FC065A4AFB834E84" box="[952,1044,367,392]" captionStart="FIGURE 3" captionStartId="4.[151,250,1755,1777]" captionTargetBox="[177,1422,887,1728]" captionTargetId="figure@4.[151,1436,877,1734]" captionTargetPageId="4" captionText="FIGURE 3. A. Altavelia montana, thorax and base of abdomen, ventral view; white arrows show position of central tubercles. B. Altavelia flavomarginata, female, apex of thorax and abdomen, dorsal view." httpUri="https://zenodo.org/record/2637346/files/figure.png" pageId="2" pageNumber="297">Fig. 3A</figureCitation>
). Metasternal scent gland opening (omphalium) obscure.
</paragraph>
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<emphasis id="36C0EAB42857E965FF295C8EFE8248CC" bold="true" box="[151,277,1963,1985]" pageId="2" pageNumber="297">FIGURE 1.</emphasis>
A. 
<taxonomicName id="C3B44D252857E965FEFE5C8EFD8548CD" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[320,530,1963,1985]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="altoandina">
<emphasis id="36C0EAB42857E965FEFE5C8EFD8548CD" box="[320,530,1963,1985]" italics="true" pageId="2" pageNumber="297">Altavelia altoandina</emphasis>
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, male holotype, dorsal view. B. 
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<emphasis id="36C0EAB42857E965FCD75C8EFBAC48CD" box="[873,1083,1963,1985]" italics="true" pageId="2" pageNumber="297">Altavelia altoandina</emphasis>
</taxonomicName>
, female paratype, dorsal view. C. 
<taxonomicName id="C3B44D252857E965FF295CEFFECD48EC" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[151,346,1994,2016]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="antioquia">
<emphasis id="36C0EAB42857E965FF295CEFFECD48EC" box="[151,346,1994,2016]" italics="true" pageId="2" pageNumber="297">Altavelia antioquia</emphasis>
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, male paratype, dorsal view. D. 
<taxonomicName id="C3B44D252857E965FD1F5CEFFCF348EC" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[673,868,1994,2016]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="297" phylum="Arthropoda" rank="species" species="antioquia">
<emphasis id="36C0EAB42857E965FD1F5CEFFCF348EC" box="[673,868,1994,2016]" italics="true" pageId="2" pageNumber="297">Altavelia antioquia</emphasis>
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, female paratype, dorsal view.
</paragraph>
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<paragraph id="040B36A62856E964FF295C8DFD0048D1" blockId="3.[151,1436,1960,2013]" pageId="3" pageNumber="298">
<emphasis id="36C0EAB42856E964FF295C8DFE8F48B1" bold="true" box="[151,280,1960,1982]" pageId="3" pageNumber="298">FIGURE 2.</emphasis>
A. 
<taxonomicName id="C3B44D252856E964FEF65C8DFD9F48B2" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[328,520,1960,1982]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="36C0EAB42856E964FEF65C8DFD9F48B2" box="[328,520,1960,1982]" italics="true" pageId="3" pageNumber="298">Altavelia montana</emphasis>
</taxonomicName>
, male paratype, dorsal view. B. 
<taxonomicName id="C3B44D252856E964FCD05C8DFBB948B2" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[878,1070,1960,1982]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="36C0EAB42856E964FCD05C8DFBB948B2" box="[878,1070,1960,1982]" italics="true" pageId="3" pageNumber="298">Altavelia montana</emphasis>
</taxonomicName>
, female paratype, dorsal view. C. 
<taxonomicName id="C3B44D252856E964FF295CE2FEC848D1" authorityName="Molano &amp; Morales. Additionally" authorityYear="2019" box="[151,351,1991,2013]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="298" phylum="Arthropoda" rank="species" species="quindiana">
<emphasis id="36C0EAB42856E964FF295CE2FEC848D1" box="[151,351,1991,2013]" italics="true" pageId="3" pageNumber="298">Altavelia quindiana</emphasis>
</taxonomicName>
, female holotype, dorsal view.
</paragraph>
</caption>
<paragraph id="040B36A62851E963FF795BB2FBA94E11" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="299">
Abdomen lacking silvery setae or pruinose areas; prominent paired longitudinal carine present on mediotergites I–II and basally on mediotergite III of macropters (visible only after removal of wings, 
<figureCitation id="9C8F2A232851E963FA8B5B99FA194FD8" box="[1333,1422,188,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1755,1777]" captionTargetBox="[177,1422,887,1728]" captionTargetId="figure@4.[151,1436,877,1734]" captionTargetPageId="4" captionText="FIGURE 3. A. Altavelia montana, thorax and base of abdomen, ventral view; white arrows show position of central tubercles. B. Altavelia flavomarginata, female, apex of thorax and abdomen, dorsal view." httpUri="https://zenodo.org/record/2637346/files/figure.png" pageId="4" pageNumber="299">Fig. 3B</figureCitation>
), absent in apterous form. Hair-free longitudinal striae or elongate lacunae absent between abdominal mediosternites and parasternites; small lacunae present at sutures between individual parasternites.
</paragraph>
<paragraph id="040B36A62851E963FF795A0DFAA74EDC" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="299">Legs very long and slender, length of hind leg exceeding length of body; uniformly dark colored, lacking pale annuli; all segments unarmed, lacking prominent spines or teeth; male fore tibial grasping comb well-developed and prominent, extending for 1/6 to over half of length of tibia; females without grasping comb; all tarsi threesegmented, article I shortest, article II shorter than III on fore leg, article II subequal to or shorter than III on middle leg, article II usually longer than article III on hind leg; pretarsal claws moderately long, evenly tapering.</paragraph>
<paragraph id="040B36A62851E963FF795AF9FE614D89" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="299">
Male abdominal sternum VII without projections; terminalia very large; proctiger bearing prominent setal tufts or sclerotized wing-like processes; parameres symmetrical, very long and slender, lacking a prominent basal inflection, basal lobe weakly developed. Female abdominal tergum VIII deflected downward, not lying on same horizontal plane as tergum VII (
<figureCitation id="9C8F2A232851E963FDA25962FDED4D6C" box="[540,634,583,608]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1810]" captionTargetBox="[149,1436,337,1668]" captionTargetId="figure@9.[149,1437,337,1767]" captionTargetPageId="9" captionText="FIGURE 6. A. Altavelia boliviana, male, dorsal view. B. Altavelia boliviana, female, apex of abdomen, left posterolateral view. C. Altavelia columbiensis, male, dorsal view. D. Altavelia columbiensis, male, lateral view." httpUri="https://zenodo.org/record/2637352/files/figure.png" pageId="4" pageNumber="299">Fig. 6B</figureCitation>
); first gonocoxae partially exposed, plate-like; tergum IX elongate, strongly protruding posteriorly.
</paragraph>
</subSubSection>
<subSubSection id="4CAE652D2851E963FF7959AAFC9C4DA4" box="[199,779,655,680]" pageId="4" pageNumber="299" type="materials_examined">
<paragraph id="040B36A62851E963FF7959AAFC9C4DA4" blockId="4.[151,1437,151,824]" box="[199,779,655,680]" pageId="4" pageNumber="299">
<emphasis id="36C0EAB42851E963FF7959AAFECC4DA4" bold="true" box="[199,347,655,680]" pageId="4" pageNumber="299">
<typeStatus id="DB0F88042851E963FF7959AAFE974DA4" box="[199,256,655,680]" pageId="4" pageNumber="299">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="C3B44D252851E963FED759AAFC914DA4" authority="Kirkaldy, 1899" authorityName="Kirkaldy" authorityYear="1899" box="[361,774,655,680]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42851E963FED759AAFDC34DA4" box="[361,596,655,680]" italics="true" pageId="4" pageNumber="299">Paravelia inveruglas</emphasis>
<bibRefCitation id="60254B572851E963FDE259AAFC914DA4" author="Kirkaldy, G. W." box="[604,774,655,680]" pageId="4" pageNumber="299" pagination="1 - 9" refId="ref11699" refString="Kirkaldy, G. W. (1899) Viaggio del Dr. Enrico Festa nell'Ecuador e regioni vicine. XIX. Aquatic Rhynchota. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 14 (350), 1 - 9." type="journal article" year="1899">Kirkaldy, 1899</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="4CAE652D2851E963FF795991FC854DFC" pageId="4" pageNumber="299" type="etymology">
<paragraph id="040B36A62851E963FF795991FC854DFC" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="299">
<emphasis id="36C0EAB42851E963FF795991FED14DC1" bold="true" box="[199,326,692,717]" pageId="4" pageNumber="299">Etymology</emphasis>
. The name “ 
<emphasis id="36C0EAB42851E963FE625991FDA94DC1" box="[476,574,692,717]" italics="true" pageId="4" pageNumber="299">altavelia</emphasis>
” refers to the high elevations at which species of this genus occur (“ 
<emphasis id="36C0EAB42851E963FAEA5991FA174DC1" box="[1364,1408,692,717]" italics="true" pageId="4" pageNumber="299">alta</emphasis>
-”, Latin, nominative feminine singular, meaning high, tall).
</paragraph>
</subSubSection>
<subSubSection id="4CAE652D2851E963FF7959D9FE154C34" pageId="4" pageNumber="299" type="distribution">
<paragraph id="040B36A62851E963FF7959D9FE154C34" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="299">
<emphasis id="36C0EAB42851E963FF7959D9FECE4C19" bold="true" box="[199,345,764,789]" pageId="4" pageNumber="299">Distribution</emphasis>
. Known from elevations between 1500 and 
<quantity id="C34C9B432851E963FCF559D9FC374C18" box="[843,928,764,789]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="299" unit="m" value="3600.0">3600 m</quantity>
in the Andes of 
<collectingCountry id="7CA376362851E963FBE559D9FB274C19" box="[1115,1200,764,789]" name="Bolivia" pageId="4" pageNumber="299">Bolivia</collectingCountry>
, 
<collectingCountry id="7CA376362851E963FB0259D9FABA4C19" box="[1212,1325,764,789]" name="Colombia" pageId="4" pageNumber="299">Colombia</collectingCountry>
, 
<collectingCountry id="7CA376362851E963FA8759D9FA0F4C19" box="[1337,1432,764,789]" name="Ecuador" pageId="4" pageNumber="299">Ecuador</collectingCountry>
, 
<collectingCountry id="7CA376362851E963FF295805FF5A4C3B" box="[151,205,800,823]" name="Peru" pageId="4" pageNumber="299">Peru</collectingCountry>
, and 
<collectingCountry id="7CA376362851E963FEB6583AFEE84C34" box="[264,383,799,824]" name="Venezuela" pageId="4" pageNumber="299">Venezuela</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="50CB662E2851E963FF295DFEFC20481C" httpUri="https://zenodo.org/record/2637346/files/figure.png" pageId="4" pageNumber="299" startId="4.[151,250,1755,1777]" targetBox="[177,1422,887,1728]" targetPageId="4">
<paragraph id="040B36A62851E963FF295DFEFC20481C" blockId="4.[151,1437,1755,1808]" pageId="4" pageNumber="299">
<emphasis id="36C0EAB42851E963FF295DFEFE8449FC" bold="true" box="[151,275,1755,1777]" pageId="4" pageNumber="299">FIGURE 3.</emphasis>
A. 
<taxonomicName id="C3B44D252851E963FE865DFEFE6449FD" authorityName="Polhemus &amp; Molano &amp; Morales &amp; Moreira &amp; Floriano" authorityYear="2019" box="[312,499,1755,1777]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="36C0EAB42851E963FE865DFEFE6449FD" box="[312,499,1755,1777]" italics="true" pageId="4" pageNumber="299">Altavelia montana</emphasis>
</taxonomicName>
, thorax and base of abdomen, ventral view; white arrows show position of central tubercles. B. 
<taxonomicName id="C3B44D252851E963FF0A5DDFFE26481C" baseAuthorityName="Hungerford" baseAuthorityYear="1930" box="[180,433,1786,1808]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="flavomarginata">
<emphasis id="36C0EAB42851E963FF0A5DDFFE26481C" box="[180,433,1786,1808]" italics="true" pageId="4" pageNumber="299">Altavelia flavomarginata</emphasis>
</taxonomicName>
, female, apex of thorax and abdomen, dorsal view.
</paragraph>
</caption>
<subSubSection id="4CAE652D2851E962FF795C12FE454BAC" lastPageId="5" lastPageNumber="300" pageId="4" pageNumber="299" type="discussion">
<paragraph id="040B36A62851E962FF795C12FAC04E4C" blockId="4.[151,1437,1847,2016]" lastBlockId="5.[151,1437,151,1184]" lastPageId="5" lastPageNumber="300" pageId="4" pageNumber="299">
<emphasis id="36C0EAB42851E963FF795C12FEDF485C" bold="true" box="[199,328,1847,1872]" pageId="4" pageNumber="299">Comments</emphasis>
. Previous students of Neotropical 
<taxonomicName id="C3B44D252851E963FD5F5C12FCAA485C" box="[737,829,1847,1872]" class="Insecta" family="Veliidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="family">Veliidae</taxonomicName>
have long recognized the distinctive attributes of a subgroup of elongate, large-sized species of 
<taxonomicName id="C3B44D252851E963FD325C7EFD6D4878" authorityName="Breddin" authorityYear="1898" box="[652,762,1883,1908]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="genus">
<emphasis id="36C0EAB42851E963FD325C7EFD6D4878" box="[652,762,1883,1908]" italics="true" pageId="4" pageNumber="299">Paravelia</emphasis>
</taxonomicName>
occurring in the Andes, which were treated in part as the 
<taxonomicName id="C3B44D252851E963FA395C78FE994894" authorityName="Kirkaldy" authorityYear="1899" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="299" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42851E963FA395C78FE994894" italics="true" pageId="4" pageNumber="299">P. inveruglas</emphasis>
</taxonomicName>
group by 
<bibRefCitation id="60254B572851E963FE345C5AFDF34894" author="Hungerford, H. B." box="[394,612,1919,1944]" pageId="4" pageNumber="299" pagination="120 - 125" refId="ref11652" refString="Hungerford, H. B. (1930) Concerning Lelia inveruglas Kirkaldy and related forms. (Veliidae-Hemiptera). Annals of the Entomological Society of America, 23, 120 - 125. https: // doi. org / 10.1093 / aesa / 23.1.120" type="journal article" year="1930">Hungerford (1930)</bibRefCitation>
, based on the characteristics of that nominate species. As noted by 
<bibRefCitation id="60254B572851E963FA3E5CA5FECD48B0" author="Polhemus, D. A." pageId="4" pageNumber="299" pagination="151 - 162" refId="ref11987" refString="Polhemus, D. A. (2014) Two new Peruvian species of Paravelia (Heteropera: Veliidae) from water-filled bamboo internodes, and distributional notes for other Paravelia species. Tijdschrift voor Entomologie, 157, 151 - 162. https: // doi. org / 10.1163 / 22119434 - 00002043" type="journal article" year="2014">D. Polhemus (2014)</bibRefCitation>
, species strictly conforming to this group concept display the following features: 1) development of prominent setal tufts or sclerotized wing-like processes on the male proctiger; 2) elongate body shapes, with length-to-width ratios ranging from ~3.0 to 4.5; 3) a uniformly blackish forewing coloration, lacking pale markings; 4) antennae with segment I longest; 5) very long and slender legs in relation to the body, with the length of hind leg exceeding that of the body; and 6) the female abdomen with tergum VIII deflected downward and not lying on same horizontal plane as tergum VII, thereby displacing the female proctiger downward to a position below the other abdominal terga. The latter character is particularly evident when viewed laterally (
<figureCitation id="9C8F2A232850E962FB4F5A02FADE4E4C" box="[1265,1353,295,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1788,1810]" captionTargetBox="[149,1436,337,1668]" captionTargetId="figure@9.[149,1437,337,1767]" captionTargetPageId="9" captionText="FIGURE 6. A. Altavelia boliviana, male, dorsal view. B. Altavelia boliviana, female, apex of abdomen, left posterolateral view. C. Altavelia columbiensis, male, dorsal view. D. Altavelia columbiensis, male, lateral view." httpUri="https://zenodo.org/record/2637352/files/figure.png" pageId="5" pageNumber="300">Fig. 6B</figureCitation>
).
</paragraph>
<paragraph id="040B36A62850E962FF795A68FCB34D89" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="300">
In the current paper, species possessing the above characteristics are removed from 
<taxonomicName id="C3B44D252850E962FBC65A69FB714E69" authorityName="Breddin" authorityYear="1898" box="[1144,1254,332,357]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="36C0EAB42850E962FBC65A69FB714E69" box="[1144,1254,332,357]" italics="true" pageId="5" pageNumber="300">Paravelia</emphasis>
</taxonomicName>
and assigned to the new genus 
<emphasis id="36C0EAB42850E962FEFC5A4AFE304E84" box="[322,423,367,392]" italics="true" pageId="5" pageNumber="300">Altavelia</emphasis>
. Ecologically, the members of this group occur at high elevations in the Andes (between 1500 and 
<quantity id="C34C9B432850E962FEB85AB1FEC94EA0" box="[262,350,404,429]" metricMagnitude="3" metricUnit="m" metricValue="3.6" pageId="5" pageNumber="300" unit="m" value="3600.0">3600 m</quantity>
), and possess semi-terrestrial habits that are notably different from those of other species held in 
<taxonomicName id="C3B44D252850E962FF295A92FE924EDC" authorityName="Breddin" authorityYear="1898" box="[151,261,439,464]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="36C0EAB42850E962FF295A92FE924EDC" box="[151,261,439,464]" italics="true" pageId="5" pageNumber="300">Paravelia</emphasis>
</taxonomicName>
(
<bibRefCitation id="60254B572850E962FEA85A92FDCC4EDC" author="Padilla-Gil, D. N. &amp; Moreira, F. F. F." box="[278,603,439,464]" pageId="5" pageNumber="300" pagination="14 - 26" refId="ref11935" refString="Padilla-Gil, D. N. &amp; Moreira, F. F. F. (2011) A new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3065, 14 - 26." type="journal article" year="2011">Padilla-Gil &amp; Moreira 2011</bibRefCitation>
, 
<bibRefCitation id="60254B572850E962FDD75A9DFCD44EDC" author="Polhemus, D. A." box="[617,835,439,464]" pageId="5" pageNumber="300" pagination="151 - 162" refId="ref11987" refString="Polhemus, D. A. (2014) Two new Peruvian species of Paravelia (Heteropera: Veliidae) from water-filled bamboo internodes, and distributional notes for other Paravelia species. Tijdschrift voor Entomologie, 157, 151 - 162. https: // doi. org / 10.1163 / 22119434 - 00002043" type="journal article" year="2014">D. Polhemus 2014</bibRefCitation>
). All of the included species have very long legs in relation to the size of their bodies, which appear to aid in terrestrial locomotion. Notes on distribution, habitat preferences, and nomenclatural history were provided for 
<taxonomicName id="C3B44D252850E962FC9A5925FBD34D14" authority="(Drake, 1957)" baseAuthorityName="Drake" baseAuthorityYear="1957" box="[804,1092,511,536]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="amoena">
<emphasis id="36C0EAB42850E962FC9A5925FC0A4D14" box="[804,925,512,536]" italics="true" pageId="5" pageNumber="300">A. amoena</emphasis>
(
<bibRefCitation id="60254B572850E962FC125ADAFBAA4D14" author="Drake, C. J." box="[940,1085,511,536]" pageId="5" pageNumber="300" pagination="138 - 140" refId="ref11500" refString="Drake, C. J. (1957) Notes on the genus Lelia (Hemiptera: Veliidae). Journal of the Kansas Entomological Society, 30 (4), 138 - 140." type="journal article" year="1957">Drake, 1957</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="C3B44D252850E962FBEE5925FA004D14" authority="(Breddin, 1898)" baseAuthorityName="Breddin" baseAuthorityYear="1898" box="[1104,1431,511,536]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="boliviana">
<emphasis id="36C0EAB42850E962FBEE5925FB4D4D14" box="[1104,1242,511,536]" italics="true" pageId="5" pageNumber="300">A. boliviana</emphasis>
(
<bibRefCitation id="60254B572850E962FB575ADAFA184D14" author="Breddin, G." box="[1257,1423,511,536]" pageId="5" pageNumber="300" pagination="149 - 163" refId="ref11473" refString="Breddin, G. (1898) Studia hemipterologica. Jahresbericht und Abhandlungen des Naturwissenschaftlichen Lereins in Magdeburg, 1896 - 1898, 149 - 163." type="journal article" year="1898">Breddin, 1898</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="C3B44D252850E962FF295901FDF44D30" authority="(Padilla-Gil &amp; Moreira, 2011)" baseAuthorityName="Padilla-Gil &amp; Moreira" baseAuthorityYear="2011" box="[151,611,548,573]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="daza">
<emphasis id="36C0EAB42850E962FF295901FF664D31" box="[151,241,548,573]" italics="true" pageId="5" pageNumber="300">A. daza</emphasis>
(
<bibRefCitation id="60254B572850E962FEBB5901FDCC4D30" author="Padilla-Gil, D. N. &amp; Moreira, F. F. F." box="[261,603,548,573]" pageId="5" pageNumber="300" pagination="14 - 26" refId="ref11935" refString="Padilla-Gil, D. N. &amp; Moreira, F. F. F. (2011) A new species of Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to Colombian Paravelia species. Zootaxa, 3065, 14 - 26." type="journal article" year="2011">Padilla-Gil &amp; Moreira, 2011</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="C3B44D252850E962FDCA5901FBA14D30" authority="(Hungerford, 1930)" baseAuthorityName="Hungerford" baseAuthorityYear="1930" box="[628,1078,548,573]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="flavomarginata">
<emphasis id="36C0EAB42850E962FDCA5901FCD14D31" box="[628,838,548,573]" italics="true" pageId="5" pageNumber="300">A. flavomarginata</emphasis>
(
<bibRefCitation id="60254B572850E962FCE45901FBBA4D30" author="Hungerford, H. B." box="[858,1069,548,573]" pageId="5" pageNumber="300" pagination="120 - 125" refId="ref11652" refString="Hungerford, H. B. (1930) Concerning Lelia inveruglas Kirkaldy and related forms. (Veliidae-Hemiptera). Annals of the Entomological Society of America, 23, 120 - 125. https: // doi. org / 10.1093 / aesa / 23.1.120" type="journal article" year="1930">Hungerford, 1930</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="C3B44D252850E962FBF65901FB744D31" authorityName="Kirkaldy" authorityYear="1899" box="[1096,1251,548,573]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42850E962FBF65901FB744D31" box="[1096,1251,548,573]" italics="true" pageId="5" pageNumber="300">A. inveruglas</emphasis>
</taxonomicName>
, 
<taxonomicName id="C3B44D252850E962FB495901FEC04D6C" authority="(Kirkaldy, 1909)" baseAuthorityName="Kirkaldy" baseAuthorityYear="1909" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="osborniana">
<emphasis id="36C0EAB42850E962FB495901FA0C4D31" box="[1271,1435,548,573]" italics="true" pageId="5" pageNumber="300">A. osborniana</emphasis>
(Kirkaldy, 1909)
</taxonomicName>
, and 
<taxonomicName id="C3B44D252850E962FE24596DFC8F4D6C" authority="(Drake &amp; Harris, 1940)" baseAuthorityName="Drake &amp; Harris" baseAuthorityYear="1940" box="[410,792,583,608]" class="Insecta" family="Veliidae" genus="Altavelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="willei">
<emphasis id="36C0EAB42850E962FE24596DFE6D4D6C" box="[410,506,583,608]" italics="true" pageId="5" pageNumber="300">A. willei</emphasis>
(
<bibRefCitation id="60254B572850E962FDB35962FC984D6C" author="Drake, C. J. &amp; Harris, H. M." box="[525,783,583,608]" pageId="5" pageNumber="300" pagination="31" refId="ref11536" refString="Drake, C. J. &amp; Harris, H. M. (1940) A new Lelia from Peru (Hemiptera, Veliidae). Pan-Pacific Entomologist, 16 (1), 31." type="journal article" year="1940">Drake &amp; Harris, 1940</bibRefCitation>
)
</taxonomicName>
by 
<bibRefCitation id="60254B572850E962FCF7596DFBA14D6C" author="Polhemus, D. A." box="[841,1078,583,608]" pageId="5" pageNumber="300" pagination="151 - 162" refId="ref11987" refString="Polhemus, D. A. (2014) Two new Peruvian species of Paravelia (Heteropera: Veliidae) from water-filled bamboo internodes, and distributional notes for other Paravelia species. Tijdschrift voor Entomologie, 157, 151 - 162. https: // doi. org / 10.1163 / 22119434 - 00002043" type="journal article" year="2014">D. Polhemus (2014)</bibRefCitation>
, 
<bibRefCitation id="60254B572850E962FBFB5962FAF34D6C" author="Rodrigues, H. D. D. &amp; Moreira, F. F. F. &amp; Nieser, N. &amp; Chen, P. P. &amp; Melo, A. L. &amp; Dias-Silva, K. &amp; Giehl, N. F. S." box="[1093,1380,583,608]" pageId="5" pageNumber="300" pagination="1 - 47" refId="ref12291" refString="Rodrigues, H. D. D., Moreira, F. F. F., Nieser, N., Chen, P. P., Melo, A. L., Dias-Silva, K. &amp; Giehl, N. F. S. (2014 b) The genus Paravelia Breddin, 1898 (Hemiptera: Heteroptera: Veliidae) in Brazil, with descriptions of eight new species. Zootaxa, 3784 (1), 1 - 47. http: // dx. doi. org / 10.11646 / zootaxa. 3784.1.1" type="journal article" year="2014">
Rodrigues 
<emphasis id="36C0EAB42850E962FB7D596CFB684D6C" box="[1219,1279,583,608]" italics="true" pageId="5" pageNumber="300">et al.</emphasis>
(2014b)
</bibRefCitation>
, and 
<bibRefCitation id="60254B572850E962FF295949FE714D89" author="Rodrigues, H. D. D. &amp; Moreira, F. F. F." box="[151,486,620,645]" pageId="5" pageNumber="300" pagination="642 - 667" refId="ref12184" refString="Rodrigues, H. D. D. &amp; Moreira, F. F. F. (2016 a) Four new species of Paravelia (Hemiptera: Heteroptera: Veliidae) from Brazil, with notes on previously described species. Canadian Entomologist, 148, 642 - 667." type="journal article" year="2016">Rodrigues &amp; Moreira (2016a)</bibRefCitation>
and are not repeated herein.
</paragraph>
<paragraph id="040B36A62850E962FF7959AAFE454BAC" blockId="5.[151,1437,151,1184]" pageId="5" pageNumber="300">
Although 
<bibRefCitation id="60254B572850E962FE8459AAFD9C4DA4" author="Hungerford, H. B." box="[314,523,655,680]" pageId="5" pageNumber="300" pagination="759 - 761" refId="ref11603" refString="Hungerford, H. B. (1929) A new Lelia from Arizona with notes on other species (Hemiptera-Veliidae). Annals of the Entomological Society of America, 22, 759 - 761. https: // doi. org / 10.1093 / aesa / 22.4.759" type="journal article" year="1929">Hungerford (1929</bibRefCitation>
, 
<bibRefCitation id="60254B572850E962FDA659B5FDC64DA4" author="Hungerford, H. B." box="[536,593,656,680]" pageId="5" pageNumber="300" pagination="120 - 125" refId="ref11652" refString="Hungerford, H. B. (1930) Concerning Lelia inveruglas Kirkaldy and related forms. (Veliidae-Hemiptera). Annals of the Entomological Society of America, 23, 120 - 125. https: // doi. org / 10.1093 / aesa / 23.1.120" type="journal article" year="1930">1930</bibRefCitation>
) based his 
<taxonomicName id="C3B44D252850E962FD6959AAFCDA4DA4" authorityName="Kirkaldy" authorityYear="1899" box="[727,845,655,680]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42850E962FD6959AAFCDA4DA4" box="[727,845,655,680]" italics="true" pageId="5" pageNumber="300">inveruglas</emphasis>
</taxonomicName>
species group concept on 
<taxonomicName id="C3B44D252850E962FB3D59AAFAE64DA4" authorityName="Kirkaldy" authorityYear="1899" box="[1155,1393,655,680]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42850E962FB3D59AAFAE64DA4" box="[1155,1393,655,680]" italics="true" pageId="5" pageNumber="300">Paravelia inveruglas</emphasis>
</taxonomicName>
, he included within his group two species, 
<taxonomicName id="C3B44D252850E962FDF55990FC4C4DC0" authority="(Kirkaldy, 1899)" baseAuthorityName="Kirkaldy" baseAuthorityYear="1899" box="[587,987,692,717]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="albotrimaculata">
<emphasis id="36C0EAB42850E962FDF55990FC8D4DC1" box="[587,794,692,717]" italics="true" pageId="5" pageNumber="300">P. albotrimaculata</emphasis>
(
<bibRefCitation id="60254B572850E962FC965991FC444DC0" author="Kirkaldy, G. W." box="[808,979,692,717]" pageId="5" pageNumber="300" pagination="1 - 9" refId="ref11699" refString="Kirkaldy, G. W. (1899) Viaggio del Dr. Enrico Festa nell'Ecuador e regioni vicine. XIX. Aquatic Rhynchota. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 14 (350), 1 - 9." type="journal article" year="1899">Kirkaldy, 1899</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="C3B44D252850E962FBAE5990FAF14DC0" authority="(Hungerford, 1929)" baseAuthorityName="Hungerford" baseAuthorityYear="1929" box="[1040,1382,692,717]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="36C0EAB42850E962FBAE5990FB144DC1" box="[1040,1155,692,717]" italics="true" pageId="5" pageNumber="300">P. helenae</emphasis>
(
<bibRefCitation id="60254B572850E962FB2F5991FAC84DC0" author="Hungerford, H. B." box="[1169,1375,692,717]" pageId="5" pageNumber="300" pagination="759 - 761" refId="ref11603" refString="Hungerford, H. B. (1929) A new Lelia from Arizona with notes on other species (Hemiptera-Veliidae). Annals of the Entomological Society of America, 22, 759 - 761. https: // doi. org / 10.1093 / aesa / 22.4.759" type="journal article" year="1929">Hungerford, 1929</bibRefCitation>
)
</taxonomicName>
, that do not possess all of the distinctive character states displayed by 
<taxonomicName id="C3B44D252850E962FCD059FCFB974DFC" authorityName="Kirkaldy" authorityYear="1899" box="[878,1024,727,752]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42850E962FCD059FCFB974DFC" box="[878,1024,727,752]" italics="true" pageId="5" pageNumber="300">P. inveruglas</emphasis>
</taxonomicName>
and other species now transferred to 
<emphasis id="36C0EAB42850E962FF2959D9FF6B4C19" box="[151,252,764,789]" italics="true" pageId="5" pageNumber="300">Altavelia</emphasis>
. In the case of 
<taxonomicName id="C3B44D252850E962FE1B59D8FDE24C19" baseAuthorityName="Kirkaldy" baseAuthorityYear="1899" box="[421,629,764,789]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="albotrimaculata">
<emphasis id="36C0EAB42850E962FE1B59D8FDE24C19" box="[421,629,764,789]" italics="true" pageId="5" pageNumber="300">P. albotrimaculata</emphasis>
</taxonomicName>
, this species is known from only a single macropterous female 
<typeStatus id="DB0F88042850E962FA8459D9FA0B4C19" box="[1338,1436,764,789]" pageId="5" pageNumber="300" type="holotype">holotype</typeStatus>
from 
<collectingCountry id="7CA376362850E962FF68583AFEDA4C34" box="[214,333,799,824]" name="Venezuela" pageId="5" pageNumber="300">Venezuela</collectingCountry>
; therefore, assessment of the character states in the male terminalia is not possible. However, on the basis of its large size, elongate body shape, hind leg armature and a forewing pattern consisting of two pale spots basally and a large pale spot distally (see figures 12 and 
<quantity id="C34C9B432850E962FCE7584DFC054C8C" box="[857,914,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="5" pageNumber="300" unit="in" value="13.0">13 in</quantity>
<bibRefCitation id="60254B572850E962FC245842FBC84C8C" author="Hungerford, H. B." box="[922,1119,871,896]" pageId="5" pageNumber="300" pagination="120 - 125" refId="ref11652" refString="Hungerford, H. B. (1930) Concerning Lelia inveruglas Kirkaldy and related forms. (Veliidae-Hemiptera). Annals of the Entomological Society of America, 23, 120 - 125. https: // doi. org / 10.1093 / aesa / 23.1.120" type="journal article" year="1930">Hungerford 1930</bibRefCitation>
, and note that this character state was misinterpreted by 
<bibRefCitation id="60254B572850E962FE6C58A8FD344CA8" author="Polhemus, D. A." box="[466,675,908,933]" pageId="5" pageNumber="300" pagination="151 - 162" refId="ref11987" refString="Polhemus, D. A. (2014) Two new Peruvian species of Paravelia (Heteropera: Veliidae) from water-filled bamboo internodes, and distributional notes for other Paravelia species. Tijdschrift voor Entomologie, 157, 151 - 162. https: // doi. org / 10.1163 / 22119434 - 00002043" type="journal article" year="2014">
D. Polhemus 
<quantity id="C34C9B432850E962FDD558A9FD344CA8" box="[619,675,908,932]" metricMagnitude="1" metricUnit="m" metricValue="5.11556" pageId="5" pageNumber="300" unit="in" value="2014.0">2014</quantity>
</bibRefCitation>
in the context of his discussion of Hungerford’s 
<taxonomicName id="C3B44D252850E962FB7258A9FAD54CA9" authorityName="Kirkaldy" authorityYear="1899" box="[1228,1346,908,933]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="inveruglas">
<emphasis id="36C0EAB42850E962FB7258A9FAD54CA9" box="[1228,1346,908,933]" italics="true" pageId="5" pageNumber="300">inveruglas</emphasis>
</taxonomicName>
group), this taxon appears to be more closely allied to a group of large-sized species breeding in phytotelmata, including 
<taxonomicName id="C3B44D252850E962FA395894FE464CE0" authority="D. Polhemus, 2014" authorityName="D. Polhemus" authorityYear="2014" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="loutoni">
<emphasis id="36C0EAB42850E962FA395894FF714CE1" italics="true" pageId="5" pageNumber="300">P. loutoni</emphasis>
<bibRefCitation id="60254B572850E962FF4E58F0FE464CE0" author="Polhemus, D. A." box="[240,465,980,1005]" pageId="5" pageNumber="300" pagination="151 - 162" refId="ref11987" refString="Polhemus, D. A. (2014) Two new Peruvian species of Paravelia (Heteropera: Veliidae) from water-filled bamboo internodes, and distributional notes for other Paravelia species. Tijdschrift voor Entomologie, 157, 151 - 162. https: // doi. org / 10.1163 / 22119434 - 00002043" type="journal article" year="2014">D. Polhemus, 2014</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="C3B44D252850E962FE6058F0FCA44CE0" authority="(Hungerford, 1931)" baseAuthorityName="Hungerford" baseAuthorityYear="1931" box="[478,819,980,1005]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="myersi">
<emphasis id="36C0EAB42850E962FE6058F0FDDF4CE0" box="[478,584,981,1004]" italics="true" pageId="5" pageNumber="300">P. myersi</emphasis>
(Hungerford, 1931)
</taxonomicName>
, and 
<taxonomicName id="C3B44D252850E962FCCA58F0FB474CE0" authority="D. Polhemus, 2014" authorityName="D. Polhemus" authorityYear="2014" box="[884,1232,980,1005]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="reclusa">
<emphasis id="36C0EAB42850E962FCCA58F0FC724CE1" box="[884,997,980,1005]" italics="true" pageId="5" pageNumber="300">P. reclusa</emphasis>
<bibRefCitation id="60254B572850E962FC5158F0FB474CE0" author="Polhemus, D. A." box="[1007,1232,980,1005]" pageId="5" pageNumber="300" pagination="151 - 162" refId="ref11987" refString="Polhemus, D. A. (2014) Two new Peruvian species of Paravelia (Heteropera: Veliidae) from water-filled bamboo internodes, and distributional notes for other Paravelia species. Tijdschrift voor Entomologie, 157, 151 - 162. https: // doi. org / 10.1163 / 22119434 - 00002043" type="journal article" year="2014">D. Polhemus, 2014</bibRefCitation>
</taxonomicName>
. In particular, the multiple pale spots in the forewing represent a character state inconsistent with the uniformly dark coloration present in all species of 
<emphasis id="36C0EAB42850E962FE175F39FD994B39" box="[425,526,1052,1077]" italics="true" pageId="5" pageNumber="300">Altavelia</emphasis>
. In the case of 
<taxonomicName id="C3B44D252850E962FD035F38FCA74B39" baseAuthorityName="Hungerford" baseAuthorityYear="1929" box="[701,816,1052,1077]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="species" species="helenae">
<emphasis id="36C0EAB42850E962FD035F38FCA74B39" box="[701,816,1052,1077]" italics="true" pageId="5" pageNumber="300">P. helenae</emphasis>
</taxonomicName>
, this species also has three pale spots on the forewing, lacks lateral projections on the male proctiger, and inhabits bromeliads. As such, it also does not conform to the morphological or ecological character states definitive for 
<emphasis id="36C0EAB42850E962FC955F41FC074B71" box="[811,912,1124,1149]" italics="true" pageId="5" pageNumber="300">Altavelia</emphasis>
. Both of the above species have been retained in 
<taxonomicName id="C3B44D252850E962FF0A5FA2FEB54BAC" authorityName="Breddin" authorityYear="1898" box="[180,290,1159,1184]" class="Insecta" family="Veliidae" genus="Paravelia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="36C0EAB42850E962FF0A5FA2FEB54BAC" box="[180,290,1159,1184]" italics="true" pageId="5" pageNumber="300">Paravelia</emphasis>
</taxonomicName>
for the present.
</paragraph>
</subSubSection>
</treatment>
</document>