259 lines
30 KiB
XML
259 lines
30 KiB
XML
<document id="5103441E8FEB61AAB56E57CBB7F0F24B" ID-CLB-Dataset="2949" ID-DOI="10.11646/zootaxa.5082.1.1" ID-GBIF-Dataset="b1305c10-d429-4335-aa4b-20786c960972" ID-ISSN="1175-5326" ID-Zenodo-Dep="5783128" ID-ZooBank="F4F057C6-3207-4430-A88E-4F4513D803C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1639564321074" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2021" docId="397C5D1DFFC3FFD3FF6FB545FB6FFD3F" docLanguage="en" docName="zootaxa.5082.1.1.pdf" docOrigin="Zootaxa 5082 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum telnovi Shavrin 2021, sp. n." docType="treatment" docVersion="7" lastPageNumber="13" masterDocId="C5452565FFC9FFDFFFF8B17BFF99FF95" masterDocTitle="New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="14" masterPageNumber="1" pageNumber="11" updateTime="1699472531112" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9065CC3A385AFC4BE60F46EA49F5CD27">New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
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<mods:namePart id="4A5F17F93415CEE5A62B4A6DE91B07F7">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="651890E942C4BB6B2E7DA28FAF3DB1FE">Zootaxa</mods:title>
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<mods:part id="D59241398C411B80AB536BD72129CF26">
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<mods:date id="0FB4305AF4002DAC358FD5F9B724F0B8">2021</mods:date>
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<mods:number id="BD391BF97D467C1510FDB5C25FCE1EEE">2021-12-15</mods:number>
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<mods:identifier id="C8EC1B1E9EB6C1A87F69C0EA2520E97C" type="DOI">10.11646/zootaxa.5082.1.1</mods:identifier>
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<treatment id="397C5D1DFFC3FFD3FF6FB545FB6FFD3F" ID-DOI="http://doi.org/10.5281/zenodo.5783615" ID-GBIF-Taxon="191647226" ID-Zenodo-Dep="5783615" LSID="urn:lsid:plazi:treatment:397C5D1DFFC3FFD3FF6FB545FB6FFD3F" httpUri="http://treatment.plazi.org/id/397C5D1DFFC3FFD3FF6FB545FB6FFD3F" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
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<subSubSection id="F9CFBF80FFC3FFD5FF6FB545FDC5FBCC" box="[151,604,1086,1113]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="B16AEC0BFFC3FFD5FF6FB545FDC5FBCC" blockId="10.[151,604,1086,1149]" box="[151,604,1086,1113]" pageId="10" pageNumber="11">
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<heading id="EA225B67FFC3FFD5FF6FB545FDC5FBCC" bold="true" box="[151,604,1086,1113]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="83A13019FFC3FFD5FF6FB545FDC5FBCC" bold="true" box="[151,604,1086,1113]" pageId="10" pageNumber="11">
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<taxonomicName id="76D59788FFC3FFD5FF6FB545FD95FBCD" ID-CoL="9GCPV" authority="Shavrin" authorityName="Shavrin" authorityYear="2021" box="[151,524,1086,1113]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
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<emphasis id="83A13019FFC3FFD5FF6FB545FE39FBCC" bold="true" box="[151,416,1086,1113]" italics="true" pageId="10" pageNumber="11">Amphichroum telnovi</emphasis>
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Shavrin
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="98928D62FFC3FFD5FDE1B544FDC5FBCC" box="[537,604,1087,1113]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F9CFBF80FFC3FFD5FF6FB518FEEDFBE8" box="[151,372,1122,1149]" pageId="10" pageNumber="11" type="description">
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<paragraph id="B16AEC0BFFC3FFD5FF6FB518FEEDFBE8" blockId="10.[151,604,1086,1149]" box="[151,372,1122,1149]" pageId="10" pageNumber="11">
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(
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<figureCitation id="29EEF08EFFC3FFD5FF67B518FF76FBE8" box="[159,239,1123,1149]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1773,1797]" captionTargetBox="[159,1425,482,1752]" captionTargetId="figure-189@2.[151,1436,474,1764]" captionTargetPageId="2" captionText="FIGURES 1–7. Habitus of Amphichroum (males): 1—A. ahrensi (paratype), 2—A. altivagans (holotype), 3—A. anthobioides (lectotype), 4—A. monticola (holotype), 5—A. nepalicum (Nepal, Manaslu Himal), 6—A. pindarense (syntype), 7—A. telnovi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5783132" httpUri="https://zenodo.org/record/5783132/files/figure.png" pageId="10" pageNumber="11">Figs. 7</figureCitation>
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,
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<figureCitation id="29EEF08EFFC3FFD5FF03B518FE80FBE8" box="[251,281,1123,1149]" captionStart="FIGURES 15–17" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1920]" captionTargetId="figure-27@6.[277,1310,181,1920]" captionTargetPageId="6" captionText="FIGURES 15–17. Distribution of Amphichroum species in the Himalayan Region and Tibet: 15—A. ahrensi (circle), A. altivagans (triangle), A. anthobioides (squares), 16—A. monticola (circles), A. pindarense (triangles), A. telnovi (diamond), 17—A. nepalicum." figureDoi="http://doi.org/10.5281/zenodo.5783140" httpUri="https://zenodo.org/record/5783140/files/figure.png" pageId="10" pageNumber="11">16</figureCitation>
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,
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<figureCitation id="29EEF08EFFC3FFD5FEDDB519FEF4FBE8" box="[293,365,1122,1149]" captionStart="FIGURES 25–26" captionStartId="12.[151,264,1975,1999]" captionTargetBox="[169,1422,715,1925]" captionTargetId="figure-255@12.[151,1436,705,1952]" captionTargetPageId="12" captionText="FIGURES 25–26. Aedeagus of Amphichroum telnovi (holotype): 25—parameral view, 26—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5783150" httpUri="https://zenodo.org/record/5783150/files/figure.png" pageId="10" pageNumber="11">25–26</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F9CFBF80FFC3FFD5FF6FB5D0FE86FAEC" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="B16AEC0BFFC3FFD5FF6FB5D0FB36FA98" blockId="10.[151,1437,1194,1977]" pageId="10" pageNumber="11">
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<emphasis id="83A13019FFC3FFD5FF6FB5D0FE5BFB51" bold="true" box="[151,450,1194,1221]" pageId="10" pageNumber="11">Type material examined:</emphasis>
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<materialsCitation id="01BDE656FFC3FFD5FE31B5D0FB36FA98" ID-GBIF-Occurrence="3418584312" collectingDate="2001-07-07" collectionCode="NME" collectorName="A. Weigel" country="Nepal" elevation="2121" latitude="29.845001" location="Bajura" longLatPrecision="20" longitude="81.790276" municipality="Bajura" pageId="10" pageNumber="11" specimenCount="1" specimenCount-male="1" stateProvince="Seti" typeStatus="holotype">
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<typeStatus id="6E6E52A9FFC3FFD5FE31B5D0FDADFB50" box="[457,564,1195,1221]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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,
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<specimenCount id="A7D32782FFC3FFD5FDC7B5D0FDCDFB50" box="[575,596,1195,1221]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
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(dissected): ‘
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<collectingCountry id="C9C2AC9BFFC3FFD5FD1FB5D0FCDAFB50" box="[743,835,1195,1221]" name="Nepal" pageId="10" pageNumber="11">NEPAL</collectingCountry>
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P:
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<collectingRegion id="731122E9FFC3FFD5FC91B5D0FC0CFB50" box="[873,917,1195,1221]" country="Nepal" name="Seti" pageId="10" pageNumber="11">Seti</collectingRegion>
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D:
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<collectingMunicipality id="510E7671FFC3FFD5FC46B5D0FB90FB50" box="[958,1033,1195,1221]" pageId="10" pageNumber="11">Bajura</collectingMunicipality>
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<locationDeviation id="A0D8F529FFC3FFD5FBE5B5D1FB4FFB50" box="[1053,1238,1194,1221]" pageId="10" pageNumber="11">
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<quantity id="762D41EEFFC3FFD5FBE5B5D1FBC4FB50" box="[1053,1117,1194,1221]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="10" pageNumber="11" unit="km" value="15.0">15km</quantity>
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S Simikot
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</locationDeviation>
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, N pass | slope W Malikasthan’ <printed>, ‘
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<quantity id="762D41EEFFC3FFD5FE4AB5B4FDB5FB7C" box="[434,556,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.1205" metricValueMax="4.2" metricValueMin="0.040999999999999995" pageId="10" pageNumber="11" unit="m" value="2120.5" valueMax="4200.0" valueMin="41.0">
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<elevation id="3AF80B38FFC3FFD5FE4AB5B4FDB5FB7C" box="[434,556,1231,1257]" metricMagnitude="3" metricUnit="m" metricValue="2.1205" metricValueMax="4.2" metricValueMin="0.040999999999999995" pageId="10" pageNumber="11" unit="m" value="2120.5" valueMax="4200.0" valueMin="41.0">41-4200m</elevation>
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</quantity>
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,
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<geoCoordinate id="D4E18ACCFFC3FFD5FDC0B5B5FD5FFB7C" box="[568,710,1230,1257]" degrees="29" direction="north" minutes="50" orientation="latitude" pageId="10" pageNumber="11" precision="15" seconds="42" value="29.845001">29°50’42’’N</geoCoordinate>
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<geoCoordinate id="D4E18ACCFFC3FFD5FD22B5B5FCFDFB7C" box="[730,868,1230,1257]" degrees="81" direction="east" minutes="47" orientation="longitude" pageId="10" pageNumber="11" precision="15" seconds="25" value="81.790276">81°47’25’’E</geoCoordinate>
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<date id="C56BCACBFFC3FFD5FC94B5B4FC6CFB7C" box="[876,1013,1231,1257]" pageId="10" pageNumber="11" value="2001-07-07">
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||
<collectingDate id="D52F3323FFC3FFD5FC94B5B4FC6CFB7C" box="[876,1013,1231,1257]" pageId="10" pageNumber="11" value="2001-07-07">07.VII.2001</collectingDate>
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</date>
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| leg.
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<collectorName id="1C2089DDFFC3FFD5FBD7B5B4FB05FB7C" box="[1071,1180,1231,1257]" pageId="10" pageNumber="11">A.Weigel</collectorName>
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stone-deb.’ <printed>, ‘HOLOTYPE |
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<taxonomicName id="76D59788FFC3FFD5FEB0B588FE7FFA98" authorityName="Kraatz" authorityYear="1857" box="[328,486,1267,1293]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="83A13019FFC3FFD5FEB0B588FE7FFA98" box="[328,486,1267,1293]" italics="true" pageId="10" pageNumber="11">Amphichroum</emphasis>
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</taxonomicName>
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|
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<taxonomicName id="76D59788FFC3FFD5FE01B588FDDCFA98" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[505,581,1267,1293]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
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<emphasis id="83A13019FFC3FFD5FE01B588FDDCFA98" box="[505,581,1267,1293]" italics="true" pageId="10" pageNumber="11">telnovi</emphasis>
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</taxonomicName>
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<emphasis id="83A13019FFC3FFD5FDB4B588FD15FA98" bold="true" box="[588,652,1267,1293]" pageId="10" pageNumber="11">
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<taxonomicNameLabel id="98928D62FFC3FFD5FDB4B588FD15FA98" box="[588,652,1267,1293]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.V. des. 2021’ <red, printed> (
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<collectionCode id="D7C474CEFFC3FFD5FBA7B588FB3AFA98" box="[1119,1187,1267,1293]" country="Germany" name="Sammlung des Naturkundemseum Erfurt" pageId="10" pageNumber="11">NME</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="B16AEC0BFFC3FFD5FF3FB46CFE86FAEC" blockId="10.[151,1437,1194,1977]" pageId="10" pageNumber="11">
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<materialsCitation id="01BDE656FFC3FFD5FF3FB46CFE7DFAC0" ID-GBIF-Occurrence="3418584313" pageId="10" pageNumber="11" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
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||
<typeStatus id="6E6E52A9FFC3FFD5FF3FB46CFEA1FAA4" box="[199,312,1303,1329]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
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||
:
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<specimenCount id="A7D32782FFC3FFD5FEBBB46CFEF6FAA4" box="[323,367,1303,1329]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
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||
,
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<specimenCount id="A7D32782FFC3FFD5FE83B46CFE23FAA4" box="[379,442,1303,1329]" pageId="10" pageNumber="11" type="female">2 ♀♀</specimenCount>
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(apical and preapical antennomeres of both females are missing): same data as the holotype, with additional red label
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</materialsCitation>
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‘
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<typeStatus id="6E6E52A9FFC3FFD5FE17B440FDE6FAC0" box="[495,639,1339,1365]" pageId="10" pageNumber="11" type="paratype">PARATYPE</typeStatus>
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||
|
|
||
<taxonomicName id="76D59788FFC3FFD5FD6AB440FCA9FAC0" authorityName="Kraatz" authorityYear="1857" box="[658,816,1339,1365]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="83A13019FFC3FFD5FD6AB440FCA9FAC0" box="[658,816,1339,1365]" italics="true" pageId="10" pageNumber="11">Amphichroum</emphasis>
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||
</taxonomicName>
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|
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||
<taxonomicName id="76D59788FFC3FFD5FCBAB440FC17FAC0" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[834,910,1339,1365]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
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<emphasis id="83A13019FFC3FFD5FCBAB440FC17FAC0" box="[834,910,1339,1365]" italics="true" pageId="10" pageNumber="11">telnovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="83A13019FFC3FFD5FC6CB440FC4AFAC0" bold="true" box="[916,979,1339,1365]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="98928D62FFC3FFD5FC6CB440FC4AFAC0" box="[916,979,1339,1365]" pageId="10" pageNumber="11" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
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||
| Shavrin A.V. des. 2021’ (
|
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<specimenCount id="A7D32782FFC3FFD5FB06B440FAB1FAC0" box="[1278,1320,1339,1365]" pageId="10" pageNumber="11" type="male">1 ♂</specimenCount>
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||
,
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<specimenCount id="A7D32782FFC3FFD5FACBB440FAC6FAC0" box="[1331,1375,1339,1365]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
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||
: cSh,
|
||
<specimenCount id="A7D32782FFC3FFD5FF6FB424FF5AFAEC" box="[151,195,1375,1401]" pageId="10" pageNumber="11" type="female">1 ♀</specimenCount>
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||
: NME).
|
||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="F9CFBF80FFC3FFD3FF3FB4F9FEFFFDA8" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="B16AEC0BFFC3FFD5FF3FB4F9FB1BFA70" blockId="10.[151,1437,1194,1977]" pageId="10" pageNumber="11">
|
||
<emphasis id="83A13019FFC3FFD5FF3FB4F9FEC0FA09" bold="true" box="[199,345,1410,1436]" pageId="10" pageNumber="11">Description.</emphasis>
|
||
Measurements (n=4): HL: 0.37–0.41; HW: 0.59–0.62; AL (
|
||
<typeStatus id="6E6E52A9FFC3FFD5FBF9B4F8FBF3FA0B" box="[1025,1130,1411,1438]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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||
): 1.44; OL: 0.17–0.21; PL: 0.61–0.65; PW: 0.81–0.92; ESL: 1.12–1.20; EW: 1.16–1.26; AW: 0.90–1.12; MTbL (
|
||
<typeStatus id="6E6E52A9FFC3FFD5FBABB4DCFB25FA57" box="[1107,1212,1447,1474]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
): 0.67; MTrL (
|
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<typeStatus id="6E6E52A9FFC3FFD5FAA7B4DCFF54FA70" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
): 0.54 (MTrL 1–4: 0.29; MTrL 5: 0.25); AedL: 0.65; TL: 3.56–3.95 (
|
||
<typeStatus id="6E6E52A9FFC3FFD5FC36B4B0FBAFFA70" box="[974,1078,1483,1509]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
: 3.70).
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFC3FFD5FF3FB494FD4AF890" blockId="10.[151,1437,1194,1977]" pageId="10" pageNumber="11">
|
||
Habitus as in
|
||
<figureCitation id="29EEF08EFFC3FFD5FE91B494FE29F99C" box="[361,432,1519,1545]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1773,1797]" captionTargetBox="[159,1425,482,1752]" captionTargetId="figure-189@2.[151,1436,474,1764]" captionTargetPageId="2" captionText="FIGURES 1–7. Habitus of Amphichroum (males): 1—A. ahrensi (paratype), 2—A. altivagans (holotype), 3—A. anthobioides (lectotype), 4—A. monticola (holotype), 5—A. nepalicum (Nepal, Manaslu Himal), 6—A. pindarense (syntype), 7—A. telnovi (paratype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5783132" httpUri="https://zenodo.org/record/5783132/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
|
||
. Body, antennomeres 4–11 and legs yellow-brown (head and disc of pronotum slightly darker); mouthparts, antennomeres 1–3 and tarsi yellow. Head with distinct and dense microsculpture, transverse on clypeus, finer in middle and isodiametric on infraorbital ridges; pronotum without visible microreticulation except for medioapical portion with very fine transverse meshes. Head with irregular, fine and sparse punctation, denser between anteocellar foveae; punctation of pronotum moderately dense, slightly larger and deeper than that in middle of head, sparser in middle and/or mediobasal portion; punctation of elytra larger and deeper than that on pronotum, coarser in basal portion and finer along suture; abdominal tergites with indistinct, sparse and fine punctation. Pronotum and elytra with dense and short pubescence.
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFC3FFD5FF3FB674FF34F82C" blockId="10.[151,1437,1194,1977]" pageId="10" pageNumber="11">Head 1.5 times as broad as long, with narrow or moderately wide and deep anteocellar foveae, reaching level of middle or anterior margins of eyes. Ocelli very large, distance between ocelli about as long as distance between ocellus and posterior margin of eye. Antennomere 3 slightly longer and narrower than 2, 4 distinctly shorter than 3, 5–6 slightly longer and broader than 4, 7 slightly longer than 6, 8 slightly shorter than 7, 9–10 slightly longer than 8.</paragraph>
|
||
<caption id="E5AABC83FFC2FFD4FF6FB6ECFC44F862" ID-DOI="http://doi.org/10.5281/zenodo.5783144" ID-Zenodo-Dep="5783144" httpUri="https://zenodo.org/record/5783144/files/figure.png" pageId="11" pageNumber="12" startId="11.[151,264,1943,1967]" targetBox="[166,1425,189,1913]" targetPageId="11">
|
||
<paragraph id="B16AEC0BFFC2FFD4FF6FB6ECFC44F862" blockId="11.[151,1437,1943,2040]" pageId="11" pageNumber="12">
|
||
<emphasis id="83A13019FFC2FFD4FF6FB6ECFEC1F83A" bold="true" box="[151,344,1943,1967]" pageId="11" pageNumber="12">FIGURES 18–24.</emphasis>
|
||
Aedeagus of
|
||
<taxonomicName id="76D59788FFC2FFD4FE15B6E3FD72F825" authorityName="Shavrin" authorityYear="2021" box="[493,747,1944,1968]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="monticola">
|
||
<emphasis id="83A13019FFC2FFD4FE15B6E3FD72F825" box="[493,747,1944,1968]" italics="true" pageId="11" pageNumber="12">Amphichroum monticola</emphasis>
|
||
</taxonomicName>
|
||
(18, 20: Nepal, Deorali, 19: Nepal, Khola Valley),
|
||
<taxonomicName id="76D59788FFC2FFD4FAEBB6E3FA04F825" authorityName="Coiffait" authorityYear="1982" box="[1299,1437,1944,1968]" class="Insecta" family="Staphylinidae" genus="Arpedium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="nepalicum">
|
||
<emphasis id="83A13019FFC2FFD4FAEBB6E3FABBF83A" box="[1299,1314,1944,1967]" italics="true" pageId="11" pageNumber="12">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC2FFD4FACBB6E3FA04F825" box="[1331,1437,1944,1968]" italics="true" pageId="11" pageNumber="12">nepalicum</emphasis>
|
||
</taxonomicName>
|
||
(21–22: Nepal, Manaslu Himal) and
|
||
<taxonomicName id="76D59788FFC2FFD4FDEBB6C7FD38F841" authorityName="Champion" authorityYear="1920" box="[531,673,1980,2004]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pindarense">
|
||
<emphasis id="83A13019FFC2FFD4FDEBB6C7FDBBF846" box="[531,546,1980,2003]" italics="true" pageId="11" pageNumber="12">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC2FFD4FDC9B6C7FD38F841" box="[561,673,1980,2004]" italics="true" pageId="11" pageNumber="12">pindarense</emphasis>
|
||
</taxonomicName>
|
||
(23–24: Tibet, Rongshar Valley): 18, 21, 23—parameral view, 19—apical portion of aedeagus, parameral view, 20, 22, 24—lateral view. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB1ECFE9FFF40" blockId="12.[151,1436,151,682]" pageId="12" pageNumber="13">Pronotum 1.3–1.4 times as broad as long and head, widest in about middle, evenly narrowed both anteriad and posteriad.</paragraph>
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB1A4FAC6FEF0" blockId="12.[151,1436,151,682]" pageId="12" pageNumber="13">
|
||
Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII rounded. Aedeagus (
|
||
<figureCitation id="29EEF08EFFC5FFD3FF0BB078FEDAFE88" box="[243,323,258,285]" captionStart="FIGURES 25–26" captionStartId="12.[151,264,1975,1999]" captionTargetBox="[169,1422,715,1925]" captionTargetId="figure-255@12.[151,1436,705,1952]" captionTargetPageId="12" captionText="FIGURES 25–26. Aedeagus of Amphichroum telnovi (holotype): 25—parameral view, 26—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5783150" httpUri="https://zenodo.org/record/5783150/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
|
||
) with wide basal portion, gradually narrowed toward subacute apex; parameres very narrow, slightly exceeding apex of median lobe, with one short apical and preapical setae; internal sac narrow, long, with a pair of subapical and median and four basal sclerotized, elongate structures. Lateral aspect of aedeagus as in
|
||
<figureCitation id="29EEF08EFFC5FFD3FAF1B030FAC2FEF0" box="[1289,1371,331,357]" captionStart="FIGURES 25–26" captionStartId="12.[151,264,1975,1999]" captionTargetBox="[169,1422,715,1925]" captionTargetId="figure-255@12.[151,1436,705,1952]" captionTargetPageId="12" captionText="FIGURES 25–26. Aedeagus of Amphichroum telnovi (holotype): 25—parameral view, 26—lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5783150" httpUri="https://zenodo.org/record/5783150/files/figure.png" pageId="12" pageNumber="13">Fig. 26</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB014FB85FE1C" blockId="12.[151,1436,151,682]" box="[199,1052,367,393]" pageId="12" pageNumber="13">Female. Apical margins of abdominal tergite VIII and sternite VIII rounded.</paragraph>
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB0E9FEFFFDA8" blockId="12.[151,1436,151,682]" pageId="12" pageNumber="13">
|
||
<emphasis id="83A13019FFC5FFD3FF3FB0E9FE28FE38" bold="true" box="[199,433,402,429]" pageId="12" pageNumber="13">Comparative notes.</emphasis>
|
||
Based on the general shapes of the apical portion of the median lobe and the parameres,
|
||
<taxonomicName id="76D59788FFC5FFD3FF6FB0CCFE9BFE44" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[151,258,439,465]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="telnovi" status="sp. nov.">
|
||
<emphasis id="83A13019FFC5FFD3FF6FB0CCFF31FE44" box="[151,168,439,465]" italics="true" pageId="12" pageNumber="13">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC5FFD3FF4EB0CCFE9BFE44" box="[182,258,439,465]" italics="true" pageId="12" pageNumber="13">telnovi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="83A13019FFC5FFD3FEF1B0CCFED1FE44" bold="true" box="[265,328,439,465]" pageId="12" pageNumber="13">
|
||
<taxonomicNameLabel id="98928D62FFC5FFD3FEF1B0CCFED1FE44" box="[265,328,439,465]" pageId="12" pageNumber="13" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="76D59788FFC5FFD3FE21B0CCFDF4FE44" authorityName="Cameron" authorityYear="1941" box="[473,621,439,465]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="altivagans">
|
||
<emphasis id="83A13019FFC5FFD3FE21B0CCFE73FE44" box="[473,490,439,465]" italics="true" pageId="12" pageNumber="13">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC5FFD3FE00B0CCFDF4FE44" box="[504,621,439,465]" italics="true" pageId="12" pageNumber="13">altivagans</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="76D59788FFC5FFD3FD5AB0CCFCCBFE44" authorityName="Champion" authorityYear="1925" box="[674,850,439,465]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anthobioides">
|
||
<emphasis id="83A13019FFC5FFD3FD5AB0CCFD2AFE44" box="[674,691,439,465]" italics="true" pageId="12" pageNumber="13">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC5FFD3FD39B0CCFCCBFE44" box="[705,850,439,465]" italics="true" pageId="12" pageNumber="13">anthobioides</emphasis>
|
||
</taxonomicName>
|
||
. From
|
||
<taxonomicName id="76D59788FFC5FFD3FC5BB0CCFBAEFE44" authorityName="Cameron" authorityYear="1941" box="[931,1079,439,465]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="altivagans">
|
||
<emphasis id="83A13019FFC5FFD3FC5BB0CCFC2DFE44" box="[931,948,439,465]" italics="true" pageId="12" pageNumber="13">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC5FFD3FC3AB0CCFBAEFE44" box="[962,1079,439,465]" italics="true" pageId="12" pageNumber="13">altivagans</emphasis>
|
||
</taxonomicName>
|
||
it differs by the larger body, the shape of the pronotum, from middle evenly narrowed both anteriad and posteriad, and shorter elytra, and from
|
||
<taxonomicName id="76D59788FFC5FFD3FA7CB0A0FEB1FD8C" authorityName="Champion" authorityYear="1925" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="anthobioides">
|
||
<emphasis id="83A13019FFC5FFD3FA7CB0A0FA0CFE60" box="[1412,1429,475,501]" italics="true" pageId="12" pageNumber="13">A</emphasis>
|
||
.
|
||
<emphasis id="83A13019FFC5FFD3FF6FB084FEB1FD8C" box="[151,296,511,537]" italics="true" pageId="12" pageNumber="13">anthobioides</emphasis>
|
||
</taxonomicName>
|
||
by the paler pronotum and elytra. From both species it can be distinguished by details of the structure of the internal sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F9CFBF80FFC5FFD3FF3FB33DFBA0FD10" pageId="12" pageNumber="13" type="distribution">
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB33DFB6AFDF7" blockId="12.[151,1436,151,682]" box="[199,1267,582,610]" pageId="12" pageNumber="13">
|
||
<emphasis id="83A13019FFC5FFD3FF3FB33DFEF8FDF5" bold="true" box="[199,353,582,608]" pageId="12" pageNumber="13">Distribution.</emphasis>
|
||
The new species is known only from the
|
||
<typeStatus id="6E6E52A9FFC5FFD3FCCBB33CFCFAFDF4" box="[819,867,583,609]" pageId="12" pageNumber="13">type</typeStatus>
|
||
locality in western
|
||
<collectingCountry id="C9C2AC9BFFC5FFD3FBB8B33CFB1AFDF4" box="[1088,1155,583,609]" name="Nepal" pageId="12" pageNumber="13">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="29EEF08EFFC5FFD3FB6BB33CFB7FFDF7" box="[1171,1254,583,610]" captionStart="FIGURES 15–17" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[277,1310,181,1920]" captionTargetId="figure-27@6.[277,1310,181,1920]" captionTargetPageId="6" captionText="FIGURES 15–17. Distribution of Amphichroum species in the Himalayan Region and Tibet: 15—A. ahrensi (circle), A. altivagans (triangle), A. anthobioides (squares), 16—A. monticola (circles), A. pindarense (triangles), A. telnovi (diamond), 17—A. nepalicum." figureDoi="http://doi.org/10.5281/zenodo.5783140" httpUri="https://zenodo.org/record/5783140/files/figure.png" pageId="12" pageNumber="13">Fig. 16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB311FBA0FD10" blockId="12.[151,1436,151,682]" box="[199,1081,618,645]" pageId="12" pageNumber="13">
|
||
<emphasis id="83A13019FFC5FFD3FF3FB311FED2FD11" bold="true" box="[199,331,618,644]" pageId="12" pageNumber="13">Bionomics.</emphasis>
|
||
Specimens were collected at elevations from
|
||
<quantity id="762D41EEFFC5FFD3FCB5B310FC64FD10" box="[845,1021,619,645]" metricMagnitude="3" metricUnit="m" metricValue="4.15" metricValueMax="4.2" metricValueMin="4.1" pageId="12" pageNumber="13" unit="m" value="4150.0" valueMax="4200.0" valueMin="4100.0">4100 to 4200 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F9CFBF80FFC5FFD3FF3FB3F4FB6FFD3F" box="[199,1270,655,682]" pageId="12" pageNumber="13" type="etymology">
|
||
<paragraph id="B16AEC0BFFC5FFD3FF3FB3F4FB6FFD3F" blockId="12.[151,1436,151,682]" box="[199,1270,655,682]" pageId="12" pageNumber="13">
|
||
<emphasis id="83A13019FFC5FFD3FF3FB3F4FED7FD3C" bold="true" box="[199,334,655,681]" pageId="12" pageNumber="13">Etymology.</emphasis>
|
||
Patronymic, the species is named to honor my colleague, Dmitry Telnov (London).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |