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<document id="8FB3DDA982EEC986EBD518EB4851E118" ID-CLB-Dataset="123209" ID-DOI="10.35929/RSZ.0031" ID-GBIF-Dataset="72faffc3-802a-4d40-944d-4f6e5f002bbb" ID-ISSN="0035-418" ID-Zenodo-Dep="5743952" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638277245777" checkinUser="carolina" docAuthor="Schwendinger, Peter J., Lehmann-Graber, Christina, Hongpadharakiree, Komson &amp; Syuhadah, Nurul" docDate="2020" docId="6D7987B4C624FF9813EEF9185790FA2C" docLanguage="en" docName="RevueSuisseZool.127.2.423-453.pdf" docOrigin="Revue suisse de Zoologie 127 (2)" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Malayathele Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah 2020, gen. nov." docType="treatment" docVersion="9" lastPageNumber="437" masterDocId="9140FFCCC629FF971070FFDD567CFFEF" masterDocTitle="New euagrid spider species from Thailand and Malaysia, and new localities of Leptothele bencha (Arachnida: Araneae)" masterLastPageNumber="453" masterPageNumber="423" pageNumber="435" updateTime="1699471559310" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="DCC8D92AB4FF08E413E6238D857DF3A5">New euagrid spider species from Thailand and Malaysia, and new localities of Leptothele bencha (Arachnida: Araneae)</mods:title>
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<mods:name id="F788044D3287194386AD1F395D5F20A3" type="personal">
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<mods:namePart id="90FA86E5192E3B0762D56BDF2ECF6AB2">Schwendinger, Peter J.</mods:namePart>
<mods:affiliation id="3D6ABBA9C721A3E3AD1DA3F6F7A68A43">Muséum dhistoire naturelle de Genève, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="A04E089CCF8289674FDB34547C06D26A">Lehmann-Graber, Christina</mods:namePart>
<mods:affiliation id="18BE8A051D904F1D3311788E13751F32">Muséum dhistoire naturelle de Genève, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="8CE3E659450A0036ABF908DD24313694">Hongpadharakiree, Komson</mods:namePart>
<mods:affiliation id="1474FC61E36AF8143C9DC721AC3ADA32">Sirinath Rajini Mangrove Ecosystem Learning Center, Pranburi, Prachuab Khiri Khan, Thailand</mods:affiliation>
</mods:name>
<mods:name id="35E08BB6AA7685A21F20C1F65A3EBDCF" type="personal">
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<mods:namePart id="7051474D29A82B3DD62696CB8691C47A">Syuhadah, Nurul</mods:namePart>
<mods:affiliation id="06B2A82086273FC311224F1170BFF2C1">The Liphistius Project, Petaling Jaya, Malaysia</mods:affiliation>
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<mods:typeOfResource id="30B85FE424F378FF0FB5D61BE4B95A7A">text</mods:typeOfResource>
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<treatment id="6D7987B4C624FF9813EEF9185790FA2C" ID-DOI="http://doi.org/10.5281/zenodo.7004841" ID-GBIF-Taxon="199502568" ID-Zenodo-Dep="7004841" LSID="urn:lsid:plazi:treatment:6D7987B4C624FF9813EEF9185790FA2C" httpUri="http://treatment.plazi.org/id/6D7987B4C624FF9813EEF9185790FA2C" lastPageId="15" lastPageNumber="437" pageId="13" pageNumber="435">
<subSubSection id="ADCA6529C624FF9A13EEF9185342F932" box="[926,1342,1733,1757]" pageId="13" pageNumber="435" type="nomenclature">
<paragraph id="E56F36A2C624FF9A13EEF9185342F932" blockId="13.[926,1342,1733,1757]" box="[926,1342,1733,1757]" pageId="13" pageNumber="435">
<heading id="BE2781CEC624FF9A13EEF9185342F932" bold="true" box="[926,1342,1733,1757]" centered="true" fontSize="10" level="2" pageId="13" pageNumber="435" reason="2">
<emphasis id="D7A4EAB0C624FF9A13EEF9185342F932" bold="true" box="[926,1342,1733,1757]" pageId="13" pageNumber="435">
<taxonomicName id="22D04D21C624FF9A13EEF91852B2F932" ID-CoL="98X62" authority="Schwendinger" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[926,1230,1733,1757]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="435" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C624FF9A13EEF918525AF932" bold="true" box="[926,1062,1733,1757]" italics="true" pageId="13" pageNumber="435">Malayathele</emphasis>
Schwendinger
</taxonomicName>
,
<taxonomicNameLabel id="CC9757CBC624FF9A14AAF91B5342F932" box="[1242,1342,1734,1757]" pageId="13" pageNumber="435" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="ADCA6529C624FF9A134FF9285580F822" pageId="13" pageNumber="435" type="etymology">
<paragraph id="E56F36A2C624FF9A134FF9285580F822" blockId="13.[831,1437,1781,1997]" pageId="13" pageNumber="435">
<emphasis id="D7A4EAB0C624FF9A134FF92855B9F8E3" bold="true" box="[831,965,1781,1804]" pageId="13" pageNumber="435">Etymology:</emphasis>
“Malaya” is one of the old names for
<collectingCountry id="9DC77632C624FF9A134FF8CB55D8F8C2" box="[831,932,1814,1837]" name="Malaysia" pageId="13" pageNumber="435">Malaysia</collectingCountry>
, the only county from where the new spider genus is known with certainty; “thele” (Greek for “nipple”; figurative for nipple-shaped or wart-shaped structures), refers to the long posterior lateral spinnerets of these spiders and to the close relationship with the genus
<taxonomicName id="22D04D21C624FF9A13F4F86B5589F822" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[900,1013,1974,1997]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C624FF9A13F4F86B5589F822" box="[900,1013,1974,1997]" italics="true" pageId="13" pageNumber="435">Leptothele</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="ADCA6529C624FF9A134FF838526AF7F2" pageId="13" pageNumber="435" type="type_taxon">
<paragraph id="E56F36A2C624FF9A134FF838526AF7F2" blockId="13.[831,1436,2021,2077]" pageId="13" pageNumber="435">
<emphasis id="D7A4EAB0C624FF9A134FF8385598F813" bold="true" box="[831,996,2021,2044]" pageId="13" pageNumber="435">
<typeStatus id="3A6B8800C624FF9A134FF8385504F813" box="[831,888,2021,2044]" pageId="13" pageNumber="435">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="22D04D21C624FF9A1387F8385289F813" ID-CoL="99C9Q" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1015,1269,2021,2044]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="435" phylum="Arthropoda" rank="species" species="kanching" status="sp. nov.">
<emphasis id="D7A4EAB0C624FF9A1387F8385289F813" box="[1015,1269,2021,2044]" italics="true" pageId="13" pageNumber="435">Malayathele kanching</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC624FF9A1578F8385317F813" box="[1288,1387,2021,2044]" pageId="13" pageNumber="435" rank="species">sp. nov.</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="ADCA6529C627FF9910E7FEDF5416FCD5" pageId="14" pageNumber="436" type="diagnosis">
<paragraph id="E56F36A2C627FF9910E7FEDF5416FCD5" blockId="14.[151,757,258,826]" pageId="14" pageNumber="436">
<emphasis id="D7A4EAB0C627FF9910E7FEDF576DFEF6" bold="true" box="[151,273,258,281]" pageId="14" pageNumber="436">Diagnosis:</emphasis>
Distinguished from
<taxonomicName id="22D04D21C627FF991254FEDF54BEFEF6" authorityName="Simon" authorityYear="1889" box="[548,706,258,281]" class="Arachnida" family="Dipluridae" genus="Phyxioschema" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C627FF991254FEDF54BEFEF6" box="[548,706,258,281]" italics="true" pageId="14" pageNumber="436">Phyxioschema</emphasis>
</taxonomicName>
by characters given in the key to
<taxonomicName id="22D04D21C627FF9911ADFEFE5435FED5" authorityName="Raven" authorityYear="1979" box="[477,585,291,314]" class="Arachnida" family="Euagridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="family">Euagridae</taxonomicName>
genera of Asia, most prominently by small body size in both sexes, and by tibia II of males without elongate spinules on prolateral side (
<figureCitation id="7DEB2A27C627FF991134FE5E57D0FE75" box="[324,428,387,410]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">Figs 10B</figureCitation>
,
<figureCitation id="7DEB2A27C627FF9911CAFE5E5796FE75" box="[442,490,387,410]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">12G</figureCitation>
) and with only a single megaspine on a low ventromedian spur (
<figureCitation id="7DEB2A27C627FF991203FE7E54C0FE55" box="[627,700,419,442]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Figs 8</figureCitation>
H-K, 10B, 11G-J, 12G, 13J-M, 15F-H). Distinguished from
<taxonomicName id="22D04D21C627FF9910E7FE3E5775FE15" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[151,265,483,506]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C627FF9910E7FE3E5775FE15" box="[151,265,483,506]" italics="true" pageId="14" pageNumber="436">Leptothele</emphasis>
</taxonomicName>
by metatarsal preening combs present on legs II-IV in both sexes. Males with only one megaspine and no transversal subdistal ridge on ventral side of tibia II, and mostly with two small conical processes (a retroventral and a proventral one,
<figureCitation id="7DEB2A27C627FF991238FDBE54C8FD95" box="[584,692,611,634]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Figs 12G</figureCitation>
,
<figureCitation id="7DEB2A27C627FF9912B6FDBE549CFD95" box="[710,736,611,634]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">13</figureCitation>
J- M, 15F-H; the proventral one absent in
<taxonomicName id="22D04D21C627FF991209FD5E54BAFD75" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[633,710,643,666]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="species" species="ulu" status="sp. nov.">
<emphasis id="D7A4EAB0C627FF991209FD5E54BAFD75" box="[633,710,643,666]" italics="true" pageId="14" pageNumber="436">M. ulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF9912A5FD5E56BBFD55" pageId="14" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
,
<figureCitation id="7DEB2A27C627FF9910A6FD7E576BFD55" box="[214,279,675,698]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Fig. 8</figureCitation>
I-J, the retroventral one keel-shaped in
<taxonomicName id="22D04D21C627FF9912A7FD7E5755FD35" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="species" species="cameronensis" status="sp. nov.">
<emphasis id="D7A4EAB0C627FF9912A7FD7E5755FD35" italics="true" pageId="14" pageNumber="436">M. cameronensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF991142FD1E57F6FD35" box="[306,394,707,730]" pageId="14" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
,
<figureCitation id="7DEB2A27C627FF9911E7FD1E5782FD35" box="[407,510,707,730]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">Figs 10B</figureCitation>
,
<figureCitation id="7DEB2A27C627FF99127BFD1E545FFD35" box="[523,547,707,730]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">11</figureCitation>
G-J) on metatarsus II; tarsi I-II ventrally bulged (
<figureCitation id="7DEB2A27C627FF99127DFD3E5400FD15" box="[525,636,739,762]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">Fig. 10D</figureCitation>
). Females distinguished by possessing median receptacular stalks with sclerotised walls (
<figureCitation id="7DEB2A27C627FF9911FCFCFE57ADFCD5" box="[396,465,803,826]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="14" pageNumber="436">Figs 9</figureCitation>
,
<figureCitation id="7DEB2A27C627FF9911ADFCFE5470FCD5" box="[477,524,803,826]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">11K</figureCitation>
,
<figureCitation id="7DEB2A27C627FF991267FCFE5448FCD5" box="[535,564,803,826]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="14" pageNumber="436">14</figureCitation>
,
<figureCitation id="7DEB2A27C627FF99124FFCFE5421FCD5" box="[575,605,803,826]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="14" pageNumber="436">16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="ADCA6529C627FF9810E7FC8F5790FA2C" lastPageId="15" lastPageNumber="437" pageId="14" pageNumber="436" type="description">
<paragraph id="E56F36A2C627FF9910E7FC8F574EFB65" blockId="14.[151,757,850,1226]" pageId="14" pageNumber="436">
<emphasis id="D7A4EAB0C627FF9910E7FC8F5758FC86" bold="true" box="[151,292,850,873]" pageId="14" pageNumber="436">Description:</emphasis>
Body very small (carapace length in males 1.11-1.44), with relatively thin hair cover on carapace, no colour pattern (
<figureCitation id="7DEB2A27C627FF99111BFC4E57C7FC45" box="[363,443,915,938]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12</figureCitation>
E-F, H-I) or a dark mottled pattern on hirsute opisthosoma (
<figureCitation id="7DEB2A27C627FF991260FC6E541FFC25" box="[528,611,947,970]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12</figureCitation>
A-D). Fovea pitlike, with a pair of long straight bristles anterior of it (
<figureCitation id="7DEB2A27C627FF9910ABFC2E57CCFBE5" box="[219,432,1011,1034]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12A, C, E, H</figureCitation>
). Eye group distinctly wider than long, anterior eye row slightly or not procurved, posterior eye row moderately or slightly recurved. Clypeus present, shorter than eye group length (
<figureCitation id="7DEB2A27C627FF9912B8FB8E575AFB65" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12A, C, E, H</figureCitation>
).
</paragraph>
<paragraph id="E56F36A2C627FF9910E7FB4E5489FB25" blockId="14.[151,757,850,1226]" pageId="14" pageNumber="436">Chelicerae small, without rastellum, with teeth only on promargin of groove; intercheliceral tumescence absent.</paragraph>
<paragraph id="E56F36A2C627FF99134FFEDF524AFDBB" blockId="14.[831,1437,258,1225]" pageId="14" pageNumber="436">
Palpal coxae rectangular, quite wide and posteriorly arched, with a bulging ventral surface, its prolateral zone glabrous and lightly pigmented; no cuspules present (
<figureCitation id="7DEB2A27C627FF991501FE9C55D6FE98" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12B, F, I</figureCitation>
); anterior lobe indistinct, carrying a serrula formed by a narrow band of tiny denticles (
<figureCitation id="7DEB2A27C627FF991548FE5D53F2FE78" box="[1336,1422,384,407]" captionStart="Fig" captionStartId="2.[151,186,1849,1870]" captionTargetBox="[161,1430,259,1801]" captionTargetId="figure-12@2.[153,1438,254,1809]" captionTargetPageId="2" captionText="Fig. 1. Photos of unpaired leg claws, lateral view (A-J), and of serrula on palpal coxa, anteroventral view (K-L). (A) Masteria cf. cavicola (Simon, 1892), female (in MHNG; 13.IV.1977; leg. P. Strinati &amp; V. Aellen) from a cave near Bagio, Luzon, the Phillipines, leg II. (B) Same, leg IV.(C) Phyxioschema huberi Schwendinger, 2009, exuvia of female paratype, leg I. (D) Same, leg IV. (E) Leptothele bencha, female from Ko Sirey, leg I. (F) Same, leg III. (G) Leptothele chang sp. nov., exuvia of female paratype, leg I. (H) Same, leg III. (I) Malayathele ulu sp. nov., exuvia of female paratype, leg I. (J) Same, leg IV. (K) Leptothele chang sp. nov., exuvia of female paratype. (L) Malayathele kanching sp. nov., exuvia of female paratype. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743954" httpUri="https://zenodo.org/record/5743954/files/figure.png" pageId="14" pageNumber="436">Fig. 1L</figureCitation>
). Labium much wider than long, without cuspules; anterior edge carrying a row of bristles followed by a pallid zone without setae; posterior part pigmented, with few short setae. Sternum cordate; post-labial sigilla medially fused with quite long labiosternal suture; remaining three pairs of sigilla indiscernible.
</paragraph>
<paragraph id="E56F36A2C627FF99134FFD8154FDFAF2" blockId="14.[831,1437,258,1225]" lastBlockId="14.[151,641,1285,1309]" pageId="14" pageNumber="436">
Legs moderately long and thin, with spines on patellae to metatarsi; all tarsi aspinose and not pseudosegmented; tarsi I-II of males more or less distinctly bulged on ventral side and carrying a few scopuliform hairs (
<figureCitation id="7DEB2A27C627FF991501FD675525FD1E" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">Fig. 10</figureCitation>
C-E); metatarsi II-IV of both sexes with preening combs; filiform trichobothria in two dorsal rows on tibiae, and in one row on metatarsi and tarsi; tarsal organ inconspicuous, cowpad-shaped, quite remote from distal margin of tarsus. Leg I of males with only two (
<figureCitation id="7DEB2A27C627FF991548FC8553FDFC80" box="[1336,1409,856,879]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">Fig. 11</figureCitation>
E- F) or 6-7 (
<figureCitation id="7DEB2A27C627FF9913C6FCAA5587FC61" box="[950,1019,887,910]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Figs 8</figureCitation>
F-G, 13H-I, 15D-E) ventral spines on cylindrical, not incrassate tibia; patella with a series of bristles (
<figureCitation id="7DEB2A27C627FF9913EEFC6B558FFC22" box="[926,1011,950,973]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Figs 8F</figureCitation>
,
<figureCitation id="7DEB2A27C627FF991471FC6B524DFC22" box="[1025,1073,950,973]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">10A</figureCitation>
,
<figureCitation id="7DEB2A27C627FF99144FFC6B5214FC22" box="[1087,1128,950,973]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">11F</figureCitation>
,
<figureCitation id="7DEB2A27C627FF991406FC6B52E7FC22" box="[1142,1179,950,973]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">13I</figureCitation>
) or with a single spine (
<figureCitation id="7DEB2A27C627FF991337FC0B55D5FC02" box="[839,937,982,1005]" captionStart="Fig" captionStartId="27.[151,186,1849,1870]" captionTargetBox="[157,1426,263,1804]" captionTargetId="figure-83@27.[151,1432,257,1810]" captionTargetPageId="27" captionText="Fig. 15. Malayathele kanching sp. nov.; male holotype (A-B, D-G) and male paratype (C, H). (A) Distal part of right palp, ventral view. (B-C) Same of left palp, prolateral view. (D) Patella to metatarsus of left leg I, prolateral view. (E) Patella and tibia of left leg I, ventral view. (F) Patella to metatarsus of left leg II, ventral view. (G-H) Same, prolateral view. Arrow indicates single spine retroventrally-distally on patella I. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743987" httpUri="https://zenodo.org/record/5743987/files/figure.png" pageId="14" pageNumber="436">Fig. 15E</figureCitation>
) retroventrally-distally. Leg II of males with a small conical proventral process and a small conical retroventral processes (
<figureCitation id="7DEB2A27C627FF99144BFBC852DFFBC3" box="[1083,1187,1045,1068]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Figs 12G</figureCitation>
,
<figureCitation id="7DEB2A27C627FF9914C0FBC852B5FBC3" box="[1200,1225,1045,1068]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">13</figureCitation>
J-K, M, 15F-H), or with only a small conical retroventral process (
<figureCitation id="7DEB2A27C627FF991542FBE95313FBA4" box="[1330,1391,1076,1099]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Fig. 8</figureCitation>
I-J), or with a small conical proventral process and a longer keel-shaped retroventral process (
<figureCitation id="7DEB2A27C627FF9914C4FBAE5362FB65" box="[1204,1310,1139,1162]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">Figs 10B</figureCitation>
,
<figureCitation id="7DEB2A27C627FF99155DFBAE5339FB65" box="[1325,1349,1139,1162]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">11</figureCitation>
G-J) on metatarsus; tibia with a single, slightly curved or slightly sigmoid megaspine on a low medioventral spur; area
<emphasis id="D7A4EAB0C627FF9910E7FADB54FDFAF2" bold="true" box="[151,641,1285,1309]" pageId="14" pageNumber="436">
Key to the species of
<taxonomicName id="22D04D21C627FF9911F5FAD85471FAF2" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[389,525,1285,1309]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="D7A4EAB0C627FF9911F5FAD85471FAF2" bold="true" box="[389,525,1285,1309]" italics="true" pageId="14" pageNumber="436">Malayathele</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF991264FADB5400FAF2" box="[532,636,1286,1309]" pageId="14" pageNumber="436" rank="genus">gen. nov.</taxonomicNameLabel>
:
</emphasis>
</paragraph>
<paragraph id="E56F36A2C627FF9910E7FA9B53E1FA52" blockId="14.[151,1437,1350,1597]" pageId="14" pageNumber="436">
<key id="91416193C627FF9910E7FA9B53E1FA52" pageId="14" pageNumber="436">
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1A Males with corkscrew-shaped embolus (
<figureCitation id="7DEB2A27C627FF9912FDFA9B54AAFAB2" box="[653,726,1350,1373]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">Fig. 11</figureCitation>
A-D), with only two ventral spines on metatarsus I (
<figureCitation id="7DEB2A27C627FF99148AFA9B5323FAB2" box="[1274,1375,1350,1373]" captionStart="Fig" captionStartId="20.[151,186,2012,2033]" captionTargetBox="[157,1434,1019,1973]" captionTargetId="figure-450@20.[153,1438,1015,1976]" captionTargetPageId="20" captionText="Fig. 10. Malayathele cameronensis sp. nov., male holotype. (A) Left leg I, retrolateral view; one ventral spine on tibia fallen off. (B) Left leg II, prolateral view. (C) Unpaired claw and scopuliform hairs of left leg II, prolateral view. (D) Ventrally bulged tarsus of left leg II, prolateral view. (E) Scopuliform hair on left tarsus II. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743975" httpUri="https://zenodo.org/record/5743975/files/figure.png" pageId="14" pageNumber="436">Figs 10A</figureCitation>
,
<figureCitation id="7DEB2A27C627FF991519FA9B53FDFAB2" box="[1385,1409,1350,1373]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">11</figureCitation>
E- F) and with retroventral process on metatarsus II developed as a short blunt keel (
<figureCitation id="7DEB2A27C627FF99143EFABB52E5FA92" box="[1102,1177,1382,1405]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">Fig. 11</figureCitation>
G-J). Females with very wide, quite short spermathecal trunks bearing rudimentary (reduced to a boss) lateral receptacles (
<figureCitation id="7DEB2A27C627FF991487FA5B532BFA72" box="[1271,1367,1414,1437]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="14" pageNumber="436">Fig. 11K</figureCitation>
) ....... ............................................................................................................................................
<taxonomicName id="22D04D21C627FF9914FEFA7B5338FA52" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1166,1348,1446,1469]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="species" species="cameronensis" status="sp. nov.">
<emphasis id="D7A4EAB0C627FF9914FEFA7B5338FA52" box="[1166,1348,1446,1469]" italics="true" pageId="14" pageNumber="436">M. cameronensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF99153AFA7B53E1FA52" box="[1354,1437,1446,1469]" pageId="14" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
<paragraph id="E56F36A2C627FF9910E7FA1B53E0F9D2" blockId="14.[151,1437,1350,1597]" pageId="14" pageNumber="436">
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<keyStep id="5E242E07C627FF9910E7FA1B53E0F9D2" pageId="14" pageNumber="436">
1B Males with more or less strongly curved embolus (
<figureCitation id="7DEB2A27C627FF99129EFA1B5552FA32" box="[750,814,1478,1501]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Figs 8</figureCitation>
A-E, 13A-G, 15A-C), with 6-7 ventral spines on metatarsus I (
<figureCitation id="7DEB2A27C627FF991089FA3B5747FA12" box="[249,315,1510,1533]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Figs 8</figureCitation>
F-G, 13H-I, 15D-E) and with retroventral process on metatarsus II developed as a small cone (
<figureCitation id="7DEB2A27C627FF991553FA3B5319FA12" box="[1315,1381,1510,1533]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Figs 8</figureCitation>
H-K, 13J-M, 15F-H). Females with narrower spermathecal trunks and with well-developed lateral receptacles (
<figureCitation id="7DEB2A27C627FF991521F9DB53EBF9F2" box="[1361,1431,1542,1565]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="14" pageNumber="436">Figs 9</figureCitation>
,
<figureCitation id="7DEB2A27C627FF991093F9FB577CF9D2" box="[227,256,1574,1597]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="14" pageNumber="436">14</figureCitation>
,
<figureCitation id="7DEB2A27C627FF99117BF9FB575AF9D2" box="[267,294,1574,1597]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="14" pageNumber="436">16</figureCitation>
) ...................................................................................................................................................................... 2
</keyStep>
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<keyStep id="5E242E07C627FF9910E7F9BB53E0F8F2" pageId="14" pageNumber="436">
<paragraph id="E56F36A2C627FF9910E7F9BB53E1F952" blockId="14.[151,1437,1638,1821]" pageId="14" pageNumber="436">
<keyLead id="5E219597C627FF9910E7F9BB53E1F952" pageId="14" pageNumber="436">
2A Males with only slightly curved embolus (
<figureCitation id="7DEB2A27C627FF9912D9F9BB549BF992" box="[681,743,1638,1661]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Fig. 8</figureCitation>
A-E) and with only one ventral process (the retroventral one) on metatarsus II (
<figureCitation id="7DEB2A27C627FF99110BF95B57C5F972" box="[379,441,1670,1693]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="14" pageNumber="436">Fig. 8</figureCitation>
H-K). Females without dark pattern on opisthosoma; stalks of median receptacles longer than corresponding heads (
<figureCitation id="7DEB2A27C627FF9911BDF97B5471F952" box="[461,525,1702,1725]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="14" pageNumber="436">Fig. 9</figureCitation>
) .............................................................................................................
<taxonomicName id="22D04D21C627FF99148EF97B533FF952" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1278,1347,1702,1725]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="species" species="ulu" status="sp. nov.">
<emphasis id="D7A4EAB0C627FF99148EF97B533FF952" box="[1278,1347,1702,1725]" italics="true" pageId="14" pageNumber="436">M. ulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF99153AF97B53E1F952" box="[1354,1437,1702,1725]" pageId="14" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="E56F36A2C627FF9910E7F91B53E0F8F2" blockId="14.[151,1437,1638,1821]" pageId="14" pageNumber="436">
<keyLead id="5E219597C627FF9910E7F91B53E0F8F2" pageId="14" pageNumber="436">
2B Males with slightly or strongly curved embolus (
<figureCitation id="7DEB2A27C627FF991290F91B5551F932" box="[736,813,1734,1757]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">Figs 13</figureCitation>
A-G, 15A-C) and with two ventral processes on metatarsus II (
<figureCitation id="7DEB2A27C627FF991172F93B5714F912" box="[258,360,1766,1789]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Figs 12G</figureCitation>
,
<figureCitation id="7DEB2A27C627FF991103F93B57F0F912" box="[371,396,1766,1789]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">13</figureCitation>
J-M, 15F-H). Females with more or less distinct dark pattern on opisthosoma (
<figureCitation id="7DEB2A27C627FF9914BEF93B5365F912" box="[1230,1305,1766,1789]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12</figureCitation>
A-D); stalks of median receptacles as long as or shorter than corresponding heads (
<figureCitation id="7DEB2A27C627FF9913BEF8DB525DF8F2" box="[974,1057,1798,1821]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="14" pageNumber="436">Figs 14</figureCitation>
,
<figureCitation id="7DEB2A27C627FF99145CF8DB523BF8F2" box="[1068,1095,1798,1821]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="14" pageNumber="436">16</figureCitation>
) .............................................. 3
</keyLead>
</paragraph>
</keyStep>
<paragraph id="E56F36A2C627FF9910E7F89B53E1F872" blockId="14.[151,1437,1862,2077]" pageId="14" pageNumber="436">
<keyStep id="5E242E07C627FF9910E7F89B53E1F872" pageId="14" pageNumber="436">
<keyLead id="5E219597C627FF9910E7F89B53E1F872" pageId="14" pageNumber="436">
3A Males with distally strongly curved embolus (
<figureCitation id="7DEB2A27C627FF9912AFF89B5551F8B2" box="[735,813,1862,1885]" captionStart="Fig" captionStartId="27.[151,186,1849,1870]" captionTargetBox="[157,1426,263,1804]" captionTargetId="figure-83@27.[151,1432,257,1810]" captionTargetPageId="27" captionText="Fig. 15. Malayathele kanching sp. nov.; male holotype (A-B, D-G) and male paratype (C, H). (A) Distal part of right palp, ventral view. (B-C) Same of left palp, prolateral view. (D) Patella to metatarsus of left leg I, prolateral view. (E) Patella and tibia of left leg I, ventral view. (F) Patella to metatarsus of left leg II, ventral view. (G-H) Same, prolateral view. Arrow indicates single spine retroventrally-distally on patella I. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743987" httpUri="https://zenodo.org/record/5743987/files/figure.png" pageId="14" pageNumber="436">Fig. 15</figureCitation>
A-C); patella I retroventrally-distally with a single spine (
<figureCitation id="7DEB2A27C627FF99109BF8BB5737F892" box="[235,331,1894,1917]" captionStart="Fig" captionStartId="27.[151,186,1849,1870]" captionTargetBox="[157,1426,263,1804]" captionTargetId="figure-83@27.[151,1432,257,1810]" captionTargetPageId="27" captionText="Fig. 15. Malayathele kanching sp. nov.; male holotype (A-B, D-G) and male paratype (C, H). (A) Distal part of right palp, ventral view. (B-C) Same of left palp, prolateral view. (D) Patella to metatarsus of left leg I, prolateral view. (E) Patella and tibia of left leg I, ventral view. (F) Patella to metatarsus of left leg II, ventral view. (G-H) Same, prolateral view. Arrow indicates single spine retroventrally-distally on patella I. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743987" httpUri="https://zenodo.org/record/5743987/files/figure.png" pageId="14" pageNumber="436">Fig. 15E</figureCitation>
). Females with dark pattern of opisthosoma indistinct and restricted to dorsal side; spermathecal trunks rectangular, lateral receptacles bent ventrad, their stalks not constricted (
<figureCitation id="7DEB2A27C627FF991397F85B524FF872" box="[999,1075,1926,1949]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="14" pageNumber="436">Fig. 16</figureCitation>
) ..................
<taxonomicName id="22D04D21C627FF9914CEF85B5338F872" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1214,1348,1926,1949]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="species" species="kanching" status="sp. nov.">
<emphasis id="D7A4EAB0C627FF9914CEF85B5338F872" box="[1214,1348,1926,1949]" italics="true" pageId="14" pageNumber="436">M. kanching</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF99153AF85B53E1F872" box="[1354,1437,1926,1949]" pageId="14" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="E56F36A2C627FF9810E7F87B56A8FE38" blockId="14.[151,1437,1862,2077]" lastBlockId="15.[151,757,258,1234]" lastPageId="15" lastPageNumber="437" pageId="14" pageNumber="436">
<keyLead id="5E219597C627FF9910E7F87B53E1F7F2" pageId="14" pageNumber="436">
3B Males with only slightly curved embolus (
<figureCitation id="7DEB2A27C627FF9912C3F87B557CF852" box="[691,768,1958,1981]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">Fig. 13</figureCitation>
A-G); patella I retroventrally-distally with a series of bristles (
<figureCitation id="7DEB2A27C627FF99109BF81B5738F832" box="[235,324,1990,2013]" captionStart="Fig" captionStartId="24.[149,184,1934,1955]" captionTargetBox="[152,1427,583,1872]" captionTargetId="figure-267@24.[149,1433,576,1879]" captionTargetPageId="24" captionText="Fig. 13. Malayathele maculosa sp. nov., male holotype (A-C, H-L) and two male paratypes (D-E; F-G, M). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C, E, G) Palpal organ of right palp, prolateral view. (D, F) Palpal organ of left palp, prolateral view. (H) Patella to metatarsus of left leg I, prolateral view. (I) Patella to tibia of left leg I, ventral view. (J) Patella to metatarsus of right leg II, ventral view. (K) Same of left leg II, prolateral view. (L) Same of right leg II, retrolateral view. (M) Same of left leg II, retrodorsal view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743983" httpUri="https://zenodo.org/record/5743983/files/figure.png" pageId="14" pageNumber="436">Fig. 13I</figureCitation>
). Females with dark pattern of opisthosoma distinct and extending to posteroventral side (
<figureCitation id="7DEB2A27C627FF99157FF81B5320F832" box="[1295,1372,1990,2013]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="14" pageNumber="436">Fig. 12</figureCitation>
A-D); spermathecal trunks conical, lateral receptacles directed anteriad, their stalks clearly constricted (
<figureCitation id="7DEB2A27C627FF99149EF83B5347F812" box="[1262,1339,2022,2045]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="14" pageNumber="436">Fig. 14</figureCitation>
) ........... ..................................................................................................................................................
<taxonomicName id="22D04D21C627FF9914CAF7DB533FF7F2" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[1210,1347,2054,2077]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="436" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D7A4EAB0C627FF9914CAF7DB533FF7F2" box="[1210,1347,2054,2077]" italics="true" pageId="14" pageNumber="436">M. maculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC627FF99153AF7DB53E1F7F2" box="[1354,1437,2054,2077]" pageId="14" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
between ventral spur and distoventral margin of tibia slightly flattened and wrinkled (
<figureCitation id="7DEB2A27C626FF98119FFEFF544FFED6" box="[495,563,290,313]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="15" pageNumber="437">Figs 8</figureCitation>
H-K, 10B, 11H-J, 12G, 13J-M; indistinct wrinkles in
<taxonomicName id="22D04D21C626FF98127FFE9C54E9FEB7" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[527,661,321,344]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="437" phylum="Arthropoda" rank="species" species="kanching" status="sp. nov.">
<emphasis id="D7A4EAB0C626FF98127FFE9C54E9FEB7" box="[527,661,321,344]" italics="true" pageId="15" pageNumber="437">M. kanching</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC626FF9812EBFE9C548CFEB7" box="[667,752,321,344]" pageId="15" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
,
<figureCitation id="7DEB2A27C626FF9810E7FEBC569EFE97" box="[151,226,353,376]" captionStart="Fig" captionStartId="27.[151,186,1849,1870]" captionTargetBox="[157,1426,263,1804]" captionTargetId="figure-83@27.[151,1432,257,1810]" captionTargetPageId="27" captionText="Fig. 15. Malayathele kanching sp. nov.; male holotype (A-B, D-G) and male paratype (C, H). (A) Distal part of right palp, ventral view. (B-C) Same of left palp, prolateral view. (D) Patella to metatarsus of left leg I, prolateral view. (E) Patella and tibia of left leg I, ventral view. (F) Patella to metatarsus of left leg II, ventral view. (G-H) Same, prolateral view. Arrow indicates single spine retroventrally-distally on patella I. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743987" httpUri="https://zenodo.org/record/5743987/files/figure.png" pageId="15" pageNumber="437">Fig. 15</figureCitation>
F-H). Paired leg claws with a single row of 4-12 teeth; unpaired claws with 1-3 quite long teeth, the basal ones on posterior legs set on a common base (
<figureCitation id="7DEB2A27C626FF9812FCFE7C54B1FE57" box="[652,717,417,440]" captionStart="Fig" captionStartId="2.[151,186,1849,1870]" captionTargetBox="[161,1430,259,1801]" captionTargetId="figure-12@2.[153,1438,254,1809]" captionTargetPageId="2" captionText="Fig. 1. Photos of unpaired leg claws, lateral view (A-J), and of serrula on palpal coxa, anteroventral view (K-L). (A) Masteria cf. cavicola (Simon, 1892), female (in MHNG; 13.IV.1977; leg. P. Strinati &amp; V. Aellen) from a cave near Bagio, Luzon, the Phillipines, leg II. (B) Same, leg IV.(C) Phyxioschema huberi Schwendinger, 2009, exuvia of female paratype, leg I. (D) Same, leg IV. (E) Leptothele bencha, female from Ko Sirey, leg I. (F) Same, leg III. (G) Leptothele chang sp. nov., exuvia of female paratype, leg I. (H) Same, leg III. (I) Malayathele ulu sp. nov., exuvia of female paratype, leg I. (J) Same, leg IV. (K) Leptothele chang sp. nov., exuvia of female paratype. (L) Malayathele kanching sp. nov., exuvia of female paratype. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743954" httpUri="https://zenodo.org/record/5743954/files/figure.png" pageId="15" pageNumber="437">Figs 1</figureCitation>
I-J, 10C).
</paragraph>
<paragraph id="E56F36A2C626FF9810E7FE3D54DCFD59" blockId="15.[151,757,258,1234]" pageId="15" pageNumber="437">
Palpal tarsus of males with two apical lobes, the retrodorsal lobe larger than the prodorsal one and carrying 2-3 distal spines (as in
<taxonomicName id="22D04D21C626FF981252FDFD54EFFDD8" authorityName="Raven &amp; Schwendinger" authorityYear="1995" box="[546,659,544,567]" class="Arachnida" family="Dipluridae" genus="Leptothele" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="437" phylum="Arthropoda" rank="genus">
<emphasis id="D7A4EAB0C626FF981252FDFD54EFFDD8" box="[546,659,544,567]" italics="true" pageId="15" pageNumber="437">Leptothele</emphasis>
</taxonomicName>
); palpal organ with short lenticular subtegulum and wide bulbous tegular part, embolic part thin and apically pointed, either slightly curved (
<figureCitation id="7DEB2A27C626FF981123FDA257E0FD79" box="[339,412,639,662]" captionStart="Fig" captionStartId="16.[151,186,1938,1959]" captionTargetBox="[157,1430,263,1909]" captionTargetId="figure-100@16.[151,1435,257,1914]" captionTargetPageId="16" captionText="Fig. 8. Malayathele ulu sp. nov.; male paratype (A-B, J-K) and male holotype (C-I). (A, C) Distal part of left palp, prolateral view. (B, D) Same of right palp, prolateral view. (E) Distal part of left palp, ventral view. (F) Patella to metatarsus of right leg I, prolateral view. (G) Right tibia I, ventral view. (H) Patella to metatarsus of right leg II, prolateral view. (I) Same, retroventral view. (J) Patella to metatarsus of left leg II, ventral view. (K) Same, prolateral view. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743970" httpUri="https://zenodo.org/record/5743970/files/figure.png" pageId="15" pageNumber="437">Figs 8</figureCitation>
A-E, 13A-G), strongly curved (
<figureCitation id="7DEB2A27C626FF9810EFFD425695FD59" box="[159,233,671,694]" captionStart="Fig" captionStartId="27.[151,186,1849,1870]" captionTargetBox="[157,1426,263,1804]" captionTargetId="figure-83@27.[151,1432,257,1810]" captionTargetPageId="27" captionText="Fig. 15. Malayathele kanching sp. nov.; male holotype (A-B, D-G) and male paratype (C, H). (A) Distal part of right palp, ventral view. (B-C) Same of left palp, prolateral view. (D) Patella to metatarsus of left leg I, prolateral view. (E) Patella and tibia of left leg I, ventral view. (F) Patella to metatarsus of left leg II, ventral view. (G-H) Same, prolateral view. Arrow indicates single spine retroventrally-distally on patella I. Scale line 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5743987" httpUri="https://zenodo.org/record/5743987/files/figure.png" pageId="15" pageNumber="437">Fig. 15</figureCitation>
A-C) or corkscrew-shaped (
<figureCitation id="7DEB2A27C626FF981265FD425423FD59" box="[533,607,671,694]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="15" pageNumber="437">Fig. 11</figureCitation>
A-D).
</paragraph>
<paragraph id="E56F36A2C626FF9810E7FD6356C6FC9B" blockId="15.[151,757,258,1234]" pageId="15" pageNumber="437">
Four widely separated spinnerets; no australotheline crescent at base of PLS; PMS short and with only one article; PLS long and composed of three articles (
<figureCitation id="7DEB2A27C626FF9812E2FD2356D0FCDA" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="15" pageNumber="437">Fig. 12B, D</figureCitation>
, E-F, H-I); distal article of PLS not pseudosegmented, with a reduced pigmentation in posterior half (
<figureCitation id="7DEB2A27C626FF9812E3FCE3549CFCBA" box="[659,736,830,853]" captionStart="Fig" captionStartId="23.[151,186,1913,1934]" captionTargetBox="[157,1433,261,1852]" captionTargetId="figure-85@23.[153,1438,257,1862]" captionTargetPageId="23" captionText="Fig. 12. Malayathele maculosa sp. nov., female paratype from Genting Highlands (A-D), M. kanching sp. nov., male holotype (E-G), female allotype (H-I) and male paratype (J-K). (A, E, H) Habitus, dorsal view. (B, F, I) Habitus, ventral view. (C) Habitus, dorsolateral view. (D) Opisthosoma, posteroventral view. (G) Tibia and metatarsus of right leg II, prolateral view. (J) Distal part of proximal article and entire median and distal articles of PLS, lateral view. (K) Same, ventral view. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743981" httpUri="https://zenodo.org/record/5743981/files/figure.png" pageId="15" pageNumber="437">Fig. 12</figureCitation>
J- K).
</paragraph>
<paragraph id="E56F36A2C626FF9810E7FCA05696FB3D" blockId="15.[151,757,258,1234]" pageId="15" pageNumber="437">
Vulva with two spermathecae, each carrying two receptacles; lateral receptacles anteriad-directed (
<figureCitation id="7DEB2A27C626FF9812B5FC4056DAFC3B" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="15" pageNumber="437">Figs 9</figureCitation>
,
<figureCitation id="7DEB2A27C626FF9810C4FC60569FFC3B" box="[180,227,957,980]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="15" pageNumber="437">11K</figureCitation>
,
<figureCitation id="7DEB2A27C626FF981081FC605770FC3B" box="[241,268,957,980]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="15" pageNumber="437">14</figureCitation>
) or ventrad-directed (
<figureCitation id="7DEB2A27C626FF98118CFC605431FC3B" box="[508,589,957,980]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="15" pageNumber="437">Fig. 16</figureCitation>
), perforated by numerous gland pores for their entire length, with more or less distinctly constricted, unsclerotised stalks (
<figureCitation id="7DEB2A27C626FF9812B5FC2156DAFBDC" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="15" pageNumber="437">Figs 9</figureCitation>
,
<figureCitation id="7DEB2A27C626FF9810C4FBC156ADFBDC" box="[180,209,1052,1075]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="15" pageNumber="437">14</figureCitation>
), without discernible stalks (
<figureCitation id="7DEB2A27C626FF98127BFBC15426FBDC" box="[523,602,1052,1075]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="15" pageNumber="437">Fig. 16</figureCitation>
) or altogether strongly reduced (
<figureCitation id="7DEB2A27C626FF98112BFBE157BCFBBC" box="[347,448,1084,1107]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="15" pageNumber="437">Fig. 11K</figureCitation>
); median receptacles always clearly separated into a more or less strongly curved or twisted stalk with sclerotised walls and a bulbous or globular head perforated by few or no pores (
<figureCitation id="7DEB2A27C626FF981202FB4654CAFB5D" box="[626,694,1179,1202]" captionStart="Fig" captionStartId="17.[151,186,1715,1736]" captionTargetBox="[154,1431,263,1659]" captionTargetId="figure-71@17.[151,1435,259,1666]" captionTargetPageId="17" captionText="Fig. 9. Malayathele ulu sp. nov.; vulvae of seven females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B-C) Paratypes from the type locality (Ulu Groh). (D-G) Paratypes from the Cameron Highlands: Parit Falls, 26.IV.2009 (D); Tanah Rata, 7.VIII.1972 (E); Gunung Perdah, 2.-14.V.2009 (F); trails 4 &amp; 13, 23.III.1993, heads of median receptacles collapsed and right lateral receptacle torn off (G). Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743972" httpUri="https://zenodo.org/record/5743972/files/figure.png" pageId="15" pageNumber="437">Figs 9</figureCitation>
,
<figureCitation id="7DEB2A27C626FF9812B0FB465493FB5D" box="[704,751,1179,1202]" captionStart="Fig" captionStartId="21.[149,184,1911,1932]" captionTargetBox="[173,1419,262,1860]" captionTargetId="figure-123@21.[168,1427,254,1866]" captionTargetPageId="21" captionText="Fig. 11. Malayathele cameronensis sp. nov., male holotype (A-J) and female allotype (K). (A) Distal part of left palp, prolateral view. (B) Same, ventral view. (C) Distal part of right palp, proventral and slightly distal view. (D) Same, prolateral view. (E) Patella to metatarsus of right leg I, prolateral view. (F) Patella to tibia of right leg I, ventral view. (G) Patella to metatarsus of right leg II, prolateral view. (H) Same of left leg II, prolateral view. (I) Patella to metatarsus of left leg II, ventral view. (J) Same of right leg II, ventral view. (K) Vulva, dorsal view. Arrows indicate keel-shaped retroventral process on metatarsus II. Scale lines 0.5 mm (A-J), 0.1 mm (K)." figureDoi="http://doi.org/10.5281/zenodo.5743977" httpUri="https://zenodo.org/record/5743977/files/figure.png" pageId="15" pageNumber="437">11K</figureCitation>
,
<figureCitation id="7DEB2A27C626FF9810E7FB6656C8FB3D" box="[151,180,1211,1234]" captionStart="Fig" captionStartId="25.[151,186,2027,2048]" captionTargetBox="[155,1418,1378,1967]" captionTargetId="figure-608@25.[151,1422,1374,1976]" captionTargetPageId="25" captionText="Fig. 14. Malayathele maculosa sp. nov., vulvae of three females (all drawn from dissected specimens), dorsal view. (A) Allotype. (B) Paratype from Genting Highlands. (C) Juvenile female paratype from Frasers Hill. Scale lines 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743985" httpUri="https://zenodo.org/record/5743985/files/figure.png" pageId="15" pageNumber="437">14</figureCitation>
,
<figureCitation id="7DEB2A27C626FF9810CFFB6656A1FB3D" box="[191,221,1211,1234]" captionStart="Fig" captionStartId="28.[152,187,2025,2046]" captionTargetBox="[157,1428,1213,1987]" captionTargetId="figure-544@28.[151,1431,1208,1994]" captionTargetPageId="28" captionText="Fig. 16. Malayathele kanching sp. nov.; vulvae of four female paratypes, dorsal (A-B) to increasingly more postero-dorsal view (C- D). (A, C-D) Specimens from the Templer Park. (B) Specimen from Ulu Gombak. Scale lines 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.5743991" httpUri="https://zenodo.org/record/5743991/files/figure.png" pageId="15" pageNumber="437">16</figureCitation>
).
</paragraph>
<paragraph id="E56F36A2C626FF9810E7FB375790FA2C" blockId="15.[151,757,1258,1475]" pageId="15" pageNumber="437">
<emphasis id="D7A4EAB0C626FF9810E7FB3757BDFAEE" bold="true" box="[151,449,1258,1281]" pageId="15" pageNumber="437">Species and distribution:</emphasis>
<taxonomicName id="22D04D21C626FF9811BEFB375488FAEE" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[462,756,1258,1281]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="437" phylum="Arthropoda" rank="species" species="cameronensis" status="sp. nov.">
<emphasis id="D7A4EAB0C626FF9811BEFB375488FAEE" box="[462,756,1258,1281]" italics="true" pageId="15" pageNumber="437">Malayathele cameronensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC626FF9810E7FAD7568EFACE" box="[151,242,1290,1313]" pageId="15" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="22D04D21C626FF981171FAD757F0FACE" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[257,396,1290,1313]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="437" phylum="Arthropoda" rank="species" species="kanching" status="sp. nov.">
<emphasis id="D7A4EAB0C626FF981171FAD757F0FACE" box="[257,396,1290,1313]" italics="true" pageId="15" pageNumber="437">M. kanching</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC626FF9811E8FAD7578EFACE" box="[408,498,1290,1313]" pageId="15" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="22D04D21C626FF981271FAD754EDFACE" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[513,657,1290,1313]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="437" phylum="Arthropoda" rank="species" species="maculosa" status="sp. nov.">
<emphasis id="D7A4EAB0C626FF981271FAD754EDFACE" box="[513,657,1290,1313]" italics="true" pageId="15" pageNumber="437">M. maculosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC626FF9812ECFAD75489FACE" box="[668,757,1290,1313]" pageId="15" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="22D04D21C626FF9810B8FAF6576CFAAD" authority="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah, 2020" authorityName="Schwendinger &amp; Lehmann-Graber &amp; Hongpadharakiree &amp; Syuhadah" authorityYear="2020" box="[200,272,1323,1346]" class="Arachnida" family="Euagridae" genus="Malayathele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="437" phylum="Arthropoda" rank="species" species="ulu" status="sp. nov.">
<emphasis id="D7A4EAB0C626FF9810B8FAF6576CFAAD" box="[200,272,1323,1346]" italics="true" pageId="15" pageNumber="437">M. ulu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9757CBC626FF98116AFAF6570FFAAD" box="[282,371,1323,1346]" pageId="15" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
, all from the Titiwangsa Mountain Range (= Main Range, Banjaran Besar) of Peninsular
<collectingCountry id="9DC77632C626FF9810E7FAB65685FA6D" box="[151,249,1387,1410]" name="Malaysia" pageId="15" pageNumber="437">Malaysia</collectingCountry>
and its foothills (
<figureCitation id="7DEB2A27C626FF9811B3FAB65474FA6D" box="[451,520,1387,1410]" captionStart="Fig" captionStartId="4.[155,190,1853,1874]" captionTargetBox="[154,1433,260,1820]" captionTargetId="figure-12@4.[151,1437,256,1824]" captionTargetPageId="4" captionText="Fig. 2. Localities of Leptothele spp. and Malayathele spp. on the Thai-Malay Peninsula. The coast of Sumatra is omitted. 1 - Ko Sirey (L. bencha); 2 - Ko Yao Noi (L. bencha); 3 - Khao Phanom Bencha (L. bencha, type locality); 4 - Thab Khaek - Hang Nak Hill and Khlong Jilat (L. bencha); 5 - Ban Chong Phlie (L. bencha); 6 - Ko Lanta (L. bencha); 7 - Ko Muk (L. bencha); 8 - Ko Libong (L. bencha); 9 - Ko Sukon (L. bencha); 10 - Ko Tarutao (L. bencha); 11 - Tham Khao Chang Hai (L. chang sp. nov., type locality); 12 - Ulu Groh (M. ulu sp. nov., type locality); 13 - Chenderiang (Malayathele sp.); 14 - Cameron Highlands (M. cameronensis sp. nov., type locality; M. ulu sp. nov.); 15 - Frasers Hill (M. maculosa sp. nov., type locality); 16 - Genting Highlands (M. maculosa sp. nov.); 17 - Templer Park (M. kanching sp. nov.; type locality); 18 - Ulu Gombak (M. kanching sp. nov.); 19 - Jeram Pasu Waterfall (Malayathele sp.); 20 - Singapore (Euagridae/Dipluridae gen. sp.)." figureDoi="http://doi.org/10.5281/zenodo.5743958" httpUri="https://zenodo.org/record/5743958/files/figure.png" pageId="15" pageNumber="437">Fig. 2</figureCitation>
, localities 12-18). At least one undescribed species in the northeastern part of the country (
<figureCitation id="7DEB2A27C626FF98116FFA715723FA2C" box="[287,351,1452,1475]" captionStart="Fig" captionStartId="4.[155,190,1853,1874]" captionTargetBox="[154,1433,260,1820]" captionTargetId="figure-12@4.[151,1437,256,1824]" captionTargetPageId="4" captionText="Fig. 2. Localities of Leptothele spp. and Malayathele spp. on the Thai-Malay Peninsula. The coast of Sumatra is omitted. 1 - Ko Sirey (L. bencha); 2 - Ko Yao Noi (L. bencha); 3 - Khao Phanom Bencha (L. bencha, type locality); 4 - Thab Khaek - Hang Nak Hill and Khlong Jilat (L. bencha); 5 - Ban Chong Phlie (L. bencha); 6 - Ko Lanta (L. bencha); 7 - Ko Muk (L. bencha); 8 - Ko Libong (L. bencha); 9 - Ko Sukon (L. bencha); 10 - Ko Tarutao (L. bencha); 11 - Tham Khao Chang Hai (L. chang sp. nov., type locality); 12 - Ulu Groh (M. ulu sp. nov., type locality); 13 - Chenderiang (Malayathele sp.); 14 - Cameron Highlands (M. cameronensis sp. nov., type locality; M. ulu sp. nov.); 15 - Frasers Hill (M. maculosa sp. nov., type locality); 16 - Genting Highlands (M. maculosa sp. nov.); 17 - Templer Park (M. kanching sp. nov.; type locality); 18 - Ulu Gombak (M. kanching sp. nov.); 19 - Jeram Pasu Waterfall (Malayathele sp.); 20 - Singapore (Euagridae/Dipluridae gen. sp.)." figureDoi="http://doi.org/10.5281/zenodo.5743958" httpUri="https://zenodo.org/record/5743958/files/figure.png" pageId="15" pageNumber="437">Fig. 2</figureCitation>
, locality 19).
</paragraph>
</subSubSection>
</treatment>
</document>