treatments-xml/data/22/78/87/227887BEFFDDFF88250DF419FDF6FFE0.xml
2024-06-21 12:31:24 +02:00

224 lines
25 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="1D5A96BC68A0DAD4CA0F208E8513955F" ID-DOI="10.11646/zootaxa.1099.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5051674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625110034160" checkinUser="felipe" docAuthor="Ball, George E. &amp; Shpeley, Danny" docDate="2005" docId="227887BEFFDDFF88250DF419FDF6FFE0" docLanguage="en" docName="zt01099p060.pdf" docOrigin="Zootaxa 1099 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1099.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cymbionotum (Procoscinia) negrei Perrault 1994" docType="treatment" docVersion="4" lastPageNumber="120" masterDocId="DE41FFC6FFE4FFB12405F505FFA2FA39" masterDocTitle="Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution" masterLastPageNumber="120" masterPageNumber="1" pageNumber="1" updateTime="1698981561892" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="157BC564ABF89E64680360BA7352A3C1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E1156BF6D4EFAA340E9DB18C85FEE24E">
<mods:title id="33E266B01C003B3FC56AA8AF093D323A">Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution</mods:title>
</mods:titleInfo>
<mods:name id="8BAF8C465FF2FE48D89D4C42F738F99A" type="personal">
<mods:role id="0AA0676218904B48175C7E6EB6415AFD">
<mods:roleTerm id="569BC0CA52868B9F4782919E0B2266EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="853790B495C39A62C0BC41F99C9079CA">Ball, George E.</mods:namePart>
</mods:name>
<mods:name id="FAAB7C04A8EFC642B41B830A4A001495" type="personal">
<mods:role id="A98F18791DD5BCF760A4CAA1E54CE3DB">
<mods:roleTerm id="506FD10B73DC10365863C46E93F553AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="298D69FA66E63479050EA3DA39E80442">Shpeley, Danny</mods:namePart>
</mods:name>
<mods:typeOfResource id="0042D257B232D248CE479C15D510AEEE">text</mods:typeOfResource>
<mods:relatedItem id="C6DF26FA281702AF9EF660709E228F4E" type="host">
<mods:titleInfo id="970F893D80B598180A62F16E186C30F3">
<mods:title id="7DFBB1E2AAF7E4C1FCEC6521F78D95EF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="94CCF706BC7D886BB7A6CBC0E76806FB">
<mods:date id="AD4CD207D648AF10D11ACB97CFB28225">2005</mods:date>
<mods:detail id="49A4D0AF7547D2BC190111AF21F0A93E" type="pubDate">
<mods:number id="AD9E4321AF1482C787C23ECD1DC4456F">2005-12-19</mods:number>
</mods:detail>
<mods:detail id="96D6E649CC90EB19D5F18AB7BCDCF1B9" type="volume">
<mods:number id="90B740D88D18D638F32BF1D124E6423D">1099</mods:number>
</mods:detail>
<mods:detail id="E84B66FF1E91AD3D570A6380135DCA8A" type="issue">
<mods:number id="FB225F817F8488FB120611EEB40AB969">1</mods:number>
</mods:detail>
<mods:extent id="FBE072C49202BFF0977CB617B88C5CC3" unit="page">
<mods:start id="4E3742FA9F0D7CCF75445539545CC4B1">1</mods:start>
<mods:end id="4B5FAA9A6BB9D1AF9079D711F5FAE8CB">120</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="974C04A64A536C410A628683334C273B">
<mods:url id="2E32B08CC31E878FA3A4688D6463B13F">https://biotaxa.org/Zootaxa/article/view/zootaxa.1099.1.1</mods:url>
</mods:location>
<mods:classification id="F9215996899DF8194F282FAED51F7D11">journal article</mods:classification>
<mods:identifier id="07C4C83D7467376FA96474194B70A158" type="DOI">10.11646/zootaxa.1099.1.1</mods:identifier>
<mods:identifier id="8482C9CC16EA248433DEC363DD2392AF" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="179F09286DBBC66588350C7EB58A97D4" type="Zenodo-Dep">5051674</mods:identifier>
<mods:identifier id="B51389ADC74DBF0C30F0E7337865ED8D" type="ZooBank">CA0A0ED3-D49D-4198-A409-407EF86A164E</mods:identifier>
</mods:mods>
<treatment id="227887BEFFDDFF88250DF419FDF6FFE0" ID-DOI="http://doi.org/10.5281/zenodo.5054428" ID-Zenodo-Dep="5054428" LSID="urn:lsid:plazi:treatment:227887BEFFDDFF88250DF419FDF6FFE0" httpUri="http://treatment.plazi.org/id/227887BEFFDDFF88250DF419FDF6FFE0" pageId="57">
<subSubSection id="E2CB6523FFDDFF88250DF419FCC6FB0C" box="[264,868,283,310]" pageId="57" type="nomenclature">
<paragraph id="AA6E36A8FFDDFF88250DF419FCC6FB0C" blockId="57.[264,868,283,345]" box="[264,868,283,310]" pageId="57">
<heading id="F12681C4FFDDFF88250DF419FCC6FB0C" bold="true" box="[264,868,283,310]" fontSize="11" level="1" pageId="57" reason="1">
<taxonomicName id="6DD14D2BFFDDFF88250DF419FCC6FB0C" ID-CoL="6C776" authority="Perrault 1994" authorityName="Perrault" authorityYear="1994" box="[264,868,283,310]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="negrei" subGenus="Procoscinia">
<emphasis id="98A5EABAFFDDFF88250DF419FCC6FB0C" bold="true" box="[264,868,283,310]" pageId="57">
<emphasis id="98A5EABAFFDDFF88250DF419FD12FB0F" bold="true" box="[264,688,284,310]" italics="true" pageId="57">Cymbionotum (Procoscinia) negrei</emphasis>
Perrault 1994
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E2CB6523FFDDFF88250DF43AFD42F899" pageId="57" type="description">
<paragraph id="AA6E36A8FFDDFF88250DF43AFE0AFB60" blockId="57.[264,868,283,345]" box="[264,424,319,345]" pageId="57">
(
<figureCitation id="32EA2A2DFFDDFF882514F43AFED7FB60" box="[273,373,319,345]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (AE) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; CD left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and BD and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="57">Fig. 15F</figureCitation>
,
<figureCitation id="32EA2A2DFFDDFF882584F43AFE3DFB60" box="[385,415,319,345]" captionStart="FIGURE 16" captionStartId="59.[264,368,1187,1211]" captionTargetBox="[284,1298,525,1160]" captionTargetId="figure-227@59.[284,1302,524,1161]" captionTargetPageId="59" captionText="FIGURE 16. Geographical range of the species of Cymbionotum (Procoscinia), and of Cymbionotum (s. str.) basale species group illustrated on an outline map of South America and of southwestern Asia, southern Europe and Africa." figureDoi="http://doi.org/10.5281/zenodo.5051718" httpUri="https://zenodo.org/record/5051718/files/figure.png" pageId="57">16</figureCitation>
)
</paragraph>
<paragraph id="AA6E36A8FFDDFF88250DF48FFD42F899" blockId="57.[264,1323,393,672]" pageId="57">
<taxonomicName id="6DD14D2BFFDDFF88250DF48FFD71FB99" authority="Perrault 1994: 208" authorityName="Perrault" authorityPageNumber="208" authorityYear="1994" box="[264,723,393,417]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="98A5EABAFFDDFF88250DF48FFE4CFB98" box="[264,494,394,417]" italics="true" pageId="57">Cymbionotum negrei</emphasis>
Perrault 1994: 208
</taxonomicName>
. TYPE MATERIAL:
<specimenCount id="BCD7FD21FFDDFF8827D5F48CFB27FB99" box="[976,1157,393,416]" pageId="57" type="generic">three specimens</specimenCount>
, card­mounted (MNHP).
<materialsCitation id="1AB93CF5FFDDFF8825A0F4ACFA85FBD9" country="Venezuela" location="BAS SARARE" pageId="57" specimenCount="2" stateProvince="BAS SARARE" typeStatus="holotype">
<typeStatus id="756A880AFFDDFF8825A0F4ACFD93FBF9" box="[421,561,425,448]" pageId="57" type="holotype">HOLOTYPE</typeStatus>
male, labeled: &quot;MUSEUM PARIS/
<collectingCountry id="D2C67638FFDDFF8827B0F4ACFBF1FBF9" box="[949,1107,425,448]" name="Venezuela" pageId="57">VENEZUELA</collectingCountry>
/
<collectingRegion id="6815F84AFFDDFF882066F4ACFAA5FBF9" box="[1123,1287,425,448]" country="Venezuela" name="BAS SARARE" pageId="57">BAS SARARE</collectingRegion>
/ F. GEAY 33­96&quot;; &quot;Sous les écorce/ darbres&quot; [handwritten]; &quot;
<typeStatus id="756A880AFFDDFF8827F0F4CCFB25FBD9" box="[1013,1159,457,480]" pageId="57" type="holotype">HOLOTYPE</typeStatus>
&quot; [red paper]
</materialsCitation>
.
<materialsCitation id="1AB93CF5FFDDFF88253DF4ECFD7EF899" collectorName="S. Fern. &amp; L. Laglaize &amp; R. Oberthur" country="Venezuela" determinerName="G. G. Perrault" location="MUSEUM PARIS" pageId="57" specimenCount="5" stateProvince="MUSEUM PARIS" typeStatus="paratype">
<typeStatus id="756A880AFFDDFF88253DF4ECFE6CF839" box="[312,462,489,512]" pageId="57" type="paratype">PARATYPES</typeStatus>
, male and female. Male, labeled: &quot;
<collectingRegion id="6815F84AFFDDFF882756F4ECFBBCF839" box="[851,1054,489,512]" country="Venezuela" name="MUSEUM PARIS" pageId="57">MUSEUM PARIS</collectingRegion>
/
<collectingCountry id="D2C67638FFDDFF882035F4ECFB6CF839" box="[1072,1230,489,512]" name="Venezuela" pageId="57">VENEZUELA</collectingCountry>
/ HAUT
<collectingRegion id="6815F84AFFDDFF88253DF70CFE2DF819" box="[312,399,521,544]" country="Venezuela" name="Apure" pageId="57">APURE</collectingRegion>
/ F. GEAY 33­96&quot;; &quot;
<typeStatus id="756A880AFFDDFF882666F70CFD4CF819" box="[611,750,521,544]" pageId="57" type="paratype">PARATYPE</typeStatus>
&quot; [red paper];
<taxonomicName id="6DD14D2BFFDDFF88277AF70CFBBBF819" authorityName="Baudi di Selve. The" authorityYear="1864" box="[895,1049,521,544]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="genus">Cymbionotum</taxonomicName>
/ negrei m. [handwritten]/
<typeStatus id="756A880AFFDDFF88253DF72CFE1EF879" box="[312,444,553,576]" pageId="57" type="paratype">PARATYPE</typeStatus>
/
<determinerName id="5FB2DA6CFFDDFF8825CFF72CFDFAF879" box="[458,600,553,576]" pageId="57">G. G. Perrault</determinerName>
det. [printed] 1992&quot; [handwritten]. Female, &quot;Venezuela,
<collectorName id="0724537EFFDDFF8820B3F72CFAA8F879" box="[1206,1290,553,576]" pageId="57">S. Fern.</collectorName>
do
<collectingRegion id="6815F84AFFDDFF88253DF74CFEDFF859" box="[312,381,585,608]" country="Venezuela" name="Apure" pageId="57">Apure</collectingRegion>
,
<collectorName id="0724537EFFDDFF882589F74CFDA8F859" box="[396,522,585,608]" pageId="57">L. Laglaize</collectorName>
1896&quot;; &quot;
<collectingRegion id="6815F84AFFDDFF88266AF74CFC99F859" box="[623,827,585,608]" country="Venezuela" name="MUSEUM PARIS" pageId="57">MUSEUM PARIS</collectingRegion>
/ ex. Coll./
<collectorName id="0724537EFFDDFF8827C5F74CFBE4F859" box="[960,1094,585,608]" pageId="57">R. Oberthur</collectorName>
&quot; [pale green paper]; &quot;
<typeStatus id="756A880AFFDDFF882545F76CFE69F8B9" box="[320,459,617,640]" pageId="57" type="paratype">PARATYPE</typeStatus>
&quot; [red paper]; &quot;
<typeStatus id="756A880AFFDDFF88266EF76CFD50F8B9" box="[619,754,617,640]" pageId="57" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="6DD14D2BFFDDFF882706F76CFC3FF8B9" authorityName="Baudi di Selve. The" authorityYear="1864" box="[771,925,617,640]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="genus">Cymbionotum</taxonomicName>
/negrei m. [handwritten]/
<determinerName id="5FB2DA6CFFDDFF8820C5F76CFECBF899" pageId="57">G. G. Perrault</determinerName>
det. [printed] 1992&quot; [handwritten]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E2CB6523FFDDFF88250DF7C3FB38F9E8" pageId="57" type="materials_examined">
<paragraph id="AA6E36A8FFDDFF88250DF7C3FD02F8D8" blockId="57.[264,1323,710,1497]" box="[264,672,710,737]" pageId="57">
<materialsCitation id="1AB93CF5FFDDFF88250DF7C3FD3EF8D8" box="[264,668,710,737]" country="Venezuela" location="Bas Sarare" pageId="57" specimenCount="1" typeStatus="holotype">
<emphasis id="98A5EABAFFDDFF88250DF7C3FE28F8D9" bold="true" box="[264,394,710,736]" pageId="57">
<typeStatus id="756A880AFFDDFF88250DF7C3FEE7F8D9" box="[264,325,710,736]" pageId="57">Type</typeStatus>
area.
</emphasis>
<collectingCountry id="D2C67638FFDDFF882597F7C2FDADF8D8" box="[402,527,711,737]" name="Venezuela" pageId="57">Venezuela</collectingCountry>
:
<location id="AF0E6073FFDDFF88261FF7C2FD3EF8D8" LSID="urn:lsid:plazi:treatment:227887BEFFDDFF88250DF419FDF6FFE0:AF0E6073FFDDFF88261FF7C2FD3EF8D8" box="[538,668,711,737]" country="Venezuela" name="Bas Sarare" pageId="57">Bas Sarare</location>
</materialsCitation>
.
</paragraph>
<paragraph id="AA6E36A8FFDDFF88253DF7EBFB38F9E8" blockId="57.[264,1323,710,1497]" pageId="57">
<emphasis id="98A5EABAFFDDFF88253DF7EBFE77F931" bold="true" box="[312,469,750,776]" pageId="57">Recognition.</emphasis>
Adults of this species are markedly similar to those of
<emphasis id="98A5EABAFFDDFF882087F7EAFA88F931" box="[1154,1322,751,776]" italics="true" pageId="57">
<taxonomicName id="6DD14D2BFFDDFF882087F7EAFA84F931" authorityName="Ball &amp; Shpeley" authorityYear="2005" box="[1154,1318,751,776]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="fernandezi">C. fernandezi</taxonomicName>
,
</emphasis>
differing in absence of microlines from most of the dorsal surface, and by average smaller size, narrower vertex, narrower head in relation to pronotum, and narrower pronotal base (lower values for PWB/PWA,
<tableCitation id="E7530313FFDDFF88267EF662FD45F9B8" box="[635,743,871,897]" captionStart="TABLE 12" captionStartId="56.[264,352,1123,1147]" captionText="TABLE 12. Data about diagnostically useful measurements and ratios for the species of Procoscinia (C. fernandezi, n. sp. [N = 37] and C. negrei Perrault [N = 3])." pageId="57">Table 12</tableCitation>
). The phalli of these two species are similar in form, but that of
<taxonomicName id="6DD14D2BFFDDFF8825D2F68AFDE7F991" box="[471,581,911,936]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="98A5EABAFFDDFF8825D2F68AFDE7F991" box="[471,581,911,936]" italics="true" pageId="57">C. negrei</emphasis>
</taxonomicName>
(
<figureCitation id="32EA2A2DFFDDFF882652F68AFD1CF990" box="[599,702,911,937]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (AE) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; CD left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and BD and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="57">Fig. 15F</figureCitation>
) is narrower, with the preapical portion of the shaft bent more distinctly ventrad, and ventral surface generally pale (cf.
<figureCitation id="32EA2A2DFFDDFF882027F6B2FB2EF9E8" box="[1058,1164,951,977]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (AE) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; CD left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and BD and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="57">Fig. 15B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E2CB6523FFDDFF88253DF6DBFDCBFFB0" pageId="57" type="description">
<paragraph id="AA6E36A8FFDDFF88253DF6DBFAA9FE18" blockId="57.[264,1323,710,1497]" pageId="57">
<emphasis id="98A5EABAFFDDFF88253DF6DBFE72F9C1" bold="true" box="[312,464,990,1016]" pageId="57">Description.</emphasis>
With recognition features as specified above, and in the subgeneric treatment; similar generally to
<taxonomicName id="6DD14D2BFFDDFF882674F102FCADFE19" authorityName="Ball &amp; Shpeley" authorityYear="2005" box="[625,783,1031,1056]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="98A5EABAFFDDFF882674F102FCADFE19" box="[625,783,1031,1056]" italics="true" pageId="57">C. fernandezi</emphasis>
</taxonomicName>
. Male genitalia with phallus as in
<figureCitation id="32EA2A2DFFDDFF8820A6F102FAA5FE18" box="[1187,1287,1031,1057]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (AE) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; CD left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and BD and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="57">
Fig.
<specimenCount id="BCD7FD21FFDDFF8820DDF102FAA5FE18" box="[1240,1287,1031,1057]" pageId="57" type="generic">15F</specimenCount>
</figureCitation>
.
</paragraph>
<paragraph id="AA6E36A8FFDDFF88253DF12BFCBEFEF8" blockId="57.[264,1323,710,1497]" pageId="57">
<emphasis id="98A5EABAFFDDFF88253DF12BFE3DFE71" bold="true" box="[312,415,1070,1096]" pageId="57">Habitat.</emphasis>
The
<typeStatus id="756A880AFFDDFF8825E4F12AFDE5FE70" box="[481,583,1071,1097]" pageId="57" type="holotype">holotype</typeStatus>
was collected from under bark of a (presumably fallen) tree. From the locality data, we infer that
<taxonomicName id="6DD14D2BFFDDFF8826CAF152FC9CFE49" box="[719,830,1111,1136]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="98A5EABAFFDDFF8826CAF152FC9CFE49" box="[719,830,1111,1136]" italics="true" pageId="57">C. negrei</emphasis>
</taxonomicName>
is mesophilous, living in the rain forest vegetation zone (Table 17), at an altitude of less than
<specimenCount id="BCD7FD21FFDDFF882795F17AFC46FEA0" box="[912,996,1151,1177]" pageId="57" type="generic">
<quantity id="6D299B4DFFDDFF882795F17AFC42FEA0" box="[912,992,1151,1177]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="57" unit="m" value="200.0">200 m</quantity>
.
</specimenCount>
We do not believe that the species is subcortical, but that is a possibility.
</paragraph>
<paragraph id="AA6E36A8FFDDFF88253DF1CBFD71FF28" blockId="57.[264,1323,710,1497]" pageId="57">
<emphasis id="98A5EABAFFDDFF88253DF1CBFDDFFED1" bold="true" box="[312,637,1230,1256]" pageId="57">Geographical distribution</emphasis>
(
<figureCitation id="32EA2A2DFFDDFF88268AF1CAFD45FED0" box="[655,743,1231,1257]" captionStart="FIGURE 16" captionStartId="59.[264,368,1187,1211]" captionTargetBox="[284,1298,525,1160]" captionTargetId="figure-227@59.[284,1302,524,1161]" captionTargetPageId="59" captionText="FIGURE 16. Geographical range of the species of Cymbionotum (Procoscinia), and of Cymbionotum (s. str.) basale species group illustrated on an outline map of South America and of southwestern Asia, southern Europe and Africa." figureDoi="http://doi.org/10.5281/zenodo.5051718" httpUri="https://zenodo.org/record/5051718/files/figure.png" pageId="57">Fig. 16</figureCitation>
). This species is known only from two localities in the
<collectingRegion id="6815F84AFFDDFF882554F1F2FE38FF28" box="[337,410,1271,1297]" country="Venezuela" name="Apure" pageId="57">Apure</collectingRegion>
River Basin of
<collectingCountry id="D2C67638FFDDFF882656F1F2FD6DFF28" box="[595,719,1271,1297]" name="Venezuela" pageId="57">Venezuela</collectingCountry>
.
</paragraph>
<paragraph id="AA6E36A8FFDDFF88253DF01BFDCBFFB0" blockId="57.[264,1323,710,1497]" pageId="57">
<emphasis id="98A5EABAFFDDFF88253DF01BFC98FF01" bold="true" box="[312,826,1310,1336]" pageId="57">Relationships and chorological affinities.</emphasis>
Based on shared morphological features, geographical proximity, and allopatric ranges,
<taxonomicName id="6DD14D2BFFDDFF882729F042FC3AFF59" box="[812,920,1351,1376]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="98A5EABAFFDDFF882729F042FC3AFF59" box="[812,920,1351,1376]" italics="true" pageId="57">C. negrei</emphasis>
</taxonomicName>
is inferred to be the adelphotaxon of
<taxonomicName id="6DD14D2BFFDDFF88252DF06AFE67FFB1" authority="Ball &amp; Shpeley, 2005" authorityName="Ball &amp; Shpeley" authorityYear="2005" box="[296,453,1391,1416]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="57" phylum="Arthropoda" rank="species" species="fernandezi" status="sp. nov.">
<emphasis id="98A5EABAFFDDFF88252DF06AFE67FFB1" box="[296,453,1391,1416]" italics="true" pageId="57">C. fernandezi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="839657C1FFDDFF8825D0F06AFDC7FFB0" box="[469,613,1391,1417]" pageId="57" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="E2CB6523FFDDFF88253DF093FDF6FFE0" pageId="57" type="materials_examined">
<paragraph id="AA6E36A8FFDDFF88253DF093FDF6FFE0" blockId="57.[264,1323,710,1497]" pageId="57">
<emphasis id="98A5EABAFFDDFF88253DF093FD8CFF89" bold="true" box="[312,558,1430,1456]" pageId="57">Material examined</emphasis>
. We have seen the
<typeStatus id="756A880AFFDDFF882741F092FCD4FF88" box="[836,886,1431,1457]" pageId="57">type</typeStatus>
series, only. For details, see the synonymical section, above.
</paragraph>
</subSubSection>
</treatment>
</document>