220 lines
24 KiB
XML
220 lines
24 KiB
XML
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<mods:title id="58F2B7698DA5818F31964D33B944C7B3">Identification of Mediterranean marine gobies (Actinopterygii: Gobiidae) of the continental shelf from photographs of in situ individuals</mods:title>
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<mods:namePart id="ACFBE726F55FDB88B3560ACA842E57EA">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="FEAA576F7827FAE12F45C018F3C4EF9F">Renoult, Julien P.</mods:namePart>
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<mods:namePart id="44291FBEAAC7B03279FC71F14981B7D7">Svensen, Rudolf</mods:namePart>
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<mods:namePart id="255A7D7ED9A3ABA6CC5AAB8D604D687B">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="F3FFC8AC69957AEBA8288DB066CD99A6">Engin, Semih</mods:namePart>
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<mods:namePart id="99085A9FAE532E97D4837607F7C8114E">Louisy, Patrick</mods:namePart>
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<mods:title id="5CCA937F24B71F7E0559FF055C5544BD">Zootaxa</mods:title>
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<mods:date id="671105153E69CDF6E6FDEB4292A6E92B">2022</mods:date>
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<mods:number id="04BBE0939F41AEBD507E8931E894070A">2022-06-01</mods:number>
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<treatment id="616687CB3F09FF9CFF76F88FFD0FFE4C" ID-DOI="http://doi.org/10.5281/zenodo.6601547" ID-GBIF-Taxon="198574969" ID-Zenodo-Dep="6601547" LSID="urn:lsid:plazi:treatment:616687CB3F09FF9CFF76F88FFD0FFE4C" httpUri="http://treatment.plazi.org/id/616687CB3F09FF9CFF76F88FFD0FFE4C" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61">
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<subSubSection id="A1D565563F09FF9EFF76F88FFCB7F8C2" box="[151,855,1801,1828]" pageId="60" pageNumber="61" type="nomenclature">
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<heading id="B23881B13F09FF9EFF76F88FFCB7F8C2" bold="true" box="[151,855,1801,1828]" fontSize="11" level="2" pageId="60" pageNumber="61" reason="3">
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<emphasis id="DBBBEACF3F09FF9EFF76F88FFCB7F8C2" bold="true" box="[151,855,1801,1828]" pageId="60" pageNumber="61">
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<taxonomicName id="2ECF4D5E3F09FF9EFF76F88FFDDEF8C2" ID-CoL="3GQ39" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[151,574,1801,1828]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="paganellus">
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<emphasis id="DBBBEACF3F09FF9EFF76F88FFE99F8C5" bold="true" box="[151,377,1801,1828]" italics="true" pageId="60" pageNumber="61">Gobius paganellus</emphasis>
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<bibRefCitation id="8D5E4B2C3F09FF9EFE61F88FFDDEF8C2" author="Linnaeus, C." box="[384,574,1801,1827]" pageId="60" pageNumber="61" refId="ref60330" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Impensis Direct. Laurentii Salvii, Holmiae, ii + 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
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</taxonomicName>
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(
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<figureCitation id="71F42A583F09FF9EFDACF88FFD43F8C2" box="[589,675,1801,1827]" captionStart="FIGURE 57" captionStartId="61.[151,250,1770,1794]" captionTargetBox="[151,1436,273,1747]" captionTargetId="figure-72@61.[151,1436,273,1747]" captionTargetPageId="61" captionText="FIGURE 57. Live coloration in its natural habitat of Gobius paganellus. A) Light colored individual, photo by M. Kovačić, Bol, island of Brač, Croatia, central Adriatic Sea. B) Dark-colored male, photo by G. Kunz, Donji Kraj, Istria, Croatia, northern Adriatic Sea." figureDoi="http://doi.org/10.5281/zenodo.6601745" httpUri="https://zenodo.org/record/6601745/files/figure.png" pageId="60" pageNumber="61">Fig. 57</figureCitation>
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)—Rock Goby
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E97036DD3F09FF9EFF76F8C8FC6AF887" blockId="60.[151,906,1870,1894]" box="[151,906,1870,1894]" pageId="60" pageNumber="61">
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<treatmentCitation id="686E10CC3F09FF9EFF76F8C8FDCEF887" ID-DOI="http://doi.org/10.5281/zenodo.3913174" ID-Zenodo-Dep="3913174" LSID="urn:lsid:zoobank.org:act:C7BC417A-B3F3-419C-A487-0DEF517479A5" author="Linnaeus, C." box="[151,558,1870,1894]" httpUri="http://treatment.plazi.org/id/ACAB471AC9EE923F31D3B40F8AAB9E37" page="263" pageId="60" pageNumber="61" year="1758">
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<taxonomicName id="2ECF4D5E3F09FF9EFF76F8C8FDCEF887" ID-CoL="3GQ39" authority="Linnaeus, 1758: 263" authorityName="Linnaeus" authorityPageNumber="263" authorityYear="1758" box="[151,558,1870,1894]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="60" pageNumber="61" phylum="Chordata" rank="species" species="paganellus">
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<emphasis id="DBBBEACF3F09FF9EFF76F8C8FEB6F887" box="[151,342,1870,1894]" italics="true" pageId="60" pageNumber="61">Gobius paganellus</emphasis>
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<bibRefCitation id="8D5E4B2C3F09FF9EFEBDF8C8FDCEF887" author="Linnaeus, C." box="[348,558,1870,1894]" pageId="60" pageNumber="61" refId="ref60330" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Impensis Direct. Laurentii Salvii, Holmiae, ii + 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758: 263</bibRefCitation>
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</taxonomicName>
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,
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<typeStatus id="3674887F3F09FF9EFDD9F8C8FD84F887" box="[568,612,1870,1894]" pageId="60" pageNumber="61">type</typeStatus>
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locality: Mediterranean Sea.
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</paragraph>
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<paragraph id="E97036DD3F09FF9EFF76F812FD58F84F" blockId="60.[151,1436,1940,2038]" box="[151,696,1940,1967]" pageId="60" pageNumber="61">
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Size. Known adult size about
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<quantity id="2E379B383F09FF9EFE07F812FDCBF84E" box="[486,555,1940,1967]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="60" pageNumber="61" unit="cm" value="12.0">12 cm</quantity>
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total length.
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<subSubSection id="A1D565563F09FF9CFF26F83EFD0FFE4C" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61" type="description">
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<paragraph id="E97036DD3F09FF9FFF26F83EFB0DFF18" blockId="60.[151,1436,1940,2038]" lastBlockId="61.[151,1437,151,249]" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61">
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Morphology. D VI + I,12–15; A I,10–13; P 15–20. Medium-sized goby with a proportionally short and stocky body. Head wide and moderately depressed. Caudal peduncle deep, but lower than body. Dorsal fins of similar height, the first dorsal fin may be slightly higher, with a more or less rounded distal edge and no elongate spines.
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<emphasis id="DBBBEACF3F08FF9FFF76FF3CFD6CFF34" bold="true" box="[151,652,186,213]" pageId="61" pageNumber="62">Pectoral-fin free rays very well developed.</emphasis>
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Caudal fin rounded.
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<emphasis id="DBBBEACF3F08FF9FFC99FF3DFEC6FF18" bold="true" pageId="61" pageNumber="62">Predorsal area and nape entirely covered with large scales.</emphasis>
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Scales visible on body and usually also on predorsal area (Miller 1986;
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<bibRefCitation id="8D5E4B2C3F08FF9FFBB1FF59FB02FF18" author="Louisy, P." box="[1104,1250,222,249]" pageId="61" pageNumber="62" refId="ref60639" refString="Louisy, P. (2015) Europe and Mediterranean Marine Fish Identification Guide. 3 rd Edition. Eugen Ulmer Ed., Paris, 512 pp." type="book" year="2015">Louisy 2015</bibRefCitation>
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).
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</paragraph>
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||
<caption id="BDB066553F08FF9FFF76F96CFEFCF8AA" ID-DOI="http://doi.org/10.5281/zenodo.6601745" ID-Zenodo-Dep="6601745" httpUri="https://zenodo.org/record/6601745/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1770,1794]" targetBox="[151,1436,273,1747]" targetPageId="61">
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<paragraph id="E97036DD3F08FF9FFF76F96CFEFCF8AA" blockId="61.[151,1436,1770,1867]" pageId="61" pageNumber="62">
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<emphasis id="DBBBEACF3F08FF9FFF76F96CFEC2F8E2" bold="true" box="[151,290,1770,1795]" pageId="61" pageNumber="62">FIGURE 57.</emphasis>
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Live coloration in its natural habitat of
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<taxonomicName id="2ECF4D5E3F08FF9FFD23F96DFC62F8E2" authorityName="Linnaeus" authorityYear="1758" box="[706,898,1771,1795]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="61" pageNumber="62" phylum="Chordata" rank="species" species="paganellus">
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<emphasis id="DBBBEACF3F08FF9FFD23F96DFC62F8E2" box="[706,898,1771,1795]" italics="true" pageId="61" pageNumber="62">Gobius paganellus</emphasis>
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</taxonomicName>
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||
. A) Light colored individual, photo by M. Kovačić, Bol, island of Brač, Croatia, central Adriatic Sea. B) Dark-colored male, photo by G. Kunz, Donji Kraj, Istria, Croatia, northern Adriatic Sea.
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||
</paragraph>
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||
</caption>
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||
<paragraph id="E97036DD3F08FF9CFF26F8F5FD73FF34" blockId="61.[151,1436,1907,2042]" lastBlockId="62.[151,1436,150,429]" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">
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||
Live coloration. Background coloration highly variable, ranging from light beige to dark brown (
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||
<figureCitation id="71F42A583F08FF9FFACDF8F5FA70F86F" box="[1324,1424,1907,1934]" captionStart="FIGURE 57" captionStartId="61.[151,250,1770,1794]" captionTargetBox="[151,1436,273,1747]" captionTargetId="figure-72@61.[151,1436,273,1747]" captionTargetPageId="61" captionText="FIGURE 57. Live coloration in its natural habitat of Gobius paganellus. A) Light colored individual, photo by M. Kovačić, Bol, island of Brač, Croatia, central Adriatic Sea. B) Dark-colored male, photo by G. Kunz, Donji Kraj, Istria, Croatia, northern Adriatic Sea." figureDoi="http://doi.org/10.5281/zenodo.6601745" httpUri="https://zenodo.org/record/6601745/files/figure.png" pageId="61" pageNumber="62">Fig. 57a</figureCitation>
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). Nesting males occasionally entirely black, with just the first dorsal fin margin colored (
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<figureCitation id="71F42A583F08FF9FFB62F811FB07F853" box="[1155,1255,1943,1970]" captionStart="FIGURE 57" captionStartId="61.[151,250,1770,1794]" captionTargetBox="[151,1436,273,1747]" captionTargetId="figure-72@61.[151,1436,273,1747]" captionTargetPageId="61" captionText="FIGURE 57. Live coloration in its natural habitat of Gobius paganellus. A) Light colored individual, photo by M. Kovačić, Bol, island of Brač, Croatia, central Adriatic Sea. B) Dark-colored male, photo by G. Kunz, Donji Kraj, Istria, Croatia, northern Adriatic Sea." figureDoi="http://doi.org/10.5281/zenodo.6601745" httpUri="https://zenodo.org/record/6601745/files/figure.png" pageId="61" pageNumber="62">Fig. 57b</figureCitation>
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). Most frequent pattern: broad dark bars and thin alternate white saddles on the back, medium-sized dark blotches along lateral midline, and small dark markings on a light background below.
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<emphasis id="DBBBEACF3F08FF9CFC9CF866FEBCFF50" bold="true" lastPageId="62" lastPageNumber="63" pageId="61" pageNumber="62">Usually a pale to white bar between eye and corner of mouth</emphasis>
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; it is often preceded by a dark preorbital bar.
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<emphasis id="DBBBEACF3F0BFF9CFC88FF10FE7BFF34" bold="true" pageId="62" pageNumber="63">First dorsal fin usually dark with an orange to yellow marginal band</emphasis>
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(
|
||
<bibRefCitation id="8D5E4B2C3F0BFF9CFE48FF3DFD67FF34" author="Renoult, J. P. & Pillon, R. & Kovacic, M. & Louisy, P." box="[425,647,187,213]" pageId="62" pageNumber="63" refId="ref63723" refString="Renoult, J. P., Pillon, R., Kovacic, M. & Louisy, P. (2022) Frontiers in Fishwatching Series - Gobies of North-eastern Atlantic and the Mediterranean: Gobius and Thorogobius. Les cahiers de la fondation Biotope. [in press]" type="book" year="2022">
|
||
Renoult
|
||
<emphasis id="DBBBEACF3F0BFF9CFDEAFF3DFDDCFF34" box="[523,572,187,213]" italics="true" pageId="62" pageNumber="63">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E97036DD3F0BFF9CFF26FF59FAFDFF18" blockId="62.[151,1436,150,429]" box="[199,1309,223,249]" pageId="62" pageNumber="63">
|
||
Similar species.
|
||
<taxonomicName id="2ECF4D5E3F0BFF9CFE9EFF59FDE4FF19" authorityName="Bellotti" authorityYear="1888" box="[383,516,223,249]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="ater">
|
||
<emphasis id="DBBBEACF3F0BFF9CFE9EFF59FDE4FF19" box="[383,516,223,249]" italics="true" pageId="62" pageNumber="63">Gobius ater</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2ECF4D5E3F0BFF9CFDF3FF59FD60FF18" authorityName="Pallas" authorityYear="1814" box="[530,640,223,249]" class="Actinopterygii" family="Gobiidae" genus="Gobius" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="cobitis">
|
||
<emphasis id="DBBBEACF3F0BFF9CFDF3FF59FDC6FF18" box="[530,550,223,249]" italics="true" pageId="62" pageNumber="63">G</emphasis>
|
||
.
|
||
<emphasis id="DBBBEACF3F0BFF9CFDD4FF59FD60FF18" box="[565,640,223,249]" italics="true" pageId="62" pageNumber="63">cobitis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2ECF4D5E3F0BFF9CFD6CFF59FC37FF18" authorityName="Benthic" baseAuthorityName="Kolombatovic" baseAuthorityYear="1891" box="[653,983,223,249]" class="Actinopterygii" family="Gobiidae" genus="Millerigobius" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="macrocephalus">
|
||
<emphasis id="DBBBEACF3F0BFF9CFD6CFF59FC37FF18" box="[653,983,223,249]" italics="true" pageId="62" pageNumber="63">Millerigobius macrocephalus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2ECF4D5E3F0BFF9CFC04FF59FB63FF18" authorityName="Benthic" baseAuthorityName="Risso" baseAuthorityYear="1827" box="[997,1155,223,249]" class="Actinopterygii" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="zebrus">
|
||
<emphasis id="DBBBEACF3F0BFF9CFC04FF59FB63FF18" box="[997,1155,223,249]" italics="true" pageId="62" pageNumber="63">Zebrus zebrus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="2ECF4D5E3F0BFF9CFB70FF59FAF5FF18" authorityName="Kovacic, Sanda & Vukic" authorityYear="2021" box="[1169,1301,223,249]" class="Actinopterygii" family="Gobiidae" genus="Zebrus" kingdom="Animalia" order="Perciformes" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="pallaoroi">
|
||
<emphasis id="DBBBEACF3F0BFF9CFB70FF59FB41FF19" box="[1169,1185,223,248]" italics="true" pageId="62" pageNumber="63">Z</emphasis>
|
||
.
|
||
<emphasis id="DBBBEACF3F0BFF9CFB4FFF59FAF5FF18" box="[1198,1301,223,249]" italics="true" pageId="62" pageNumber="63">pallaoroi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="E97036DD3F0BFF9CFF26FE85FDCAFEA3" blockId="62.[151,1436,150,429]" pageId="62" pageNumber="63">
|
||
Habitat. Infralittoral species known from
|
||
<quantity id="2E379B383F0BFF9CFD75FE84FD1DFEFC" box="[660,765,258,285]" metricMagnitude="1" metricUnit="m" metricValue="1.025" metricValueMax="2.0" metricValueMin="0.05" pageId="62" pageNumber="63" unit="m" value="10.25" valueMax="20.0" valueMin="0.5">0.5–20 m</quantity>
|
||
depth on very diverse hard or mixed bottoms (
|
||
<bibRefCitation id="8D5E4B2C3F0BFF9CFAE0FE85FA77FEFC" author="Patzner, R. A." box="[1281,1431,259,285]" pageId="62" pageNumber="63" refId="ref62855" refString="Patzner, R. A. (2021) Gobiidae. Available from: http: // www. patzner. sbg. ac. at / Gobiidae / GobiidaeSpecies. htm (accessed 1 October 2021)" type="url" year="2021">Patzner 2021</bibRefCitation>
|
||
; P. Louisy unpublished observation).
|
||
</paragraph>
|
||
<paragraph id="E97036DD3F0BFF9CFF26FECDFD0FFE4C" blockId="62.[151,1436,150,429]" pageId="62" pageNumber="63">
|
||
Geographic distribution. The whole of the Mediterranean (Miller 1986), Black Sea (
|
||
<bibRefCitation id="8D5E4B2C3F0BFF9CFB8DFECDFA86FE84" author="Engin, S. & Seyhan, K." box="[1132,1382,331,357]" pageId="62" pageNumber="63" pagination="111 - 118" refId="ref53766" refString="Engin, S. & Seyhan, K. (2009) Biological characteristics of rock goby Gobius paganellus in the South eastern Black Sea. Acta Ichthyologica et Piscatoria, 39, 111 - 118. https: // doi. org / 10.3750 / AIP 2009.39.2.05" type="journal article" year="2009">Engin & Seyhan 2009</bibRefCitation>
|
||
) and Eastern Atlantic from
|
||
<collectingCountry id="91D8764D3F0BFF9CFE6EFEE9FDC2FE68" box="[399,546,367,393]" name="United Kingdom" pageId="62" pageNumber="63">Great Britain</collectingCountry>
|
||
to
|
||
<collectingCountry id="91D8764D3F0BFF9CFDA7FEE9FD7FFE68" box="[582,671,367,393]" name="Senegal" pageId="62" pageNumber="63">Senegal</collectingCountry>
|
||
(Miller 1986), including the Azores (
|
||
<bibRefCitation id="8D5E4B2C3F0BFF9CFBA0FEE9FAEFFE68" author="Santos, R. S. & Porteiro, F. M. & Barreiros, J. P." box="[1089,1295,367,393]" pageId="62" pageNumber="63" pagination="1 - 244" refId="ref64282" refString="Santos, R. S., Porteiro, F. M. & Barreiros, J. P. (1997) Marine fishes of the Azores: annotated checklist and bibliography. Bulletin of the University of Azores, Supplement 1, 1 - 244." type="journal article" year="1997">
|
||
Santos
|
||
<emphasis id="DBBBEACF3F0BFF9CFB75FEE9FB25FE68" box="[1172,1221,367,393]" italics="true" pageId="62" pageNumber="63">et al</emphasis>
|
||
. 1997
|
||
</bibRefCitation>
|
||
), the Canary Islands (
|
||
<bibRefCitation id="8D5E4B2C3F0BFF9CFF14FE15FE95FE4C" author="Wirtz, P." box="[245,373,403,429]" pageId="62" pageNumber="63" refId="ref66060" refString="Wirtz, P. (1994) Underwater Guide. Fish of Madeira, Canary Islands and Azores. Verlag Stephanie Naglschmid, Stuttgart, 126 pp." type="book" year="1994">Wirtz 1994</bibRefCitation>
|
||
) and Madeira (
|
||
<bibRefCitation id="8D5E4B2C3F0BFF9CFDC1FE15FD03FE4C" author="Wirtz, P. & Fricke, R. & Biscoito, M. J." box="[544,739,403,429]" pageId="62" pageNumber="63" pagination="1 - 26" refId="ref66145" refString="Wirtz, P., Fricke, R. & Biscoito, M. J. (2008) The coastal fishes of Madeira Island - new records and an annotated check-list. Zootaxa, 1715 (1), 1 - 26. https: // doi. org / 10.11646 / zootaxa. 1715.1.1" type="journal article" year="2008">
|
||
Wirtz
|
||
<emphasis id="DBBBEACF3F0BFF9CFD86FE15FD79FE4C" box="[615,665,403,429]" italics="true" pageId="62" pageNumber="63">et al</emphasis>
|
||
. 2008
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |