280 lines
36 KiB
XML
280 lines
36 KiB
XML
<document id="0D11815DD7C13933C767D27D0006D056" ID-CLB-Dataset="2949" ID-DOI="10.11646/zootaxa.5082.1.1" ID-GBIF-Dataset="b1305c10-d429-4335-aa4b-20786c960972" ID-ISSN="1175-5326" ID-Zenodo-Dep="5783128" ID-ZooBank="F4F057C6-3207-4430-A88E-4F4513D803C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639564321074" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2021" docId="397C5D1DFFC8FFDCFF6FB545FEEDFBA0" docLanguage="en" docName="zootaxa.5082.1.1.pdf" docOrigin="Zootaxa 5082 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum ahrensi Shavrin 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="C5452565FFC9FFDFFFF8B17BFF99FF95" masterDocTitle="New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="2" updateTime="1699472531112" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6B6EA533DCB31778F8D7610483381EAE">New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="BC22CD4A1685639E13F983C431FF2D0D">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="99488810B48BDDD8C79EF8DED42EAC5C">Zootaxa</mods:title>
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<mods:part id="7B09C75C3201255087EAAA42C7BBC099">
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<mods:date id="9E94A0E0380140DF30328E8CEA16FF4B">2021</mods:date>
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<mods:number id="B001F2B1665A18390398BDBFA2233C54">2021-12-15</mods:number>
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<mods:number id="A8708E8FCFD4C752A6B5C78876A89821">5082</mods:number>
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<mods:classification id="558A954CA376DD269EAA6C5A7EA6A6E4">journal article</mods:classification>
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<mods:identifier id="01F5AAC614FA6C27C3F064544B6800BE" type="DOI">10.11646/zootaxa.5082.1.1</mods:identifier>
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<mods:identifier id="6186D349983A6B7DAE9DEE27759AF8C7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="397C5D1DFFC8FFDCFF6FB545FEEDFBA0" ID-DOI="http://doi.org/10.5281/zenodo.5783152" ID-GBIF-Taxon="191647225" ID-Zenodo-Dep="5783152" LSID="urn:lsid:plazi:treatment:397C5D1DFFC8FFDCFF6FB545FEEDFBA0" httpUri="http://treatment.plazi.org/id/397C5D1DFFC8FFDCFF6FB545FEEDFBA0" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="F9CFBF80FFC8FFDEFF6FB545FDFFFBCC" box="[151,614,1086,1113]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="B16AEC0BFFC8FFDEFF6FB545FDFFFBCC" blockId="1.[151,614,1086,1149]" box="[151,614,1086,1113]" pageId="1" pageNumber="2">
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<heading id="EA225B67FFC8FFDEFF6FB545FDFFFBCC" bold="true" box="[151,614,1086,1113]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<emphasis id="83A13019FFC8FFDEFF6FB545FDFFFBCC" bold="true" box="[151,614,1086,1113]" pageId="1" pageNumber="2">
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<taxonomicName id="76D59788FFC8FFDEFF6FB545FD8FFBCD" ID-CoL="9GCPW" authority="Shavrin" authorityName="Shavrin" authorityYear="2021" box="[151,534,1086,1113]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ahrensi" status="sp. nov.">
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<emphasis id="83A13019FFC8FFDEFF6FB545FE33FBCC" bold="true" box="[151,426,1086,1113]" italics="true" pageId="1" pageNumber="2">Amphichroum ahrensi</emphasis>
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Shavrin
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</taxonomicName>
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,
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<taxonomicNameLabel id="98928D62FFC8FFDEFDDBB544FDFFFBCC" box="[547,614,1087,1113]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F9CFBF80FFC8FFDEFF6FB518FEC1FBE8" box="[151,344,1122,1149]" pageId="1" pageNumber="2" type="description">
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<paragraph id="B16AEC0BFFC8FFDEFF6FB518FEC1FBE8" blockId="1.[151,614,1086,1149]" box="[151,344,1122,1149]" pageId="1" pageNumber="2">
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(
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<figureCitation id="29EEF08EFFC8FFDEFF67B518FF74FBE8" box="[159,237,1123,1149]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1773,1797]" captionTargetBox="[159,1425,482,1752]" captionTargetId="figure-189@2.[151,1436,474,1764]" captionTargetPageId="2" captionText="FIGURES 1–7. Habitus of Amphichroum (males): 1—A. ahrensi (paratype), 2—A. altivagans (holotype), 3—A. anthobioides (lectotype), 4—A. monticola (holotype), 5—A. nepalicum (Nepal, Manaslu Himal), 6—A. pindarense (syntype), 7—A. telnovi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5783132" httpUri="https://zenodo.org/record/5783132/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
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,
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<figureCitation id="29EEF08EFFC8FFDEFF03B518FEBEFBE8" box="[251,295,1123,1149]" captionStart="FIGURES 8–14" captionStartId="5.[151,264,1907,1931]" captionTargetBox="[168,1418,194,1872]" captionTargetId="figure-15@5.[151,1436,181,1884]" captionTargetPageId="5" captionText="FIGURES 8–14. Aedeagus of Amphichroum ahrensi (8–9: paratype), A. altivagans (10–11: Kashmir, Khelanmarg) and A. anthobioides (12, 14: syntype; 13: Nepal, Bajura): 8, 10, 12—parameral view, 9, 11, 14—lateral view, 13—apical portion of aedeagus, parameral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5783134" httpUri="https://zenodo.org/record/5783134/files/figure.png" pageId="1" pageNumber="2">8–9</figureCitation>
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,
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<figureCitation id="29EEF08EFFC8FFDEFECBB519FEC9FBE8" box="[307,336,1122,1149]" captionStart="FIGURES 15–17" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1920]" captionTargetId="figure-27@6.[277,1310,181,1920]" captionTargetPageId="6" captionText="FIGURES 15–17. Distribution of Amphichroum species in the Himalayan Region and Tibet: 15—A. ahrensi (circle), A. altivagans (triangle), A. anthobioides (squares), 16—A. monticola (circles), A. pindarense (triangles), A. telnovi (diamond), 17—A. nepalicum." figureDoi="http://doi.org/10.5281/zenodo.5783140" httpUri="https://zenodo.org/record/5783140/files/figure.png" pageId="1" pageNumber="2">15</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F9CFBF80FFC8FFDEFF6FB5D0FAFFFAC0" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="B16AEC0BFFC8FFDEFF6FB5D0FECFFA98" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="2">
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<emphasis id="83A13019FFC8FFDEFF6FB5D0FE5DFB51" bold="true" box="[151,452,1194,1221]" pageId="1" pageNumber="2">Type material examined:</emphasis>
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<materialsCitation id="01BDE656FFC8FFDEFE34B5D0FECFFA98" ID-GBIF-Occurrence="3418584314" collectingDate="1998-05-19" collectingDateMax="1998-05-23" collectingDateMin="1998-05-19" collectorName="Fabrizi & Ahrens'" country="India" elevation="2800" location="Tonglu" municipality="Darjeeling" pageId="1" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="West Bengal" typeStatus="holotype">
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<typeStatus id="6E6E52A9FFC8FFDEFE34B5D0FDAEFB50" box="[460,567,1195,1221]" pageId="1" pageNumber="2" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="A7D32782FFC8FFDEFDBCB5D0FDC5FB50" box="[580,604,1195,1221]" pageId="1" pageNumber="2" type="male">♂</specimenCount>
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: ‘
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<collectingCountry id="C9C2AC9BFFC8FFDEFD95B5D0FD59FB50" box="[621,704,1195,1221]" name="India" pageId="1" pageNumber="2">INDIA</collectingCountry>
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,
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<collectingRegion id="731122E9FFC8FFDEFD3CB5D0FCCEFB50" box="[708,855,1195,1221]" country="India" name="West Bengal" pageId="1" pageNumber="2">West Bengal</collectingRegion>
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<collectingMunicipality id="510E7671FFC8FFDEFC96B5D0FC7EFB50" box="[878,999,1195,1221]" pageId="1" pageNumber="2">Darjeeling</collectingMunicipality>
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,
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<location id="B40ABAD0FFC8FFDEFC0BB5D0FBDBFB50" LSID="urn:lsid:plazi:treatment:397C5D1DFFC8FFDCFF6FB545FEEDFBA0:B40ABAD0FFC8FFDEFC0BB5D0FBDBFB50" box="[1011,1090,1195,1221]" country="India" municipality="Darjeeling" name="Tonglu" pageId="1" pageNumber="2" stateProvince="West Bengal">Tonglu</location>
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<date id="C56BCACBFFC8FFDEFBA0B5D0FB6EFB50" box="[1112,1271,1195,1221]" pageId="1" pageNumber="2" value="1998-05-19" valueMax="1998-05-23" valueMin="1998-05-19">
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<collectingDate id="D52F3323FFC8FFDEFBA0B5D0FB6EFB50" box="[1112,1271,1195,1221]" pageId="1" pageNumber="2" value="1998-05-19" valueMax="1998-05-23" valueMin="1998-05-19">19.-23.V.1998</collectingDate>
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</date>
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<quantity id="762D41EEFFC8FFDEFAF5B5D0FA05FB50" box="[1293,1436,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="1" pageNumber="2" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">
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<elevation id="3AF80B38FFC8FFDEFAF5B5D0FA05FB50" box="[1293,1436,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="1" pageNumber="2" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">2600-3000m</elevation>
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</quantity>
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NN | leg.
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<collectorName id="1C2089DDFFC8FFDEFEF9B5B4FE50FB7C" box="[257,457,1231,1257]" pageId="1" pageNumber="2">Fabrizi & Ahrens’</collectorName>
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<printed>, ‘HOLOTYPE |
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<taxonomicName id="76D59788FFC8FFDEFD03B5B4FC00FB7C" authorityName="Kraatz" authorityYear="1857" box="[763,921,1231,1257]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="83A13019FFC8FFDEFD03B5B4FC00FB7C" box="[763,921,1231,1257]" italics="true" pageId="1" pageNumber="2">Amphichroum</emphasis>
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</taxonomicName>
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<taxonomicName id="76D59788FFC8FFDEFC53B5B4FC67FB7C" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[939,1022,1231,1257]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ahrensi" status="sp. nov.">
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<emphasis id="83A13019FFC8FFDEFC53B5B4FC67FB7C" box="[939,1022,1231,1257]" italics="true" pageId="1" pageNumber="2">ahrensi</emphasis>
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</taxonomicName>
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<emphasis id="83A13019FFC8FFDEFBFCB5B4FBDAFB7C" bold="true" box="[1028,1091,1231,1257]" pageId="1" pageNumber="2">
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<taxonomicNameLabel id="98928D62FFC8FFDEFBFCB5B4FBDAFB7C" box="[1028,1091,1231,1257]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2021’ <red, printed> (NME).
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</materialsCitation>
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</paragraph>
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<paragraph id="B16AEC0BFFC8FFDEFF3FB46CFAFFFAC0" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="2">
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<typeStatus id="6E6E52A9FFC8FFDEFF3FB46CFEA1FAA4" box="[199,312,1303,1329]" pageId="1" pageNumber="2" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="A7D32782FFC8FFDEFEBBB46CFE1AFAA4" box="[323,387,1303,1329]" pageId="1" pageNumber="2" type="male">7 ♂♂</specimenCount>
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(
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<specimenCount id="A7D32782FFC8FFDEFE6AB46CFDA3FAA4" box="[402,570,1303,1329]" pageId="1" pageNumber="2" type="generic">two specimens</specimenCount>
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dissected),
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<specimenCount id="A7D32782FFC8FFDEFD38B46CFC96FAA4" box="[704,783,1303,1329]" pageId="1" pageNumber="2" type="female">11 ♀♀</specimenCount>
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: same data as the holotype. (
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<specimenCount id="A7D32782FFC8FFDEFBADB46DFB0EFAA4" box="[1109,1175,1302,1329]" pageId="1" pageNumber="2" type="male">5 ♂♂</specimenCount>
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,
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<specimenCount id="A7D32782FFC8FFDEFB5CB46CFB7CFAA4" box="[1188,1253,1303,1329]" pageId="1" pageNumber="2" type="female">7 ♀♀</specimenCount>
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: NME;
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<specimenCount id="A7D32782FFC8FFDEFAB8B46CFA1BFAA4" box="[1344,1410,1303,1329]" pageId="1" pageNumber="2" type="male">2 ♂♂</specimenCount>
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,
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<specimenCount id="A7D32782FFC8FFDEFA76B46CFF5AFAC0" pageId="1" pageNumber="2" type="female">4 ♀♀</specimenCount>
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: cSh). All
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<typeStatus id="6E6E52A9FFC8FFDEFEC3B440FE3FFAC0" box="[315,422,1339,1365]" pageId="1" pageNumber="2" type="paratype">paratypes</typeStatus>
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with red label ‘
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<typeStatus id="6E6E52A9FFC8FFDEFDAAB440FD7BFAC0" box="[594,738,1339,1365]" pageId="1" pageNumber="2" type="paratype">PARATYPE</typeStatus>
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<taxonomicName id="76D59788FFC8FFDEFD0EB440FC0DFAC0" authorityName="Kraatz" authorityYear="1857" box="[758,916,1339,1365]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="83A13019FFC8FFDEFD0EB440FC0DFAC0" box="[758,916,1339,1365]" italics="true" pageId="1" pageNumber="2">Amphichroum</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="76D59788FFC8FFDEFC5FB440FC63FAC0" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[935,1018,1339,1365]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="ahrensi" status="sp. nov.">
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<emphasis id="83A13019FFC8FFDEFC5FB440FC63FAC0" box="[935,1018,1339,1365]" italics="true" pageId="1" pageNumber="2">ahrensi</emphasis>
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</taxonomicName>
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<emphasis id="83A13019FFC8FFDEFBF9B440FBD9FAC0" bold="true" box="[1025,1088,1339,1365]" pageId="1" pageNumber="2">
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||
<taxonomicNameLabel id="98928D62FFC8FFDEFBF9B440FBD9FAC0" box="[1025,1088,1339,1365]" pageId="1" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2021’.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F9CFBF80FFC8FFDCFF3FB425FF5BFC30" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
|
||
<paragraph id="B16AEC0BFFC8FFDEFF3FB425FB58FA54" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="2">
|
||
<emphasis id="83A13019FFC8FFDEFF3FB425FEC0FAED" bold="true" box="[199,345,1374,1400]" pageId="1" pageNumber="2">Description.</emphasis>
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Measurements (n=19): HL: 0.30–0.43; HW: 0.55–0.70; AL (
|
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<typeStatus id="6E6E52A9FFC8FFDEFBFDB424FBF7FAEF" box="[1029,1134,1375,1402]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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): 1.54; OL: 0.19–0.25; PL: 0.53–0.72; PW: 0.88–1.00; ESL: 1.05–1.25; EW: 1.18–1.32; AW: 1.16–1.30; MTbL (
|
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<typeStatus id="6E6E52A9FFC8FFDEFBABB4F8FB25FA0B" box="[1107,1212,1411,1438]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
|
||
): 0.57; MTrL (
|
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<typeStatus id="6E6E52A9FFC8FFDEFAA7B4F8FF54FA54" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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||
): 0.52 (MTrL 1–4: 0.30; MTrL 5: 0.22); AedL: 0.70–0.75; TL: 2.65–3.75 (
|
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<typeStatus id="6E6E52A9FFC8FFDEFBF5B4DCFBECFA54" box="[1037,1141,1447,1473]" pageId="1" pageNumber="2" type="holotype">holotype</typeStatus>
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||
: 3.70).
|
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</paragraph>
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<paragraph id="B16AEC0BFFC8FFDEFF3FB4B0FAC3F82C" blockId="1.[151,1437,1194,2013]" pageId="1" pageNumber="2">
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Habitus as in
|
||
<figureCitation id="29EEF08EFFC8FFDEFE9EB4B0FE32FA70" box="[358,427,1483,1509]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1773,1797]" captionTargetBox="[159,1425,482,1752]" captionTargetId="figure-189@2.[151,1436,474,1764]" captionTargetPageId="2" captionText="FIGURES 1–7. Habitus of Amphichroum (males): 1—A. ahrensi (paratype), 2—A. altivagans (holotype), 3—A. anthobioides (lectotype), 4—A. monticola (holotype), 5—A. nepalicum (Nepal, Manaslu Himal), 6—A. pindarense (syntype), 7—A. telnovi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5783132" httpUri="https://zenodo.org/record/5783132/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
|
||
. Head, mouthparts, antennomeres 4–11, disc of pronotum, scutellum, legs and abdomen yellow-brown; antennomeres 1–3, lateral and basal portion of pronotum, elytra and paratergites yellow. Forebody glossy; frontal portion of head with dense transverse microsculpture, middle portion of head without or with very fine transverse microreticulation, infraorbital ridges with fine, rounded or longitudinal meshes; neck with dense transverse microsculpture; pronotum with distinct isodiametric microreticulation, without meshes in mediobasal third; exposed part of scutellum with dense, transverse sculpture; abdominal tergites with very dense, fine, isodiametric microreticulation. Head with very fine and sparse punctation, slightly denser in middle portion and with several moderately large punctures between anteocellar foveae and eyes; punctation of pronotum distinctly larger, denser and deeper than that in middle portion of head, sometimes finer and sparser in medioapical portion or without punctures in middle and/or mediobasal portions; punctation of elytra large, distinctly denser, coarser and deeper in basal portion, finer and sparser in middle along suture; abdominal tergites with dense, fine punctures, coarser and denser on abdominal tergite VII. Forebody without setation except for head with several long setae on frontal portion and two to four moderately long and erect setae between anteocellar foveae and eyes, and lateral margins of pronotum with four moderately long setae; abdominal tergites with regular, short, moderately dense setation.
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFC8FFDDFF3FB6B8FACBFE44" blockId="1.[151,1437,1194,2013]" lastBlockId="2.[151,1437,151,465]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">Head subtrapezoidal, transverse, 1.6–1.8 times as broad as long, flattened in middle and slightly convex on posterior portions of infraorbital ridges; anteocellar foveae narrow, deep and long, diagonally stretching toward antennal bases about to level of anterior margins of eyes. Eyes very large, convex. Ocelli large, located at level of posterior margins of eyes near deep occipital furrow, distance between ocelli as long as or slightly shorter than distance between ocellus and posterior margin of eye. Apical maxillary palpomere 1.3 times as long as preceding segment, gradually narrowed from basal third toward subacute apex. Antenna moderately long, reaching basal part of elytra when reflexed, from antennomere 5 progressively widened apicad; basal antennomere oblong, about three times as long as broad, antennomere 2 distinctly shorter than and about as broad as basal antennomere, 3 slightly longer than 2, 4 slightly shorter than 3, 5–6 slightly broader than 4, 7–8 slightly broader than 6, 9–10 slightly broader than 8, apical antennomere about 1.3 times as long as 10, sharply narrowed from apical third toward subacute apex.</paragraph>
|
||
<caption id="E5AABC83FFCBFFDDFF6FB796FE4CF8AA" ID-DOI="http://doi.org/10.5281/zenodo.5783132" ID-Zenodo-Dep="5783132" httpUri="https://zenodo.org/record/5783132/files/figure.png" pageId="2" pageNumber="3" startId="2.[151,264,1773,1797]" targetBox="[159,1425,482,1752]" targetPageId="2">
|
||
<paragraph id="B16AEC0BFFCBFFDDFF6FB796FE4CF8AA" blockId="2.[151,1436,1773,1856]" pageId="2" pageNumber="3">
|
||
<emphasis id="83A13019FFCBFFDDFF6FB796FEA2F890" bold="true" box="[151,315,1773,1797]" pageId="2" pageNumber="3">FIGURES 1–7.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="76D59788FFCBFFDDFE4CB796FDDAF890" authorityName="Kraatz" authorityYear="1857" box="[436,579,1773,1797]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="83A13019FFCBFFDDFE4CB796FDDAF890" box="[436,579,1773,1797]" italics="true" pageId="2" pageNumber="3">Amphichroum</emphasis>
|
||
</taxonomicName>
|
||
(males): 1—
|
||
<taxonomicName id="76D59788FFCBFFDDFD31B796FCA8F890" authorityName="Shavrin" authorityYear="2021" box="[713,817,1773,1797]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="ahrensi">
|
||
<emphasis id="83A13019FFCBFFDDFD31B796FD41F891" box="[713,728,1773,1796]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFD1EB796FCA8F890" box="[742,817,1773,1797]" italics="true" pageId="2" pageNumber="3">ahrensi</emphasis>
|
||
</taxonomicName>
|
||
(paratype), 2—
|
||
<taxonomicName id="76D59788FFCBFFDDFC2BB796FBC0F890" authorityName="Cameron" authorityYear="1941" box="[979,1113,1773,1797]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="altivagans">
|
||
<emphasis id="83A13019FFCBFFDDFC2BB796FC7BF891" box="[979,994,1773,1796]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFC17B796FBC0F890" box="[1007,1113,1773,1797]" italics="true" pageId="2" pageNumber="3">altivagans</emphasis>
|
||
</taxonomicName>
|
||
(holotype), 3—
|
||
<taxonomicName id="76D59788FFCBFFDDFB05B796FA05F890" authorityName="Champion" authorityYear="1925" box="[1277,1436,1773,1797]" class="Insecta" family="Staphylinidae" genus="Arpedium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="anthobioides">
|
||
<emphasis id="83A13019FFCBFFDDFB05B796FA95F891" box="[1277,1292,1773,1796]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFAE1B796FA05F890" box="[1305,1436,1773,1797]" italics="true" pageId="2" pageNumber="3">anthobioides</emphasis>
|
||
</taxonomicName>
|
||
(lectotype), 4—
|
||
<taxonomicName id="76D59788FFCBFFDDFEC0B670FE23F8B7" authorityName="Cameron" authorityYear="1928" box="[312,442,1802,1826]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="monticola">
|
||
<emphasis id="83A13019FFCBFFDDFEC0B670FEDEF8B7" box="[312,327,1803,1826]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFEADB671FE23F8B7" box="[341,442,1802,1826]" italics="true" pageId="2" pageNumber="3">monticola</emphasis>
|
||
</taxonomicName>
|
||
(holotype), 5—
|
||
<taxonomicName id="76D59788FFCBFFDDFDA5B670FD7DF8B7" authorityName="Shavrin" authorityYear="2021" baseAuthorityName="Coiffait" baseAuthorityYear="1982" box="[605,740,1802,1826]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="83A13019FFCBFFDDFDA5B670FDF5F8B7" box="[605,620,1803,1826]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFD82B671FD7DF8B7" box="[634,740,1802,1826]" italics="true" pageId="2" pageNumber="3">nepalicum</emphasis>
|
||
</taxonomicName>
|
||
(Nepal, Manaslu Himal), 6—
|
||
<taxonomicName id="76D59788FFCBFFDDFBEDB670FB3BF8B7" authorityName="Champion" authorityYear="1920" box="[1045,1186,1802,1826]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pindarense">
|
||
<emphasis id="83A13019FFCBFFDDFBEDB670FBBDF8B7" box="[1045,1060,1803,1826]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFBCAB671FB3BF8B7" box="[1074,1186,1802,1826]" italics="true" pageId="2" pageNumber="3">pindarense</emphasis>
|
||
</taxonomicName>
|
||
(syntype), 7—
|
||
<taxonomicName id="76D59788FFCBFFDDFAC3B670FA05F8B7" authorityName="Shavrin" authorityYear="2021" box="[1339,1436,1802,1826]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="telnovi">
|
||
<emphasis id="83A13019FFCBFFDDFAC3B670FAD3F8B7" box="[1339,1354,1803,1826]" italics="true" pageId="2" pageNumber="3">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCBFFDDFAAFB671FA05F8B7" box="[1367,1436,1802,1826]" italics="true" pageId="2" pageNumber="3">telnovi</emphasis>
|
||
</taxonomicName>
|
||
(paratype). Scale bars: 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B16AEC0BFFCBFFDDFF3FB614FC00F861" blockId="2.[151,1437,1902,2037]" pageId="2" pageNumber="3">Pronotum 1.3–1.6 times as broad as long, 1.4–1.6 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; anterior angles widely rounded, slightly protruded anteriad; posterior angles widely rounded; laterobasal margins slightly sinuate in some specimens; lateral portions flattened, moderately wide, slightly explanate, explanate area sometimes gradually widened posteriad.</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB1ECFA0FFF40" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">Elytra slightly broader than long and slightly widened apicad, reaching apical margin of abdominal tergite IV or V, slightly less than twice as long as pronotum, with widely rounded or straight apical margins. Hind wings fully developed.</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB1A4FE08FE88" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">Legs moderately long; metatibia slightly longer than metatarsus, with long strong thorns on inner and long setation on outer surfaces.</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB05CFB03FEF0" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">Abdomen slightly narrower than elytra, with a pair of very large transverse tomentose spots (wing-folding patches) in middle of tergite IV, with narrow palisade fringe on apical margin of tergite VII.</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB014FA84FD10" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">
|
||
Male. Protarsomeres 1–4 very wide. Medial margin of inner side of protibia with two parallel rows of eight to twelve short peg setae; mesotibia strongly curved mediad, with dense row of numerous very short and wide thorns beginning from medial side of bend and stretching to apex of mesotibia; lateral margins of mesotibia with very long and strong, sparse thorns and apical margins with several strong and long thorns. Apical margin of abdominal tergite VIII straight or rounded. Apical margin of abdominal sternite VIII widely rounded. Aedeagus (
|
||
<figureCitation id="29EEF08EFFCAFFDCFB46B084FA99FD8C" box="[1214,1280,511,537]" captionStart="FIGURES 8–14" captionStartId="5.[151,264,1907,1931]" captionTargetBox="[168,1418,194,1872]" captionTargetId="figure-15@5.[151,1436,181,1884]" captionTargetPageId="5" captionText="FIGURES 8–14. Aedeagus of Amphichroum ahrensi (8–9: paratype), A. altivagans (10–11: Kashmir, Khelanmarg) and A. anthobioides (12, 14: syntype; 13: Nepal, Bajura): 8, 10, 12—parameral view, 9, 11, 14—lateral view, 13—apical portion of aedeagus, parameral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5783134" httpUri="https://zenodo.org/record/5783134/files/figure.png" pageId="3" pageNumber="4">Fig. 8</figureCitation>
|
||
) with median lobe widest at basal bulb, gradually narrowed apicad; parameres moderately narrow, distinctly exceeding apex of median lobe, each with two moderately short preapical setae; internal sac long, with two fields of long thorns in apical and two complicated sclerotized structures in basal portions. Lateral aspect of aedeagus as in
|
||
<figureCitation id="29EEF08EFFCAFFDCFB2DB310FA80FD10" box="[1237,1305,619,645]" captionStart="FIGURES 8–14" captionStartId="5.[151,264,1907,1931]" captionTargetBox="[168,1418,194,1872]" captionTargetId="figure-15@5.[151,1436,181,1884]" captionTargetPageId="5" captionText="FIGURES 8–14. Aedeagus of Amphichroum ahrensi (8–9: paratype), A. altivagans (10–11: Kashmir, Khelanmarg) and A. anthobioides (12, 14: syntype; 13: Nepal, Bajura): 8, 10, 12—parameral view, 9, 11, 14—lateral view, 13—apical portion of aedeagus, parameral view. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5783134" httpUri="https://zenodo.org/record/5783134/files/figure.png" pageId="3" pageNumber="4">Fig. 9</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB3F4FD09FD64" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">Female. Protarsomeres 1–4 narrow. Medial margin of protibia without peg setae; mesotibia without modifications, slender, with long strong thorn along entire length of lateral margin. Abdominal tergite VIII and sternite VIII with small rounded or straight apical margin.</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB381FF5BFC30" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">
|
||
<emphasis id="83A13019FFCAFFDCFF3FB381FE29FC80" bold="true" box="[199,432,762,789]" pageId="3" pageNumber="4">Comparative notes.</emphasis>
|
||
<taxonomicName id="76D59788FFCAFFDCFE4FB380FD36FC80" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[439,687,763,789]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ahrensi" status="sp. nov.">
|
||
<emphasis id="83A13019FFCAFFDCFE4FB380FD36FC80" box="[439,687,763,789]" italics="true" pageId="3" pageNumber="4">Amphichroum ahrensi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="83A13019FFCAFFDCFD4EB380FD6CFC80" bold="true" box="[694,757,763,789]" pageId="3" pageNumber="4">
|
||
<taxonomicNameLabel id="98928D62FFCAFFDCFD4EB380FD6CFC80" box="[694,757,763,789]" pageId="3" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the remaining species of the Himalayan Region by the absence of pubescence on the disc of pronotum and elytra. Based on the shape of the median lobe, it somewhat similar to the Chinese
|
||
<taxonomicName id="76D59788FFCAFFDCFE25B238FC28FCC8" authority="Shavrin & Smetana, 2018" authorityName="Shavrin & Smetana" authorityYear="2018" box="[477,945,835,861]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="maculosum">
|
||
<emphasis id="83A13019FFCAFFDCFE25B238FE77FCC8" box="[477,494,835,861]" italics="true" pageId="3" pageNumber="4">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCAFFDCFE07B238FDE6FCC8" box="[511,639,835,861]" italics="true" pageId="3" pageNumber="4">maculosum</emphasis>
|
||
<bibRefCitation id="D54491FAFFCAFFDCFD70B238FC28FCC8" author="Shavrin, A. V. & Smetana, A." box="[648,945,835,861]" pageId="3" pageNumber="4" pagination="377 - 402" refId="ref9408" refString="Shavrin, A. V. & Smetana, A. (2018) New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China. Zootaxa, 4508 (3), 377 - 402. https: // doi. org / 10.11646 / zootaxa. 4508.3.3" type="journal article" year="2018">Shavrin & Smetana, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="76D59788FFCAFFDCFC15B238FF4BFC14" authority="Shavrin & Smetana, 2018" authorityName="Shavrin & Smetana" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="subelongatum">
|
||
<emphasis id="83A13019FFCAFFDCFC15B238FC67FCC8" box="[1005,1022,835,861]" italics="true" pageId="3" pageNumber="4">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFCAFFDCFBF6B238FB32FCC8" box="[1038,1195,835,861]" italics="true" pageId="3" pageNumber="4">subelongatum</emphasis>
|
||
<bibRefCitation id="D54491FAFFCAFFDCFB4DB238FF4BFC14" author="Shavrin, A. V. & Smetana, A." pageId="3" pageNumber="4" pagination="377 - 402" refId="ref9408" refString="Shavrin, A. V. & Smetana, A. (2018) New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China. Zootaxa, 4508 (3), 377 - 402. https: // doi. org / 10.11646 / zootaxa. 4508.3.3" type="journal article" year="2018">Shavrin & Smetana, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
, but differs from them by the paler coloration of the body and completely different morphology of the internal sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F9CFBF80FFCAFFDCFF3FB2D5FBA0FC78" pageId="3" pageNumber="4" type="distribution">
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB2D5FA02FC5C" blockId="3.[151,1437,151,1077]" box="[199,1435,942,969]" pageId="3" pageNumber="4">
|
||
<emphasis id="83A13019FFCAFFDCFF3FB2D5FEF8FC5D" bold="true" box="[199,353,942,968]" pageId="3" pageNumber="4">Distribution.</emphasis>
|
||
The new species is at present known only from the
|
||
<typeStatus id="6E6E52A9FFCAFFDCFC5EB2D4FC4FFC5C" box="[934,982,943,969]" pageId="3" pageNumber="4">type</typeStatus>
|
||
locality (
|
||
<figureCitation id="29EEF08EFFCAFFDCFBB9B2D4FB0BFC5C" box="[1089,1170,942,969]" captionStart="FIGURES 15–17" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1920]" captionTargetId="figure-27@6.[277,1310,181,1920]" captionTargetPageId="6" captionText="FIGURES 15–17. Distribution of Amphichroum species in the Himalayan Region and Tibet: 15—A. ahrensi (circle), A. altivagans (triangle), A. anthobioides (squares), 16—A. monticola (circles), A. pindarense (triangles), A. telnovi (diamond), 17—A. nepalicum." figureDoi="http://doi.org/10.5281/zenodo.5783140" httpUri="https://zenodo.org/record/5783140/files/figure.png" pageId="3" pageNumber="4">Fig. 15</figureCitation>
|
||
) in
|
||
<collectingRegion id="731122E9FFCAFFDCFB46B2D4FAD7FC5C" box="[1214,1358,943,969]" country="India" name="West Bengal" pageId="3" pageNumber="4">West Bengal</collectingRegion>
|
||
,
|
||
<collectingCountry id="C9C2AC9BFFCAFFDCFAA1B2D4FA0FFC5C" box="[1369,1430,943,969]" name="India" pageId="3" pageNumber="4">India</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB2A9FBA0FC78" blockId="3.[151,1437,151,1077]" box="[199,1081,978,1005]" pageId="3" pageNumber="4">
|
||
<emphasis id="83A13019FFCAFFDCFF3FB2A9FED2FC79" bold="true" box="[199,331,978,1004]" pageId="3" pageNumber="4">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="762D41EEFFCAFFDCFCB5B2A8FC64FC78" box="[845,1021,979,1005]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="3" pageNumber="4" unit="m" value="2800.0" valueMax="3000.0" valueMin="2600.0">2600 to 3000 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F9CFBF80FFCAFFDCFF3FB28CFEEDFBA0" pageId="3" pageNumber="4" type="etymology">
|
||
<paragraph id="B16AEC0BFFCAFFDCFF3FB28CFEEDFBA0" blockId="3.[151,1437,151,1077]" pageId="3" pageNumber="4">
|
||
<emphasis id="83A13019FFCAFFDCFF3FB28CFED7FB84" bold="true" box="[199,334,1015,1041]" pageId="3" pageNumber="4">Etymology.</emphasis>
|
||
Patronymic, the species is named to honor Dirk Ahrens (Bonn,
|
||
<collectingCountry id="C9C2AC9BFFCAFFDCFBE2B28CFB1EFB84" box="[1050,1159,1015,1041]" name="Germany" pageId="3" pageNumber="4">Germany</collectingCountry>
|
||
), one of the collectors of the
|
||
<typeStatus id="6E6E52A9FFCAFFDCFF38B560FF69FBA0" box="[192,240,1051,1077]" pageId="3" pageNumber="4">type</typeStatus>
|
||
specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |