treatments-xml/data/22/78/87/227887BEFFD3FF89250DF621FE72F9E7.xml

<document id="52BC0BEE8CF6CD0CCE21F4CB29CFC595" ID-DOI="10.11646/zootaxa.1099.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5051674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625110034160" checkinUser="felipe" docAuthor="Ball, George E. &amp; Shpeley, Danny" docDate="2005" docId="227887BEFFD3FF89250DF621FE72F9E7" docLanguage="en" docName="zt01099p060.pdf" docOrigin="Zootaxa 1099 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1099.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Cymbionotum (Procoscinia) fernandezi , Ball and Shpeley 2005, n. sp." docType="treatment" docVersion="4" lastPageNumber="120" masterDocId="DE41FFC6FFE4FFB12405F505FFA2FA39" masterDocTitle="Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution" masterLastPageNumber="120" masterPageNumber="1" pageNumber="1" updateTime="1698981561892" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7B2FEC964641EF36DC972E96D1CA5390">Taxonomic review of the Tribe Melaenini (Coleoptera: Carabidae), with observations on morphological, ecological and chorological evolution</mods:title>
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<mods:namePart id="817E9694952B67573B8343626A896F50">Ball, George E.</mods:namePart>
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<mods:namePart id="0C0B3DF1B3B8A1DD96B595ABCBA077B5">Shpeley, Danny</mods:namePart>
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<mods:date id="06921CE7CC0F33726F24128519A5D4DE">2005</mods:date>
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<mods:number id="1F132E41EDACE39411E07B4C8F568C27">2005-12-19</mods:number>
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<paragraph id="AA6E36A8FFD3FF86250DF621FBB7F907" blockId="55.[264,1045,804,865]" box="[264,1045,804,830]" pageId="55">
<heading id="F12681C4FFD3FF86250DF621FBB7F907" bold="true" box="[264,1045,804,830]" fontSize="11" level="1" pageId="55" reason="1">
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<taxonomicName id="6DD14D2BFFD3FF86250DF621FC64F907" ID-CoL="336P9" authority="Ball and Shpeley" authorityName=", Ball and Shpeley" authorityYear="2005" box="[264,966,804,830]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="55" phylum="Arthropoda" rank="species" species="fernandezi" status="sp. nov." subGenus="Procoscinia">
<emphasis id="98A5EABAFFD3FF86250DF621FD4FF907" bold="true" box="[264,749,804,830]" italics="true" pageId="55">Cymbionotum (Procoscinia) fernandezi,</emphasis>
Ball and Shpeley
</taxonomicName>
, 
<taxonomicNameLabel id="839657C1FFD3FF8627D6F621FBB7F907" box="[979,1045,804,830]" pageId="55" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="AA6E36A8FFD3FF86250DF642FC31F958" blockId="55.[264,1045,804,865]" box="[264,915,839,865]" pageId="55">
(
<figureCitation id="32EA2A2DFFD3FF862514F642FED5F958" box="[273,375,839,865]" captionStart="FIGURE 1" captionStartId="2.[264,367,1842,1866]" captionTargetBox="[267,1320,828,1823]" captionTargetId="figure-115@2.[264,1323,816,1827]" captionTargetPageId="2" captionText="FIGURE 1. Photographs illustrating dorsal habitus: A, Melaenus piger Fabricius (SBL 7.33 mm); B, Cymbionotum fernandezi, new species (SBL 3.96 mm)." figureDoi="http://doi.org/10.5281/zenodo.5051676" httpUri="https://zenodo.org/record/5051676/files/figure.png" pageId="55">Figs. 1B</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF862581F642FE7BF958" box="[388,473,839,865]" captionStart="FIGURE 2" captionStartId="10.[264,368,1069,1093]" captionTargetBox="[387,1205,291,1040]" captionTargetId="figure-282@10.[384,1208,289,1045]" captionTargetPageId="10" captionText="FIGURE 2. Elytra of Cymbionotum species. SEM photographs illustrating setal length. A, left elytron, dorsal aspect, of C. fernandezi, new species. B–F, portions of left elytra, dorsolateral aspect, of: B, C. pictulum Bates; C, C. semelederi Chaudoir; D, C. striatum Reitter, E, C. fernandezi, new species; and F, C. semirubricum Reitter. Scale bars: A = 1.0 mm; B–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051680" httpUri="https://zenodo.org/record/5051680/files/figure.png" pageId="55">2A, 2E</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF8625E0F642FDA6F958" box="[485,516,839,865]" captionStart="FIGURE 3" captionStartId="11.[264,367,1358,1382]" captionTargetBox="[457,1143,284,1314]" captionTargetId="figure-170@11.[454,1146,281,1317]" captionTargetPageId="11" captionText="FIGURE 3. Elytra of Cymbionotum species. SEM photographs of sections of elytral discs in anterior one third, illustrating density of elytral punctation: A, C. striatum Reitter; B, C. semelederi Chaudoir; C, C. schueppelii Dejean; D, C. pictulum Bates; E, C. semirubricum Reitter; and F, C. fernandezi, new species. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051682" httpUri="https://zenodo.org/record/5051682/files/figure.png" pageId="55">3F</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF862615F642FDFEF958" box="[528,604,839,865]" captionStart="FIGURE 12" captionStartId="50.[264,368,1370,1394]" captionTargetBox="[264,1323,449,1344]" captionTargetId="figure-297@50.[264,1323,449,1344]" captionTargetPageId="50" captionText="FIGURE 12. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A–H, mandibles: A, D, lateral aspect, left and right mandible, respectively; B, C, dorsal aspect, left and right mandible, respectively; E, H, occlusal aspect, left and right mandible, respectively; and F, G, ventral aspect, left and right mandible, respectively. I, J, labrum, dorsal and ventral aspect, respectively. K,L, left maxilla, ventral and dorsal aspects, respectively. Legend: aog, anterior occlusal groove; art, anterior retinacular tooth; as, apical seta; b; basal portion of mandible; bb; basal brush; bemr, basal extension of molar ridge; c, cardo; cr, crepis; cs, coeloconic sensillum; dgs, dorsal groove of scrobe; dls, dorsal lobe of stipes; eps, extra parapedial seta; g1, galeomere 1; g2, galeomere 2; it, incisor tooth; lc, lacinia; lct, lacinial tooth; lps, lateral parapedial seta; mp2, maxillary palpomere 2; mp3, maxillary palpomere 3; mp4, maxillary palpomere 4; mps, medial parapedial seta; mr, molar ridge; mt, molar tooth; mtr, microtrichia; omr, occlusal part of molar ridge; ped, pedium; pf, palpifer; pfs, palpiferal seta; pmr, premolar ridge; pmt, premolar tooth; pog, posterior occlusal groove; pp, parapedial projection; pr, parapedial ridge; prt, posterior retinacular tooth; pss, primary scrobal seta; rr, retinacular ridge; s, scrobe; sss, secondary scrobal seta; svg, secondary ventral groove; t, terebra; tr, terebral ridge; tt, terebral tooth; vd, ventral depression; vg, ventral groove; vls, ventral lobe of stipes. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051706" httpUri="https://zenodo.org/record/5051706/files/figure.png" pageId="55">12A–I</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF86266CF642FD1FF958" box="[617,701,839,865]" captionStart="FIGURE 13" captionStartId="51.[264,368,1369,1393]" captionTargetBox="[264,1323,288,1345]" captionTargetId="figure-261@51.[264,1323,288,1345]" captionTargetPageId="51" captionText="FIGURE 13. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A, labium, ventral aspect; B. paraglossa, dorsal aspect; C, labium, dorsal aspect; D, middle tibia, apical portion, anterior aspect; E, middle tarsomere 5, apical portion, and claws, ventral aspect; F–J, ovipositor, left gonocoxa––F, gonocoxa 1 and 2, lateral aspect; G–I, gonocoxite 1, apical portion and gonocoxite 2, lateral, dorsal and ventral aspects, respectively; J, gonocoxite 2, apical portion, oblique aspect. Legend: cs, coeloconic sensillum; ats, anterior tibial spur; avl, apical ventral lobe of t­5; css, clamshell sensillum; dts, dorsal trichoid seta; e, epilobe of mentum; es, ensiform seta; gc1, gonocoxite 1; gc2, gonocoxite 2; gs, glossal apical seta; gsap, glossal sclerite, apical portion; gsbp, glossal sclerite, basal portion; gsc, glossal sclerite; gsmp, glossal sclerite, medial portion; ifs, inner face sculpticell; ll, lateral lobe of mentum; lp1, labial palpomere 1; lp2, labial palpomere 2; lp3, labial palpomere 3; m, mentum; ms, paramedial seta of mentum; m­sms, mental­submental suture; mt, mental tooth; mtb, middle tibial brush; mtr, microtrichia; ns, nematiform seta; pg, paraglossa; pge, palpiger; psms, primary submental seta; sm, submentum; ssf, subapical sensory furrow; ssms, secondary submental seta; tc, tarsal claw; ts, trichoid seta; up, unguitractor plate; vls; ventrolateral seta. Scale bars: A, C– F, and I = 100 µm; B and J = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5051708" httpUri="https://zenodo.org/record/5051708/files/figure.png" pageId="55">13 A–J</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF8626CFF642FCA2F958" box="[714,768,839,865]" captionStart="FIGURE 14" captionStartId="52.[264,368,1082,1106]" captionTargetBox="[269,1315,595,1062]" captionTargetId="figure-321@52.[238,1357,569,1081]" captionTargetPageId="52" captionText="FIGURE 14. Pronotum, dorsal aspect, of Cymbionotum species, illustrating subgeneric differences with outline drawings. A, C. (Procoscinia) fernandezi, new species. B, C. (Cymbionotum) semelederi Chaudoir. Legend: pep, proepipleuron. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5051712" httpUri="https://zenodo.org/record/5051712/files/figure.png" pageId="55">14A</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF86270EF642FCFDF958" box="[779,863,839,865]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="55">15A–E</figureCitation>
, 
<figureCitation id="32EA2A2DFFD3FF862769F642FC28F958" box="[876,906,839,865]" captionStart="FIGURE 16" captionStartId="59.[264,368,1187,1211]" captionTargetBox="[284,1298,525,1160]" captionTargetId="figure-227@59.[284,1302,524,1161]" captionTargetPageId="59" captionText="FIGURE 16. Geographical range of the species of Cymbionotum (Procoscinia), and of Cymbionotum (s. str.) basale species group illustrated on an outline map of South America and of southwestern Asia, southern Europe and Africa." figureDoi="http://doi.org/10.5281/zenodo.5051718" httpUri="https://zenodo.org/record/5051718/files/figure.png" pageId="55">16</figureCitation>
)
</paragraph>
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<paragraph id="AA6E36A8FFD3FF86250DF693FB90FF80" blockId="55.[264,1325,918,1905]" pageId="55">
<emphasis id="98A5EABAFFD3FF86250DF693FE63F989" bold="true" box="[264,449,918,944]" pageId="55">Type material.</emphasis>
<materialsCitation id="1AB93CF5FFD3FF8625C8F692FB8CFF80" collectingDate="1999-05" collectingDateMax="1999-05-15" collectingDateMin="1999-05" collectorName="H. Moreno" country="Colombia" elevation="20" location="Thirty" municipality="Eighteen" pageId="55" specimenCount="73" specimenCount-female="32" specimenCount-male="20" typeStatus="holotype">
<location id="AF0E6073FFD3FF8625C8F692FDB4F988" LSID="urn:lsid:plazi:treatment:227887BEFFD3FF89250DF621FE72F9E7:AF0E6073FFD3FF8625C8F692FDB4F988" box="[461,534,919,945]" country="Colombia" municipality="Eighteen" name="Thirty" pageId="55">Thirty</location>
<specimenCount id="BCD7FD21FFD3FF862627F692FD43F988" box="[546,737,919,945]" pageId="55" type="generic">eight specimens</specimenCount>
(
<specimenCount id="BCD7FD21FFD3FF8626F0F692FCC1F988" box="[757,867,919,945]" pageId="55" type="male">20 males</specimenCount>
and 
<specimenCount id="BCD7FD21FFD3FF8627A1F692FB8FF988" box="[932,1069,919,945]" pageId="55" type="female">17 females</specimenCount>
). 
<typeStatus id="756A880AFFD3FF862040F692FB42F988" box="[1093,1248,919,945]" pageId="55" type="holotype">HOLOTYPE</typeStatus>
male, card­mounted, labeled: &quot; 
<collectingCountry id="D2C67638FFD3FF862647F6BAFD40F9E0" box="[578,738,959,985]" name="Colombia" pageId="55">COLOMBIA</collectingCountry>
Bolivar/ Zambrano bosque seco/ Lote Papayo cultivo/ 
<taxonomicName id="6DD14D2BFFD3FF862568F6E2FD9BFE38" box="[365,569,999,1025]" class="Magnoliopsida" family="Lamiaceae" genus="Gmelina" kingdom="Plantae" order="Lamiales" pageId="55" phylum="Tracheophyta" rank="species" species="arborea">Gmelina arborea</taxonomicName>
. Luz/ 
<date id="DE6F1068FFD3FF86268EF6E2FCE7FE38" box="[651,837,999,1025]" pageId="55" value="1999-05-10" valueMax="1999-05-15" valueMin="1999-05-10">
<collectingDate id="CE2BE980FFD3FF86268EF6E2FCE7FE38" box="[651,837,999,1025]" pageId="55" value="1999-05-10" valueMax="1999-05-15" valueMin="1999-05-10">10–15. V. 1999</collectingDate>
</date>
/Col. H.M.­M.A.­M.G./ No. 967597&quot;. A microvial, pinned beneath the whole specimen, contains its genitalia (UNCB). 
<typeStatus id="756A880AFFD3FF86250DF132FE0FFE68" box="[264,429,1079,1105]" pageId="55" type="paratype">PARATYPES</typeStatus>
, 4 labeled same as 
<typeStatus id="756A880AFFD3FF8626B5F132FCB9FE68" box="[688,795,1079,1105]" pageId="55" type="holotype">holotype</typeStatus>
: male, point­mounted, head and prothorax missing (broken and parts lost, following for habitus picture, 
<figureCitation id="32EA2A2DFFD3FF862003F15AFBC6FE40" box="[1030,1124,1119,1145]" captionStart="FIGURE 1" captionStartId="2.[264,367,1842,1866]" captionTargetBox="[267,1320,828,1823]" captionTargetId="figure-115@2.[264,1323,816,1827]" captionTargetPageId="2" captionText="FIGURE 1. Photographs illustrating dorsal habitus: A, Melaenus piger Fabricius (SBL 7.33 mm); B, Cymbionotum fernandezi, new species (SBL 3.96 mm)." figureDoi="http://doi.org/10.5281/zenodo.5051676" httpUri="https://zenodo.org/record/5051676/files/figure.png" pageId="55">Fig. 1B</figureCitation>
); female, pointmounted, reproductive tract removed, in microvial pinned beneath whole specimen; female, card­mounted, with left elytron removed, and attached to card; and female, cardmounted, disassembled for SEM photographs of mouthparts and ovipositor sclerites). 
<collectingMunicipality id="4A0AACD2FFD3FF86250DF1FAFED2FF20" box="[264,368,1279,1305]" pageId="55">Eighteen</collectingMunicipality>
males, 
<specimenCount id="BCD7FD21FFD3FF8625C8F1FAFDF3FF20" box="[461,593,1279,1305]" pageId="55" type="female">15 females</specimenCount>
, labeled: &quot; 
<collectingCountry id="D2C67638FFD3FF8626C9F1FAFCCEFF20" box="[716,876,1279,1305]" name="Colombia" pageId="55">COLOMBIA</collectingCountry>
Bolivar Zam­/brano. Hda. Monterrey For­/estal BsT 9¼42'N, 74¼53'W/ 
<specimenCount id="BCD7FD21FFD3FF8626B0F022FD53FF78" box="[693,753,1319,1345]" pageId="55" type="generic">
<quantity id="6D299B4DFFD3FF8626B0F022FD4FFF78" box="[693,749,1319,1345]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="55" unit="m" value="20.0">
<elevation id="21FCD19BFFD3FF8626B0F022FD4FFF78" box="[693,749,1319,1345]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="55" unit="m" value="20.0">20m</elevation>
</quantity>
.
</specimenCount>
<date id="DE6F1068FFD3FF8626F9F022FCDCFF78" box="[764,894,1319,1345]" pageId="55" value="1999-05">
<collectingDate id="CE2BE980FFD3FF8626F9F022FCDCFF78" box="[764,894,1319,1345]" pageId="55" value="1999-05">May 1999</collectingDate>
</date>
&quot;; &quot;Lote Papayo Interior de/ cultivo 
<taxonomicName id="6DD14D2BFFD3FF86250DF04AFE73FF51" box="[264,465,1359,1384]" class="Magnoliopsida" family="Lamiaceae" genus="Gmelina" kingdom="Plantae" order="Lamiales" pageId="55" phylum="Tracheophyta" rank="species" species="arborea">
<emphasis id="98A5EABAFFD3FF86250DF04AFE73FF51" box="[264,465,1359,1384]" italics="true" pageId="55">Gmelina arborea</emphasis>
</taxonomicName>
/ trampa de luz,/ 
<collectorName id="0724537EFFD3FF86269DF04AFCBBFF50" box="[664,793,1359,1385]" pageId="55">H. Moreno</collectorName>
y M. çvila/ No. 967543&quot; With the agreement of 
<collectorName id="0724537EFFD3FF86252FF072FE12FFA8" box="[298,432,1399,1425]" pageId="55">H. Moreno</collectorName>
, the 
<typeStatus id="756A880AFFD3FF8625EEF072FDF9FFA8" box="[491,603,1399,1425]" pageId="55" type="paratype">paratypes</typeStatus>
were distributed as follows: BMNH, 2; CASC, 2; CNIF, 7; IAVH, 7; MNHP, 2; MRAC, 2; UASM, 2; UNCB, 11; and UASM, 2
</materialsCitation>
.
</paragraph>
<paragraph id="AA6E36A8FFD3FF86253DF0C3FC7BFC08" blockId="55.[264,1325,918,1905]" pageId="55">
<emphasis id="98A5EABAFFD3FF86253DF0C3FDA0FFD9" bold="true" box="[312,514,1478,1504]" pageId="55">Specific epithet.</emphasis>
This Latinized eponym, genitive case, is based on the surname of Prof. Fernando Fernandez, who loaned the 
<typeStatus id="756A880AFFD3FF862703F0EAFC9AFC30" box="[774,824,1519,1545]" pageId="55">type</typeStatus>
specimens to us, and with whom we have cooperated in extending knowledge of Colombian 
<taxonomicName id="6DD14D2BFFD3FF86275FF312FC77FC08" authorityName="Latreille" authorityYear="1802" box="[858,981,1559,1585]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="55" phylum="Arthropoda" rank="family">Carabidae</taxonomicName>
.
</paragraph>
<paragraph id="AA6E36A8FFD3FF86253DF33BFCECFC60" blockId="55.[264,1325,918,1905]" box="[312,846,1598,1625]" pageId="55">
<materialsCitation id="1AB93CF5FFD3FF86253DF33BFCE8FC60" box="[312,842,1598,1625]" country="Colombia" location="Bolivar" municipality="Zambrano" pageId="55" specimenCount="1" typeStatus="holotype">
<emphasis id="98A5EABAFFD3FF86253DF33BFE7FFC61" bold="true" box="[312,477,1598,1624]" pageId="55">
<typeStatus id="756A880AFFD3FF86253DF33BFED7FC61" box="[312,373,1598,1624]" pageId="55">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D2C67638FFD3FF8625E1F33AFDFEFC60" box="[484,604,1599,1625]" name="Colombia" pageId="55">Colombia</collectingCountry>
: 
<location id="AF0E6073FFD3FF862662F33AFD66FC60" LSID="urn:lsid:plazi:treatment:227887BEFFD3FF89250DF621FE72F9E7:AF0E6073FFD3FF862662F33AFD66FC60" box="[615,708,1599,1625]" country="Colombia" municipality="Zambrano" name="Bolivar" pageId="55">Bolivar</location>
: 
<collectingMunicipality id="4A0AACD2FFD3FF8626CBF33AFCE8FC60" box="[718,842,1599,1625]" pageId="55">Zambrano</collectingMunicipality>
</materialsCitation>
.
</paragraph>
<paragraph id="AA6E36A8FFD3FF86253DF363FBC0FD18" blockId="55.[264,1325,918,1905]" pageId="55">
<emphasis id="98A5EABAFFD3FF86253DF363FE77FCB9" bold="true" box="[312,469,1638,1664]" pageId="55">Recognition.</emphasis>
Compared to 
<taxonomicName id="6DD14D2BFFD3FF862686F362FD53FCB9" box="[643,753,1639,1664]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="55" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="98A5EABAFFD3FF862686F362FD53FCB9" box="[643,753,1639,1664]" italics="true" pageId="55">C. negrei</emphasis>
</taxonomicName>
, adults of 
<taxonomicName id="6DD14D2BFFD3FF862770F362FBB4FCB9" authorityName=", Ball and Shpeley" authorityYear="2005" box="[885,1046,1639,1664]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="55" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="98A5EABAFFD3FF862770F362FBB4FCB9" box="[885,1046,1639,1664]" italics="true" pageId="55">C. fernandezi</emphasis>
</taxonomicName>
are on average, larger, both in length and width, with wider vertex, wider head in relation to pronotum, and wider pronotal base (
<tableCitation id="E7530313FFD3FF8625B7F3B2FDB9FCE8" box="[434,539,1719,1745]" captionStart="TABLE 12" captionStartId="56.[264,352,1123,1147]" captionText="TABLE 12. Data about diagnostically useful measurements and ratios for the species of Procoscinia (C. fernandezi, n. sp. [N = 37] and C. negrei Perrault [N = 3])." pageId="55">Table 12</tableCitation>
). The phalli (
<figureCitation id="32EA2A2DFFD3FF8626B3F3B2FCE8FCE8" box="[694,842,1719,1745]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="55">Figs. 15D, F</figureCitation>
) of these two species are similar in form, but that of 
<taxonomicName id="6DD14D2BFFD3FF86258CF3DAFD8AFCC1" authorityName=", Ball and Shpeley" authorityYear="2005" box="[393,552,1759,1784]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="55" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="98A5EABAFFD3FF86258CF3DAFD8AFCC1" box="[393,552,1759,1784]" italics="true" pageId="55">C. fernandezi</emphasis>
</taxonomicName>
(
<figureCitation id="32EA2A2DFFD3FF86263DF3DAFD02FCC0" box="[568,672,1759,1785]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="55">Fig. 15B</figureCitation>
) is broader, with the preapical portion of the shaft bent more gradually ventrad, and ventral surface generally dark (cf. 
<figureCitation id="32EA2A2DFFD3FF8627EBF202FBF6FD18" box="[1006,1108,1799,1825]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="55">Fig. 15F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="E2CB6523FFD3FF89253DF22BFE7CF9B7" lastPageId="56" pageId="55" type="description">
<paragraph id="AA6E36A8FFD3FF89253DF22BFBD6FBEF" blockId="55.[264,1325,918,1905]" lastBlockId="56.[264,1323,284,990]" lastPageId="56" pageId="55">
<emphasis id="98A5EABAFFD3FF86253DF22BFE72FD71" bold="true" box="[312,464,1838,1864]" pageId="55">Description.</emphasis>
With recognition features as noted above, and other details specified in the subgeneric description, and illustrations, as follows: habitus, 
<figureCitation id="32EA2A2DFFD3FF862010F252FBD3FD48" box="[1045,1137,1879,1905]" captionStart="FIGURE 1" captionStartId="2.[264,367,1842,1866]" captionTargetBox="[267,1320,828,1823]" captionTargetId="figure-115@2.[264,1323,816,1827]" captionTargetPageId="2" captionText="FIGURE 1. Photographs illustrating dorsal habitus: A, Melaenus piger Fabricius (SBL 7.33 mm); B, Cymbionotum fernandezi, new species (SBL 3.96 mm)." figureDoi="http://doi.org/10.5281/zenodo.5051676" httpUri="https://zenodo.org/record/5051676/files/figure.png" pageId="55">Fig. 1B</figureCitation>
; integumentary setal length, 
<figureCitation id="32EA2A2DFFDCFF8925A5F419FE59FB0F" box="[416,507,284,310]" captionStart="FIGURE 2" captionStartId="10.[264,368,1069,1093]" captionTargetBox="[387,1205,291,1040]" captionTargetId="figure-282@10.[384,1208,289,1045]" captionTargetPageId="10" captionText="FIGURE 2. Elytra of Cymbionotum species. SEM photographs illustrating setal length. A, left elytron, dorsal aspect, of C. fernandezi, new species. B–F, portions of left elytra, dorsolateral aspect, of: B, C. pictulum Bates; C, C. semelederi Chaudoir; D, C. striatum Reitter, E, C. fernandezi, new species; and F, C. semirubricum Reitter. Scale bars: A = 1.0 mm; B–F = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051680" httpUri="https://zenodo.org/record/5051680/files/figure.png" pageId="56">Fig. 2E</figureCitation>
; density of elytral punctation, 
<figureCitation id="32EA2A2DFFDCFF892762F419FC1DFB0F" box="[871,959,284,310]" captionStart="FIGURE 3" captionStartId="11.[264,367,1358,1382]" captionTargetBox="[457,1143,284,1314]" captionTargetId="figure-170@11.[454,1146,281,1317]" captionTargetPageId="11" captionText="FIGURE 3. Elytra of Cymbionotum species. SEM photographs of sections of elytral discs in anterior one third, illustrating density of elytral punctation: A, C. striatum Reitter; B, C. semelederi Chaudoir; C, C. schueppelii Dejean; D, C. pictulum Bates; E, C. semirubricum Reitter; and F, C. fernandezi, new species. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051682" httpUri="https://zenodo.org/record/5051682/files/figure.png" pageId="56">Fig. 3F</figureCitation>
; labrum and epipharynx, 
<figureCitation id="32EA2A2DFFDCFF8920F4F419FEDCFB67" captionStart="FIGURE 12" captionStartId="50.[264,368,1370,1394]" captionTargetBox="[264,1323,449,1344]" captionTargetId="figure-297@50.[264,1323,449,1344]" captionTargetPageId="50" captionText="FIGURE 12. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A–H, mandibles: A, D, lateral aspect, left and right mandible, respectively; B, C, dorsal aspect, left and right mandible, respectively; E, H, occlusal aspect, left and right mandible, respectively; and F, G, ventral aspect, left and right mandible, respectively. I, J, labrum, dorsal and ventral aspect, respectively. K,L, left maxilla, ventral and dorsal aspects, respectively. Legend: aog, anterior occlusal groove; art, anterior retinacular tooth; as, apical seta; b; basal portion of mandible; bb; basal brush; bemr, basal extension of molar ridge; c, cardo; cr, crepis; cs, coeloconic sensillum; dgs, dorsal groove of scrobe; dls, dorsal lobe of stipes; eps, extra parapedial seta; g1, galeomere 1; g2, galeomere 2; it, incisor tooth; lc, lacinia; lct, lacinial tooth; lps, lateral parapedial seta; mp2, maxillary palpomere 2; mp3, maxillary palpomere 3; mp4, maxillary palpomere 4; mps, medial parapedial seta; mr, molar ridge; mt, molar tooth; mtr, microtrichia; omr, occlusal part of molar ridge; ped, pedium; pf, palpifer; pfs, palpiferal seta; pmr, premolar ridge; pmt, premolar tooth; pog, posterior occlusal groove; pp, parapedial projection; pr, parapedial ridge; prt, posterior retinacular tooth; pss, primary scrobal seta; rr, retinacular ridge; s, scrobe; sss, secondary scrobal seta; svg, secondary ventral groove; t, terebra; tr, terebral ridge; tt, terebral tooth; vd, ventral depression; vg, ventral groove; vls, ventral lobe of stipes. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051706" httpUri="https://zenodo.org/record/5051706/files/figure.png" pageId="56">Figs. 12I and J</figureCitation>
, respectively; mandibles, 
<figureCitation id="32EA2A2DFFDCFF8926B9F441FCFBFB67" box="[700,857,324,350]" captionStart="FIGURE 12" captionStartId="50.[264,368,1370,1394]" captionTargetBox="[264,1323,449,1344]" captionTargetId="figure-297@50.[264,1323,449,1344]" captionTargetPageId="50" captionText="FIGURE 12. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A–H, mandibles: A, D, lateral aspect, left and right mandible, respectively; B, C, dorsal aspect, left and right mandible, respectively; E, H, occlusal aspect, left and right mandible, respectively; and F, G, ventral aspect, left and right mandible, respectively. I, J, labrum, dorsal and ventral aspect, respectively. K,L, left maxilla, ventral and dorsal aspects, respectively. Legend: aog, anterior occlusal groove; art, anterior retinacular tooth; as, apical seta; b; basal portion of mandible; bb; basal brush; bemr, basal extension of molar ridge; c, cardo; cr, crepis; cs, coeloconic sensillum; dgs, dorsal groove of scrobe; dls, dorsal lobe of stipes; eps, extra parapedial seta; g1, galeomere 1; g2, galeomere 2; it, incisor tooth; lc, lacinia; lct, lacinial tooth; lps, lateral parapedial seta; mp2, maxillary palpomere 2; mp3, maxillary palpomere 3; mp4, maxillary palpomere 4; mps, medial parapedial seta; mr, molar ridge; mt, molar tooth; mtr, microtrichia; omr, occlusal part of molar ridge; ped, pedium; pf, palpifer; pfs, palpiferal seta; pmr, premolar ridge; pmt, premolar tooth; pog, posterior occlusal groove; pp, parapedial projection; pr, parapedial ridge; prt, posterior retinacular tooth; pss, primary scrobal seta; rr, retinacular ridge; s, scrobe; sss, secondary scrobal seta; svg, secondary ventral groove; t, terebra; tr, terebral ridge; tt, terebral tooth; vd, ventral depression; vg, ventral groove; vls, ventral lobe of stipes. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051706" httpUri="https://zenodo.org/record/5051706/files/figure.png" pageId="56">Figs. 12A–H</figureCitation>
; maxillae, 
<figureCitation id="32EA2A2DFFDCFF8927E7F441FBDEFB67" box="[994,1148,324,350]" captionStart="FIGURE 12" captionStartId="50.[264,368,1370,1394]" captionTargetBox="[264,1323,449,1344]" captionTargetId="figure-297@50.[264,1323,449,1344]" captionTargetPageId="50" captionText="FIGURE 12. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A–H, mandibles: A, D, lateral aspect, left and right mandible, respectively; B, C, dorsal aspect, left and right mandible, respectively; E, H, occlusal aspect, left and right mandible, respectively; and F, G, ventral aspect, left and right mandible, respectively. I, J, labrum, dorsal and ventral aspect, respectively. K,L, left maxilla, ventral and dorsal aspects, respectively. Legend: aog, anterior occlusal groove; art, anterior retinacular tooth; as, apical seta; b; basal portion of mandible; bb; basal brush; bemr, basal extension of molar ridge; c, cardo; cr, crepis; cs, coeloconic sensillum; dgs, dorsal groove of scrobe; dls, dorsal lobe of stipes; eps, extra parapedial seta; g1, galeomere 1; g2, galeomere 2; it, incisor tooth; lc, lacinia; lct, lacinial tooth; lps, lateral parapedial seta; mp2, maxillary palpomere 2; mp3, maxillary palpomere 3; mp4, maxillary palpomere 4; mps, medial parapedial seta; mr, molar ridge; mt, molar tooth; mtr, microtrichia; omr, occlusal part of molar ridge; ped, pedium; pf, palpifer; pfs, palpiferal seta; pmr, premolar ridge; pmt, premolar tooth; pog, posterior occlusal groove; pp, parapedial projection; pr, parapedial ridge; prt, posterior retinacular tooth; pss, primary scrobal seta; rr, retinacular ridge; s, scrobe; sss, secondary scrobal seta; svg, secondary ventral groove; t, terebra; tr, terebral ridge; tt, terebral tooth; vd, ventral depression; vg, ventral groove; vls, ventral lobe of stipes. Scale bars = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.5051706" httpUri="https://zenodo.org/record/5051706/files/figure.png" pageId="56">Figs. 12K–L</figureCitation>
; labium, 
<figureCitation id="32EA2A2DFFDCFF8920F4F441FEFCFBBF" captionStart="FIGURE 13" captionStartId="51.[264,368,1369,1393]" captionTargetBox="[264,1323,288,1345]" captionTargetId="figure-261@51.[264,1323,288,1345]" captionTargetPageId="51" captionText="FIGURE 13. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A, labium, ventral aspect; B. paraglossa, dorsal aspect; C, labium, dorsal aspect; D, middle tibia, apical portion, anterior aspect; E, middle tarsomere 5, apical portion, and claws, ventral aspect; F–J, ovipositor, left gonocoxa––F, gonocoxa 1 and 2, lateral aspect; G–I, gonocoxite 1, apical portion and gonocoxite 2, lateral, dorsal and ventral aspects, respectively; J, gonocoxite 2, apical portion, oblique aspect. Legend: cs, coeloconic sensillum; ats, anterior tibial spur; avl, apical ventral lobe of t­5; css, clamshell sensillum; dts, dorsal trichoid seta; e, epilobe of mentum; es, ensiform seta; gc1, gonocoxite 1; gc2, gonocoxite 2; gs, glossal apical seta; gsap, glossal sclerite, apical portion; gsbp, glossal sclerite, basal portion; gsc, glossal sclerite; gsmp, glossal sclerite, medial portion; ifs, inner face sculpticell; ll, lateral lobe of mentum; lp1, labial palpomere 1; lp2, labial palpomere 2; lp3, labial palpomere 3; m, mentum; ms, paramedial seta of mentum; m­sms, mental­submental suture; mt, mental tooth; mtb, middle tibial brush; mtr, microtrichia; ns, nematiform seta; pg, paraglossa; pge, palpiger; psms, primary submental seta; sm, submentum; ssf, subapical sensory furrow; ssms, secondary submental seta; tc, tarsal claw; ts, trichoid seta; up, unguitractor plate; vls; ventrolateral seta. Scale bars: A, C– F, and I = 100 µm; B and J = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5051708" httpUri="https://zenodo.org/record/5051708/files/figure.png" pageId="56">Figs. 13A–C</figureCitation>
; pronotum, dorsal aspect, 
<figureCitation id="32EA2A2DFFDCFF8926B5F469FC81FBBF" box="[688,803,364,390]" captionStart="FIGURE 14" captionStartId="52.[264,368,1082,1106]" captionTargetBox="[269,1315,595,1062]" captionTargetId="figure-321@52.[238,1357,569,1081]" captionTargetPageId="52" captionText="FIGURE 14. Pronotum, dorsal aspect, of Cymbionotum species, illustrating subgeneric differences with outline drawings. A, C. (Procoscinia) fernandezi, new species. B, C. (Cymbionotum) semelederi Chaudoir. Legend: pep, proepipleuron. Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5051712" httpUri="https://zenodo.org/record/5051712/files/figure.png" pageId="56">Fig. 14A</figureCitation>
; hind wings, 
<figureCitation id="32EA2A2DFFDCFF8927D3F469FBEAFBBF" box="[982,1096,364,390]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="56">Fig. 15A</figureCitation>
; middle tibia and tarsomere 5, 
<figureCitation id="32EA2A2DFFDCFF8925A0F491FDCFFB97" box="[421,621,404,430]" captionStart="FIGURE 13" captionStartId="51.[264,368,1369,1393]" captionTargetBox="[264,1323,288,1345]" captionTargetId="figure-261@51.[264,1323,288,1345]" captionTargetPageId="51" captionText="FIGURE 13. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A, labium, ventral aspect; B. paraglossa, dorsal aspect; C, labium, dorsal aspect; D, middle tibia, apical portion, anterior aspect; E, middle tarsomere 5, apical portion, and claws, ventral aspect; F–J, ovipositor, left gonocoxa––F, gonocoxa 1 and 2, lateral aspect; G–I, gonocoxite 1, apical portion and gonocoxite 2, lateral, dorsal and ventral aspects, respectively; J, gonocoxite 2, apical portion, oblique aspect. Legend: cs, coeloconic sensillum; ats, anterior tibial spur; avl, apical ventral lobe of t­5; css, clamshell sensillum; dts, dorsal trichoid seta; e, epilobe of mentum; es, ensiform seta; gc1, gonocoxite 1; gc2, gonocoxite 2; gs, glossal apical seta; gsap, glossal sclerite, apical portion; gsbp, glossal sclerite, basal portion; gsc, glossal sclerite; gsmp, glossal sclerite, medial portion; ifs, inner face sculpticell; ll, lateral lobe of mentum; lp1, labial palpomere 1; lp2, labial palpomere 2; lp3, labial palpomere 3; m, mentum; ms, paramedial seta of mentum; m­sms, mental­submental suture; mt, mental tooth; mtb, middle tibial brush; mtr, microtrichia; ns, nematiform seta; pg, paraglossa; pge, palpiger; psms, primary submental seta; sm, submentum; ssf, subapical sensory furrow; ssms, secondary submental seta; tc, tarsal claw; ts, trichoid seta; up, unguitractor plate; vls; ventrolateral seta. Scale bars: A, C– F, and I = 100 µm; B and J = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5051708" httpUri="https://zenodo.org/record/5051708/files/figure.png" pageId="56">Figs. 13D and E</figureCitation>
, respectively; phallus and endophallic armature, 
<figureCitation id="32EA2A2DFFDCFF8920BCF491FA87FB97" box="[1209,1317,404,430]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="56">Fig. 15B</figureCitation>
; gonocoxa of ovipositor, 
<figureCitation id="32EA2A2DFFDCFF892620F4B9FD12FBEF" box="[549,688,444,470]" captionStart="FIGURE 13" captionStartId="51.[264,368,1369,1393]" captionTargetBox="[264,1323,288,1345]" captionTargetId="figure-261@51.[264,1323,288,1345]" captionTargetPageId="51" captionText="FIGURE 13. Structural features of Cymbionotum fernandezi, new species, illustrated with SEM photographs. A, labium, ventral aspect; B. paraglossa, dorsal aspect; C, labium, dorsal aspect; D, middle tibia, apical portion, anterior aspect; E, middle tarsomere 5, apical portion, and claws, ventral aspect; F–J, ovipositor, left gonocoxa––F, gonocoxa 1 and 2, lateral aspect; G–I, gonocoxite 1, apical portion and gonocoxite 2, lateral, dorsal and ventral aspects, respectively; J, gonocoxite 2, apical portion, oblique aspect. Legend: cs, coeloconic sensillum; ats, anterior tibial spur; avl, apical ventral lobe of t­5; css, clamshell sensillum; dts, dorsal trichoid seta; e, epilobe of mentum; es, ensiform seta; gc1, gonocoxite 1; gc2, gonocoxite 2; gs, glossal apical seta; gsap, glossal sclerite, apical portion; gsbp, glossal sclerite, basal portion; gsc, glossal sclerite; gsmp, glossal sclerite, medial portion; ifs, inner face sculpticell; ll, lateral lobe of mentum; lp1, labial palpomere 1; lp2, labial palpomere 2; lp3, labial palpomere 3; m, mentum; ms, paramedial seta of mentum; m­sms, mental­submental suture; mt, mental tooth; mtb, middle tibial brush; mtr, microtrichia; ns, nematiform seta; pg, paraglossa; pge, palpiger; psms, primary submental seta; sm, submentum; ssf, subapical sensory furrow; ssms, secondary submental seta; tc, tarsal claw; ts, trichoid seta; up, unguitractor plate; vls; ventrolateral seta. Scale bars: A, C– F, and I = 100 µm; B and J = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5051708" httpUri="https://zenodo.org/record/5051708/files/figure.png" pageId="56">Figs. 13F–J</figureCitation>
; internal genitalia of female, 
<figureCitation id="32EA2A2DFFDCFF89200DF4B9FBD2FBEF" box="[1032,1136,444,470]" captionStart="FIGURE 15" captionStartId="53.[264,367,1547,1571]" captionTargetBox="[287,1298,410,1527]" captionTargetId="figure-193@53.[263,1333,377,1541]" captionTargetPageId="53" captionText="FIGURE 15. Structural features of Cymbionotum fernandezi, new species (A–E) and C. negrei Perrault (F) illustrated with line drawings. A, left hind wing, dorsal aspect. B, male genitalia: phallus, left lateral aspect, with endophallus partially everted; C–D left and right paramere; respectively, ventral aspect. E, female genitalia, ventral aspect. F, male genitalia, left lateral aspect. Legend: wing veins and cells­­ AA1+ 2, anal vein 1 + 2; MP3 and MP4, posterior median veins 3 and 4, respectively; o, oblongum cell; RA4, anterior radial vein 4; RP2, posterior radial vein. Male genitalia: eph, endophallus; ph, phallus; x, sclerite x of endophallus. Female genitalia: bc, bursa copulatrix; co, common oviduct; di, spermathecal diverticulum; gc 1 and 2, gonocoxite 1 and 2, respectively; lt, laterotergite of abdominal segment IX; sp, spermatheca; spd, spermathecal duct. Scale bars: A and E = 1.0 mm; and B–D and F, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5051714" httpUri="https://zenodo.org/record/5051714/files/figure.png" pageId="56">Fig. 15E</figureCitation>
.
</paragraph>
<paragraph id="AA6E36A8FFDCFF89253DF4E6FEE3F8FF" blockId="56.[264,1323,284,990]" pageId="56">
<emphasis id="98A5EABAFFDCFF89253DF4E6FCE2FBC4" bold="true" box="[312,832,483,509]" pageId="56">Habitat, habits, and seasonal occurrence.</emphasis>
All specimens were collected in May, at an ultraviolet light, within a plantation of an exotic tree 
<emphasis id="98A5EABAFFDCFF8927B4F709FC18F81C" box="[945,954,524,549]" italics="true" pageId="56">(</emphasis>
<taxonomicName id="6DD14D2BFFDCFF8927BFF709FB79F81F" authority="Roxb." authorityName="Roxb." box="[954,1243,524,550]" class="Magnoliopsida" family="Lamiaceae" genus="Gmelina" kingdom="Plantae" order="Lamiales" pageId="56" phylum="Tracheophyta" rank="species" species="arborea">
<emphasis id="98A5EABAFFDCFF8927BFF709FB25F81C" box="[954,1159,524,549]" italics="true" pageId="56">Gmelina arborea</emphasis>
Roxb.
</taxonomicName>
, white teak, family 
<taxonomicName id="6DD14D2BFFDCFF8925ADF731FDE3F877" box="[424,577,564,590]" class="Magnoliopsida" family="Verbenaceae" kingdom="Plantae" order="Lamiales" pageId="56" phylum="Tracheophyta" rank="family">Verbenaceae</taxonomicName>
), a species indigenous in the Oriental Region and southern 
<collectingCountry id="D2C67638FFDCFF89250DF759FEF3F84F" box="[264,337,604,630]" name="China" pageId="56">China</collectingCountry>
. We infer that 
<taxonomicName id="6DD14D2BFFDCFF8925F8F759FD39F84C" authorityName=", Ball and Shpeley" authorityYear="2005" box="[509,667,604,629]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="56" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="98A5EABAFFDCFF8925F8F759FD39F84C" box="[509,667,604,629]" italics="true" pageId="56">C. fernandezi</emphasis>
</taxonomicName>
is mesophilic, occupying disturbed areas in the tropical rain forest zone (Table 17, Fig 24), at less than 
<quantity id="6D299B4DFFDCFF892744F781FC2FF8A7" box="[833,909,644,670]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="56" unit="m" value="200.0">200 m</quantity>
altitude with adults active in midyear.
</paragraph>
<paragraph id="AA6E36A8FFDCFF89253DF7D1FDC3F92F" blockId="56.[264,1323,284,990]" pageId="56">
<emphasis id="98A5EABAFFDCFF89253DF7D1FD26F8D7" bold="true" box="[312,644,724,750]" pageId="56">Geographical distribution</emphasis>
(
<figureCitation id="32EA2A2DFFDCFF892699F7D1FD59F8D7" box="[668,763,724,750]" captionStart="FIGURE 16" captionStartId="59.[264,368,1187,1211]" captionTargetBox="[284,1298,525,1160]" captionTargetId="figure-227@59.[284,1302,524,1161]" captionTargetPageId="59" captionText="FIGURE 16. Geographical range of the species of Cymbionotum (Procoscinia), and of Cymbionotum (s. str.) basale species group illustrated on an outline map of South America and of southwestern Asia, southern Europe and Africa." figureDoi="http://doi.org/10.5281/zenodo.5051718" httpUri="https://zenodo.org/record/5051718/files/figure.png" pageId="56">Fig. 16</figureCitation>
). This species is known only from the Rio 
<collectingRegion id="6815F84AFFDCFF89250DF7F9FE28F92F" box="[264,394,764,790]" country="Colombia" name="Magdalena" pageId="56">Magdalena</collectingRegion>
valley, 
<collectingCountry id="D2C67638FFDCFF8925E3F7F9FDFFF92F" box="[486,605,764,790]" name="Colombia" pageId="56">Colombia</collectingCountry>
.
</paragraph>
<paragraph id="AA6E36A8FFDCFF89253DF621FE7CF9B7" blockId="56.[264,1323,284,990]" pageId="56">
<emphasis id="98A5EABAFFDCFF89253DF621FC98F907" bold="true" box="[312,826,804,830]" pageId="56">Relationships and chorological affinities.</emphasis>
Based on shared morphological features, geographical distribution and allopatric ranges, 
<taxonomicName id="6DD14D2BFFDCFF89273FF649FC7BF95C" authorityName=", Ball and Shpeley" authorityYear="2005" box="[826,985,844,869]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="56" phylum="Arthropoda" rank="species" species="fernandezi">
<emphasis id="98A5EABAFFDCFF89273FF649FC7BF95C" box="[826,985,844,869]" italics="true" pageId="56">C. fernandezi</emphasis>
</taxonomicName>
and 
<taxonomicName id="6DD14D2BFFDCFF892017F649FBDDF95C" box="[1042,1151,844,869]" class="Insecta" family="Carabidae" genus="Cymbionotum" kingdom="Animalia" order="Coleoptera" pageId="56" phylum="Arthropoda" rank="species" species="negrei">
<emphasis id="98A5EABAFFDCFF892017F649FBDDF95C" box="[1042,1151,844,869]" italics="true" pageId="56">C. negrei</emphasis>
</taxonomicName>
are postulated to be adelphotaxa.
</paragraph>
</subSubSection>
<subSubSection id="E2CB6523FFDCFF89253DF699FE72F9E7" pageId="56" type="materials_examined">
<paragraph id="AA6E36A8FFDCFF89253DF699FE72F9E7" blockId="56.[264,1323,284,990]" pageId="56">
<emphasis id="98A5EABAFFDCFF89253DF699FD91F98F" bold="true" box="[312,563,924,950]" pageId="56">Material examined.</emphasis>
We have seen the 
<typeStatus id="756A880AFFDCFF892736F699FCC7F98F" box="[819,869,924,950]" pageId="56">type</typeStatus>
series, only. For details, see &quot; 
<typeStatus id="756A880AFFDCFF8920E8F699FA89F98F" box="[1261,1323,924,950]" pageId="56">Type</typeStatus>
material&quot;, above.
</paragraph>
</subSubSection>
</treatment>
</document>