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<mods:title id="823CCEC2CE459F4752A1688E29778245">A review of the Caryanda tamdaoensis species-group (Orthoptera: Acrididae), with description of a new species from Yunnan, China</mods:title>
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<mods:affiliation id="F0714515B45B22A413E75580C9492546">College of Agriculture and Biological Science, Dali University, Dali, Yunnan 671003, China. &amp; Research Center for Northwest Yunnan Biodiversity, Dali University, Dali, Yunnan 671003, China.</mods:affiliation>
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Genus
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<emphasis id="3E89515AFFF8FF8AD5D1FD98FEBDFD69" bold="true" box="[238,357,613,640]" italics="true" pageId="1" pageNumber="413">Caryanda</emphasis>
Stål, 1878
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Species group
<taxonomicName id="CBFDF6CBFFF8FF8AD473FD50FDB8FD21" authorityName="Storozhenko" authorityYear="1992" box="[332,608,685,712]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="3E89515AFFF8FF8AD473FD50FDB8FD21" bold="true" box="[332,608,685,712]" italics="true" pageId="1" pageNumber="413">Caryanda tamdaoensis</emphasis>
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<emphasis id="3E89515AFFF8FF8AD5A8FD08FE7BFCE6" bold="true" box="[151,419,757,783]" pageId="1" pageNumber="413">Differential diagnosis.</emphasis>
<taxonomicName id="CBFDF6CBFFF8FF8AD491FD0BFD6EFCF9" authorityName="Storozhenko" authorityYear="1992" box="[430,694,758,784]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="3E89515AFFF8FF8AD491FD0BFD6EFCF9" box="[430,694,758,784]" italics="true" pageId="1" pageNumber="413">Caryanda tamdaoensis</emphasis>
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species-group shares the following diagnostic characters: male supra-anal plate nearly pentagonal; male cerci conical; apical penis valves of phallic complex lathy, upcurved and with a beak-shaped apex in lateral view; in females ventral basivalvular sclerite with inner margins always contiguous with each other.
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<emphasis id="3E89515AFFF8FF8AD5F8FC78FEBFFC76" bold="true" box="[199,359,901,927]" pageId="1" pageNumber="413">Composition.</emphasis>
This species group consists of five species. A key to species and subspecies is given below.
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<heading id="570A3A24FFF8FF8AD5A8FC0FFBFCFBE2" bold="true" box="[151,1060,1009,1036]" fontSize="11" level="1" pageId="1" pageNumber="413" reason="1">
<emphasis id="3E89515AFFF8FF8AD5A8FC0FFBFCFBE2" bold="true" box="[151,1060,1009,1036]" pageId="1" pageNumber="413">
Key to the species and subspecies of
<taxonomicName id="CBFDF6CBFFF8FF8AD762FC0CFCA9FBE5" ID-CoL="RHW8" authorityName="Storozhenko" authorityYear="1992" box="[605,881,1009,1036]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="3E89515AFFF8FF8AD762FC0CFCA9FBE5" bold="true" box="[605,881,1009,1036]" italics="true" pageId="1" pageNumber="413">Caryanda tamdaoensis</emphasis>
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species-group
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1. Hind tibiae entirely black in male or blue in female; posterior margin of female subgenital plate concave in middle with two obtuse dentes..............................................................
<taxonomicName id="CBFDF6CBFFF8FF8AD169FBA9FA44FB80" ID-CoL="RHVH" authority="Mao, Xu &amp; Li, 2017" authorityName="Mao, Xu &amp; Li" authorityYear="2017" box="[1110,1436,1108,1130]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="nigrotibia">
<emphasis id="3E89515AFFF8FF8AD169FBA9FB09FB80" box="[1110,1233,1108,1129]" italics="true" pageId="1" pageNumber="413">C. nigrotibia</emphasis>
<bibRefCitation id="686CF0B9FFF8FF8AD1E8FBA8FA44FB80" author="Mao, B. Y. &amp; Xu, J. S. &amp; Li, M." box="[1239,1436,1108,1130]" pageId="1" pageNumber="413" pagination="85 - 93" refId="ref6502" refString="Mao, B. Y., Xu, J. S. &amp; Li, M. (2017) Proposal of the Caryanda aurata species group (Orthoptera: Acrididae) with descriptions of two new species from China. Entomotaxonomia, 39 (2), 85 - 93." type="journal article" year="2017">Mao, Xu &amp; Li, 2017</bibRefCitation>
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<keyLead id="B70C2E7DFFF8FF8AD5A8FB8FFA44FB6E" box="[151,1436,1137,1159]" pageId="1" pageNumber="413">- Hind tibiae blue except base black in both sexes; posterior margin of female subgenital plate not as above.............. 2</keyLead>
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<keyLead id="B70C2E7DFFF8FF8AD5A8FB72FA44FB4D" box="[151,1436,1167,1189]" pageId="1" pageNumber="413">2. Hind femora partially red and partially yellow; outer lophi of epiphallus parallelogram in posterior view................ 3</keyLead>
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<paragraph id="0C428D48FFF8FF8AD5A8FB50FA44FB2B" blockId="1.[151,1437,1079,1717]" box="[151,1436,1196,1218]" pageId="1" pageNumber="413">
<keyLead id="B70C2E7DFFF8FF8AD5A8FB50FA44FB2B" box="[151,1436,1196,1218]" pageId="1" pageNumber="413">- Hind femora mainly green; outer lophi of epiphallus triangular in posterior view, distally tapering..................... 5</keyLead>
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<paragraph id="0C428D48FFF8FF8AD5A8FB37FA44FAF0" blockId="1.[151,1437,1079,1717]" pageId="1" pageNumber="413">
<keyLead id="B70C2E7DFFF8FF8AD5A8FB37FA44FAF0" pageId="1" pageNumber="413">
3. Hind femora with yellowish green basal half and reddish apical half in both sexes; posterior margin of female subgenital plate roundly projected at the middle; apical valves of penis and cingular valves long, curved upward, S-shaped.......................................................................................
<taxonomicName id="CBFDF6CBFFF8FF8AD172FAF9FA44FAF0" ID-CoL="RHW8" authority="Storozhenko, 1992" authorityName="Storozhenko" authorityYear="1992" box="[1101,1436,1284,1306]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="tamdaoensis">
<emphasis id="3E89515AFFF8FF8AD172FAF9FB39FAF0" box="[1101,1249,1284,1305]" italics="true" pageId="1" pageNumber="413">C. tamdaoensis</emphasis>
<bibRefCitation id="686CF0B9FFF8FF8AD1D8FAF9FA44FAF0" author="Storozhenko, S. Y." box="[1255,1436,1284,1306]" pageId="1" pageNumber="413" pagination="35 - 45" refId="ref6555" refString="Storozhenko, S. Y. (1992) To the knowledge of grasshoppers (Orthoptera, Acrididae) of Vietnam. Trudy Zoologitscheskogo Instituta Akademi Nauk SSSR, 245, 35 - 45. [in Russian]" type="journal article" year="1992">Storozhenko, 1992</bibRefCitation>
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<keyLead id="B70C2E7DFFF8FF8AD5A8FADFFA44FA98" pageId="1" pageNumber="413">- Hind femora either yellowish brown (rare grayish green) or almost red in both sexes; female subgenital plate with posterior margin undulated, medially concaved with two blunt teeth; apical valves of penis and cingular valves relatively short, curved upward, but not S-shaped............................................................................... 4</keyLead>
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4. Body size larger, body length
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in male or
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in female; hind femora yellowish brown (rare grayish green) without any red or pink parts in both sexes..................
<taxonomicName id="CBFDF6CBFFF8FF8AD667FA6AFAEFFA45" ID-CoL="RHTX" authority="Gorochov &amp; Storozhenko, 1994" authorityName="Gorochov &amp; Storozhenko" authorityYear="1994" baseAuthorityName="Gorochov &amp; Storozhenko" baseAuthorityYear="1994" box="[856,1335,1431,1453]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="subSpecies" species="azurea" status="stat. nov." subSpecies="azurea">
<emphasis id="3E89515AFFF8FF8AD667FA6AFC22FA45" box="[856,1018,1431,1452]" italics="true" pageId="1" pageNumber="413">C. azurea azurea</emphasis>
<bibRefCitation id="686CF0B9FFF8FF8AD13EFA6AFAEFFA45" author="Gorochov, A. V. &amp; Storozhenko, S. Y." box="[1025,1335,1431,1453]" pageId="1" pageNumber="413" pagination="55 - 58" refId="ref6015" refString="Gorochov, A. V. &amp; Storozhenko, S. Y. (1994) A new species of the genus Caryanda Stal (Orthoptera, Acrididae) from Vietnam. Trudy Zoologitscheskogo Instituta Akademi Nauk SSSR, 257, 55 - 58. [in Russian]" type="journal article" year="1994">Gorochov &amp; Storozhenko, 1994</bibRefCitation>
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,
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<taxonomicNameLabel id="25BAEC21FFF8FF8AD07EFA6AFA44FA45" box="[1345,1436,1431,1452]" pageId="1" pageNumber="413">stat. nov.</taxonomicNameLabel>
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- Body size smaller, body length
<quantity id="CB0520ADFFF8FF8AD72FFA49FD43FA23" box="[528,667,1460,1482]" metricMagnitude="-2" metricUnit="m" metricValue="1.7799999999999998" metricValueMax="1.7899999999999998" metricValueMin="1.77" pageId="1" pageNumber="413" unit="mm" value="17.799999999999997" valueMax="17.9" valueMin="17.7">17.717.9 mm</quantity>
in male or
<quantity id="CB0520ADFFF8FF8AD637FA49FC85FA23" box="[776,861,1460,1482]" metricMagnitude="-2" metricUnit="m" metricValue="2.07" pageId="1" pageNumber="413" unit="mm" value="20.7">20.7 mm</quantity>
in female; hind femora almost red but base yellowish green in both sexes..............................................
<taxonomicName id="CBFDF6CBFFF8FF8AD6ACFA2FFAEFFA0E" ID-CoL="5XDGL" authority="Mao, Ren &amp; Ou, 2011" authorityName="Mao, Ren &amp; Ou" authorityYear="2011" baseAuthorityName="Mao, Ren &amp; Ou" baseAuthorityYear="2011" box="[915,1335,1489,1511]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="subSpecies" species="azurea" status="stat. nov." subSpecies="colourfula">
<emphasis id="3E89515AFFF8FF8AD6ACFA2FFB81FA0F" box="[915,1113,1489,1511]" italics="true" pageId="1" pageNumber="413">C. azurea colourfula</emphasis>
<bibRefCitation id="686CF0B9FFF8FF8AD160FA2FFAEFFA0E" author="Mao, B. Y. &amp; Ren, G. D. &amp; Ou, X. H." box="[1119,1335,1490,1511]" pageId="1" pageNumber="413" refId="ref6458" refString="Mao, B. Y., Ren, G. D. &amp; Ou, X. H. (2011) Fauna, distribution pattern and adaptability on Acridoidea from Yunnan. China Forestry Publishing House, Beijing, 336 pp." type="book" year="2011">Mao, Ren &amp; Ou, 2011</bibRefCitation>
</taxonomicName>
,
<emphasis id="3E89515AFFF8FF8AD07EFA2FFA44FA0E" bold="true" box="[1345,1436,1490,1511]" pageId="1" pageNumber="413">
<taxonomicNameLabel id="25BAEC21FFF8FF8AD07EFA2FFA44FA0E" box="[1345,1436,1490,1511]" pageId="1" pageNumber="413">stat. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="B70995EDFFF8FF8AD5A8FA12FA44F95D" pageId="1" pageNumber="413">
<paragraph id="0C428D48FFF8FF8AD5A8FA12FA44F9D6" blockId="1.[151,1437,1079,1717]" pageId="1" pageNumber="413">
<keyLead id="B70C2E7DFFF8FF8AD5A8FA12FA44F9D6" pageId="1" pageNumber="413">
5. Abdominal tergites darkish green with 2 black longitudinal stripes in male; hind femora with 3/5 basal parts yellowish green and the apical 2/5 orange; female subgenital plate with posterior margin roundly projected at the middle......................................................................................
<taxonomicName id="CBFDF6CBFFF8FF8AD115F9D7FA44F9D6" ID-CoL="RHWQ" authority="Mao, Xu &amp; Li, 2017" authorityName="Mao, Xu &amp; Li" authorityYear="2017" box="[1066,1436,1577,1599]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="zhenyuanensis">
<emphasis id="3E89515AFFF8FF8AD115F9D7FB09F9D7" box="[1066,1233,1577,1599]" italics="true" pageId="1" pageNumber="413">C. zhenyuanensis</emphasis>
<bibRefCitation id="686CF0B9FFF8FF8AD1E8F9D7FA44F9D6" author="Mao, B. Y. &amp; Xu, J. S. &amp; Li, M." box="[1239,1436,1577,1599]" pageId="1" pageNumber="413" pagination="85 - 93" refId="ref6502" refString="Mao, B. Y., Xu, J. S. &amp; Li, M. (2017) Proposal of the Caryanda aurata species group (Orthoptera: Acrididae) with descriptions of two new species from China. Entomotaxonomia, 39 (2), 85 - 93." type="journal article" year="2017">Mao, Xu &amp; Li, 2017</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="0C428D48FFF8FF8AD5A8F9BAFA44F95D" blockId="1.[151,1437,1079,1717]" pageId="1" pageNumber="413">
<keyLead id="B70C2E7DFFF8FF8AD5A8F9BAFA44F95D" pageId="1" pageNumber="413">
- Abdominal tergites black with 3 yellow longitudinal stripes in male; hind femora green with or without a yellow preapical ring in male, or green with or without a red preapical ring in female; female subgenital plate broad, near rectangular, median area concaved, posterior margin nearly straight but weakly excised near both sides.......................................................................................................
<taxonomicName id="CBFDF6CBFFF8FF8AD13EF962FB2CF95C" authority="Mao, Yin" authorityName="Mao, Yin" box="[1025,1268,1695,1717]" class="Insecta" family="Acrididae" genus="Caryanda" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="413" phylum="Arthropoda" rank="species" species="hongheensis">
<emphasis id="3E89515AFFF8FF8AD13EF962FB4CF95D" box="[1025,1172,1695,1716]" italics="true" pageId="1" pageNumber="413">C. hongheensis</emphasis>
Mao, Yin
</taxonomicName>
&amp; Song,
<emphasis id="3E89515AFFF8FF8AD06FF962FA44F95D" bold="true" box="[1360,1436,1695,1716]" pageId="1" pageNumber="413">sp. nov.</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>