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<document id="356865FC67272A3EE82CEE04FE30B023" ID-CLB-Dataset="270324" ID-DOI="10.5252/geodiversitas2023v45a17" ID-GBIF-Dataset="96784ccf-a749-4494-bfb4-f514eea7e824" ID-ISSN="1638-9395" ID-Zenodo-Dep="8399030" ID-ZooBank="urn:lsid:zoobank.org:pub:EE7AA253-A023-4011-B7E1-5820F8118ED3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1695890445085" checkinUser="plazi" docAuthor="Béthoux, Olivier &amp; Anderson, John M." docDate="2023" docId="313072224B53FFB1FC3AF97394D7E573" docLanguage="en" docName="geodiversitas.2023.45.17.pdf" docOrigin="Geodiversitas 45 (17)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Nototriadophlebia pritykinae Béthoux &amp; Anderson 2023, n. gen., n. sp." docType="treatment" docVersion="3" lastPageNumber="494" masterDocId="CD090A5A4B5DFFA0FFB9FFE59428E17C" masterDocTitle="New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata)" masterLastPageNumber="496" masterPageNumber="479" pageNumber="491" updateTime="1696249958766" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="48A729D438CCF57C9E124833C328237F">New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata)</mods:title>
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<mods:namePart id="3A9F12659B9F2E72846AEE4F826356A3">Béthoux, Olivier</mods:namePart>
<mods:affiliation id="3A78F76B5F6068160D23C21CC403C09F">CR 2 P (CNRS, MNHN, Sorbonne Université), Muséum national dHistoire naturelle, case postale 48, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) obethoux @ mnhn. fr</mods:affiliation>
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<mods:affiliation id="71E829E2845C91119B0652E3DEE0BDFC">Evolutionary Studies Institute, Witwatersrand University, 1 Jan Smuts Ave., Braamfontein, Johannesburg 2000, South Africa jmanderson. gondwana @ gmail. com</mods:affiliation>
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<emphasis id="8BED1F264B53FFAEFC3AF9739097E7CC" bold="true" box="[899,1215,1686,1712]" italics="true" pageId="14" pageNumber="491">Nototriadophlebia pritykinae</emphasis>
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(
<figureCitation id="21A2DFB14B53FFAEFBFBF95090B1E7B3" box="[1090,1177,1717,1743]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="14.[813,824,830,847]" captionStartId-1="15.[132,143,1917,1934]" captionStartId-2="16.[132,143,886,903]" captionTargetBox-0="[813,1456,215,787]" captionTargetBox-1="[155,1401,1611,1807]" captionTargetBox-2="[308,1280,215,844]" captionTargetId-0="figure-641@14.[813,1456,215,787]" captionTargetId-1="graphics-19@15.[152,1396,1608,1795]" captionTargetId-2="figure-636@16.[308,1280,450,844]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionText-0="FIG. 7. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a (preserving the right forewing, isolated, in ventral aspect, and other wings,overlapping,in dorsal aspect),photograph (dry).Scale bar:10 mm." captionText-1="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." captionText-2="FIG. 9. — Nototriadophlebia pritykinae n. gen., n. sp., paratype specimen, BP/2/20986b, left wing (large red arrows indicating the CuP+AA stem): A, B, overview; A, drawing of venation; B, photograph (light-mirrored); C, D, detail of cubito-anal area, location as indicated on B (large red arrows, CuP+AA stem); C, drawing of venation; D, photograph (light-mirrored). Scale bars: A, B, 10 mm; C, D, 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8399048" figureDoi-1="http://doi.org/10.5281/zenodo.8399054" figureDoi-2="http://doi.org/10.5281/zenodo.8399056" httpUri-0="https://zenodo.org/record/8399048/files/figure.png" httpUri-1="https://zenodo.org/record/8399054/files/figure.png" httpUri-2="https://zenodo.org/record/8399056/files/figure.png" pageId="14" pageNumber="491">Figs 7-9</figureCitation>
)
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. —
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/2/20950ab, adpression preserving four wings (
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collection,
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,
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,
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of the
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)
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.
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RA RP1
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-rp
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<paragraph id="B926C3344B52FFAFFD8BF9DD94B4E727" blockId="15.[140,1453,1368,1815]" pageId="15" pageNumber="492">ScP F</paragraph>
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. 8. —
<taxonomicName id="7E99B8B74B52FFAFFF6AF8989595E6F2" authority="Béthoux &amp; Anderson, 2023" authorityName="Béthoux &amp; Anderson" authorityYear="2023" box="[211,445,1917,1934]" class="Insecta" family="Permophlebiidae" genus="Nototriadophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="492" phylum="Arthropoda" rank="species" species="pritykinae" status="n. gen., n. sp.">
<emphasis id="8BED1F264B52FFAFFF6AF8989595E6F2" box="[211,445,1917,1934]" italics="true" pageId="15" pageNumber="492">Nototriadophlebia pritykinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DEA25D4B52FFAFFE7BF898961FE6F2" box="[450,567,1917,1934]" pageId="15" pageNumber="492" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, holotype, specimen BP/2/20950a:
<emphasis id="8BED1F264B52FFAFFCE4F8989742E6F2" bold="true" box="[861,874,1917,1934]" pageId="15" pageNumber="492">A</emphasis>
,
<emphasis id="8BED1F264B52FFAFFCCDF89897A9E6F2" bold="true" box="[884,897,1917,1934]" pageId="15" pageNumber="492">B</emphasis>
, right forewing (
<emphasis id="8BED1F264B52FFAFFBBEF8989019E6F2" bold="true" box="[1031,1073,1917,1934]" pageId="15" pageNumber="492">RFW</emphasis>
);
<emphasis id="8BED1F264B52FFAFFBF9F8989065E6F2" bold="true" box="[1088,1101,1917,1934]" pageId="15" pageNumber="492">A</emphasis>
, drawing of venation, with tentative reconstruction of wing base;
<emphasis id="8BED1F264B52FFAFFEF0F871957EE6D9" bold="true" box="[329,342,1940,1957]" pageId="15" pageNumber="492">B</emphasis>
, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally);
<emphasis id="8BED1F264B52FFAFFB16F8719095E6D9" bold="true" box="[1199,1213,1940,1957]" pageId="15" pageNumber="492">C</emphasis>
,
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, overlapping left forewing (
<emphasis id="8BED1F264B52FFAFFF30F84E9499E6C0" bold="true" box="[137,177,1963,1980]" pageId="15" pageNumber="492">LFW</emphasis>
), right hind wing (
<emphasis id="8BED1F264B52FFAFFEF3F84E955FE6C0" bold="true" box="[330,375,1963,1980]" pageId="15" pageNumber="492">RHW</emphasis>
) and left hind wing (
<emphasis id="8BED1F264B52FFAFFD9DF84E9667E6C0" bold="true" box="[548,591,1963,1980]" pageId="15" pageNumber="492">LHW</emphasis>
;
<emphasis id="8BED1F264B52FFAFFDE3F84E96BEE6C0" bold="true" box="[602,662,1963,1980]" pageId="15" pageNumber="492">in blue</emphasis>
);
<emphasis id="8BED1F264B52FFAFFD1FF84E969CE6C0" bold="true" box="[678,692,1963,1980]" pageId="15" pageNumber="492">C</emphasis>
, drawing of venation;
<emphasis id="8BED1F264B52FFAFFCC8F84E9757E6C0" bold="true" box="[881,895,1963,1980]" pageId="15" pageNumber="492">D</emphasis>
, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally);
<emphasis id="8BED1F264B52FFAFFE30F82795BDE6AF" bold="true" box="[393,405,1986,2003]" pageId="15" pageNumber="492">E</emphasis>
, left forewing (
<emphasis id="8BED1F264B52FFAFFDA9F8279610E6AF" bold="true" box="[528,568,1986,2003]" pageId="15" pageNumber="492">LFW</emphasis>
), drawing of venation isolated from
<emphasis id="8BED1F264B52FFAFFCDBF8279758E6AF" bold="true" box="[866,880,1986,2003]" pageId="15" pageNumber="492">C</emphasis>
, with tentative reconstruction of the posterior area;
<emphasis id="8BED1F264B52FFAFFAA6F8279102E6AF" bold="true" box="[1311,1322,1986,2003]" pageId="15" pageNumber="492">F</emphasis>
, right hind wing (
<emphasis id="8BED1F264B52FFAFFF30F83D949EE695" bold="true" box="[137,182,2008,2025]" pageId="15" pageNumber="492">RHW</emphasis>
), drawing of venation isolated from
<emphasis id="8BED1F264B52FFAFFE58F83D95C7E695" bold="true" box="[481,495,2008,2025]" pageId="15" pageNumber="492">C</emphasis>
(large red arrow, CuP+AA stem). Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="EDE693BC4B4DFFB0FF3DFC939701E2C8" ID-DOI="http://doi.org/10.5281/zenodo.8399056" ID-Zenodo-Dep="8399056" httpUri="https://zenodo.org/record/8399056/files/figure.png" pageId="16" pageNumber="493" startId="16.[132,143,886,903]" targetBox="[308,1280,215,844]" targetPageId="16" targetType="figure">
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. 9. —
<taxonomicName id="7E99B8B74B4DFFB0FF6AFC939595E2FB" authority="Béthoux &amp; Anderson, 2023" authorityName="Béthoux &amp; Anderson" authorityYear="2023" box="[211,445,886,903]" class="Insecta" family="Permophlebiidae" genus="Nototriadophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="16" pageNumber="493" phylum="Arthropoda" rank="species" species="pritykinae" status="n. gen., n. sp.">
<emphasis id="8BED1F264B4DFFB0FF6AFC939595E2FB" box="[211,445,886,903]" italics="true" pageId="16" pageNumber="493">Nototriadophlebia pritykinae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DEA25D4B4DFFB0FE7BFC93961EE2FB" box="[450,566,886,903]" pageId="16" pageNumber="493" rank="species">n. gen., n. sp.</taxonomicNameLabel>
, paratype specimen, BP/2/20986b, left wing (large red arrows indicating the CuP+AA stem):
<emphasis id="8BED1F264B4DFFB0FA8BFC939117E2FB" bold="true" box="[1330,1343,886,903]" pageId="16" pageNumber="493">A</emphasis>
,
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, overview;
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, drawing of venation;
<emphasis id="8BED1F264B4DFFB0FEF2FC689570E2E2" bold="true" box="[331,344,909,926]" pageId="16" pageNumber="493">B</emphasis>
, photograph (light-mirrored);
<emphasis id="8BED1F264B4DFFB0FDF7FC689674E2E2" bold="true" box="[590,604,909,926]" pageId="16" pageNumber="493">C</emphasis>
,
<emphasis id="8BED1F264B4DFFB0FDDFFC68965CE2E2" bold="true" box="[614,628,909,926]" pageId="16" pageNumber="493">D</emphasis>
, detail of cubito-anal area, location as indicated on
<emphasis id="8BED1F264B4DFFB0FB9CFC68901AE2E2" bold="true" box="[1061,1074,909,926]" pageId="16" pageNumber="493">B</emphasis>
(large red arrows, CuP+AA stem);
<emphasis id="8BED1F264B4DFFB0FAEDFC68914AE2E2" bold="true" box="[1364,1378,909,926]" pageId="16" pageNumber="493">C</emphasis>
, drawing of venation;
<emphasis id="8BED1F264B4DFFB0FF52FC4694D1E2C8" bold="true" box="[235,249,931,948]" pageId="16" pageNumber="493">D</emphasis>
, photograph (light-mirrored). Scale bars: A, B, 10 mm; C, D, 2 mm.
</paragraph>
</caption>
<subSubSection id="F18390BF4B4DFFB0FF3DFC1196B3E5A3" pageId="16" pageNumber="493" type="diagnosis">
<paragraph id="B926C3344B4DFFB0FF3DFC1196B3E5A3" blockId="16.[132,776,1011,1247]" pageId="16" pageNumber="493">
<smallCapsWord id="BFC055E84B4DFFB0FF3DFC1194C7E576" baselines="1029,1030" box="[132,239,1012,1035]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="16" pageNumber="493">DIAGNOSIS</smallCapsWord>
. — Area between the anterior wing margin and RA (distal to the nodus) widening gradually to ª/5 of wing length, where it is as wide as the RA-RP1 area (most prominent in forewing); first I- in the RP3+4-MA area (i.e. RP3+4 pseudo-fork; large orange arrows on
<figureCitation id="21A2DFB14B4DFFB0FF55FBB89578E509" box="[236,336,1117,1142]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="16" pageNumber="493">Fig. 8A, F</figureCitation>
) arising before RP3+4 mid-length; CuP+AA stem splitting into CuA and CuP+AA (as opposed to Cu and AA; large red arrow on
<figureCitation id="21A2DFB14B4DFFB0FEFAFB7795A4E5D6" box="[323,396,1170,1194]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="16" pageNumber="493">Figs 8F</figureCitation>
,
<figureCitation id="21A2DFB14B4DFFB0FE2DFB7795DBE5D6" box="[404,499,1170,1194]" captionStart="FIG" captionStartId="16.[132,143,886,903]" captionTargetBox="[308,1280,215,844]" captionTargetId="figure-636@16.[308,1280,450,844]" captionTargetPageId="16" captionText="FIG. 9. — Nototriadophlebia pritykinae n. gen., n. sp., paratype specimen, BP/2/20986b, left wing (large red arrows indicating the CuP+AA stem): A, B, overview; A, drawing of venation; B, photograph (light-mirrored); C, D, detail of cubito-anal area, location as indicated on B (large red arrows, CuP+AA stem); C, drawing of venation; D, photograph (light-mirrored). Scale bars: A, B, 10 mm; C, D, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8399056" httpUri="https://zenodo.org/record/8399056/files/figure.png" pageId="16" pageNumber="493">9A, C, D</figureCitation>
); AA reaching the posterior wing margin basal to the end of ScP; between the endings of MA and AA, posterior wing margin comparatively straight.
</paragraph>
</subSubSection>
<subSubSection id="F18390BF4B4DFFB0FF3DFB1295C9E438" pageId="16" pageNumber="493" type="etymology">
<paragraph id="B926C3344B4DFFB0FF3DFB1295C9E438" blockId="16.[131,775,1271,1348]" pageId="16" pageNumber="493">
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. — The species epithet honours Lyudmila N. Pritykina, for her contribution to our knowledge of fossil
<taxonomicName id="7E99B8B74B4DFFB0FDD9FAF49692E455" authorityName="Fabricius, 1793 DISCOIDALIA Bechly, 1996 TRIADOPHLEBIOMORPHA Pritykina" authorityYear="1981" box="[608,698,1297,1321]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Odonata" pageId="16" pageNumber="493" phylum="Tracheophyta" rank="order">Odonata</taxonomicName>
, and of Triadophlebiomorpha in particular.
</paragraph>
</subSubSection>
<subSubSection id="F18390BF4B4DFFB0FF3DFAB9903FE610" pageId="16" pageNumber="493" type="materials_examined">
<paragraph id="B926C3344B4DFFB0FF3DFAB995D0E4BE" blockId="16.[131,775,1371,1475]" pageId="16" pageNumber="493">
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<collectionCode id="DF885BF14B4DFFB0FF3DFAB9949DE40D" box="[132,181,1372,1395]" pageId="16" pageNumber="493">TYPE</collectionCode>
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<collectionCode id="DF885BF14B4DFFB0FF03FA84953DE40E" box="[186,277,1377,1394]" pageId="16" pageNumber="493">LOCALITY</collectionCode>
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<collectionCode id="DF885BF14B4DFFB0FEA0FA84956AE40E" box="[281,322,1377,1394]" pageId="16" pageNumber="493">AND</collectionCode>
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<collectionCode id="DF885BF14B4DFFB0FEFEFA8495FAE40E" box="[327,466,1377,1394]" pageId="16" pageNumber="493">STRATIGRAPHY</collectionCode>
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. — Both
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and
<typeStatus id="66227D964B4DFFB0FD0DFAB9972FE408" box="[692,775,1372,1396]" pageId="16" pageNumber="493">paratype</typeStatus>
were collected at
<location id="BC4695EF4B4DFFB0FE9BFA9395A3E4F2" LSID="urn:lsid:plazi:treatment:313072224B53FFB1FC3AF97394D7E573:BC4695EF4B4DFFB0FE9BFA9395A3E4F2" box="[290,395,1398,1422]" country="South Africa" name="Birds River" pageId="16" pageNumber="493">Birds River</location>
(locality code Bir 111; see
<bibRefCitation id="DD08BEC54B4DFFB0FD36FA939531E4D4" author="ANDERSON J. M. &amp; ANDERSON H. M." pageId="16" pageNumber="493" pagination="39 - 59" refId="ref13024" refString="ANDERSON J. M. &amp; ANDERSON H. M. 1984. - The fossil content of the Upper Triassic Molteno Formation, South Africa. Palaeontologia Africana 25: 39 - 59. http: // hdl. handle. net / 10539 / 16124" type="journal article" year="1984">Anderson &amp; Anderson 1984</bibRefCitation>
),
<collectingCountry id="C18E83A44B4DFFB0FE92FA75958FE4D4" box="[299,423,1424,1448]" name="South Africa" pageId="16" pageNumber="493">South Africa</collectingCountry>
; Molteno Formation;
<location id="BC4695EF4B4DFFB0FDC1FA75972CE4D4" LSID="urn:lsid:plazi:treatment:313072224B53FFB1FC3AF97394D7E573:BC4695EF4B4DFFB0FDC1FA75972CE4D4" box="[632,772,1424,1448]" country="South Africa" municipality="Upper Triassic" name="lower Carnian" pageId="16" pageNumber="493">lower Carnian</location>
,
<collectingMunicipality id="5942594E4B4DFFB0FF3AFA4E9526E4BF" box="[131,270,1451,1475]" pageId="16" pageNumber="493">Upper Triassic</collectingMunicipality>
(
<bibRefCitation id="DD08BEC54B4DFFB0FEA7FA4E95C3E4BF" author="ANDERSON J. M. &amp; ANDERSON H. M. &amp; CRUICKSHANK A. R. I." box="[286,491,1451,1475]" pageId="16" pageNumber="493" pagination="387 - 421" refId="ref13071" refString="ANDERSON J. M., ANDERSON H. M. &amp; CRUICKSHANK A. R. I. 1998. - Late Triassic ecosystems of the Molteno / Lower Elliot Biome of Southern Africa. Palaeontology 41: 387 - 421. https: // www. biodiversitylibrary. org / page / 49739480" type="journal article" year="1998">
Anderson
<emphasis id="8BED1F264B4DFFB0FE3BFA4E9599E4BF" box="[386,433,1451,1475]" italics="true" pageId="16" pageNumber="493">et al.</emphasis>
1998
</bibRefCitation>
).
</paragraph>
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</paragraph>
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Fore- and hindwings subequal, very elongated, broadest at mid-length; wing length about
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(as derived from
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specimen; possibly down to
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), maximum width about 11.0 mm (possibly down to
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), width opposite nodus
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); petiole long (about
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) and narrow (width about
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); ScP reaching the anterior wing margin basal to the first third of wing length; distal to the nodus, area between anterior wing margin and RA broadening gradually, at its broadest opposite the end of Irp
<subScript id="251DC1714B4DFFB0FDBAF8FA9608E64C" attach="left" box="[515,544,1823,1840]" fontSize="7" pageId="16" pageNumber="493">1+2</subScript>
-rp
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; structure of the nodus and origin of RP+MA, RP and MA unknown; RP1/ RP2 fork in a position more distal than that of the point of origin of the first I- in the RP3+4 area; Irp
<subScript id="251DC1714B4DFFB0FDEDF89B9676E6F3" attach="left" box="[596,606,1918,1935]" fontSize="7" pageId="16" pageNumber="493">1</subScript>
-rp
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weakly differentiated, zigzagging along RP1; occurrence of 3 I- (and I+ between them) forming RP2 posterior pseudo-branches in the distal part of the area between RP2 and Irp
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-rp
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; RP3+4 with a marked angle opposite the origin of the first I- (between RP3+4 and MA; large orange arrows on
<figureCitation id="21A2DFB14B4DFFB0FB64FBF19163E552" box="[1245,1355,1044,1071]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="16" pageNumber="493">Fig. 8A, F</figureCitation>
); the two first I- in the area between RP3+4 and MA branched; in this area, total of about 8 I- reaching the posterior wing margin; MA simple, rectilinear, very close to RP3+
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its basal third; marked inflexion of the posterior wing margin opposite the ending of MA;
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originating from
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+Cu+AA with a slight obliquity; branches of
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and CuA not clearly distinguishable (probable occurrence of a CuAprlg), CuA with 2 branches at least, and with a I- distally branched between its two (first posterior) branches;
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and CuA covering more than a quarter (less than a third) of the posterior wing margin length; Cu+AA splitting into CuA and CuP+AA, CuA diverging very obliquely; CuP rectilinear, with about 4-5 posterior branches and I+ between them; AA gently curved, reaching the posterior wing margin basal to the end of ScP.
</paragraph>
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</paragraph>
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Specimen BP/2/20950 (
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;
<figureCitation id="21A2DFB14B4DFFB0FBC8F9D69057E731" box="[1137,1151,1587,1613]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="16" pageNumber="493">8</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="B926C3344B4DFFB0FC94F9B79187E7B1" blockId="16.[811,1457,1555,1741]" pageId="16" pageNumber="493">Adpression preserving four wings, with side BP/2/20950a preserving the right forewing (isolated) and left wings in ventral aspect, and right hindwing in dorsal aspect (the three latter largely overlapping); and its reverse side BP/2/20950b.</paragraph>
<paragraph id="B926C3344B4DFFB0FC94F914903FE610" blockId="16.[811,1457,1777,1900]" pageId="16" pageNumber="493">
<emphasis id="8BED1F264B4DFFB0FC94F9149058E677" bold="true" box="[813,1136,1777,1804]" pageId="16" pageNumber="493">
Right forewing (
<figureCitation id="21A2DFB14B4DFFB0FC62F9149031E670" box="[987,1049,1777,1804]" captionStart="FIG" captionStartId="14.[813,824,830,847]" captionTargetBox="[813,1456,215,787]" captionTargetId="figure-641@14.[813,1456,215,787]" captionTargetPageId="14" captionText="FIG. 7. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a (preserving the right forewing, isolated, in ventral aspect, and other wings,overlapping,in dorsal aspect),photograph (dry).Scale bar:10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399048" httpUri="https://zenodo.org/record/8399048/files/figure.png" pageId="16" pageNumber="493">Figs 7</figureCitation>
;
<figureCitation id="21A2DFB14B4DFFB0FB9CF917904BE677" box="[1061,1123,1777,1804]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="16" pageNumber="493">8A, B</figureCitation>
).
</emphasis>
Well preserved, with area basal to the nodus and small portion of apex missing; preserved length
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, maximum width
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, width opposite nodus about
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.
</paragraph>
</subSubSection>
<subSubSection id="F18390BF4B4DFFB1FC94F87494D7E573" lastPageId="17" lastPageNumber="494" pageId="16" pageNumber="493" type="description">
<paragraph id="B926C3344B4DFFB0FC94F87497F6E690" blockId="16.[813,1455,1937,2028]" pageId="16" pageNumber="493">
<emphasis id="8BED1F264B4DFFB0FC94F8749042E6D7" bold="true" box="[813,1130,1937,1963]" pageId="16" pageNumber="493">
Left forewing (
<figureCitation id="21A2DFB14B4DFFB0FC69F8749039E6D7" box="[976,1041,1937,1963]" captionStart="FIG" captionStartId="14.[813,824,830,847]" captionTargetBox="[813,1456,215,787]" captionTargetId="figure-641@14.[813,1456,215,787]" captionTargetPageId="14" captionText="FIG. 7. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a (preserving the right forewing, isolated, in ventral aspect, and other wings,overlapping,in dorsal aspect),photograph (dry).Scale bar:10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399048" httpUri="https://zenodo.org/record/8399048/files/figure.png" pageId="16" pageNumber="493">Figs 7</figureCitation>
;
<figureCitation id="21A2DFB14B4DFFB0FBA6F8749005E6D7" box="[1055,1069,1937,1963]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="16" pageNumber="493">8</figureCitation>
C-E).
</emphasis>
Only anterior-most area decipherable; preserved length
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; estimated total length about
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.
</paragraph>
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. 10. — Detail of the cubito-anal area in
<taxonomicName id="7E99B8B74B4CFFB1FE58FD82972CE304" authority="Pritykina, 1981" authorityName="Pritykina" authorityYear="1981" box="[481,772,615,632]" class="Insecta" genus="Neritophlebia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="494" phylum="Arthropoda" rank="species" species="longa">
<emphasis id="8BED1F264B4CFFB1FE58FD8296ABE304" box="[481,643,615,632]" italics="true" pageId="17" pageNumber="494">Neritophlebia longa</emphasis>
<bibRefCitation id="DD08BEC54B4CFFB1FD3EFD82972CE304" author="PRITYKINA L. N." box="[647,772,615,632]" pageId="17" pageNumber="494" pagination="5 - 42" refId="ref15101" refString="PRITYKINA L. N. 1981. - Novye triasovye strekozy srednej Azii, in VISHNIAKOVA V. N., DLUSSKY G. M. &amp; PRITYKINA L. N. (eds), Novye iskopaemye nasekomye s Terrotorii SSSR. Trudy Paleontologicheskogo instituta, Akademiya Nauk SSSR, Moscow: 5 - 42." type="book chapter" year="1981">Pritykina,1981</bibRefCitation>
</taxonomicName>
, holotype, specimen PIN 2555/614, left wing (large red arrow, CuP+AA stem). Scale bar: 4 mm. Courtesy A. S. Felker.
</paragraph>
</caption>
<paragraph id="B926C3344B4CFFB1FF3DFD10962AE2F3" blockId="17.[130,775,757,911]" pageId="17" pageNumber="494">
<emphasis id="8BED1F264B4CFFB1FF3DFD1095D8E273" bold="true" box="[132,496,757,783]" pageId="17" pageNumber="494">
Right hindwing (
<figureCitation id="21A2DFB14B4CFFB1FE83FD109550E273" box="[314,376,757,783]" captionStart="FIG" captionStartId="14.[813,824,830,847]" captionTargetBox="[813,1456,215,787]" captionTargetId="figure-641@14.[813,1456,215,787]" captionTargetPageId="14" captionText="FIG. 7. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a (preserving the right forewing, isolated, in ventral aspect, and other wings,overlapping,in dorsal aspect),photograph (dry).Scale bar:10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399048" httpUri="https://zenodo.org/record/8399048/files/figure.png" pageId="17" pageNumber="494">Figs 7</figureCitation>
;
<figureCitation id="21A2DFB14B4CFFB1FE3DFD1095CBE273" box="[388,483,757,783]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="17" pageNumber="494">8C, D, F</figureCitation>
).
</emphasis>
Well preserved, with nodal area and part of apex missing, and venation in the petiole difficult to decipher; basal to the nodus, RP and MA not distinguishable from each other (assumed to closely overlap); cubito-anal area preserved, showing a CuP+AA stem.
</paragraph>
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<emphasis id="8BED1F264B4CFFB1FF3DFC5195F2E2B2" bold="true" box="[132,474,948,975]" pageId="17" pageNumber="494">
Left hindwing (
<figureCitation id="21A2DFB14B4CFFB1FE8BFC51955CE2B3" box="[306,372,948,975]" captionStart="FIG" captionStartId="14.[813,824,830,847]" captionTargetBox="[813,1456,215,787]" captionTargetId="figure-641@14.[813,1456,215,787]" captionTargetPageId="14" captionText="FIG. 7. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a (preserving the right forewing, isolated, in ventral aspect, and other wings,overlapping,in dorsal aspect),photograph (dry).Scale bar:10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399048" httpUri="https://zenodo.org/record/8399048/files/figure.png" pageId="17" pageNumber="494">Figs 7</figureCitation>
;
<figureCitation id="21A2DFB14B4CFFB1FE3DFC5195E3E2B2" box="[388,459,948,974]" captionStart="FIG" captionStartId="15.[132,143,1917,1934]" captionTargetBox="[155,1401,1611,1807]" captionTargetId="graphics-19@15.[152,1396,1608,1795]" captionTargetPageId="15" captionText="FIG. 8. — Nototriadophlebia pritykinae n. gen., n. sp., holotype, specimen BP/2/20950a: A, B, right forewing (RFW); A, drawing of venation, with tentative reconstruction of wing base; B, photograph, location as shown on Fig. 7 (dry-ethanol composite, light-mirrored, flipped horizontally); C, D, overlapping left forewing (LFW), right hind wing (RHW) and left hind wing (LHW; in blue); C, drawing of venation; D, photograph, location as shown on Fig. 7 (dry-ethanol composite, fragments readjusted digitally); E, left forewing (LFW), drawing of venation isolated from C, with tentative reconstruction of the posterior area; F, right hind wing (RHW), drawing of venation isolated from C (large red arrow, CuP+AA stem). Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.8399054" httpUri="https://zenodo.org/record/8399054/files/figure.png" pageId="17" pageNumber="494">8C, D</figureCitation>
).
</emphasis>
Barely decipherable but for the posterior wing margin, with a sharp inflexion opposite the end of MA.
</paragraph>
</subSubSection>
</treatment>
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