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<document id="0D407669641A4C1145F174269F247EE5" ID-DOI="10.11646/zootaxa.3869.5.2" ID-GBIF-Dataset="72c3b4dc-783c-47f7-8bf0-6097f2eb2189" ID-ISSN="1175-5326" ID-Zenodo-Dep="251217" ID-ZooBank="45D2A66A-3A45-4A1E-BF65-DC9755295505" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460925384811" checkinUser="plazi" docAuthor="Oliveira, Danilo E., Cunha, Hélida F. &amp; Constantino, Reginaldo" docDate="2014" docId="2A658955C35EE753FF53F899FB5BFF52" docLanguage="en" docName="zt03869p536.pdf" docOrigin="Zootaxa 3869 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anhangatermes anhanguera Oliveira" docType="treatment" docVersion="7" lastPageNumber="527" masterDocId="D65CF12DC35CE756FFC4FFB6FF91FF87" masterDocTitle="A taxonomic revision of the soil-feeding termite genus Anhangatermes (Isoptera: Termitidae: Nasutitermitinae)" masterLastPageNumber="536" masterPageNumber="523" pageNumber="525" updateTime="1698362381006" updateUser="plazi">
<mods:mods id="82239AA486B48BA6A622C37FDF106748" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BED49F3F254675947AC635FC8E3350FE">
<mods:title id="837FB36D7E10CD5AC26B4260D75D9CB2">A taxonomic revision of the soil-feeding termite genus Anhangatermes (Isoptera: Termitidae: Nasutitermitinae)</mods:title>
</mods:titleInfo>
<mods:name id="7EBB6096BD97E1650B2F17C6F37D38C0" type="personal">
<mods:role id="EB8FB762054AEF8CC1EB7BB10BA2D3B7">
<mods:roleTerm id="59159DEF0B5F09EF3CDEB6E003DC2DC9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F650DB05075ED3E0DE548697D940720">Oliveira, Danilo E.</mods:namePart>
</mods:name>
<mods:name id="F47D262C6D4ABDDAF07B25562E6236A6" type="personal">
<mods:role id="B12DAE0A3A0F0FA5193503679B728732">
<mods:roleTerm id="1B349818781E92700587ADC164F1132F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17126648362D3B552BE8AAA8902DF928">Cunha, Hélida F.</mods:namePart>
</mods:name>
<mods:name id="393BD0D35E2CC97A15BE8982AA3F6DD3" type="personal">
<mods:role id="EBBBEBF2BB6F13047DA008CE3089628B">
<mods:roleTerm id="C26E22D54EEE9D3505C7E3C8674E3144">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B559C2E4F691A0FDFBD57FE2AA2D0176">Constantino, Reginaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="ADCA7E6243E6C37B683AE2534D54E387">text</mods:typeOfResource>
<mods:relatedItem id="F04F41EEDDC4783258E44E5C59A4F2B1" type="host">
<mods:titleInfo id="72B0F495EFA038FDEFFFAD04C91F2769">
<mods:title id="1A653E6FF8AB7DEF40F467726C5DE6C1">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="354D3DDFC3F3A5B56EF5AD244CE5C55A">
<mods:date id="3B952807721ACD35D1AF37A3339566CD">2014</mods:date>
<mods:detail id="0F0F2D114474EB691E007D7DF58DDCA9" type="volume">
<mods:number id="796E9D3D487F64BBB1CFEE954D376A6F">3869</mods:number>
</mods:detail>
<mods:detail id="A02D37EF1A1CAF642B55442D50681E24" type="issue">
<mods:number id="8204C22DB200908D657D272207B33F67">5</mods:number>
</mods:detail>
<mods:extent id="57467E399D24F041CC2B6F4246643120" unit="page">
<mods:start id="DB7AE24009554054BC8A06BFA5C80434">523</mods:start>
<mods:end id="1C13F849CA0FF8EA4EBA0D1A71ACA3DD">536</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="651B25E1D84BBB3B77742D156740368A">journal article</mods:classification>
<mods:identifier id="4A2CB628D6F041B0E988398062C1085B" type="DOI">10.11646/zootaxa.3869.5.2</mods:identifier>
<mods:identifier id="9710653BD504628780420CEB7BA116B6" type="GBIF-Dataset">72c3b4dc-783c-47f7-8bf0-6097f2eb2189</mods:identifier>
<mods:identifier id="28E3582BDDDF10904518589B9A061E80" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="4D9B60F3E6E673BF543A874C43D6E0E0" type="Zenodo-Dep">251217</mods:identifier>
<mods:identifier id="4D322A1751A3E990FCDE18913ABB2CB9" type="ZooBank">45D2A66A-3A45-4A1E-BF65-DC9755295505</mods:identifier>
</mods:mods>
<treatment id="2A658955C35EE753FF53F899FB5BFF52" ID-DOI="http://doi.org/10.5281/zenodo.6142859" ID-GBIF-Taxon="119589492" ID-Zenodo-Dep="6142859" LSID="urn:lsid:plazi:treatment:2A658955C35EE753FF53F899FB5BFF52" httpUri="http://treatment.plazi.org/id/2A658955C35EE753FF53F899FB5BFF52" lastPageId="5" lastPageNumber="527" pageId="2" pageNumber="525">
<subSubSection id="EAD66BC8C35EE754FF53F899FCEAF8CE" box="[151,891,1839,1865]" pageId="2" pageNumber="525" type="nomenclature">
<paragraph id="A2733843C35EE754FF53F899FCEAF8CE" blockId="2.[151,891,1839,1865]" box="[151,891,1839,1865]" pageId="2" pageNumber="525">
<heading id="F93B8F2FC35EE754FF53F899FCEAF8CE" bold="true" box="[151,891,1839,1865]" fontSize="11" level="1" pageId="2" pageNumber="525" reason="1">
<emphasis id="90B8E451C35EE754FF53F899FCEAF8CE" bold="true" box="[151,891,1839,1865]" pageId="2" pageNumber="525">
<taxonomicName id="65CC43C0C35EE754FF53F899FDF0F8CE" authority="Oliveira" authorityName="Oliveira" box="[151,609,1839,1865]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="2" pageNumber="525" phylum="Arthropoda" rank="species" species="anhanguera">
<emphasis id="90B8E451C35EE754FF53F899FE62F8CE" bold="true" box="[151,499,1839,1865]" italics="true" pageId="2" pageNumber="525">Anhangatermes anhanguera</emphasis>
Oliveira
</taxonomicName>
&amp;
<collectingCountry id="DADB78D3C35EE754FD43F899FD4CF8CE" box="[647,733,1839,1865]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="525">Cunha</collectingCountry>
, new species
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EAD66BC8C35EE755FF53F8C7FD25FA15" lastPageId="3" lastPageNumber="526" pageId="2" pageNumber="525" type="materials_examined">
<paragraph id="A2733843C35EE754FF53F8C7FCC8F82A" blockId="2.[151,1436,1905,2038]" pageId="2" pageNumber="525">
<emphasis id="90B8E451C35EE754FF53F8C7FE95F80D" bold="true" box="[151,260,1905,1930]" pageId="2" pageNumber="525">
<typeStatus id="7D7786E1C35EE754FF53F8C7FE95F80D" box="[151,260,1905,1930]" pageId="2" pageNumber="525" type="holotype">Holotype</typeStatus>
</emphasis>
: soldier, part of lot DZUB-10265,
<date id="D6721E83C35EE754FD4FF8C7FD66F80D" box="[651,759,1905,1930]" pageId="2" pageNumber="525" value="2013-01-10">10.i.2013</date>
, collected by Hélida F.
<collectingCountry id="DADB78D3C35EE754FC3BF8C7FBDDF80D" box="[1023,1100,1905,1930]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="2" pageNumber="525">Cunha</collectingCountry>
.
<emphasis id="90B8E451C35EE754FF03F822FE13F82A" bold="true" box="[199,386,1940,1965]" pageId="2" pageNumber="525">
<typeStatus id="7D7786E1C35EE754FF03F822FE93F82A" box="[199,258,1940,1965]" pageId="2" pageNumber="525">Type</typeStatus>
repository
</emphasis>
: Dept. of Zoology, University of
<collectingCountry id="DADB78D3C35EE754FD3FF822FCC7F82A" box="[763,854,1940,1965]" name="Brazil" pageId="2" pageNumber="525">Brasília</collectingCountry>
.
</paragraph>
<paragraph id="A2733843C35EE754FF03F80FFD15F872" blockId="2.[151,1436,1905,2038]" pageId="2" pageNumber="525">
<emphasis id="90B8E451C35EE754FF03F80FFEF3F855" bold="true" box="[199,354,1977,2002]" pageId="2" pageNumber="525">Type-locality</emphasis>
:
<collectingCountry id="DADB78D3C35EE754FEB4F80FFE48F856" box="[368,473,1977,2001]" name="Brazil" pageId="2" pageNumber="525">BRAZIL</collectingCountry>
.
<emphasis id="90B8E451C35EE754FE20F80FFD10F857" box="[484,641,1977,2001]" italics="true" pageId="2" pageNumber="525">State of Goiás</emphasis>
. Anápolis, Campus da Universidade Estadual de Goiás (UnUCET), lat
<geoCoordinate id="C7F85E84C35EE754FF53F86BFF67F871" box="[151,246,2013,2038]" direction="south" orientation="latitude" pageId="2" pageNumber="525" precision="55" value="-16.383">16.383S</geoCoordinate>
, long
<geoCoordinate id="C7F85E84C35EE754FEFEF86BFE21F871" box="[314,432,2013,2038]" direction="west" orientation="longitude" pageId="2" pageNumber="525" precision="5" value="-48.9453">48.9453W</geoCoordinate>
, elevation
<quantity id="653495A6C35EE754FDEDF86BFD11F872" box="[553,640,2013,2037]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="525" unit="m" value="1100.0">1100 m</quantity>
.
</paragraph>
<caption id="F6B368CBC35FE755FF53FB79FCD9FA83" httpUri="https://zenodo.org/record/251220/files/figure.png" pageId="3" pageNumber="526" targetBox="[151,1436,183,1210]" targetPageId="3">
<paragraph id="A2733843C35FE755FF53FB79FCD9FA83" blockId="3.[151,1435,1231,1284]" pageId="3" pageNumber="526">
<emphasis id="90B8E451C35FE755FF53FB79FE85FB63" bold="true" box="[151,276,1231,1253]" pageId="3" pageNumber="526">FIGURE 3.</emphasis>
Enteric valve armature of
<emphasis id="90B8E451C35FE755FDECFB79FD5EFB62" box="[552,719,1231,1253]" italics="true" pageId="3" pageNumber="526">
<taxonomicName id="65CC43C0C35FE755FDECFB79FD5AFB62" box="[552,715,1231,1253]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">Anhangatermes</taxonomicName>
.
</emphasis>
A,
<taxonomicName id="65CC43C0C35FE755FD31FB66FC1DFB62" box="[757,908,1231,1253]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="anhanguera" status="sp. nov.">
<emphasis id="90B8E451C35FE755FD31FB66FC1DFB62" box="[757,908,1231,1253]" italics="true" pageId="3" pageNumber="526">A. anhanguera</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C35FE755FC5EFB66FC45FB62" bold="true" box="[922,980,1232,1253]" pageId="3" pageNumber="526">
<taxonomicNameLabel id="8B8B592AC35FE755FC5EFB66FC45FB62" box="[922,980,1232,1253]" pageId="3" pageNumber="526" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; B–C:
<taxonomicName id="65CC43C0C35FE755FBE4FB66FB55FB62" box="[1056,1220,1231,1253]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="eurycephalus" status="sp. nov.">
<emphasis id="90B8E451C35FE755FBE4FB66FB55FB62" box="[1056,1220,1231,1253]" italics="true" pageId="3" pageNumber="526">A. eurycephalus</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C35FE755FB16FB66FA9DFB62" bold="true" box="[1234,1292,1232,1253]" pageId="3" pageNumber="526">
<taxonomicNameLabel id="8B8B592AC35FE755FB16FB66FA9DFB62" box="[1234,1292,1232,1253]" pageId="3" pageNumber="526" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(C is a crosssection in posterior view); D,
<taxonomicName id="65CC43C0C35FE755FE01FB59FDA1FA83" box="[453,560,1263,1284]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="juruena" status="sp. nov.">
<emphasis id="90B8E451C35FE755FE01FB59FDA1FA83" box="[453,560,1263,1284]" italics="true" pageId="3" pageNumber="526">A. juruena</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C35FE755FDF8FB59FDE7FA83" bold="true" box="[572,630,1263,1284]" pageId="3" pageNumber="526">
<taxonomicNameLabel id="8B8B592AC35FE755FDF8FB59FDE7FA83" box="[572,630,1263,1284]" pageId="3" pageNumber="526" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; E,
<taxonomicName id="65CC43C0C35FE755FD5BFB59FC92FA83" box="[671,771,1262,1284]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="pilosus" status="sp. nov.">
<emphasis id="90B8E451C35FE755FD5BFB59FC92FA83" box="[671,771,1262,1284]" italics="true" pageId="3" pageNumber="526">A. pilosus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="8B8B592AC35FE755FCCBFB59FCD9FA83" box="[783,840,1263,1284]" pageId="3" pageNumber="526" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="A2733843C35FE755FF03FA87FD25FA15" blockId="3.[151,1437,1329,2038]" pageId="3" pageNumber="526">
<emphasis id="90B8E451C35FE755FF03FA87FEAEFACD" bold="true" box="[199,319,1329,1354]" pageId="3" pageNumber="526">
<typeStatus id="7D7786E1C35FE755FF03FA87FEAEFACD" box="[199,319,1329,1354]" pageId="3" pageNumber="526" type="paratype">Paratypes</typeStatus>
</emphasis>
(all from the type-locality):
<specimenCount id="B4CAF3CAC35FE755FD4CFA87FD67FACD" box="[648,758,1329,1354]" pageId="3" pageNumber="526" type="soldier">8 soldiers</specimenCount>
and
<specimenCount id="B4CAF3CAC35FE755FCF4FA87FC3EFACD" box="[816,943,1329,1354]" pageId="3" pageNumber="526" type="worker">20 workers</specimenCount>
from the same sample as
<typeStatus id="7D7786E1C35FE755FB1AFA87FAD2FACD" box="[1246,1347,1329,1354]" pageId="3" pageNumber="526" type="holotype">holotype</typeStatus>
, (UEG- 1684, DZUB-10265); one soldier and one worker,
<date id="D6721E83C35FE755FD34FAE3FCE3FAE9" box="[752,882,1365,1390]" pageId="3" pageNumber="526" value="2012-12-12">12.xii.2012</date>
, H.F.
<collectingCountry id="DADB78D3C35FE755FC7AFAE3FB96FAE9" box="[958,1031,1365,1390]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="3" pageNumber="526">Cunha</collectingCountry>
(UEG-2869);
<specimenCount id="B4CAF3CAC35FE755FB70FAE3FAA2FAE9" box="[1204,1331,1365,1390]" pageId="3" pageNumber="526" type="soldier">60 soldiers</specimenCount>
and
<specimenCount id="B4CAF3CAC35FE755FAB7FAE0FF65FA15" pageId="3" pageNumber="526" type="worker">117 workers</specimenCount>
,
<date id="D6721E83C35FE755FF3BFACFFEE7FA15" box="[255,374,1401,1426]" pageId="3" pageNumber="526" value="2014-04-11">11.iv.2014</date>
, D.E. Oliveira (UEG-2868).
</paragraph>
</subSubSection>
<subSubSection id="EAD66BC8C35FE752FF03FA2BFCF5F83B" lastPageId="4" lastPageNumber="527" pageId="3" pageNumber="526" type="etymology">
<paragraph id="A2733843C35FE755FF03FA2BFEBAFA79" blockId="3.[151,1437,1329,2038]" pageId="3" pageNumber="526">
<emphasis id="90B8E451C35FE755FF03FA2BFEDFFA31" bold="true" box="[199,334,1437,1462]" pageId="3" pageNumber="526">Etymology.</emphasis>
From the Tupi word “
<taxonomicName id="65CC43C0C35FE755FD8DFA2BFD40FA31" box="[585,721,1437,1462]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="anhanguera">anhanguera</taxonomicName>
”, old devil, combined as a noun in apposition.
<taxonomicName id="65CC43C0C35FE755FB24FA2BFAF8FA31" box="[1248,1385,1437,1462]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="anhanguera" status="sp. nov.">Anhanguera</taxonomicName>
was also the nickname of Bartolomeu Bueno da Silva, who led the first explorations to the type-locality of this species, around 1722.
</paragraph>
<paragraph id="A2733843C35FE755FF03F9BCFE02F9A5" blockId="3.[151,1437,1329,2038]" box="[199,403,1545,1570]" pageId="3" pageNumber="526">Imago. Unknown.</paragraph>
<paragraph id="A2733843C35FE755FF03F99BFCBEF871" blockId="3.[151,1437,1329,2038]" pageId="3" pageNumber="526">
Soldier (
<figureCitation id="3AF724C6C35FE755FEEFF99BFEEAF9C1" box="[299,379,1581,1606]" captionStart="FIGURE 5" captionStartId="5.[151,250,1945,1967]" captionTargetBox="[150,1436,1595,1921]" captionTargetId="figure@5.[150,1436,1595,1923]" captionTargetPageId="5" captionText="FIGURE 5. Schematic drawings of soldiers of Anhangatermes in dorsal view, showing chaetotaxy of thoracic nota and tergites 1 and 2. A, A. anhanguera, sp. n.; B: A. eurycephalus, sp. n.; C, A. juruena, sp. n.; D, A. pilosus, sp. n. Microscopic hairs not shown." httpUri="https://zenodo.org/record/251222/files/figure.png" pageId="3" pageNumber="526">Figs. 5</figureCitation>
A, 6A, 6B, 7, 9A). Head capsule yellow-orange; nasus darker. Antennae and thoracic nota yellow. Legs yellowish. Tergites and sternites hyaline. Head capsule typically with 2 bristles on vertex, 3 on the upper part of each lateral side, 4 at base of nasus, and 2 on clypeus; number, size and arrangement of bristles somewhat variable. Head capsule with numerous microscopic hairs, more numerous on vertex and frons (
<figureCitation id="3AF724C6C35FE755FAF3F92FFAEBF936" box="[1335,1402,1689,1714]" captionStart="FIGURE 7" captionStartId="7.[151,250,1150,1172]" captionTargetBox="[151,1436,193,1128]" captionTargetId="figure@7.[151,1436,193,1129]" captionTargetPageId="7" captionText="FIGURE 7. Scanning electron micrograph of the soldier of Anhangatermes anhanguera, sp. n. A, tip of nasus; B, head in profile; C, vertex of head; D, head in dorsal view." httpUri="https://zenodo.org/record/251224/files/figure.png" pageId="3" pageNumber="526">Fig. 7</figureCitation>
C). Apical 1/6 of nasus densely covered with short inclined hairs, which become longer toward apex; 4 longer hairs at tip (
<figureCitation id="3AF724C6C35FE755FF01F957FE9BF97E" box="[197,266,1761,1786]" captionStart="FIGURE 7" captionStartId="7.[151,250,1150,1172]" captionTargetBox="[151,1436,193,1128]" captionTargetId="figure@7.[151,1436,193,1129]" captionTargetPageId="7" captionText="FIGURE 7. Scanning electron micrograph of the soldier of Anhangatermes anhanguera, sp. n. A, tip of nasus; B, head in profile; C, vertex of head; D, head in dorsal view." httpUri="https://zenodo.org/record/251224/files/figure.png" pageId="3" pageNumber="526">Fig. 7</figureCitation>
A). Pronotum with a row of about 6 bristles on anterior margin and a few or none on posterior margin (
<figureCitation id="3AF724C6C35FE755FF5BF8B3FF74F899" box="[159,229,1797,1822]" captionStart="FIGURE 5" captionStartId="5.[151,250,1945,1967]" captionTargetBox="[150,1436,1595,1921]" captionTargetId="figure@5.[150,1436,1595,1923]" captionTargetPageId="5" captionText="FIGURE 5. Schematic drawings of soldiers of Anhangatermes in dorsal view, showing chaetotaxy of thoracic nota and tergites 1 and 2. A, A. anhanguera, sp. n.; B: A. eurycephalus, sp. n.; C, A. juruena, sp. n.; D, A. pilosus, sp. n. Microscopic hairs not shown." httpUri="https://zenodo.org/record/251222/files/figure.png" pageId="3" pageNumber="526">Fig. 5</figureCitation>
A). Meso- and metanotum without bristles in most specimens; posterior margin sometimes with isolate bristles. Thoracic nota with numerous microscopic hairs. Tergites with a row of about 8 bristles on posterior margin, and one short bristle on lateral margin (
<figureCitation id="3AF724C6C35FE755FD0FF8FBFC9BF8E1" box="[715,778,1869,1894]" captionStart="FIGURE 5" captionStartId="5.[151,250,1945,1967]" captionTargetBox="[150,1436,1595,1921]" captionTargetId="figure@5.[150,1436,1595,1923]" captionTargetPageId="5" captionText="FIGURE 5. Schematic drawings of soldiers of Anhangatermes in dorsal view, showing chaetotaxy of thoracic nota and tergites 1 and 2. A, A. anhanguera, sp. n.; B: A. eurycephalus, sp. n.; C, A. juruena, sp. n.; D, A. pilosus, sp. n. Microscopic hairs not shown." httpUri="https://zenodo.org/record/251222/files/figure.png" pageId="3" pageNumber="526">Fig 5</figureCitation>
A); surface of tergites with numerous microscopic hairs. Sternites with a row of long bristles on posterior margin; surface with dense inclined hairs of variable length. Head capsule broad, with evenly convex sides. Mandibles apically rounded, without sharp points (
<figureCitation id="3AF724C6C35FE755FB6DF823FB7DF829" box="[1193,1260,1941,1966]" captionStart="FIGURE 9" captionStartId="9.[151,250,1244,1266]" captionTargetBox="[150,1432,151,1205]" captionTargetId="figure@9.[148,1433,151,1206]" captionTargetPageId="9" captionText="FIGURE 9. Soldier mandibles of Anhangatermes in dorsal view. A, A. anhanguera, sp. n.; B: A. eurycephalus, sp. n.; C, A. juruena, sp. n.; D, A. macarthuri Constantino 1990 (paratype); E, A. pilosus, sp. n." httpUri="https://zenodo.org/record/251226/files/figure.png" pageId="3" pageNumber="526">Fig. 9</figureCitation>
A). Nasus long, subcylindrical and slightly upturned. Pronotum with anterior margin convex, slightly emarginate. Relative length of antennal articles: 3&gt;6&gt;5&gt;2=4. Measurements in
<tableCitation id="EF4E0DF8C35FE755FD11F86BFCB8F871" box="[725,809,2013,2038]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Measurements (in mm) of soldiers of Anhangatermes (hol. = holotype)." pageId="3" pageNumber="526">Table 1</tableCitation>
.
</paragraph>
<caption id="F6B368CBC358E752FF53FF2EFB9CFF28" box="[151,1037,151,175]" pageId="4" pageNumber="527">
<paragraph id="A2733843C358E752FF53FF2EFB9CFF28" blockId="4.[151,1037,151,175]" box="[151,1037,151,175]" pageId="4" pageNumber="527">
<emphasis id="90B8E451C358E752FF53FF2EFE9BFF28" bold="true" box="[151,266,152,175]" pageId="4" pageNumber="527">TABLE 1.</emphasis>
Measurements (in mm) of soldiers of
<taxonomicName id="65CC43C0C358E752FD67FF21FCDBFF28" box="[675,842,151,175]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="90B8E451C358E752FD67FF21FCDBFF28" box="[675,842,151,175]" italics="true" pageId="4" pageNumber="527">Anhangatermes</emphasis>
</taxonomicName>
(hol. = holotype).
</paragraph>
</caption>
<paragraph id="A2733843C358E752FF5BFF7CFB66FF67" blockId="4.[159,1413,202,472]" box="[159,1271,202,224]" pageId="4" pageNumber="527">Species n Length of head Width of head Height of head</paragraph>
<paragraph id="A2733843C358E752FE7EFF43FB72FDC5" blockId="4.[159,1413,202,472]" lastBlockId="4.[159,1409,556,826]" pageId="4" pageNumber="527">
range mean hol. range mean hol. range mean hol.
<taxonomicName id="65CC43C0C358E752FF5BFEA8FEA7FEB4" box="[159,310,285,307]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="anhanguera">
<emphasis id="90B8E451C358E752FF5BFEA8FEA7FEB4" box="[159,310,285,307]" italics="true" pageId="4" pageNumber="527">A. anhanguera</emphasis>
</taxonomicName>
20 2.38–2.60 2.47 2.44 1.40–1.56 1.48 1.46 0.90–1.04 0.96 0.96
<taxonomicName id="65CC43C0C358E752FF5BFEF1FED3FEDB" box="[159,322,326,348]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="eurycephalus">
<emphasis id="90B8E451C358E752FF5BFEF1FED3FEDB" box="[159,322,326,348]" italics="true" pageId="4" pageNumber="527">A. eurycephalus</emphasis>
</taxonomicName>
22 2.40–2.60 2.55 2.42 1.52–1.68 1.58 1.32 0.96–1.12 1.02 0.88
<taxonomicName id="65CC43C0C358E752FF5BFEC7FE9BFE01" box="[159,266,369,390]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="juruena">
<emphasis id="90B8E451C358E752FF5BFEC7FE9BFE01" box="[159,266,369,390]" italics="true" pageId="4" pageNumber="527">A. juruena</emphasis>
</taxonomicName>
16 2.32–2.44 2.38 2.60 1.24–1.36 1.30 1.68 0.80–0.90 0.84 1.06
<taxonomicName id="65CC43C0C358E752FF5BFE2CFEBCFE28" box="[159,301,409,431]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="macarthuri">
<emphasis id="90B8E451C358E752FF5BFE2CFEBCFE28" box="[159,301,409,431]" italics="true" pageId="4" pageNumber="527">A. macarthuri</emphasis>
</taxonomicName>
15 2.16–2.40 2.29 — 1.22–1.32 1.28 — 0.80–0.85 0.83 —
<taxonomicName id="65CC43C0C358E752FF5BFE75FE92FE5F" box="[159,259,450,472]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="90B8E451C358E752FF5BFE75FE92FE5F" box="[159,259,450,472]" italics="true" pageId="4" pageNumber="527">A. pilosus</emphasis>
</taxonomicName>
30 2.34–2.52 2.41 2.44 1.36–1.52 1.45 1.46 0.90–1.00 0.95 1.00 Species Length of nasus Width of pronotum Length of tibia
</paragraph>
<caption id="F6B368CBC358E752FF53FE4FFE00FD97" box="[151,401,505,528]" pageId="4" pageNumber="527">
<paragraph id="A2733843C358E752FF53FE4FFE00FD97" blockId="4.[151,401,505,528]" box="[151,401,505,528]" pageId="4" pageNumber="527">
<emphasis id="90B8E451C358E752FF53FE4FFE9BFD97" bold="true" box="[151,266,505,528]" pageId="4" pageNumber="527">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="A2733843C358E752FEBEFDE0FB15FB7C" blockId="4.[159,1409,556,826]" lastBlockId="4.[151,1437,890,1275]" pageId="4" pageNumber="527">
range mean hol. range mean hol. range mean hol.
<taxonomicName id="65CC43C0C358E752FF5BFDC9FEA7FD13" box="[159,310,638,660]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="anhanguera">
<emphasis id="90B8E451C358E752FF5BFDC9FEA7FD13" box="[159,310,638,660]" italics="true" pageId="4" pageNumber="527">A. anhanguera</emphasis>
</taxonomicName>
1.12–1.22 1.17 1.14 0.65–0.70 0.68 0.70 1.26–1.35 1.31 1.26
<taxonomicName id="65CC43C0C358E752FF5BFD1FFED3FD39" box="[159,322,680,702]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="eurycephalus">
<emphasis id="90B8E451C358E752FF5BFD1FFED3FD39" box="[159,322,680,702]" italics="true" pageId="4" pageNumber="527">A. eurycephalus</emphasis>
</taxonomicName>
1.06–1.22 1.15 1.18 0.66–0.72 0.70 0.66 1.12–1.38 1.29 1.20
<taxonomicName id="65CC43C0C358E752FF5BFD64FE9BFD60" box="[159,266,722,743]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="juruena">
<emphasis id="90B8E451C358E752FF5BFD64FE9BFD60" box="[159,266,722,743]" italics="true" pageId="4" pageNumber="527">A. juruena</emphasis>
</taxonomicName>
1.10–1.20 1.15 1.20 0.62–0.68 0.66 0.72 1.20–1.24 1.21 1.28
<taxonomicName id="65CC43C0C358E752FF5BFD4DFEBCFC97" box="[159,301,762,784]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="macarthuri">
<emphasis id="90B8E451C358E752FF5BFD4DFEBCFC97" box="[159,301,762,784]" italics="true" pageId="4" pageNumber="527">A. macarthuri</emphasis>
</taxonomicName>
1.04–1.22 1.13 — 0.60–0.66 0.62 — 1.11–1.22 1.16 —
<taxonomicName id="65CC43C0C358E752FF5BFC92FE92FCBD" box="[159,259,804,826]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="90B8E451C358E752FF5BFC92FE92FCBD" box="[159,259,804,826]" italics="true" pageId="4" pageNumber="527">A. pilosus</emphasis>
</taxonomicName>
1.08–1.20 1.14 1.16 0.62–0.68 0.65 0.64 1.20–1.34 1.27 1.26 Worker (
<figureCitation id="3AF724C6C358E752FEEEFCCCFEEBFC14" box="[298,378,890,915]" captionStart="FIGURE 2" captionStartId="2.[151,250,1742,1764]" captionTargetBox="[151,1436,1290,1720]" captionTargetId="figure@2.[151,1436,1290,1720]" captionTargetPageId="2" captionText="FIGURE 2. Gizzard armature of Anhangatermes anhanguera, sp. n." httpUri="https://zenodo.org/record/251219/files/figure.png" pageId="4" pageNumber="527">Figs. 2</figureCitation>
,
<figureCitation id="3AF724C6C358E752FE42FCCDFE02FC14" box="[390,403,891,915]" captionStart="FIGURE 3" captionStartId="3.[151,250,1231,1253]" captionTargetBox="[151,1436,183,1210]" captionTargetId="figure@3.[151,1436,183,1210]" captionTargetPageId="3" captionText="FIGURE 3. Enteric valve armature of Anhangatermes. A, A. anhanguera, sp. n.; B – C: A. eurycephalus, sp. n. (C is a crosssection in posterior view); D, A. juruena, sp. n.; E, A. pilosus, sp. n." httpUri="https://zenodo.org/record/251220/files/figure.png" pageId="4" pageNumber="527">3</figureCitation>
A, 10A). Head capsule and postclypeus yellowish. Postclypeus strongly inflated. Epicranial suture and fontanelle spot inconspicuous. Fontanelle spot slightly depressed. Antennae with basal articles whitish, becoming yellow towards apex. Thoracic nota, legs and abdomen whitish. Head capsule with numerous bristles of variable size (
<figureCitation id="3AF724C6C358E752FEF2FC50FE19FC78" box="[310,392,998,1023]" captionStart="FIGURE 10" captionStartId="10.[151,250,1468,1490]" captionTargetBox="[154,1436,506,1425]" captionTargetId="figure@10.[154,1439,506,1425]" captionTargetPageId="10" captionText="FIGURE 10. Workers of Anhangatermes, head in dorsal view. A, A. anhanguera, sp. n.; B, A. eurycephalus, sp. n.; C, A. juruena, sp. n.; D, A. pilosus, sp. n." httpUri="https://zenodo.org/record/251227/files/figure.png" pageId="4" pageNumber="527">Fig. 10</figureCitation>
A) and many short hairs and microscopic hairs; postclypeus with many bristles and hairs of variable size. Pro-, meso- and metanota with many bristles on margins and numerous microscopic hairs on surface. Tergites with a irregular row of bristles of variable size on posterior margin, and a second, similar row near middle; surface of tergites with sparse hairs, about half the length of bristles. Pilosity of sternites similar to that of soldier. Enteric valve armature elongate, with 3 major plates alternating with 3 minor plates; minor plates narrower and a little shorter (
<figureCitation id="3AF724C6C358E752FEEBFB2CFEE0FB34" box="[303,369,1178,1203]" captionStart="FIGURE 3" captionStartId="3.[151,250,1231,1253]" captionTargetBox="[151,1436,183,1210]" captionTargetId="figure@3.[151,1436,183,1210]" captionTargetPageId="3" captionText="FIGURE 3. Enteric valve armature of Anhangatermes. A, A. anhanguera, sp. n.; B – C: A. eurycephalus, sp. n. (C is a crosssection in posterior view); D, A. juruena, sp. n.; E, A. pilosus, sp. n." httpUri="https://zenodo.org/record/251220/files/figure.png" pageId="4" pageNumber="527">Fig. 3</figureCitation>
A). Length of major plates about
<quantity id="653495A6C358E752FD2DFB2DFCAEFB35" box="[745,831,1179,1203]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="527" unit="mm" value="0.4">0.4 mm</quantity>
and of minor plates about
<quantity id="653495A6C358E752FBA8FB2DFB54FB35" box="[1132,1221,1179,1203]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="527" unit="mm" value="0.3">0.3 mm</quantity>
. All plates covered with relatively large conical spines orientated in direction of food movement; posterior end of each plate with a line of perpendicular spines. Surface between plates without spines. Measurements in
<tableCitation id="EF4E0DF8C358E752FBEFFB54FB10FB7C" box="[1067,1153,1250,1275]" captionStart="TABLE 2" captionStartId="4.[151,239,1318,1341]" captionText="TABLE 2. Measurements (in mm) of workers of Anhangatermes." pageId="4" pageNumber="527">Table 2</tableCitation>
.
</paragraph>
<caption id="F6B368CBC358E752FF53FA90FCC5FABA" box="[151,852,1317,1341]" pageId="4" pageNumber="527">
<paragraph id="A2733843C358E752FF53FA90FCC5FABA" blockId="4.[151,852,1317,1341]" box="[151,852,1317,1341]" pageId="4" pageNumber="527">
<emphasis id="90B8E451C358E752FF53FA90FE98FABA" bold="true" box="[151,265,1318,1341]" pageId="4" pageNumber="527">TABLE 2.</emphasis>
Measurements (in mm) of workers of
<taxonomicName id="65CC43C0C358E752FD62FA93FCDCFABA" box="[678,845,1317,1341]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="90B8E451C358E752FD62FA93FCDCFABA" box="[678,845,1317,1341]" italics="true" pageId="4" pageNumber="527">Anhangatermes</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="A2733843C358E752FF5BFAEFFAD3FAE8" blockId="4.[159,1396,1369,1639]" box="[159,1346,1369,1391]" pageId="4" pageNumber="527">Species n length of head width of head width of pronotum length of tibia</paragraph>
<paragraph id="A2733843C358E752FE72FA35FE36F81F" blockId="4.[159,1396,1369,1639]" lastBlockId="4.[151,1436,1703,2017]" pageId="4" pageNumber="527">
range mean range mean range mean range mean
<taxonomicName id="65CC43C0C358E752FF5BFA1AFEA7FA45" box="[159,310,1452,1474]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="anhanguera">
<emphasis id="90B8E451C358E752FF5BFA1AFEA7FA45" box="[159,310,1452,1474]" italics="true" pageId="4" pageNumber="527">A. anhanguera</emphasis>
</taxonomicName>
20 0.82–0.94 0.88 1.00–1.14 1.08 0.56–0.62 0.59 1.04–1.20 1.13
<taxonomicName id="65CC43C0C358E752FF5BFA60FED3FA6C" box="[159,322,1493,1515]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="eurycephalus">
<emphasis id="90B8E451C358E752FF5BFA60FED3FA6C" box="[159,322,1493,1515]" italics="true" pageId="4" pageNumber="527">A. eurycephalus</emphasis>
</taxonomicName>
19 0.80–0.96 0.86 0.98–1.12 1.04 0.52–0.70 0.56 1.08–1.32 1.15
<taxonomicName id="65CC43C0C358E752FF5BFA49FE9BF992" box="[159,266,1535,1557]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="juruena">
<emphasis id="90B8E451C358E752FF5BFA49FE9BF992" box="[159,266,1535,1557]" italics="true" pageId="4" pageNumber="527">A. juruena</emphasis>
</taxonomicName>
20 0.84–0.94 0.88 1.02–1.10 1.06 0.56–0.60 0.58 1.08–1.16 1.12
<taxonomicName id="65CC43C0C358E752FF5BF99FFEBCF9B9" box="[159,301,1576,1598]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="macarthuri">
<emphasis id="90B8E451C358E752FF5BF99FFEBCF9B9" box="[159,301,1576,1598]" italics="true" pageId="4" pageNumber="527">A. macarthuri</emphasis>
</taxonomicName>
3 0.78–0.82 0.79 0.96–1.00 0.98 0.52–0.56 0.54 1.06 1.06
<taxonomicName id="65CC43C0C358E752FF5BF9E4FE92F9E0" box="[159,259,1617,1639]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="pilosus">
<emphasis id="90B8E451C358E752FF5BF9E4FE92F9E0" box="[159,259,1617,1639]" italics="true" pageId="4" pageNumber="527">A. pilosus</emphasis>
</taxonomicName>
30 0.82–0.98 0.90 0.92–1.18 1.11 0.52–0.68 0.58 1.06–1.38 1.21
<emphasis id="90B8E451C358E752FF03F911FEFDF947" bold="true" box="[199,364,1703,1728]" pageId="4" pageNumber="527">Comparisons.</emphasis>
Soldiers of
<taxonomicName id="65CC43C0C358E752FE37F91EFD08F947" box="[499,665,1704,1728]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="anhanguera" status="sp. nov.">
<emphasis id="90B8E451C358E752FE37F91EFD08F947" box="[499,665,1704,1728]" italics="true" pageId="4" pageNumber="527">A. anhanguera</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C358E752FD62F911FD74F947" bold="true" box="[678,741,1703,1728]" pageId="4" pageNumber="527">
<taxonomicNameLabel id="8B8B592AC358E752FD62F911FD74F947" box="[678,741,1703,1728]" pageId="4" pageNumber="527" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from those of
<taxonomicName id="65CC43C0C358E752FB7DF91EFAFDF947" box="[1209,1388,1704,1728]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="eurycephalus" status="sp. nov.">
<emphasis id="90B8E451C358E752FB7DF91EFAFDF947" box="[1209,1388,1704,1728]" italics="true" pageId="4" pageNumber="527">A. eurycephalus</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C358E752FABEF911FF3FF962" bold="true" pageId="4" pageNumber="527">
<taxonomicNameLabel id="8B8B592AC358E752FABEF911FF3FF962" pageId="4" pageNumber="527" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="65CC43C0C358E752FF78F97AFEA5F963" box="[188,308,1740,1764]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="juruena" status="sp. nov.">
<emphasis id="90B8E451C358E752FF78F97AFEA5F963" box="[188,308,1740,1764]" italics="true" pageId="4" pageNumber="527">A. juruena</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C358E752FE87F97AFE15F962" bold="true" box="[323,388,1740,1765]" pageId="4" pageNumber="527">
<taxonomicNameLabel id="8B8B592AC358E752FE87F97AFE15F962" box="[323,388,1740,1765]" pageId="4" pageNumber="527" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="65CC43C0C358E752FE78F97AFDCAF963" box="[444,603,1740,1764]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="macarthuri">
<emphasis id="90B8E451C358E752FE78F97AFDCAF963" box="[444,603,1740,1764]" italics="true" pageId="4" pageNumber="527">A. macarthuri</emphasis>
</taxonomicName>
based on chaetotaxy: the later species have only two bristles on the head capsule, no bristles on thoracic nota, no bristles on clypeus, and a line of 4 bristles on the posterior margin of each tergite. The soldier of
<taxonomicName id="65CC43C0C358E752FE64F8A2FD83F8AB" box="[416,530,1812,1836]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="pilosus" status="sp. nov.">
<emphasis id="90B8E451C358E752FE64F8A2FD83F8AB" box="[416,530,1812,1836]" italics="true" pageId="4" pageNumber="527">A. pilosus</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C358E752FDE0F8A2FDF9F8AA" bold="true" box="[548,616,1812,1837]" pageId="4" pageNumber="527">
<taxonomicNameLabel id="8B8B592AC358E752FDE0F8A2FDF9F8AA" box="[548,616,1812,1837]" pageId="4" pageNumber="527" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs in the chaetotaxy of meso- and metanotum, which have many bristles, and of tergites, which have two rows of bristles instead of one (
<figureCitation id="3AF724C6C358E752FC06F881FB96F8D7" box="[962,1031,1847,1872]" captionStart="FIGURE 5" captionStartId="5.[151,250,1945,1967]" captionTargetBox="[150,1436,1595,1921]" captionTargetId="figure@5.[150,1436,1595,1923]" captionTargetPageId="5" captionText="FIGURE 5. Schematic drawings of soldiers of Anhangatermes in dorsal view, showing chaetotaxy of thoracic nota and tergites 1 and 2. A, A. anhanguera, sp. n.; B: A. eurycephalus, sp. n.; C, A. juruena, sp. n.; D, A. pilosus, sp. n. Microscopic hairs not shown." httpUri="https://zenodo.org/record/251222/files/figure.png" pageId="4" pageNumber="527">Fig. 5</figureCitation>
). The worker of
<taxonomicName id="65CC43C0C358E752FB01F88EFAFAF8D7" box="[1221,1387,1848,1872]" class="Insecta" family="Termitidae" genus="Anhangatermes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Isoptera" pageId="4" pageNumber="527" phylum="Arthropoda" rank="species" species="anhanguera" status="sp. nov.">
<emphasis id="90B8E451C358E752FB01F88EFAFAF8D7" box="[1221,1387,1848,1872]" italics="true" pageId="4" pageNumber="527">A. anhanguera</emphasis>
</taxonomicName>
,
<emphasis id="90B8E451C358E752FABEF881FF3FF8F2" bold="true" pageId="4" pageNumber="527">
<taxonomicNameLabel id="8B8B592AC358E752FABEF881FF3FF8F2" pageId="4" pageNumber="527" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has more bristles and hairs on the head capsule than all the other species and its enteric valve armature is longer than in all other species.
</paragraph>
<paragraph id="A2733843C358E752FF03F812FCF5F83B" blockId="4.[151,1436,1703,2017]" box="[199,868,1956,1981]" pageId="4" pageNumber="527">
<emphasis id="90B8E451C358E752FF03F812FEBBF83A" bold="true" box="[199,298,1956,1981]" pageId="4" pageNumber="527">Habitat.</emphasis>
Gallery forest and savanna (cerrado
<emphasis id="90B8E451C358E752FD03F813FCC2F83B" box="[711,851,1957,1980]" italics="true" pageId="4" pageNumber="527">sensu stricto</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="EAD66BC8C358E753FF03F87EFB5BFF52" lastPageId="5" lastPageNumber="528" pageId="4" pageNumber="527" type="discussion">
<paragraph id="A2733843C358E753FF03F87EFB5BFF52" blockId="4.[151,1436,1703,2017]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="528" pageId="4" pageNumber="527">
<emphasis id="90B8E451C358E752FF03F87EFEAAF866" bold="true" box="[199,315,1992,2017]" pageId="4" pageNumber="527">Remarks.</emphasis>
Sample UEG-032 (
<collectingCountry id="DADB78D3C358E752FDD9F87EFDF7F866" box="[541,614,1992,2017]" name="Brazil" pageId="4" pageNumber="527">Brazil</collectingCountry>
, State of Goiás, Parque Estadual da Serra de Jaraguá, lat
<geoCoordinate id="C7F85E84C358E752FB33F87EFAF0F866" box="[1271,1377,1992,2017]" direction="south" orientation="latitude" pageId="4" pageNumber="527" precision="5" value="-15.8054">15.8054S</geoCoordinate>
long
<geoCoordinate id="C7F85E84C359E753FF53FF21FE9FFF37" box="[151,270,151,176]" direction="west" orientation="longitude" pageId="5" pageNumber="528" precision="5" value="-49.3206">49.3206W</geoCoordinate>
) contains a single, dried, soldier head which seems to belong to this species. This sample was excluded from the
<typeStatus id="7D7786E1C359E753FF39FF0BFEBCFF52" box="[253,301,189,213]" pageId="5" pageNumber="528">type</typeStatus>
series because the specimen is poorly preserved and its identification is uncertain.
</paragraph>
</subSubSection>
</treatment>
</document>
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