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<document id="221D13442C36900DB799744A0097B447" ID-DOI="10.11646/zootaxa.3616.3.3" ID-GBIF-Dataset="436ae0f7-a379-4cb9-a2ef-41a28fc8a279" ID-ISSN="1175-5326" ID-Zenodo-Dep="218827" ID-ZooBank="F70AC1EE-5A70-42B8-B1F2-55C734D26FD9" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460575923727" checkinUser="plazi" docAuthor="Hisamatsu, Sadatomo" docDate="2013" docId="421787CFF660C874D9F8FF2FFE2EFDF9" docLanguage="en" docName="zt03616p267.pdf" docOrigin="Zootaxa 3616 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybocephalus (Cybocephalus) politissimus Reitter 1898" docType="treatment" docVersion="9" lastPageNumber="262" masterDocId="BE2EFFB7F666C87DD96FFFB8FFE1FFC5" masterDocTitle="A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea)" masterLastPageNumber="267" masterPageNumber="253" pageNumber="259" updateTime="1698323146566" updateUser="plazi">
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<mods:title id="689A7164277ED0B3227B67282E92797B">A review of the Japanese Cybocephalidae (Coleoptera: Cucujoidea)</mods:title>
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<mods:namePart id="54202A816AC13547075927C3B02E2081">Hisamatsu, Sadatomo</mods:namePart>
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<mods:date id="28B397985F2C82E70F73A966C1773D59">2013</mods:date>
<mods:detail id="E635BC72F1286C077A1E50AFCE05005C" type="volume">
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<mods:identifier id="2C099312B47734023E804F6650C9897E" type="DOI">10.11646/zootaxa.3616.3.3</mods:identifier>
<mods:identifier id="C3674998D4FA12FE4CB076286FF85F99" type="GBIF-Dataset">436ae0f7-a379-4cb9-a2ef-41a28fc8a279</mods:identifier>
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<treatment id="421787CFF660C874D9F8FF2FFE2EFDF9" ID-DOI="http://doi.org/10.5281/zenodo.6150611" ID-GBIF-Taxon="119560037" ID-Zenodo-Dep="6150611" LSID="urn:lsid:plazi:treatment:421787CFF660C874D9F8FF2FFE2EFDF9" httpUri="http://treatment.plazi.org/id/421787CFF660C874D9F8FF2FFE2EFDF9" lastPageId="9" lastPageNumber="262" pageId="6" pageNumber="259">
<subSubSection id="82A46552F660C87BD9F8FF2FFC29FED4" pageId="6" pageNumber="259" type="nomenclature">
<paragraph id="CA0136D9F660C87BD9F8FF2FFCA0FF74" blockId="6.[151,833,151,210]" box="[151,833,151,177]" pageId="6" pageNumber="259">
<heading id="914981B5F660C87BD9F8FF2FFCA0FF74" bold="true" box="[151,833,151,177]" fontSize="11" level="1" pageId="6" pageNumber="259" reason="1">
<taxonomicName id="0DBE4D5AF660C87BD9F8FF2FFCA0FF74" ID-CoL="8RPL7" authority="Reitter, 1898" authorityName="Reitter" authorityYear="1898" box="[151,833,151,177]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="politissimus" subGenus="Cybocephalus">
<emphasis id="F8CAEACBF660C87BD9F8FF2FFCA0FF74" bold="true" box="[151,833,151,177]" pageId="6" pageNumber="259">
<emphasis id="F8CAEACBF660C87BD9F8FF2FFEA0FF74" bold="true" box="[151,321,151,177]" italics="true" pageId="6" pageNumber="259">Cybocephalus</emphasis>
(
<emphasis id="F8CAEACBF660C87BD83EFF2FFE1BFF74" bold="true" box="[337,506,151,177]" italics="true" pageId="6" pageNumber="259">Cybocephalus</emphasis>
)
<emphasis id="F8CAEACBF660C87BDB63FF2FFD7AFF74" bold="true" box="[524,667,151,177]" italics="true" pageId="6" pageNumber="259">politissimus</emphasis>
Reitter, 1898
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="CA0136D9F660C87BD9F8FF02FE99FF17" blockId="6.[151,833,151,210]" box="[151,376,186,210]" pageId="6" pageNumber="259">
<figureCitation id="52852A5CF660C87BD9F8FF02FF38FF17" box="[151,217,186,210]" captionStart="FIGURE 1" captionStartId="2.[151,250,1886,1909]" captionTargetBox="[165,1427,1464,1851]" captionTargetId="figure@2.[151,1436,1455,1862]" captionTargetPageId="2" captionText="FIGURE 1. Lateral habitus of Cybocephalidae spp. A, Cybocephalus (C.) nipponicus Endrödy-Younga, 1971 (male, Tokunomori, Ehime Pref.); B, Cybocephalus (C.) politissimus Reitter, 1898 (male, Yoshiji-chô, Shimane Pref.); C, Cybocephalus (C.) chlorocapitis sp. nov. (holotype, male, Amami Is.); D, Pastillus eminentithorax sp. nov. (paratype, female, Araike, Aichi Pref.). Scale bars = 1 mm." httpUri="https://zenodo.org/record/218828/files/figure.png" pageId="6" pageNumber="259">Figs 1</figureCitation>
B, 3C–D, 5, 6
</paragraph>
<paragraph id="CA0136D9F660C87BD9F8FF44FC29FED4" blockId="6.[151,1435,251,427]" box="[151,968,251,274]" pageId="6" pageNumber="259">
<taxonomicName id="0DBE4D5AF660C87BD9F8FF44FD85FED4" authority="Reitter, 1898: 117" authorityName="Reitter" authorityPageNumber="117" authorityYear="1898" box="[151,612,251,274]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF660C87BD9F8FF44FE44FED7" box="[151,421,252,274]" italics="true" pageId="6" pageNumber="259">Cybocephalus politissimus</emphasis>
Reitter, 1898: 117
</taxonomicName>
.
<typeStatus id="1505887BF660C87BDB01FF45FD41FED7" box="[622,672,253,274]" pageId="6" pageNumber="259">Type</typeStatus>
locality: Irkut in Ostsibirien.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F660C87BD9F8FEA2FE06FB88" pageId="6" pageNumber="259" type="description">
<paragraph id="CA0136D9F660C87BD9F8FEA2FAACFEF5" blockId="6.[151,1435,251,427]" box="[151,1357,282,304]" pageId="6" pageNumber="259">
<taxonomicName id="0DBE4D5AF660C87BD9F8FEA2FD27FEF5" authority="Endrody-Younga, 1968: 77" authorityName="Endrody-Younga" authorityPageNumber="77" authorityYear="1968" box="[151,710,282,304]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF660C87BD9F8FEA2FE44FEF5" box="[151,421,282,304]" italics="true" pageId="6" pageNumber="259">Cybocephalus politissimus</emphasis>
: Endrödy-Younga, 1968: 77
</taxonomicName>
[redescription, figure];—Kirejtshuk, 1992: 209 [in key, figure].
</paragraph>
<paragraph id="CA0136D9F660C87BD9F8FE81FBE2FE8A" blockId="6.[151,1435,251,427]" box="[151,1027,313,335]" pageId="6" pageNumber="259">
<taxonomicName id="0DBE4D5AF660C87BD9F8FE81FC65FE8A" authority="Jelinek &amp; Audisio, 2007: 490" authorityName="Jelinek &amp; Audisio" authorityPageNumber="490" authorityYear="2007" box="[151,900,313,335]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="politissimus" subGenus="Cybocephalus">
<emphasis id="F8CAEACBF660C87BD9F8FE81FEC6FE8A" box="[151,295,313,335]" italics="true" pageId="6" pageNumber="259">Cybocephalus</emphasis>
(
<emphasis id="F8CAEACBF660C87BD85AFE81FE25FE8A" box="[309,452,313,335]" italics="true" pageId="6" pageNumber="259">Cybocephalus</emphasis>
)
<emphasis id="F8CAEACBF660C87BD8BCFE81FDADFE8A" box="[467,588,313,335]" italics="true" pageId="6" pageNumber="259">politissimus</emphasis>
: Jelínek &amp; Audisio, 2007: 490
</taxonomicName>
[catalogue].
</paragraph>
<paragraph id="CA0136D9F660C87BD9F8FEE0FDD8FE49" blockId="6.[151,1435,251,427]" pageId="6" pageNumber="259">
<taxonomicName id="0DBE4D5AF660C87BD9F8FEE0FE3EFEAB" authority="Tian" authorityName="Tian" box="[151,479,344,366]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="flavicapitus">
<emphasis id="F8CAEACBF660C87BD9F8FEE0FE46FEAB" box="[151,423,344,366]" italics="true" pageId="6" pageNumber="259">Cybocephalus flavicapitus</emphasis>
Tian
</taxonomicName>
&amp; Yu, 1994b: 209.
<typeStatus id="1505887BF660C87BDBD4FEE1FD0CFEAB" box="[699,749,345,366]" pageId="6" pageNumber="259">Type</typeStatus>
locality: Shixing, Guangdong Province,
<collectingCountry id="B2A97649F660C87BDDCFFEE0FB01FEAB" box="[1184,1248,344,366]" name="China" pageId="6" pageNumber="259">China</collectingCountry>
. Synonymized by Kirejtshuk &amp; Fallahzadeh, 2008: 85.
</paragraph>
<paragraph id="CA0136D9F660C87BD9F8FE2DFB88FE6F" blockId="6.[151,1435,251,427]" box="[151,1129,405,427]" pageId="6" pageNumber="259">
<taxonomicName id="0DBE4D5AF660C87BD9F8FE2DFD5DFE6E" authority="Smith &amp; Bailey, 2007: 504" authorityName="Smith &amp; Bailey" authorityPageNumber="504" authorityYear="2007" box="[151,700,405,427]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="flavocapitis">
<emphasis id="F8CAEACBF660C87BD9F8FE2DFE43FE6E" box="[151,418,405,427]" italics="true" pageId="6" pageNumber="259">Cybocephalus flavocapitis</emphasis>
Smith &amp; Bailey, 2007: 504
</taxonomicName>
.
<typeStatus id="1505887BF660C87BDBA9FE2EFD19FE6E" box="[710,760,406,427]" pageId="6" pageNumber="259">Type</typeStatus>
locality: Kenting,
<collectingCountry id="B2A97649F660C87BDAD9FE2DFBE2FE6E" box="[950,1027,405,427]" name="Taiwan" pageId="6" pageNumber="259">Taiwan</collectingCountry>
,
<emphasis id="F8CAEACBF660C87BDD63FE2DFB88FE6F" bold="true" box="[1036,1129,405,426]" pageId="6" pageNumber="259">syn. nov.</emphasis>
</paragraph>
<paragraph id="CA0136D9F660C87BD9F8FE68FE06FB88" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">
<emphasis id="F8CAEACBF660C87BD9F8FE68FE6EFE2C" bold="true" box="[151,399,464,489]" pageId="6" pageNumber="259">Specimens examined</emphasis>
.
<emphasis id="F8CAEACBF660C87BD8F2FE69FDE7FE2F" bold="true" box="[413,518,465,490]" italics="true" pageId="6" pageNumber="259">
<collectingCountry id="B2A97649F660C87BD8F2FE69FDE7FE2F" box="[413,518,465,490]" name="Taiwan" pageId="6" pageNumber="259">TAIWAN</collectingCountry>
</emphasis>
[
<emphasis id="F8CAEACBF660C87BDB77FE68FD65FE2C" bold="true" box="[536,644,464,489]" pageId="6" pageNumber="259">TAINAN</emphasis>
] 1 ex., Kuantzuling,
<date id="BE001019F660C87BDA1DFE69FC17FE2C" box="[882,1014,465,489]" pageId="6" pageNumber="259" value="1969-05-28">28. V. 1969</date>
, S. Hisamatsu leg. [
<emphasis id="F8CAEACBF660C87BDDBAFE68FAAFFE2C" bold="true" box="[1237,1358,464,489]" pageId="6" pageNumber="259">NANTOU</emphasis>
] 1 ex., Wushe,
<date id="BE001019F660C87BD99DFE4DFE96FDC8" box="[242,375,501,525]" pageId="6" pageNumber="259" value="1972-05-24">24. V. 1972</date>
, M. Sakai leg.
<emphasis id="F8CAEACBF660C87BDB4FFE4DFD96FDCB" bold="true" box="[544,631,501,526]" italics="true" pageId="6" pageNumber="259">
<collectingCountry id="B2A97649F660C87BDB4FFE4DFD96FDCB" box="[544,631,501,526]" name="Japan" pageId="6" pageNumber="259">JAPAN</collectingCountry>
</emphasis>
[
<emphasis id="F8CAEACBF660C87BDBE7FE4DFD15FDCB" bold="true" box="[648,756,501,526]" pageId="6" pageNumber="259">MIYAGI</emphasis>
] 1 ex., Takiyama Park, Yamoto-chô,
<date id="BE001019F660C87BDDF6FE4DFAD1FDC8" box="[1177,1328,501,525]" pageId="6" pageNumber="259" value="2009-08-02">2. VIII. 2009</date>
, T. Ozaki leg. [
<emphasis id="F8CAEACBF660C87BD9BCFDA0FEDAFDF4" bold="true" box="[211,315,536,561]" pageId="6" pageNumber="259">TOKYO</emphasis>
] 3 exs., Izu-oshima Is.,
<date id="BE001019F660C87BDB38FDA1FD01FDF4" box="[599,736,537,561]" pageId="6" pageNumber="259" value="1974-05-19">19. V. 1974</date>
, Y. Hirano leg.; 1 ex., Niisima,
<date id="BE001019F660C87BDD3AFDA1FB0DFDF4" box="[1109,1260,537,561]" pageId="6" pageNumber="259" value="1992-06-21">21. VI. 1992</date>
, Y. Hirano leg. [
<emphasis id="F8CAEACBF660C87BD9CFFD85FEA7FD93" bold="true" box="[160,326,573,598]" pageId="6" pageNumber="259">KANAGAWA</emphasis>
] 2 exs., Odawara,
<date id="BE001019F660C87BDB47FD85FD45FD90" box="[552,676,573,597]" pageId="6" pageNumber="259" value="1957-05-09">9. V. 1957</date>
, Y. Hirano leg.; 1 ex., Oishi,
<date id="BE001019F660C87BDD6DFD85FB6DFD90" box="[1026,1164,573,597]" pageId="6" pageNumber="259" value="1979-09-08">8. IX. 1979</date>
, Y. Hirano leg.; 2 exs., Sarugashima, Atsugi City,
<date id="BE001019F660C87BD8BBFDD9FD8DFDBC" box="[468,620,609,633]" pageId="6" pageNumber="259" value="1996-04-25">25. IV. 1996</date>
, Y. Hirano leg.; 1 ex., Tanzawa,
<date id="BE001019F660C87BDA91FDD9FB6CFDBC" box="[1022,1165,609,633]" pageId="6" pageNumber="259" value="1986-05-18">18. V. 1986</date>
, Y. Hirano leg.; 1 ex., Myoshintoge,
<date id="BE001019F660C87BD857FD3DFE3AFD58" box="[312,475,645,669]" pageId="6" pageNumber="259" value="1993-08-12">12. VIII. 1993</date>
, Y. Hirano leg.; 1 ex., Sengoku, Hakone,
<date id="BE001019F660C87BDAC5FD3DFBD7FD58" box="[938,1078,645,669]" pageId="6" pageNumber="259" value="1992-07-04">4. VII. 1992</date>
, Y. Hirano leg.; 1 ex., Odawara, Kuno,
<date id="BE001019F660C87BD985FD11FE8CFD04" box="[234,365,681,705]" pageId="6" pageNumber="259" value="1957-05-09">9. V. 1957</date>
, Y. Hirano leg.; 1 ex., Kamitsuruma 1 chôme, Sagamihara City,
<date id="BE001019F660C87BDDE9FD11FAC4FD04" box="[1158,1317,681,705]" pageId="6" pageNumber="259" value="1997-09-20">20. IX. 1997</date>
, R. Iwata leg. [
<emphasis id="F8CAEACBF660C87BD9BFFD75FEB7FD23" bold="true" box="[208,342,717,742]" pageId="6" pageNumber="259">SHIMANE</emphasis>
] 1 ex., Yoshiji-chô, Hamada City,
<date id="BE001019F660C87BDB85FD75FC92FD20" box="[746,883,717,741]" pageId="6" pageNumber="259" value="1996-05-14">14. V. 1996</date>
, Y. Hirano leg. [
<emphasis id="F8CAEACBF660C87BDD5AFD75FB77FD20" bold="true" box="[1077,1174,717,741]" pageId="6" pageNumber="259">EHIME</emphasis>
] 12 exs., Kurotaki, nr. Matsuyama City,
<date id="BE001019F660C87BD809FD49FE12FCCC" box="[358,499,753,777]" pageId="6" pageNumber="259" value="1958-05-18">18. V. 1958</date>
, M. Miyatake leg.; 1 ex., Mt. Sara,
<date id="BE001019F660C87BDAC9FD49FBD2FCCC" box="[934,1075,753,777]" pageId="6" pageNumber="259" value="1968-05-16">16. V. 1968</date>
, M. Sakai leg.; 1 ex., Hantaji, Matsuyama City,
<date id="BE001019F660C87BD833FCADFE31FCE8" box="[348,464,789,813]" pageId="6" pageNumber="259" value="1974-05-04">4. V. 1974</date>
, K. Ito leg.; 1 ex., Yama-jinjya, Matsuyama City,
<date id="BE001019F660C87BDA99FCADFB98FCE8" box="[1014,1145,789,813]" pageId="6" pageNumber="259" value="1973-05-23">23. V. 1973</date>
, K. Ito leg.; 1 ex., Tarumi, Matsuyama City,
<date id="BE001019F660C87BD833FC81FE14FC94" box="[348,501,825,849]" pageId="6" pageNumber="259" value="1995-07-22">22. VII. 1995</date>
, N. Ohbayashi leg. [
<emphasis id="F8CAEACBF660C87BDBB4FC81FC6DFC97" bold="true" box="[731,908,825,850]" pageId="6" pageNumber="259">KUMAMOTO</emphasis>
] 1 ex., Mt. Ichibusa,
<date id="BE001019F660C87BDD14FC81FB1DFC94" box="[1147,1276,825,849]" pageId="6" pageNumber="259" value="1967-06-09">9. VI. 1967</date>
, S. Hisamatsu leg. [
<emphasis id="F8CAEACBF660C87BD9BBFCE5FE8AFCB3" bold="true" box="[212,363,861,886]" pageId="6" pageNumber="259">NAGASAKI</emphasis>
] 1 ex., Mogi,
<date id="BE001019F660C87BDB73FCE5FD46FCB0" box="[540,679,861,885]" pageId="6" pageNumber="259" value="1971-05-20">20. V. 1971</date>
, S. Hisamatsu leg.; 1 ex., Chiziwa,
<date id="BE001019F660C87BDD3BFCE5FB0DFCB0" box="[1108,1260,861,885]" pageId="6" pageNumber="259" value="1965-11-10">10. XI. 1965</date>
, S. Nakao leg. [
<emphasis id="F8CAEACBF660C87BD9CFFC39FEB5FC5F" bold="true" box="[160,340,897,922]" pageId="6" pageNumber="259">KAGOSHIMA</emphasis>
] 2 exs., Amagi, Tokunoshima Is.,
<date id="BE001019F660C87BDBB0FC39FC88FC5C" box="[735,873,897,921]" pageId="6" pageNumber="259" value="1965-10-18">18. X. 1965</date>
, S. Nakao leg.; 2 exs., Mugio, Yaku-chô,
<date id="BE001019F660C87BDC22FC39FF33FC78" pageId="6" pageNumber="259" value="1974-03-18">18. III. 1974</date>
, TK. Ishihara leg. [
<emphasis id="F8CAEACBF660C87BD8C4FC1DFDD5FC7B" bold="true" box="[427,564,933,958]" pageId="6" pageNumber="259">OKINAWA</emphasis>
] 1 ex., Sueyoshi Park, Naha City,
<date id="BE001019F660C87BDAD8FC1DFBD9FC78" box="[951,1080,933,957]" pageId="6" pageNumber="259" value="2003-03-08">8. III. 2003</date>
, Y. Kikuhara leg.; 2 exs., Syuri,
<date id="BE001019F660C87BD9F8FC71FEFDFC24" box="[151,284,969,993]" pageId="6" pageNumber="259" value="1989-05-17">17. V. 1989</date>
, Y. Hirano leg.; 2 exs., Omoto, Ishigaki Is.,
<date id="BE001019F660C87BDA7DFC71FC78FC24" box="[786,921,969,993]" pageId="6" pageNumber="259" value="1993-02-27">27. II. 1993</date>
, Y. Hirano leg.; 2 exs., Uegusuku, Kumejima Is.,
<date id="BE001019F660C87BD9ACFC55FEBBFBC0" box="[195,346,1005,1029]" pageId="6" pageNumber="259" value="2001-09-21">21. IX. 2001</date>
, Y. Hirano leg.; 1 ex., Darumayama, Kumejima Is.,
<date id="BE001019F660C87BDAD0FC55FBB7FBC0" box="[959,1110,1005,1029]" pageId="6" pageNumber="259" value="2001-09-21">21. IX. 2001</date>
, Y. Hirano leg.; 1 ex., Oku, Kunigami-son,
<date id="BE001019F660C87BD829FBA9FE29FBEC" box="[326,456,1041,1065]" pageId="6" pageNumber="259" value="2002-04-04">4. IV. 2002</date>
, Y. Hirano leg.; 1 ex., Yamada, Onnason,
<date id="BE001019F660C87BDAC8FBA9FBD6FBEC" box="[935,1079,1041,1065]" pageId="6" pageNumber="259" value="2009-04-19">19. IV. 2009</date>
, Y. Hirano leg.; 4 exs., Miyako Is., 5~
<date id="BE001019F660C87BD9B2FB8EFEB2FB88" box="[221,339,1077,1101]" pageId="6" pageNumber="259" value="1998-05-07">7. V. 1998</date>
, M. Satô leg.
</paragraph>
</subSubSection>
<subSubSection id="82A46552F660C87BD9A6FBE1FE86FAE0" pageId="6" pageNumber="259" type="diagnosis">
<paragraph id="CA0136D9F660C87BD9A6FBE1FE61FB18" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">
<emphasis id="F8CAEACBF660C87BD9A6FBE1FEDDFBB7" bold="true" box="[201,316,1113,1138]" pageId="6" pageNumber="259">Diagnosis</emphasis>
. The species is distinguished from other Japanese congeners by the following characteristics: disc of elytra smooth or shagreened; males with yellow head and black pronotum; punctures on elytra (
<figureCitation id="52852A5CF660C87BDC64FBC5FABDFB50" box="[1291,1372,1149,1173]" captionStart="FIGURE 3" captionStartId="4.[151,250,1216,1239]" captionTargetBox="[157,1430,225,1157]" captionTargetId="figure@4.[151,1436,209,1173]" captionTargetPageId="4" captionText="FIGURE 3. Elytral punctuation of Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) spp. A, Pastillus eminentithorax sp. nov.; B, Cybocephalus (C.) nipponicus Endrödy-Younga, 1971; C, D, Cybocephalus (C.) politissimus Reitter, 1898. A = female, Mt. Haigamine, Hiroshima Pref.; B = male, Morimatsu, Ehime Pref.; C = male, Yakushima Is.; D = male, Sugitate, Ehime Pref." httpUri="https://zenodo.org/record/218830/files/figure.png" pageId="6" pageNumber="259">Figs. 3</figureCitation>
C–D) smaller than an eye-facet at middle; apical margin of median lobe (
<figureCitation id="52852A5CF660C87BDAE5FB19FC2DFB7C" box="[906,972,1185,1209]" captionStart="FIGURE 5" captionStartId="7.[151,250,1887,1910]" captionTargetBox="[162,1430,485,1865]" captionTargetId="figure@7.[151,1436,461,1882]" captionTargetPageId="7" captionText="FIGURE 5. Male genitalia of Cybocephalus (C.) politissimus Reitter, 1898. A – E, sclerite of internal sac; F – J, tegmen, ventral view; K – O, median lobe, ventral view; P – T, median lobe, lateral view. A, F, K, P = Kurotaki, Ehime Pref.; B, G, L, Q = Mt. Ichibusa, Kumamoto Pref.; C, H, M, R = Tokunoshima Is.; D, I, N, S = Onna-son, Okinawa Pref.; E, J, O, T = Kuantzuling, Taiwan. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218832/files/figure.png" pageId="6" pageNumber="259">Fig. 5</figureCitation>
K–O) in ventral view with strongly acute projection at middle.
</paragraph>
<paragraph id="CA0136D9F660C87BD9A6FB51FE86FAE0" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">
<emphasis id="F8CAEACBF660C87BD9A6FB51FE97FAC7" bold="true" box="[201,374,1257,1282]" pageId="6" pageNumber="259">Redescription.</emphasis>
Length 1.24–1.62 (1.43) mm (n = 7), width 0.70–0.82 (0.76) mm (n = 5), height 0.40–0.56 (0.49) mm (n = 7).
</paragraph>
</subSubSection>
<subSubSection id="82A46552F660C874D9A8FA89FE2EFDF9" lastPageId="9" lastPageNumber="262" pageId="6" pageNumber="259" type="description">
<paragraph id="CA0136D9F660C87BD9A8FA89FB97FA70" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">
<emphasis id="F8CAEACBF660C87BD9A8FA89FEE0FA8C" box="[199,257,1329,1353]" italics="true" pageId="6" pageNumber="259">Male</emphasis>
. Body convex; dorsal disc almost glabrous, shiny. Coloration black; dorsal- and ventral side of head (excluding blackish mandibles and apical half of genae), antennae, lateral margins of pronotum, mesoventrite, pro- and mesothoracic legs (excluding blackish mesotibiae) yellow; lateral margins and posterior corners of pronotum, anterior margins of elytra translucent, yellowish brown; hind legs dark yellowish brown.
</paragraph>
<paragraph id="CA0136D9F660C87BD9A8FA79FB11F980" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">
Head extremely wide; vertex minutely punctured; interspaces smooth or shagreened; labrum transverse, with apical margin shallowly emarginate at middle. Eyes large, visible from ventral aspect. Antennae (
<figureCitation id="52852A5CF660C87BDDBEFA5DFAF2FA38" box="[1233,1299,1509,1533]" captionStart="FIGURE 6" captionStartId="8.[151,250,1799,1822]" captionTargetBox="[175,1423,220,1758]" captionTargetId="figure@8.[151,1436,193,1778]" captionTargetPageId="8" captionText="FIGURE 6. Cybocephalus (C.) politissimus Reitter, 1898. A, spermatheca, spermathecal gland, and sperm duct; B, labrum; C, left protibia; D, left mesotibia; E, left metatibia; F, antenna; G, ovipositor; H, mentum. A, G = Mt. Sara, Ehime Pref.; C – F = Kuantzuling, Taiwan; B, H = Kurotaki, Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218833/files/figure.png" pageId="6" pageNumber="259">Fig. 6</figureCitation>
F) similar to those of
<emphasis id="F8CAEACBF660C87BD996F9B2FEEDF9E4" box="[249,268,1546,1569]" italics="true" pageId="6" pageNumber="259">C</emphasis>
. (
<emphasis id="F8CAEACBF660C87BD84BF9B2FED6F9E4" box="[292,311,1546,1569]" italics="true" pageId="6" pageNumber="259">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF660C87BD83FF9B2FE2AF9E4" box="[336,459,1546,1569]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="nipponicus">
<emphasis id="F8CAEACBF660C87BD83FF9B2FE2AF9E4" box="[336,459,1546,1569]" italics="true" pageId="6" pageNumber="259">nipponicus</emphasis>
</taxonomicName>
; club 3-segmented, apical margin of eleventh antennal segment truncate; approximate ratio of each segment (n = 1) is 2.13: 1.65: 1.87: 1.13: 1.25: 1.06: 1.06: 1.00: 1.37: 1.52: 1.40.
</paragraph>
<paragraph id="CA0136D9F660C87BD9A8F9EAFE72F974" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">Pronotum convex, strongly transverse, 1.67–1.73 (1.71) times as wide as long (n = 5); anterior angles rounded; posterior angles widely rounded; dorsal disc minutely punctated; interspaces smooth or shagreened. Scutellum with apex broadly rounded.</paragraph>
<paragraph id="CA0136D9F660C87BD9A8F905FC96F830" blockId="6.[151,1436,464,2037]" pageId="6" pageNumber="259">
Elytra conjointly 0.86–0.97 (0.92) times as long as wide (n = 5), 1.50–1.73 (1.60) times as long as pronotum (n = 5); disc (
<figureCitation id="52852A5CF660C87BD872F959FE8CF93C" box="[285,365,1761,1785]" captionStart="FIGURE 3" captionStartId="4.[151,250,1216,1239]" captionTargetBox="[157,1430,225,1157]" captionTargetId="figure@4.[151,1436,209,1173]" captionTargetPageId="4" captionText="FIGURE 3. Elytral punctuation of Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) spp. A, Pastillus eminentithorax sp. nov.; B, Cybocephalus (C.) nipponicus Endrödy-Younga, 1971; C, D, Cybocephalus (C.) politissimus Reitter, 1898. A = female, Mt. Haigamine, Hiroshima Pref.; B = male, Morimatsu, Ehime Pref.; C = male, Yakushima Is.; D = male, Sugitate, Ehime Pref." httpUri="https://zenodo.org/record/218830/files/figure.png" pageId="6" pageNumber="259">Figs. 3</figureCitation>
C–D) with two kinds of punctures, one large, but smaller than an eye-facet, the larger one possessing two lateral extensions and an associated recumbent seta; the other puncture minute; interspaces smooth or shagreened. Metaventrite convex, densely covered with long, recumbent yellowish setae, but without setae at basal areas. Inter-metacoxal distance wide, separated by 1.50 (1.50) times as wide as inter- mesocoxal distance (n = 2). Abdominal sternites densely covered with long, recumbent yellowish setae; inter-metacoxal area of abdominal sternite III with widely rounded apical margin; femoral lines joined with anterior margin of abdominal sternite III. Legs (
<figureCitation id="52852A5CF660C87BD98FF801FEC4F814" box="[224,293,1977,2001]" captionStart="FIGURE 6" captionStartId="8.[151,250,1799,1822]" captionTargetBox="[175,1423,220,1758]" captionTargetId="figure@8.[151,1436,193,1778]" captionTargetPageId="8" captionText="FIGURE 6. Cybocephalus (C.) politissimus Reitter, 1898. A, spermatheca, spermathecal gland, and sperm duct; B, labrum; C, left protibia; D, left mesotibia; E, left metatibia; F, antenna; G, ovipositor; H, mentum. A, G = Mt. Sara, Ehime Pref.; C – F = Kuantzuling, Taiwan; B, H = Kurotaki, Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218833/files/figure.png" pageId="6" pageNumber="259">Fig. 6</figureCitation>
C–E) rather slim and flat, similar to those of
<emphasis id="F8CAEACBF660C87BDA50F802FCB3F814" box="[831,850,1978,2001]" italics="true" pageId="6" pageNumber="259">C</emphasis>
. (
<emphasis id="F8CAEACBF660C87BDA03F802FC9EF814" box="[876,895,1978,2001]" italics="true" pageId="6" pageNumber="259">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF660C87BDAF5F802FBF4F814" box="[922,1045,1978,2001]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="259" phylum="Arthropoda" rank="species" species="nipponicus">
<emphasis id="F8CAEACBF660C87BDAF5F802FBF4F814" box="[922,1045,1978,2001]" italics="true" pageId="6" pageNumber="259">nipponicus</emphasis>
</taxonomicName>
; protibiae (
<figureCitation id="52852A5CF660C87BDDF1F801FB05F814" box="[1182,1252,1977,2001]" captionStart="FIGURE 6" captionStartId="8.[151,250,1799,1822]" captionTargetBox="[175,1423,220,1758]" captionTargetId="figure@8.[151,1436,193,1778]" captionTargetPageId="8" captionText="FIGURE 6. Cybocephalus (C.) politissimus Reitter, 1898. A, spermatheca, spermathecal gland, and sperm duct; B, labrum; C, left protibia; D, left mesotibia; E, left metatibia; F, antenna; G, ovipositor; H, mentum. A, G = Mt. Sara, Ehime Pref.; C – F = Kuantzuling, Taiwan; B, H = Kurotaki, Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218833/files/figure.png" pageId="6" pageNumber="259">Fig. 6</figureCitation>
C) with slightly sinuate inner margins, outer apical corner rounded. Claws simple.
</paragraph>
<paragraph id="CA0136D9F661C87AD9A6FF2FFDA0FED9" blockId="7.[151,1436,151,428]" pageId="7" pageNumber="260">
Male genitalia (
<figureCitation id="52852A5CF661C87AD813FF2FFE5EFF6A" box="[380,447,151,175]" captionStart="FIGURE 5" captionStartId="7.[151,250,1887,1910]" captionTargetBox="[162,1430,485,1865]" captionTargetId="figure@7.[151,1436,461,1882]" captionTargetPageId="7" captionText="FIGURE 5. Male genitalia of Cybocephalus (C.) politissimus Reitter, 1898. A – E, sclerite of internal sac; F – J, tegmen, ventral view; K – O, median lobe, ventral view; P – T, median lobe, lateral view. A, F, K, P = Kurotaki, Ehime Pref.; B, G, L, Q = Mt. Ichibusa, Kumamoto Pref.; C, H, M, R = Tokunoshima Is.; D, I, N, S = Onna-son, Okinawa Pref.; E, J, O, T = Kuantzuling, Taiwan. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218832/files/figure.png" pageId="7" pageNumber="260">Fig. 5</figureCitation>
) with some local variations in shape (see ‘Note’ below); tegmen strongly or uniformly attenuate, scattered with sparse setae at apical margin; median lobe in lateral view strongly thickened, more or less square in shape with strongly acute projection at middle of apical margin in ventral aspect. Internal sac sclerites of male genitalia as figured (
<figureCitation id="52852A5CF661C87AD8D5FEBCFE1AFED9" box="[442,507,260,284]" captionStart="FIGURE 5" captionStartId="7.[151,250,1887,1910]" captionTargetBox="[162,1430,485,1865]" captionTargetId="figure@7.[151,1436,461,1882]" captionTargetPageId="7" captionText="FIGURE 5. Male genitalia of Cybocephalus (C.) politissimus Reitter, 1898. A – E, sclerite of internal sac; F – J, tegmen, ventral view; K – O, median lobe, ventral view; P – T, median lobe, lateral view. A, F, K, P = Kurotaki, Ehime Pref.; B, G, L, Q = Mt. Ichibusa, Kumamoto Pref.; C, H, M, R = Tokunoshima Is.; D, I, N, S = Onna-son, Okinawa Pref.; E, J, O, T = Kuantzuling, Taiwan. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218832/files/figure.png" pageId="7" pageNumber="260">Fig. 5</figureCitation>
A–E).
</paragraph>
<paragraph id="CA0136D9F661C87AD9A8FE90FA69FE69" blockId="7.[151,1436,151,428]" pageId="7" pageNumber="260">
<emphasis id="F8CAEACBF661C87AD9A8FE90FEFAFE85" box="[199,283,296,320]" italics="true" pageId="7" pageNumber="260">Female</emphasis>
. Coloration of head black. Apical margin of abdominal tergite and sternite VII rounded. Female genitalia (
<figureCitation id="52852A5CF661C87AD863FEF4FEB5FEA1" box="[268,340,332,356]" captionStart="FIGURE 6" captionStartId="8.[151,250,1799,1822]" captionTargetBox="[175,1423,220,1758]" captionTargetId="figure@8.[151,1436,193,1778]" captionTargetPageId="8" captionText="FIGURE 6. Cybocephalus (C.) politissimus Reitter, 1898. A, spermatheca, spermathecal gland, and sperm duct; B, labrum; C, left protibia; D, left mesotibia; E, left metatibia; F, antenna; G, ovipositor; H, mentum. A, G = Mt. Sara, Ehime Pref.; C – F = Kuantzuling, Taiwan; B, H = Kurotaki, Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218833/files/figure.png" pageId="7" pageNumber="260">Fig. 6</figureCitation>
G) similar to those of
<emphasis id="F8CAEACBF661C87ADB09FEF5FD98FEA1" box="[614,633,333,356]" italics="true" pageId="7" pageNumber="260">C</emphasis>
. (
<emphasis id="F8CAEACBF661C87ADBF9FEF5FD48FEA1" box="[662,681,333,356]" italics="true" pageId="7" pageNumber="260">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF661C87ADBAAFEF5FCA1FEA1" box="[709,832,333,356]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="nipponicus">
<emphasis id="F8CAEACBF661C87ADBAAFEF5FCA1FEA1" box="[709,832,333,356]" italics="true" pageId="7" pageNumber="260">nipponicus</emphasis>
</taxonomicName>
; coxites feebly sclerotized, appearing separate and unfused, with short styli near apex; apex of stylus with 2~3 setae; 9~10 setae present along apico-lateral margin; spermatheca (
<figureCitation id="52852A5CF661C87AD85DFE2CFE95FE69" box="[306,372,404,428]" captionStart="FIGURE 6" captionStartId="8.[151,250,1799,1822]" captionTargetBox="[175,1423,220,1758]" captionTargetId="figure@8.[151,1436,193,1778]" captionTargetPageId="8" captionText="FIGURE 6. Cybocephalus (C.) politissimus Reitter, 1898. A, spermatheca, spermathecal gland, and sperm duct; B, labrum; C, left protibia; D, left mesotibia; E, left metatibia; F, antenna; G, ovipositor; H, mentum. A, G = Mt. Sara, Ehime Pref.; C – F = Kuantzuling, Taiwan; B, H = Kurotaki, Ehime Pref. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218833/files/figure.png" pageId="7" pageNumber="260">Fig. 6</figureCitation>
A) oval, sclerotized; spermathecal gland long, curved, weakly sclerotized; sperm duct coiled.
</paragraph>
<caption id="9EC16651F661C87AD9F8F8E7FE25F817" httpUri="https://zenodo.org/record/218832/files/figure.png" pageId="7" pageNumber="260" targetBox="[162,1430,485,1865]" targetPageId="7">
<paragraph id="CA0136D9F661C87AD9F8F8E7FE25F817" blockId="7.[151,1437,1887,2002]" pageId="7" pageNumber="260">
<emphasis id="F8CAEACBF661C87AD9F8F8E7FEF2F8B0" bold="true" box="[151,275,1887,1910]" pageId="7" pageNumber="260">FIGURE 5.</emphasis>
Male genitalia of
<taxonomicName id="0DBE4D5AF661C87AD8A1F8D8FDBCF8B3" box="[462,605,1888,1910]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="F8CAEACBF661C87AD8A1F8D8FDBCF8B3" box="[462,605,1888,1910]" italics="true" pageId="7" pageNumber="260">Cybocephalus</emphasis>
</taxonomicName>
(
<emphasis id="F8CAEACBF661C87ADB03F8D9FD9CF8B3" box="[620,637,1889,1910]" italics="true" pageId="7" pageNumber="260">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF661C87ADBFCF8D8FC78F8B0" authority="Reitter, 1898" authorityName="Reitter" authorityYear="1898" box="[659,921,1888,1910]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="260" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF661C87ADBFCF8D8FCEDF8B3" box="[659,780,1888,1910]" italics="true" pageId="7" pageNumber="260">politissimus</emphasis>
Reitter, 1898
</taxonomicName>
. A–E, sclerite of internal sac; F–J, tegmen, ventral view; K–O, median lobe, ventral view; P–T, median lobe, lateral view. A, F, K, P = Kurotaki, Ehime Pref.; B, G, L, Q = Mt. Ichibusa, Kumamoto Pref.; C, H, M, R = Tokunoshima Is.; D, I, N, S = Onna-son, Okinawa Pref.; E, J, O, T = Kuantzuling, Taiwan. Scale bars = 0.1 mm.
</paragraph>
</caption>
<caption id="9EC16651F66EC875D9F8F8BFFC61F89E" httpUri="https://zenodo.org/record/218833/files/figure.png" pageId="8" pageNumber="261" targetBox="[175,1423,220,1758]" targetPageId="8">
<paragraph id="CA0136D9F66EC875D9F8F8BFFC61F89E" blockId="8.[151,1436,1799,1883]" pageId="8" pageNumber="261">
<emphasis id="F8CAEACBF66EC875D9F8F8BFFEF2F8D8" bold="true" box="[151,275,1799,1822]" pageId="8" pageNumber="261">FIGURE 6.</emphasis>
<taxonomicName id="0DBE4D5AF66EC875D876F8B0FE49F8DB" box="[281,424,1800,1822]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="genus">
<emphasis id="F8CAEACBF66EC875D876F8B0FE49F8DB" box="[281,424,1800,1822]" italics="true" pageId="8" pageNumber="261">Cybocephalus</emphasis>
</taxonomicName>
(
<emphasis id="F8CAEACBF66EC875D8D7F8B1FE28F8DB" box="[440,457,1801,1822]" italics="true" pageId="8" pageNumber="261">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66EC875D8B1F8B0FD05F8D8" authority="Reitter, 1898" authorityName="Reitter" authorityYear="1898" box="[478,740,1800,1822]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF66EC875D8B1F8B0FDB6F8DB" box="[478,599,1800,1822]" italics="true" pageId="8" pageNumber="261">politissimus</emphasis>
Reitter, 1898
</taxonomicName>
. A, spermatheca, spermathecal gland, and sperm duct; B, labrum; C, left protibia; D, left mesotibia; E, left metatibia; F, antenna; G, ovipositor; H, mentum. A, G = Mt. Sara, Ehime Pref.; C–F = Kuantzuling, Taiwan; B, H = Kurotaki, Ehime Pref. Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="CA0136D9F66EC875D9A6F83FFAE4F822" blockId="8.[151,1435,1927,2024]" pageId="8" pageNumber="261">
<emphasis id="F8CAEACBF66EC875D9A6F83FFEADF865" bold="true" box="[201,332,1927,1952]" pageId="8" pageNumber="261">Bionomics.</emphasis>
According to Smith &amp; Bailey (2007), this species feeds upon
<taxonomicName id="0DBE4D5AF66EC875DD6AF830FE7EF801" authority="Takagi. Distribution. This" authorityName="Takagi. Distribution. This" class="Insecta" family="Diaspididae" genus="Aulacaspis" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="261" phylum="Arthropoda" rank="species" species="yasumatsui">
<emphasis id="F8CAEACBF66EC875DD6AF830FAE4F865" box="[1029,1285,1928,1952]" italics="true" pageId="8" pageNumber="261">Aulacaspis yasumatsui</emphasis>
Takagi.
<emphasis id="F8CAEACBF66EC875D9A6F813FE83F801" bold="true" box="[201,354,1963,1988]" pageId="8" pageNumber="261">Distribution.</emphasis>
This
</taxonomicName>
species occurs in Europe (
<collectingCountry id="B2A97649F66EC875DBB1F814FCCEF801" box="[734,815,1964,1988]" name="France" pageId="8" pageNumber="261">France</collectingCountry>
,
<collectingCountry id="B2A97649F66EC875DA51F814FC46F801" box="[830,935,1964,1988]" name="Germany" pageId="8" pageNumber="261">Germany</collectingCountry>
,
<collectingCountry id="B2A97649F66EC875DAD9F814FC0CF801" box="[950,1005,1964,1988]" name="Italy" pageId="8" pageNumber="261">Italy</collectingCountry>
),
<collectingCountry id="B2A97649F66EC875DD6AF814FBB1F801" box="[1029,1104,1964,1988]" name="Russia" pageId="8" pageNumber="261">Russia</collectingCountry>
(East and West Siberia, Far East),
<collectingCountry id="B2A97649F66EC875D9B1F877FEC0F822" box="[222,289,1999,2023]" name="China" pageId="8" pageNumber="261">China</collectingCountry>
(Guangdong) (Jelínek &amp; Audisio 2007),
<collectingCountry id="B2A97649F66EC875DB9FF877FCA3F822" box="[752,834,1999,2023]" name="Taiwan" pageId="8" pageNumber="261">Taiwan</collectingCountry>
(
<emphasis id="F8CAEACBF66EC875DA3DF868FC39F82D" bold="true" box="[850,984,2000,2024]" pageId="8" pageNumber="261">new record</emphasis>
), and
<collectingCountry id="B2A97649F66EC875DD70F868FBBEF822" box="[1055,1119,2000,2023]" name="Japan" pageId="8" pageNumber="261">Japan</collectingCountry>
(
<emphasis id="F8CAEACBF66EC875DD00F868FB14F82D" bold="true" box="[1135,1269,2000,2024]" pageId="8" pageNumber="261">new record</emphasis>
).
</paragraph>
<paragraph id="CA0136D9F66FC874D9A6FF20FA91FE42" blockId="9.[151,1437,151,572]" pageId="9" pageNumber="262">
<emphasis id="F8CAEACBF66FC874D9A6FF20FEABFF75" bold="true" box="[201,330,152,176]" pageId="9" pageNumber="262">Taxonomy.</emphasis>
<taxonomicName id="0DBE4D5AF66FC874D838FF20FE15FF75" box="[343,500,152,176]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="F8CAEACBF66FC874D838FF20FE15FF75" box="[343,500,152,176]" italics="true" pageId="9" pageNumber="262">Cybocephalus</emphasis>
</taxonomicName>
(
<emphasis id="F8CAEACBF66FC874DB66FF20FDFDFF6A" box="[521,540,152,175]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874DB56FF20FCB5FF6A" authority="Smith, 2007" authorityName="Smith" authorityYear="2007" box="[569,852,151,176]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="flavocapitis">
<emphasis id="F8CAEACBF66FC874DB56FF20FD5BFF75" box="[569,698,152,176]" italics="true" pageId="9" pageNumber="262">flavocapitis</emphasis>
Smith, 2007
</taxonomicName>
is treated herein as a junior synonym of
<emphasis id="F8CAEACBF66FC874DC25FF20FABCFF6A" box="[1354,1373,152,175]" italics="true" pageId="9" pageNumber="262">C</emphasis>
. (
<emphasis id="F8CAEACBF66FC874DC16FF20FA6DFF6A" box="[1401,1420,152,175]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874D9F8FF04FEFAFF11" box="[151,283,188,212]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF66FC874D9F8FF04FEFAFF11" box="[151,283,188,212]" italics="true" pageId="9" pageNumber="262">politissimus</emphasis>
</taxonomicName>
. Japanese and Taiwanese specimens of
<emphasis id="F8CAEACBF66FC874DBB1FF05FD10FF11" box="[734,753,189,212]" italics="true" pageId="9" pageNumber="262">C</emphasis>
. (
<emphasis id="F8CAEACBF66FC874DA66FF05FCFDFF11" box="[777,796,189,212]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874DA5BFF04FC59FF11" box="[820,952,188,212]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF66FC874DA5BFF04FC59FF11" box="[820,952,188,212]" italics="true" pageId="9" pageNumber="262">politissimus</emphasis>
</taxonomicName>
were compared with
<typeStatus id="1505887BF66FC874DDC0FF05FB3EFF10" box="[1199,1247,189,213]" pageId="9" pageNumber="262">type</typeStatus>
specimens of
<emphasis id="F8CAEACBF66FC874DCECFF05FA77FF11" box="[1411,1430,189,212]" italics="true" pageId="9" pageNumber="262">C</emphasis>
. (
<emphasis id="F8CAEACBF66FC874D9CFFF58FF52FF32" box="[160,179,224,247]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874D9A3FF58FEACFF3D" box="[204,333,224,248]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="flavocapitis">
<emphasis id="F8CAEACBF66FC874D9A3FF58FEACFF3D" box="[204,333,224,248]" italics="true" pageId="9" pageNumber="262">flavocapitis</emphasis>
</taxonomicName>
by Dr. T.R. Smith, and he agrees with this emendation (Smith pers. comm.).
<taxonomicName id="0DBE4D5AF66FC874DDA5FF58FA86FF3D" box="[1226,1383,224,248]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="genus">
<emphasis id="F8CAEACBF66FC874DDA5FF58FA86FF3D" box="[1226,1383,224,248]" italics="true" pageId="9" pageNumber="262">Cybocephalus</emphasis>
</taxonomicName>
(
<emphasis id="F8CAEACBF66FC874DC16FF58FA6DFF32" box="[1401,1420,224,247]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874D9F8FEBCFEF9FED9" box="[151,280,260,284]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="flavocapitis">
<emphasis id="F8CAEACBF66FC874D9F8FEBCFEF9FED9" box="[151,280,260,284]" italics="true" pageId="9" pageNumber="262">flavocapitis</emphasis>
</taxonomicName>
and
<emphasis id="F8CAEACBF66FC874D83FFEBDFE82FED9" box="[336,355,261,284]" italics="true" pageId="9" pageNumber="262">C</emphasis>
. (
<emphasis id="F8CAEACBF66FC874D815FEBDFE6CFED9" box="[378,397,261,284]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874D8CAFEBCFDC8FED9" box="[421,553,260,284]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF66FC874D8CAFEBCFDC8FED9" box="[421,553,260,284]" italics="true" pageId="9" pageNumber="262">politissimus</emphasis>
</taxonomicName>
share the following characteristics: head yellowish; apical margin of eleventhantennal segment truncated; tegmen in ventral view more or less attenuate; median lobe in lateral view strongly truncate, in ventral view apical margin with strongly acute projection at middle; internal sac sclerites of same shape. Based on these characteristics,
<emphasis id="F8CAEACBF66FC874D880FEC8FDE3FE42" box="[495,514,368,391]" italics="true" pageId="9" pageNumber="262">C</emphasis>
. (
<emphasis id="F8CAEACBF66FC874DB76FEC8FDCDFE42" box="[537,556,368,391]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874DB2DFEC8FD22FE4D" box="[578,707,368,392]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="flavocapitis">
<emphasis id="F8CAEACBF66FC874DB2DFEC8FD22FE4D" box="[578,707,368,392]" italics="true" pageId="9" pageNumber="262">flavocapitis</emphasis>
</taxonomicName>
should be treated as a junior synonym of
<emphasis id="F8CAEACBF66FC874DDFDFEC8FB44FE42" box="[1170,1189,368,391]" italics="true" pageId="9" pageNumber="262">C</emphasis>
. (
<emphasis id="F8CAEACBF66FC874DDD3FEC8FB2EFE42" box="[1212,1231,368,391]" italics="true" pageId="9" pageNumber="262">C</emphasis>
.)
<taxonomicName id="0DBE4D5AF66FC874DD8AFEC8FA88FE4D" box="[1253,1385,368,392]" class="Insecta" family="Cybocephalidae" genus="Cybocephalus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="262" phylum="Arthropoda" rank="species" species="politissimus">
<emphasis id="F8CAEACBF66FC874DD8AFEC8FA88FE4D" box="[1253,1385,368,392]" italics="true" pageId="9" pageNumber="262">politissimus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CA0136D9F66FC874D9A6FE2CFE2EFDF9" blockId="9.[151,1437,151,572]" pageId="9" pageNumber="262">
<emphasis id="F8CAEACBF66FC874D9A6FE2CFEE9FE69" bold="true" box="[201,264,404,428]" pageId="9" pageNumber="262">Note.</emphasis>
This species has some variation in the shape of the male genitalia (
<figureCitation id="52852A5CF66FC874DD65FE2CFBACFE69" box="[1034,1101,404,428]" captionStart="FIGURE 5" captionStartId="7.[151,250,1887,1910]" captionTargetBox="[162,1430,485,1865]" captionTargetId="figure@7.[151,1436,461,1882]" captionTargetPageId="7" captionText="FIGURE 5. Male genitalia of Cybocephalus (C.) politissimus Reitter, 1898. A – E, sclerite of internal sac; F – J, tegmen, ventral view; K – O, median lobe, ventral view; P – T, median lobe, lateral view. A, F, K, P = Kurotaki, Ehime Pref.; B, G, L, Q = Mt. Ichibusa, Kumamoto Pref.; C, H, M, R = Tokunoshima Is.; D, I, N, S = Onna-son, Okinawa Pref.; E, J, O, T = Kuantzuling, Taiwan. Scale bars = 0.1 mm." httpUri="https://zenodo.org/record/218832/files/figure.png" pageId="9" pageNumber="262">Fig. 5</figureCitation>
) and punctation of the dorsal surface (
<figureCitation id="52852A5CF66FC874D996FE0FFEDCFE0A" box="[249,317,439,463]" captionStart="FIGURE 3" captionStartId="4.[151,250,1216,1239]" captionTargetBox="[157,1430,225,1157]" captionTargetId="figure@4.[151,1436,209,1173]" captionTargetPageId="4" captionText="FIGURE 3. Elytral punctuation of Pastillus eminentithorax sp. nov. and Cybocephalus (Cybocephalus) spp. A, Pastillus eminentithorax sp. nov.; B, Cybocephalus (C.) nipponicus Endrödy-Younga, 1971; C, D, Cybocephalus (C.) politissimus Reitter, 1898. A = female, Mt. Haigamine, Hiroshima Pref.; B = male, Morimatsu, Ehime Pref.; C = male, Yakushima Is.; D = male, Sugitate, Ehime Pref." httpUri="https://zenodo.org/record/218830/files/figure.png" pageId="9" pageNumber="262">Fig. 3</figureCitation>
C–D). Nevertheless, the basic structure of the male genitalia is identical, especially the thickened median lobe (in lateral aspect) and shape of the internal sac sclerites. Therefore, these variable anatomical features should be considered as intraspecific variation. Coloration of the male pronotum is usually black, but rarely becoming yellowish brown.
</paragraph>
</subSubSection>
</treatment>
</document>
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