333 lines
44 KiB
XML
333 lines
44 KiB
XML
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<mods:title id="56D2A8285AEE6A38E0160DC040934CAC">New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata)</mods:title>
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<mods:namePart id="FDAB6815A13B08A857D859429C39F7B0">Béthoux, Olivier</mods:namePart>
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<mods:affiliation id="99DB9D3FBA3563A0BFC97F2198073376">CR 2 P (CNRS, MNHN, Sorbonne Université), Muséum national d’Histoire naturelle, case postale 48, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) obethoux @ mnhn. fr</mods:affiliation>
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<mods:namePart id="949791211F9DBE43704E7DE69029B89A">Anderson, John M.</mods:namePart>
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<mods:affiliation id="D5A43404826B8CCB23B7593AD4580DBD">Evolutionary Studies Institute, Witwatersrand University, 1 Jan Smuts Ave., Braamfontein, Johannesburg 2000, South Africa jmanderson. gondwana @ gmail. com</mods:affiliation>
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<mods:nameIdentifier id="4D05BD58D811B8A1DD20092374EAB07D" type="email">gondwana@gmail.com</mods:nameIdentifier>
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<mods:title id="9A278E82186CE2773A6DD89540777E02">Geodiversitas</mods:title>
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<mods:date id="206BDD96D5B8934C882B68D8B4AC4BDF">2023</mods:date>
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Genus
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<taxonomicName id="7E99B8B74B54FFA9FBA0FBB39138E50C" authority="Riek, 1976" authorityName="Riek" authorityYear="1976" box="[1049,1296,1110,1136]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="genus">
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<emphasis id="8BED1F264B54FFA9FBA0FBB390BAE50C" box="[1049,1170,1110,1136]" italics="true" pageId="9" pageNumber="486">Triassologus</emphasis>
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<bibRefCitation id="DD08BEC54B54FFA9FB21FBB39138E50C" author="RIEK E. F." box="[1176,1296,1110,1136]" pageId="9" pageNumber="486" pagination="791 - 820" refId="ref15208" refString="RIEK E. F. 1976. - A new collection of insects from the Upper Triassic of South Africa. Annals of the Natal Museum 22 (3): 791 - 820. https: // hdl. handle. net / 10520 / AJA 03040798 _ 608" type="journal article" year="1976">Riek, 1976</bibRefCitation>
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</taxonomicName>
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<treatmentCitation id="3838E5254B54FFA9FC94FB74906AE5D5" author="RIEK E. F." box="[813,1090,1169,1193]" page="793" pageId="9" pageNumber="486" year="1976">
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<taxonomicName id="7E99B8B74B54FFA9FC94FB74906AE5D5" authority="Riek, 1976: 793" authorityName="Riek" authorityPageNumber="793" authorityYear="1976" box="[813,1090,1169,1193]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="genus">
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<emphasis id="8BED1F264B54FFA9FC94FB7497B2E5D5" box="[813,922,1169,1193]" italics="true" pageId="9" pageNumber="486">Triassologus</emphasis>
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<bibRefCitation id="DD08BEC54B54FFA9FC19FB74906AE5D5" author="RIEK E. F." box="[928,1090,1169,1193]" pageId="9" pageNumber="486" pagination="791 - 820" refId="ref15208" refString="RIEK E. F. 1976. - A new collection of insects from the Upper Triassic of South Africa. Annals of the Natal Museum 22 (3): 791 - 820. https: // hdl. handle. net / 10520 / AJA 03040798 _ 608" type="journal article" year="1976">Riek, 1976: 793</bibRefCitation>
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. —
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species: see below.
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<paragraph id="B926C3344B54FFA9FC94FB21905AE58A" blockId="9.[813,1455,1219,1270]" pageId="9" pageNumber="486">
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<treatmentCitation id="3838E5254B54FFA9FC94FB2190BAE5A7" author="BETHOUX O. & BEATTIE R. G." box="[813,1170,1219,1243]" page="689" pageId="9" pageNumber="486" year="2010">
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<taxonomicName id="7E99B8B74B54FFA9FC94FB2190BAE5A7" authority="Bethoux & Beattie, 2010: 689" authorityName="Bethoux & Beattie" authorityPageNumber="689" authorityYear="2010" box="[813,1170,1219,1243]" class="Insecta" family="Triadotypidae" genus="Iverya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="genus">
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<emphasis id="8BED1F264B54FFA9FC94FB21974EE5A7" box="[813,870,1220,1243]" italics="true" pageId="9" pageNumber="486">Iverya</emphasis>
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<bibRefCitation id="DD08BEC54B54FFA9FCD2FB2690BAE5A7" author="BETHOUX O. & BEATTIE R. G." box="[875,1170,1219,1243]" pageId="9" pageNumber="486" pagination="688 - 692" refId="ref13268" refString="BETHOUX O. & BEATTIE R. G. 2010. - Iverya averyi gen. nov. & sp. nov., a new triadotypomorphan species from the Middle Triassic at Picton, New South Wales, Australia. Acta Geologica Sinica 84: 688 - 692. https: // doi. org / 10.1111 / j. 1755 - 6724.2010.00240. x" type="journal article" year="2010">Béthoux & Beattie, 2010: 689</bibRefCitation>
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—
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species:
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<taxonomicName id="7E99B8B74B54FFA9FA81FB21900DE58A" authority="Bethoux & Beattie, 2010" authorityName="Bethoux & Beattie" authorityYear="2010" class="Insecta" family="Triadotypidae" genus="Iverya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="aveyri">
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<emphasis id="8BED1F264B54FFA9FA81FB219187E5A7" box="[1336,1455,1220,1243]" italics="true" pageId="9" pageNumber="486">Iverya aveyri</emphasis>
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<bibRefCitation id="DD08BEC54B54FFA9FC94FB38900DE58A" author="BETHOUX O. & BEATTIE R. G." box="[813,1061,1245,1270]" pageId="9" pageNumber="486" pagination="688 - 692" refId="ref13268" refString="BETHOUX O. & BEATTIE R. G. 2010. - Iverya averyi gen. nov. & sp. nov., a new triadotypomorphan species from the Middle Triassic at Picton, New South Wales, Australia. Acta Geologica Sinica 84: 688 - 692. https: // doi. org / 10.1111 / j. 1755 - 6724.2010.00240. x" type="journal article" year="2010">Béthoux & Beattie, 2010</bibRefCitation>
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</taxonomicName>
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||
,
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<emphasis id="8BED1F264B54FFA9FB89FB3B905AE58A" bold="true" box="[1072,1138,1246,1270]" pageId="9" pageNumber="486">
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<taxonomicNameLabel id="90DEA25D4B54FFA9FB89FB3B905AE58A" box="[1072,1138,1246,1270]" pageId="9" pageNumber="486">n. syn.</taxonomicNameLabel>
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</emphasis>
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. —
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<taxonomicName id="7E99B8B74B54FFA9FC5DFAEA911FE45B" authority="Riek, 1976" authorityName="Riek" authorityYear="1976" box="[996,1335,1295,1319]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="biseriatus">
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<emphasis id="8BED1F264B54FFA9FC5DFAEA9094E45B" box="[996,1212,1295,1319]" italics="true" pageId="9" pageNumber="486">Triassologus biseriatus</emphasis>
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<bibRefCitation id="DD08BEC54B54FFA9FB7DFAEA911FE45B" author="RIEK E. F." box="[1220,1335,1295,1319]" pageId="9" pageNumber="486" pagination="791 - 820" refId="ref15208" refString="RIEK E. F. 1976. - A new collection of insects from the Upper Triassic of South Africa. Annals of the Natal Museum 22 (3): 791 - 820. https: // hdl. handle. net / 10520 / AJA 03040798 _ 608" type="journal article" year="1976">Riek, 1976</bibRefCitation>
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</taxonomicName>
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by original designation.
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. —
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<taxonomicName id="7E99B8B74B54FFA9FC56FABE9183E408" authority="(Bethoux & Beattie, 2010)" authorityName="Béthoux & Anderson" authorityYear="2023" baseAuthorityName="Bethoux & Beattie" baseAuthorityYear="2010" box="[1007,1451,1371,1396]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="aveyri" status="comb. nov.">
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<emphasis id="8BED1F264B54FFA9FC56FABE90B3E40F" box="[1007,1179,1371,1395]" italics="true" pageId="9" pageNumber="486">Triassologus aveyri</emphasis>
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(
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<bibRefCitation id="DD08BEC54B54FFA9FB1EFABE918CE408" author="BETHOUX O. & BEATTIE R. G." box="[1191,1444,1371,1396]" pageId="9" pageNumber="486" pagination="688 - 692" refId="ref13268" refString="BETHOUX O. & BEATTIE R. G. 2010. - Iverya averyi gen. nov. & sp. nov., a new triadotypomorphan species from the Middle Triassic at Picton, New South Wales, Australia. Acta Geologica Sinica 84: 688 - 692. https: // doi. org / 10.1111 / j. 1755 - 6724.2010.00240. x" type="journal article" year="2010">Béthoux & Beattie, 2010</bibRefCitation>
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)
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</taxonomicName>
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,
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||
<taxonomicNameLabel id="90DEA25D4B54FFA9FC94FA9397ADE4F2" box="[813,901,1398,1422]" pageId="9" pageNumber="486" rank="species">n. comb.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="F18390BF4B54FFABFC94FA4D958AE429" lastPageId="11" lastPageNumber="488" pageId="9" pageNumber="486" type="description">
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<paragraph id="B926C3344B54FFAAFC94FA4D9442E00A" blockId="9.[811,1457,1448,2028]" lastBlockId="10.[91,106,132,374]" lastPageId="10" lastPageNumber="487" pageId="9" pageNumber="486">
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. — Hindwing broader than forewing (1.3 broader opposite nodus); area between anterior wing margin (or ScA) and ScP very broad in forewing, moderately broad in hindwing; area between anterior wing margin and RA very narrow opposite the third quarter of wing length, before widening again (as opposed to tapering, or broadening, regularly from the nodus to the apex; uncertain in
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<taxonomicName id="7E99B8B74B54FFA9FCD4F9A3903EE722" authorityName="Béthoux & Anderson" authorityYear="2023" baseAuthorityName="Bethoux & Beattie" baseAuthorityYear="2010" box="[877,1046,1606,1630]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="aveyri" status="comb. nov.">
|
||
<emphasis id="8BED1F264B54FFA9FCD4F9A3903EE722" box="[877,1046,1606,1630]" italics="true" pageId="9" pageNumber="486">Triassologus aveyri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90DEA25D4B54FFA9FBA3F9A2905AE722" box="[1050,1138,1606,1630]" pageId="9" pageNumber="486" rank="species">n. comb.</taxonomicNameLabel>
|
||
); nodal and sub-nodal cross-veins short, moderately strong, sub-aligned, oblique (condition unknown in
|
||
<taxonomicName id="7E99B8B74B54FFA9FCFEF99E97DBE7EF" authorityName="Béthoux & Anderson" authorityYear="2023" baseAuthorityName="Bethoux & Beattie" baseAuthorityYear="2010" box="[839,1011,1659,1683]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="aveyri" status="comb. nov.">
|
||
<emphasis id="8BED1F264B54FFA9FCFEF99E97DBE7EF" box="[839,1011,1659,1683]" italics="true" pageId="9" pageNumber="486">Triassologus aveyri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90DEA25D4B54FFA9FC43F999907DE7EF" box="[1018,1109,1659,1683]" pageId="9" pageNumber="486" rank="species">n. comb.</taxonomicNameLabel>
|
||
); RP1/RP2 fork in a position more distal than that of the point of origin of the first I- in the RP3+4 area; first I- in the Irp
|
||
<subScript id="251DC1714B54FFA9FBB0F95E903AE7B0" attach="left" box="[1033,1042,1723,1740]" fontSize="7" pageId="9" pageNumber="486">1</subScript>
|
||
-rp
|
||
<subScript id="251DC1714B54FFA9FB96F95E9010E7B0" attach="left" box="[1071,1080,1723,1740]" fontSize="7" pageId="9" pageNumber="486">2</subScript>
|
||
-RP2 area seemingly originating from RP2 (i.e. RP2 pseudo-forked; large blue arrows on Fig. 4B-D, F; unknown in
|
||
<taxonomicName id="7E99B8B74B54FFA9FC1FF9009066E781" authorityName="Béthoux & Anderson" authorityYear="2023" baseAuthorityName="Bethoux & Beattie" baseAuthorityYear="2010" box="[934,1102,1765,1789]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="aveyri" status="comb. nov.">
|
||
<emphasis id="8BED1F264B54FFA9FC1FF9009066E781" box="[934,1102,1765,1789]" italics="true" pageId="9" pageNumber="486">Triassologus aveyri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90DEA25D4B54FFA9FBEAF9039085E781" box="[1107,1197,1765,1789]" pageId="9" pageNumber="486" rank="species">n. comb.</taxonomicNameLabel>
|
||
); point of origin of the first I- in the RP3+4 area basal to the RP1/RP2 fork; RP3+4 very long, parallel to the posterior wing margin for some distance (unknown in
|
||
<taxonomicName id="7E99B8B74B54FFA9FCF0F8D097D3E630" authorityName="Béthoux & Anderson" authorityYear="2023" baseAuthorityName="Bethoux & Beattie" baseAuthorityYear="2010" box="[841,1019,1845,1869]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="aveyri" status="comb. nov.">
|
||
<emphasis id="8BED1F264B54FFA9FCF0F8D097D3E630" box="[841,1019,1845,1869]" italics="true" pageId="9" pageNumber="486">Triassologus aveyri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="90DEA25D4B54FFA9FBBCF8D0904CE631" box="[1029,1124,1845,1869]" pageId="9" pageNumber="486" rank="species">n. comb.</taxonomicNameLabel>
|
||
); pons moderately long; RP/MA fork opposite that of MP/Cu+AA; at least in the basal part of the CuA-CuP area, I- rectilinear and seemingly originating from CuA (possibly as convex elements quickly turning concave; large white arrows on
|
||
<figureCitation id="21A2DFB14B54FFA9FC36F87B97EEE6CA" box="[911,966,1950,1974]" captionStart="FIG" captionStartId="14.[132,143,1174,1191]" captionTargetBox="[132,775,215,1131]" captionTargetId="figure-436@14.[132,775,215,1131]" captionTargetPageId="14" captionText="FIG. 6. — Detail of the organisation of CuA branches and of I- in the basal portion of the CuA-CuP area in Triassologus spp. (large green arrows, CuA branches; large white arrows, Icua): A, Triassologus biseriatus Riek, 1976, specimen PRE/F/5118b (wing in dorsal aspect), flipped horizontally, location as shown on Fig.4E; B, Triassologus aveyri (Béthoux & Beatie,2010),n.comb.,holotype, specimen AM F.132815. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8399050" httpUri="https://zenodo.org/record/8399050/files/figure.png" pageId="9" pageNumber="486">Fig. 6</figureCitation>
|
||
), genuine CuA branches zigzagging and seemingly originating from cross-venation (i.e. as an I+; large green arrows on
|
||
<figureCitation id="21A2DFB14B54FFA9FC95F836974CE697" box="[812,868,2003,2027]" captionStart="FIG" captionStartId="14.[132,143,1174,1191]" captionTargetBox="[132,775,215,1131]" captionTargetId="figure-436@14.[132,775,215,1131]" captionTargetPageId="14" captionText="FIG. 6. — Detail of the organisation of CuA branches and of I- in the basal portion of the CuA-CuP area in Triassologus spp. (large green arrows, CuA branches; large white arrows, Icua): A, Triassologus biseriatus Riek, 1976, specimen PRE/F/5118b (wing in dorsal aspect), flipped horizontally, location as shown on Fig.4E; B, Triassologus aveyri (Béthoux & Beatie,2010),n.comb.,holotype, specimen AM F.132815. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8399050" httpUri="https://zenodo.org/record/8399050/files/figure.png" pageId="9" pageNumber="486">Fig. 6</figureCitation>
|
||
; and see small green arrows on Figs 4D, 5B); CuP reaching GEODIVERSITAS • 2023 • 45 (17)
|
||
</paragraph>
|
||
<caption id="EDE693BC4B57FFAAFF59FAE39787E40E" ID-DOI="http://doi.org/10.5281/zenodo.8399038" ID-Zenodo-Dep="8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="10" pageNumber="487" startId="10.[224,235,1286,1303]" targetBox="[258,1996,208,1246]" targetPageId="10" targetType="figure">
|
||
<paragraph id="B926C3344B57FFAAFF59FAE39787E40E" blockId="10.[89,1968,1286,1440]" pageId="10" pageNumber="487">
|
||
<smallCapsWord id="BFC055E84B57FFAAFF59FAE394D0E46B" baselines="1299,1300" box="[224,248,1286,1303]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="10" pageNumber="487">FIG</smallCapsWord>
|
||
. 2. —
|
||
<taxonomicName id="7E99B8B74B57FFAAFE95FAE396EEE46B" authority="(Grauvogel & Laurentiaux, 1952)" authorityName="Béthoux & Anderson" authorityYear="2023" baseAuthorityName="Grauvogel & Laurentiaux" baseAuthorityYear="1952" box="[300,710,1286,1303]" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="10" pageNumber="487" phylum="Arthropoda" rank="species" species="guillaumei" status="comb. nov.">
|
||
<emphasis id="8BED1F264B57FFAAFE95FAE3959EE46B" box="[300,438,1286,1303]" italics="true" pageId="10" pageNumber="487">Reisia guillaumei</emphasis>
|
||
(
|
||
<bibRefCitation id="DD08BEC54B57FFAAFE07FAE396EAE46B" author="GRAUVOGEL L. & LAURENTIAUX D." box="[446,706,1286,1303]" pageId="10" pageNumber="487" pagination="121 - 129" refId="ref14098" refString="GRAUVOGEL L. & LAURENTIAUX D. 1952. - Un protodonate du trias des Vosges. Annales de Paleontologie, Invertebres 38: 121 - 129." type="journal article" year="1952">Grauvogel & Laurentiaux, 1952</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="90DEA25D4B57FFAAFD75FAE39732E46B" box="[716,794,1286,1303]" pageId="10" pageNumber="487" rank="species">n. comb.</taxonomicNameLabel>
|
||
,holotype,specimen 7885 (collection Louis Grauvogel at the ‘ Staatlisches Museum für Naturkunde Stuttgart’,Stuttgart,Germany), left wing: view;
|
||
<emphasis id="8BED1F264B57FFAAFEA9FAF89535E452" bold="true" box="[272,285,1309,1326]" pageId="10" pageNumber="487">A</emphasis>
|
||
, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches);
|
||
<emphasis id="8BED1F264B57FFAAF8A8FAF89336E452" bold="true" box="[1809,1822,1309,1326]" pageId="10" pageNumber="487">B</emphasis>
|
||
, photograph of fragment, dorsal aspect (flipped horizontally);
|
||
<emphasis id="8BED1F264B57FFAAFDE3FAD19640E439" bold="true" box="[602,616,1332,1349]" pageId="10" pageNumber="487">C</emphasis>
|
||
, photograph of the distal fragment, dorsal aspect (flipped horizontally);
|
||
<emphasis id="8BED1F264B57FFAAFB0EFAD190FFE439" bold="true" box="[1207,1239,1332,1349]" pageId="10" pageNumber="487">D -F</emphasis>
|
||
, detail of the wing base, location as shown on
|
||
<emphasis id="8BED1F264B57FFAAF9E3FAD1924FE439" bold="true" box="[1626,1639,1332,1349]" pageId="10" pageNumber="487">B</emphasis>
|
||
; drawing of venation (enlargement from added color-coding);
|
||
<emphasis id="8BED1F264B57FFAAFE2AFAAE95B7E420" bold="true" box="[403,415,1355,1372]" pageId="10" pageNumber="487">E</emphasis>
|
||
,
|
||
<emphasis id="8BED1F264B57FFAAFE10FAAE959CE420" bold="true" box="[425,436,1355,1372]" pageId="10" pageNumber="487">F</emphasis>
|
||
, photographs extracted from RTI file (
|
||
<emphasis id="8BED1F264B57FFAAFD48FAAE96D4E420" bold="true" box="[753,764,1355,1372]" pageId="10" pageNumber="487">F</emphasis>
|
||
, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B926C3344B57FFABFFE0FA9D958AE429" blockId="10.[89,1968,1286,1440]" lastBlockId="11.[131,777,1235,1365]" lastPageId="11" lastPageNumber="488" pageId="10" pageNumber="487">487 the posterior wing margin opposite the nodus or slightly distal to it; CuP and AA posteriorly pectinate, with a curved and sigmoid course, respectively; area between AA (or Cu+AA, or MP+Cu+AA) and the posterior wing margin, opposite and basal to the pons, narrow (i.e. not forming a lobe).</paragraph>
|
||
</subSubSection>
|
||
<caption id="EDE693BC4B56FFABFF3DFB9D97A2E5E3" ID-DOI="http://doi.org/10.5281/zenodo.8399042" ID-Zenodo-Dep="8399042" httpUri="https://zenodo.org/record/8399042/files/figure.png" pageId="11" pageNumber="488" startId="11.[132,143,1144,1161]" targetBox="[175,1415,222,1101]" targetPageId="11" targetType="figure">
|
||
<paragraph id="B926C3344B56FFABFF3DFB9D97A2E5E3" blockId="11.[132,1455,1144,1183]" pageId="11" pageNumber="488">
|
||
<smallCapsWord id="BFC055E84B56FFABFF3DFB9D94B5E5F4" baselines="1157,1157" box="[132,157,1144,1161]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="11" pageNumber="488">FIG</smallCapsWord>
|
||
. 3. —
|
||
<taxonomicName id="7E99B8B74B56FFABFF6CFB9D95F5E5F5" authority="Meunier, 1907" authorityName="Meunier" authorityYear="1907" box="[213,477,1144,1161]" class="Insecta" family="Piroutetiidae" genus="Piroutetia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="liasina">
|
||
<emphasis id="8BED1F264B56FFABFF6CFB9D9575E5F5" box="[213,349,1144,1161]" italics="true" pageId="11" pageNumber="488">Piroutetia liasina</emphasis>
|
||
<bibRefCitation id="DD08BEC54B56FFABFEDAFB9D95F5E5F5" author="MEUNIER F." box="[355,477,1144,1161]" pageId="11" pageNumber="488" pagination="521 - 523" refId="ref14658" refString="MEUNIER F. 1907. - Un Odonatoptere du Rhetien (Lias inferieur) de Fort-Mouchard pres des Arcures (Jura). Bulletin du Museum d'Histoire naturelle 13: 521 - 523. https: // www. biodiversitylibrary. org / page / 42900382" type="journal article" year="1907">Meunier, 1907</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, specimen MNHN.F.B09711:
|
||
<emphasis id="8BED1F264B56FFABFC9EFB9D971CE5F5" bold="true" box="[807,820,1144,1161]" pageId="11" pageNumber="488">A</emphasis>
|
||
, drawing of venation, with tentative reconstruction of wing base;
|
||
<emphasis id="8BED1F264B56FFABFAEEFB9D914CE5F5" bold="true" box="[1367,1380,1144,1161]" pageId="11" pageNumber="488">B</emphasis>
|
||
,
|
||
<emphasis id="8BED1F264B56FFABFAD6FB9D9155E5F5" bold="true" box="[1391,1405,1144,1161]" pageId="11" pageNumber="488">C</emphasis>
|
||
, photographs extracted from RTI file;
|
||
<emphasis id="8BED1F264B56FFABFE2EFB6B958CE5E3" bold="true" box="[407,420,1166,1183]" pageId="11" pageNumber="488">B</emphasis>
|
||
, light-mirrored;
|
||
<emphasis id="8BED1F264B56FFABFD93FB6B9610E5E3" bold="true" box="[554,568,1166,1183]" pageId="11" pageNumber="488">C</emphasis>
|
||
, normals visualisation. Scale bar: 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="F18390BF4B56FFABFF3DFA91918BE753" pageId="11" pageNumber="488" type="discussion">
|
||
<paragraph id="B926C3344B56FFABFF3DFA9194C0E4F0" blockId="11.[130,777,1396,2028]" box="[132,232,1396,1422]" pageId="11" pageNumber="488">
|
||
<smallCapsWord id="BFC055E84B56FFABFF3DFA9194C0E4F0" baselines="1415,1415" box="[132,232,1396,1422]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="11" pageNumber="488">REMARKS</smallCapsWord>
|
||
</paragraph>
|
||
<paragraph id="B926C3344B56FFABFF3DFA7195B8E7B0" blockId="11.[130,777,1396,2028]" pageId="11" pageNumber="488">
|
||
It will be shown below that ‘
|
||
<taxonomicName id="7E99B8B74B56FFABFE77FA719625E4D1" authorityName="Handlirsch" authorityYear="1912" box="[462,525,1428,1453]" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFE77FA719625E4D1" box="[462,525,1428,1453]" italics="true" pageId="11" pageNumber="488">Reisia</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="7E99B8B74B56FFABFDA4FA769605E4B1" authority="Deregnaucourt, Wappler, Anderson and Bethoux, 2017" authorityName="Deregnaucourt, Wappler, Anderson and Bethoux" authorityYear="2017" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="rieki">
|
||
<emphasis id="8BED1F264B56FFABFDA4FA769666E4D1" box="[541,590,1427,1453]" italics="true" pageId="11" pageNumber="488">rieki</emphasis>
|
||
<bibRefCitation id="DD08BEC54B56FFABFDEEFA719605E4B1" author="DEREGNAUCOURT I. & WAPPLER T. & ANDERSON J. M. & BETHOUX O." pageId="11" pageNumber="488" pagination="613 - 618" refId="ref13601" refString="DEREGNAUCOURT I., WAPPLER T., ANDERSON J. M. & BETHOUX O. 2017. - A new triadotypid insect from the Late Triassic of South Africa. Acta Palaeontologica Polonica 62: 613 - 618. https: // doi. org / 10.4202 / app. 00345.2017" type="journal article" year="2017">Deregnaucourt, Wappler, Anderson and Béthoux, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="7E99B8B74B56FFABFF1AFA369626E491" authority="Riek, 1976" authorityName="Riek" authorityYear="1976" box="[163,526,1491,1517]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="biseriatus">
|
||
<emphasis id="8BED1F264B56FFABFF1AFA3695AEE491" box="[163,390,1491,1517]" italics="true" pageId="11" pageNumber="488">Triassologus biseriatus</emphasis>
|
||
<bibRefCitation id="DD08BEC54B56FFABFE29FA369626E491" author="RIEK E. F." box="[400,526,1491,1517]" pageId="11" pageNumber="488" pagination="791 - 820" refId="ref15208" refString="RIEK E. F. 1976. - A new collection of insects from the Upper Triassic of South Africa. Annals of the Natal Museum 22 (3): 791 - 820. https: // hdl. handle. net / 10520 / AJA 03040798 _ 608" type="journal article" year="1976">Riek, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
. This could imply that
|
||
<taxonomicName id="7E99B8B74B56FFABFF3FFA1694D7E771" authorityName="Riek" authorityYear="1976" box="[134,255,1523,1549]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFF3FFA1694D7E771" box="[134,255,1523,1549]" italics="true" pageId="11" pageNumber="488">Triassologus</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName id="7E99B8B74B56FFABFE48FA169606E770" authorityName="Handlirsch" authorityYear="1912" box="[497,558,1523,1548]" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFE48FA169606E770" box="[497,558,1523,1548]" italics="true" pageId="11" pageNumber="488">Reisia</emphasis>
|
||
</taxonomicName>
|
||
. However, new data derived from additional material of
|
||
<taxonomicName id="7E99B8B74B56FFABFE43F9F796FFE750" authorityName="Riek" authorityYear="1976" box="[506,727,1554,1580]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="biseriatus">
|
||
<emphasis id="8BED1F264B56FFABFE43F9F796FFE750" box="[506,727,1554,1580]" italics="true" pageId="11" pageNumber="488">Triassologus biseriatus</emphasis>
|
||
</taxonomicName>
|
||
(see below) and on
|
||
<taxonomicName id="7E99B8B74B56FFABFE9CF9D695BDE731" box="[293,405,1587,1613]" class="Insecta" family="Triadotypidae" genus="Reisia" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="8BED1F264B56FFABFE9CF9D6954AE730" box="[293,354,1587,1612]" italics="true" pageId="11" pageNumber="488">Reisia</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(see above) allows to convincingly differentiate the two genera, including the width of the area between the anterior wing margin (or ScA) and ScP, and the particular organisation of I- and CuA branches in the CuA-CuP area, among others.
|
||
</paragraph>
|
||
<paragraph id="B926C3344B56FFABFF22F937904CE432" blockId="11.[130,777,1396,2028]" lastBlockId="11.[812,1456,1236,1587]" pageId="11" pageNumber="488">
|
||
The more complete data on
|
||
<taxonomicName id="7E99B8B74B56FFABFE5FF93796FDE790" authorityName="Riek" authorityYear="1976" box="[486,725,1746,1772]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="biseriatus">
|
||
<emphasis id="8BED1F264B56FFABFE5FF93796FDE790" box="[486,725,1746,1772]" italics="true" pageId="11" pageNumber="488">Triassologus biseriatus</emphasis>
|
||
</taxonomicName>
|
||
(see below) led us to compare it with the Australian Triassic species ‘
|
||
<emphasis id="8BED1F264B56FFABFF01F8F794D7E657" box="[184,255,1810,1835]" italics="true" pageId="11" pageNumber="488">Iveryia</emphasis>
|
||
’
|
||
<taxonomicName id="7E99B8B74B56FFABFEB2F8F79676E650" authority="Bethoux & Beattie, 2010" authorityName="Bethoux & Beattie" authorityYear="2010" box="[267,606,1810,1836]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="aveyri">
|
||
<emphasis id="8BED1F264B56FFABFEB2F8F79560E657" box="[267,328,1810,1835]" italics="true" pageId="11" pageNumber="488">aveyri</emphasis>
|
||
<bibRefCitation id="DD08BEC54B56FFABFEF6F8F79676E650" author="BETHOUX O. & BEATTIE R. G." box="[335,606,1810,1836]" pageId="11" pageNumber="488" pagination="688 - 692" refId="ref13268" refString="BETHOUX O. & BEATTIE R. G. 2010. - Iverya averyi gen. nov. & sp. nov., a new triadotypomorphan species from the Middle Triassic at Picton, New South Wales, Australia. Acta Geologica Sinica 84: 688 - 692. https: // doi. org / 10.1111 / j. 1755 - 6724.2010.00240. x" type="journal article" year="2010">Béthoux & Beattie, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="66227D964B56FFABFDD5F8F796B4E650" box="[620,668,1810,1836]" pageId="11" pageNumber="488">type</typeStatus>
|
||
species of the genus
|
||
<taxonomicName id="7E99B8B74B56FFABFF40F8D79674E630" authority="Bethoux & Beattie, 2010" authorityName="Bethoux & Beattie" authorityYear="2010" box="[249,604,1841,1868]" class="Insecta" family="Triadotypidae" genus="Iverya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFF40F8D79511E637" box="[249,313,1842,1867]" italics="true" pageId="11" pageNumber="488">Iverya</emphasis>
|
||
<bibRefCitation id="DD08BEC54B56FFABFE86F8D49674E630" author="BETHOUX O. & BEATTIE R. G." box="[319,604,1841,1868]" pageId="11" pageNumber="488" pagination="688 - 692" refId="ref13268" refString="BETHOUX O. & BEATTIE R. G. 2010. - Iverya averyi gen. nov. & sp. nov., a new triadotypomorphan species from the Middle Triassic at Picton, New South Wales, Australia. Acta Geologica Sinica 84: 688 - 692. https: // doi. org / 10.1111 / j. 1755 - 6724.2010.00240. x" type="journal article" year="2010">Béthoux & Beattie, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
). Indeed, as far as comparison can be made, the two species do not display major differences other than size-related ones, such has the overall number of vein branches (assuming that the only known specimen of ‘
|
||
<taxonomicName id="7E99B8B74B56FFABFED0F8579580E6B7" authorityName="Bethoux & Beattie" authorityYear="2010" box="[361,424,1970,1995]" class="Insecta" family="Triadotypidae" genus="Iverya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFED0F8579580E6B7" box="[361,424,1970,1995]" italics="true" pageId="11" pageNumber="488">Iverya</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="7E99B8B74B56FFABFE0EF85795DCE6B7" authorityName="Bethoux & Beattie" authorityYear="2010" box="[439,500,1970,1995]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="aveyri">
|
||
<emphasis id="8BED1F264B56FFABFE0EF85795DCE6B7" box="[439,500,1970,1995]" italics="true" pageId="11" pageNumber="488">aveyri</emphasis>
|
||
</taxonomicName>
|
||
is a forewing, it is about 1.4 times larger than in that of
|
||
<taxonomicName id="7E99B8B74B56FFABFE51F83496E2E697" authorityName="Riek" authorityYear="1976" box="[488,714,2001,2027]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="biseriatus">
|
||
<emphasis id="8BED1F264B56FFABFE51F83496E2E697" box="[488,714,2001,2027]" italics="true" pageId="11" pageNumber="488">Triassologus biseriatus</emphasis>
|
||
</taxonomicName>
|
||
). We therefore propose to newly assign ‘
|
||
<taxonomicName id="7E99B8B74B56FFABFB19FB3190F7E591" authorityName="Bethoux & Beattie" authorityYear="2010" box="[1184,1247,1236,1261]" class="Insecta" family="Triadotypidae" genus="Iverya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFB19FB3190F7E591" box="[1184,1247,1236,1261]" italics="true" pageId="11" pageNumber="488">Iverya</emphasis>
|
||
</taxonomicName>
|
||
’
|
||
<taxonomicName id="7E99B8B74B56FFABFB55FB319101E591" authorityName="Bethoux & Beattie" authorityYear="2010" box="[1260,1321,1236,1261]" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="aveyri">
|
||
<emphasis id="8BED1F264B56FFABFB55FB319101E591" box="[1260,1321,1236,1261]" italics="true" pageId="11" pageNumber="488">aveyri</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="7E99B8B74B56FFABFAF4FB319765E471" authorityName="Riek" authorityYear="1976" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8BED1F264B56FFABFAF4FB319765E471" italics="true" pageId="11" pageNumber="488">Triassologus</emphasis>
|
||
</taxonomicName>
|
||
, further exemplifying similarities between South African and Australian Triassic insect faunas (
|
||
<bibRefCitation id="DD08BEC54B56FFABFB6BFAF69180E452" author="TIERNEY A. & DEREGNAUCOURT I. & ANDERSON J. M. & TIERNEY P. & WAPPLER T. & BETHOUX O." box="[1234,1448,1299,1326]" pageId="11" pageNumber="488" pagination="279 - 285" refId="ref15344" refString="TIERNEY A., DEREGNAUCOURT I., ANDERSON J. M., TIERNEY P., WAPPLER T. & BETHOUX O. 2020. - The Triassic Mesophlebiidae, a little closer to the crown of Odonata (Insecta) than other ' triassolestids'. Alcheringa 44: 279 - 285. https: // doi. org / 10.108 0 / 03115518.2020.1730964" type="journal article" year="2020">
|
||
Tierney
|
||
<emphasis id="8BED1F264B56FFABFA89FAF1914EE451" box="[1328,1382,1299,1325]" italics="true" pageId="11" pageNumber="488">et al.</emphasis>
|
||
2020
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="DD08BEC54B56FFABFC94FAD6907DE431" author="BETHOUX O. & ANDERSON J. M." box="[813,1109,1331,1357]" pageId="11" pageNumber="488" pagination="550 - 558" refId="ref13230" refString="BETHOUX O. & ANDERSON J. M. 2021. - The Polycytellidae viewed as Gondwanan Glosselytrodea. Palaeoentomology 4: 550 - 558. https: // doi. org / 10.11646 / palaeoentomology. 4.6.5" type="journal article" year="2021">Béthoux & Anderson 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B926C3344B56FFABFCFAFAB0918BE753" blockId="11.[812,1456,1236,1587]" pageId="11" pageNumber="488">
|
||
<bibRefCitation id="DD08BEC54B56FFABFCFAFAB097F6E413" author="BECHLY G." box="[835,990,1365,1391]" pageId="11" pageNumber="488" pagination="31 - 70" refId="ref13178" refString="BECHLY G. 1997. - New fossil odonates from the Upper Triassic of Italy, with a redescription of Italophlebia gervasuttii, and a reclassification of Triassic dragonflies (Insecta: Odonata). Rivista del Museo civico di Scienze Naturali " Enrico Caffi " 19: 31 - 70." type="journal article" year="1997">Bechly (1997)</bibRefCitation>
|
||
considered that RP2 is branched in
|
||
<taxonomicName id="7E99B8B74B56FFABFACDFAB397F3E4F3" authorityName="Riek" authorityYear="1976" class="Insecta" genus="Triassologus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="11" pageNumber="488" phylum="Arthropoda" rank="species" species="biseriatus">
|
||
<emphasis id="8BED1F264B56FFABFACDFAB397F3E4F3" italics="true" pageId="11" pageNumber="488">Triassologus biseriatus</emphasis>
|
||
</taxonomicName>
|
||
. However, bearing in mind the propensity of intercalary veins to acquire a main vein habitus points to
|
||
<taxonomicName id="7E99B8B74B56FFABFC94FA5097A5E4B3" authorityName="Fabricius, 1793 DISCOIDALIA Bechly, 1996 TRIADOPHLEBIOMORPHA Pritykina" authorityYear="1981" box="[813,909,1461,1487]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Odonata" pageId="11" pageNumber="488" phylum="Tracheophyta" rank="order">Odonata</taxonomicName>
|
||
in general and Triadophlebiomorpha in particular, an alternative option, favoured herein, is that the apparent anterior branch of RP2 (large blue arrow on Fig. 4B-D, F) instead is the first I- in the area between Irp
|
||
<subScript id="251DC1714B56FFABFABBF9C79124E74F" attach="left" box="[1282,1292,1570,1587]" fontSize="7" pageId="11" pageNumber="488">1</subScript>
|
||
-rp
|
||
<subScript id="251DC1714B56FFABFA95F9C7911EE74F" attach="left" box="[1324,1334,1570,1587]" fontSize="7" pageId="11" pageNumber="488">2</subScript>
|
||
and RP2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |