170 lines
20 KiB
XML
170 lines
20 KiB
XML
<document id="84538476911CD9251A8CB6BD701FD418" ENCODING="UTF8" ID-GBIF-Dataset="d65d91f3-2aa6-4824-85a3-9ac72a57c596" ModsDocID="22820" checkinTime="1251954458996" checkinUser="christiana" docAuthor="Longino, J. T." docDate="2009" docId="17E8EF5608AC3CC4B8211652EE29481C" docLanguage="en" docName="22820_tx.xml" docOrigin="Zootaxa 2181" docSource="http://hol.osu.edu/reference-full.html?id=22820" docTitle="Pheidole eowilsoni Longino, 2009, new species" docType="treatment" docVersion="12" lastPageNumber="29" masterDocId="7FC56964C9056E7DBD20595E152A29E1" masterDocTitle="Additions to the taxonomy of New World Pheidole (Hymenoptera: Formicidae)." masterLastPageNumber="90" masterPageNumber="1" pageNumber="26" updateTime="1693272131770" updateUser="admin">
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<mods:title id="512E573EAC5DAE1EE9A8D972344BBEE0">Additions to the taxonomy of New World Pheidole (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="42E1FBB94515BE0FF5A20CE327E14A63">Longino, J. T.</mods:namePart>
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<mods:typeOfResource id="7DA36B7581DB027B4974C078A2B14BCD">text</mods:typeOfResource>
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<mods:part id="B6A54494127066C4818DF80FA21D9850">
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<mods:date id="AFE65F500B59B91989A3290395933E5B">2009</mods:date>
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<mods:classification id="0BB0559F3EA9CE7F09331E001E70B5C4">journal article</mods:classification>
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<treatment id="17E8EF5608AC3CC4B8211652EE29481C" ID-DOI="http://doi.org/10.5281/zenodo.6222825" ID-GBIF-Taxon="100121381" ID-Zenodo-Dep="6222825" LSID="urn:lsid:zoobank.org:act:EEE26378-7666-404B-93C6-71A0DED667D2" httpUri="http://treatment.plazi.org/id/17E8EF5608AC3CC4B8211652EE29481C" lastPageNumber="29" pageNumber="26">
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<subSubSection id="29EDFDDB390342BBCEB1AA53B0887B11" type="nomenclature">
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<paragraph id="204540C81F09D993540F86F1FE544F43" pageNumber="26">
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<taxonomicName id="A8328816CE9A3A8B7AF07275AFB1CDA9" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253257" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole eowilsoni Longino" order="Hymenoptera" pageNumber="26" phylum="Arthropoda" rank="species" species="eowilsoni">Pheidole eowilsoni</taxonomicName>
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<taxonomicNameLabel id="AE7C428D0A1A8882A6BE42E06F701510" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="B0FDA3D82AF4D0EA723EF64991F1393E" type="description">
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<paragraph id="297A437424E8470D6AD8123B28FDABAB" pageNumber="26">Figure 5</paragraph>
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</subSubSection>
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<subSubSection id="1D3436BC7C03DD23CBA257E7B6CD30F5" type="materials_examined">
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<paragraph id="88B8BC630BF8E393206987AC1A1385D2" pageNumber="26">
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Holotype major worker. Costa Rica, Alajuela: Casa Eladio, Rio Penas Blancas,
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<geoCoordinate id="0211C1853ECEA0B9029CD79DF3C1BE64" direction="north" orientation="latitude" value="10.31667">10.31667°N</geoCoordinate>
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<geoCoordinate id="A0BCD1371870F3E40866E87FFFEB8813" direction="west" orientation="longitude" value="-84.71667">84.71667°W</geoCoordinate>
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, ±2000m, 800m, 2 Mar 2008 (J. Longino#6149) [
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<collectionCode id="A06F7C12095068C83E2D2CCD5CEE69FB" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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, unique specimen identifier CASENT0608897].
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</paragraph>
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<paragraph id="6C31CC5168122137616E9730934508E3" pageNumber="26">
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Paratypes: major and minor workers. Same data as holotype [
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<collectionCode id="0ACBDF845B19EB92877608609E6B8B62" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]">BMNH</collectionCode>
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,
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<collectionCode id="C94037C96F967DFD2FEF022F03571B25" collectionName="USA, California, San Francisco, California Academy of Sciences">CAS</collectionCode>
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,
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<collectionCode id="98BEE9E2EDF8D94FD9594C96CBC7F1F9" collectionName="Honduras, Tegucigalpa, Escuela Agricola Panamericana">EAPZ</collectionCode>
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, ECOSCE,
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<collectionCode id="C7A2286CDA459EDE531F66B841F61E35" collectionName="USA, Illinois, Chicago, Field Museum of Natural History (also used by Finnish Museum of Natural History)">FMNH</collectionCode>
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,
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<collectionCode id="3D193CCB1CFD729BCCBE1D79DB0D2149" collectionName="Costa Rica, Santo Domingo de Heredia, Instituto Nacional de Biodiversidad (INBio)">INBC</collectionCode>
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,
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<collectionCode id="74D9A8317F4007B9C0F9FCD358EBBFDE" collectionName="John T. Longino">JTLC</collectionCode>
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,
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<collectionCode id="080EDD92447EF08352D412A235FA5595" collectionName="USA, California, Los Angeles, Los Angeles County Museum of Natural History">LACM</collectionCode>
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,
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<collectionCode id="9C9B9574CAA8424A154723BFE3FDDD74" collectionName="USA, Massachusetts, Cambridge, Harvard University, Museum of Comparative Zoology">MCZ</collectionCode>
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,
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<collectionCode id="538DFCA4D9E75E8176C5B492A6EB1FE7" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">MHNG</collectionCode>
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,
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<collectionCode id="0FE63ED88CD271F9497885F714C9B775" collectionName="Venezuela, Maracay, Museuo del Instituto de Zoologia Agricola">MIZA</collectionCode>
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,
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<collectionCode id="2E972E26FA317B3F214A6795838CDD10" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo">MZSP</collectionCode>
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, MEL,
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<collectionCode id="D98D9B98195622580D366FA2188DD5AE" collectionName="USA, California, Davis, University of California, R.M. Bohart Museum of Entomology">UCD</collectionCode>
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,
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<collectionCode id="DF5E49CD6670875044D9E39AFF6DE083" collectionName="Mexico, Mexico D.F., Universidad Nacional Autonoma de Mexico">UNAM</collectionCode>
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,
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<collectionCode id="D385AE7A52BDBF1C6D128D3ED6518729" collectionName="Colombia, Bogota, Universidad Nacional de Colombia, Insituto de Ciencias Naturales de la Universidad Nacional">ICN</collectionCode>
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,
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<collectionCode id="0F4BE545FF106088B4B98B892378C4E1" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
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,
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<collectionCode id="F89DB3EADAB14851C1A4B35EBEA395B8" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos">UVGC</collectionCode>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="2BB6C0B66A60207BECDA0E21487DE92D" type="distribution">
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<paragraph id="E2F12A73EE48A98E18408338535A4312" pageNumber="26">Geographic Range</paragraph>
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<paragraph id="AC9AA6D46B3F021272EC5EBB55F65DC6" pageNumber="26">Costa Rica.</paragraph>
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</subSubSection>
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<subSubSection id="841C2BAFE70D6239E869B845592F1FF6" type="diagnosis">
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<paragraph id="AE02363048B64E4A06AD0131E31267FC" pageNumber="26">Diagnosis</paragraph>
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<paragraph id="0BB5F058B32228B4F1B939989F10CBF9" pageNumber="26">
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The morphometric profile, sculptural details, and pilosity pattern are unique in this highly distinctive species. Minor worker: the combination of large size, generally shiny mesosoma, dark color, and long, thin, nearly perpendicular propodeal spines are distinctive. Major worker: face foveolate rugulose laterally, grading to smoother and sublucid on vertex lobes, but vertex lobes still sculptured with faint mesh of foveolae and coarse piligerous puncta, versus more uniformly foveolate rugose and not shining (
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<taxonomicName id="7E9FF397EFE684776F8E02F02F426635" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:34094" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing, matching species" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole vorax (Fabricius)" order="Hymenoptera" pageNumber="26" phylum="Arthropoda" rank="species" species="vorax">vorax</taxonomicName>
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) or entirely rugose (
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<taxonomicName id="00772BD13C23FFFA45C00710CF601ADB" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33964" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole rhea Wheeler" order="Hymenoptera" pageNumber="26" phylum="Arthropoda" rank="species" species="rhea">rhea</taxonomicName>
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); scape curved at base, base slightly flattened, about as wide at curvature as maximum width at apex, versus thin at base and less curved (astur); antennal club 3-segmented versus 4-segmented (
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<taxonomicName id="7ADB42637E47CA2A2ECA147005F32A8E" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:33718" class="Insecta" family="Formicidae" genus="Pheidole" genus.completionEvidence="missing" genus.completionRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole dwyeri Gregg" order="Hymenoptera" pageNumber="26" phylum="Arthropoda" rank="species" species="dwyeri">dwyeri</taxonomicName>
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); color dark brown and dorsal surface of mandible with erect setae versus color orange brown and dorsal surface of mandible lacking erect setae (praeses).
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</paragraph>
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</subSubSection>
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<subSubSection id="DF3EFF24569B16D3329A2732C5451FF5" type="description">
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<paragraph id="2B5FAA16B07BCB745C7E097E0C76C93A" pageNumber="26">Description of minor worker</paragraph>
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<paragraph id="1D151399B92701C4FB3FF10830498BA2" pageNumber="26">Measurements (paratype): HL 1.07, HW 0.98, HLA 0.41, SL 1.20, EL 0.18, ML 1.48, PSL 0.19, PMG 0.01, SPL 0.04, PTW 0.16, PPW 0.29, CI 92, SI 122, PSLI 17, PMGI 1, SPLI 4, PPI 184.</paragraph>
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<paragraph id="2D83845EA40B9B30711595054A15B8FC" pageNumber="26">Measurements (n=10): HL 0.65-1.13, HW 0.59-1.04, SL 0.92-1.24, CI 89-92, SI 119-156.</paragraph>
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<paragraph id="A203DDA5F6BB598BB55347FFD08C8DB0" pageNumber="26">Mandible smooth and shining; clypeus smooth and shining with median carina; face shining with very faint mesh of foveolae, only visible in certain lighting conditions; margin of vertex rounded; occipital carina narrow, visible in full face view; scape with abundant subdecumbent setae less than or equal to maximum width of scape; promesonotal groove weakly impressed, promesonotum forming more or less continuous arched convexity; pronotal humerus weakly produced as a quadrate boss; propodeal spines long, thin, nearly perpendicular to dorsal face of propodeum; mesosoma generally smooth and shiny, with very faint foveolate mesh, similar to face, on sides of pronotum, juncture of katepisternum and anepisternum, posterior katepisternum, propodeum, petiole, and postpetiole; 3 strong parallel carinae on metapleural bulla; ventral margin of postpetiole slightly produced anteriorly, forming short step anteriorly; abundant setae on promesonotal dorsum; dorsal (outer) margin of hind tibia with subdecumbent setae and longer suberect setae subequal in length to maximum width of tibia; first gastral tergum smooth and shining; gastral dorsum with abundant, long, suberect setae; color dark red brown with lighter brown legs.</paragraph>
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<paragraph id="DF8F2C67EAD892AFDC4E3F5723D6E1C9" pageNumber="27">
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<pageBreakToken id="1ED7CDD8EC6D706E3AE994323E259A79" pageNumber="27">Description</pageBreakToken>
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of major worker
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</paragraph>
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<paragraph id="7DA2F3CCB986FF03E48F7D5BA67AAF36" pageNumber="27">Measurements (holotype): HL 2.75, HW 2.58, HLA 0.72, SL 1.32, EL 0.29, ML 2.20, PSL 0.25, PMG 0.00, SPL 0.10, PTW 0.37, PPW 0.80, IHT 0.45, OHT 0.98, CI 94, SI 51, PSLI 9, PMGI 0, SPLI 4, PPI 216, HTI 46.</paragraph>
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<paragraph id="05F0492EFE136DBD695EB9692DF6F252" pageNumber="27">Mandibles smooth and shiny; medial clypeus smooth and flat with a few longitudinal rugulae laterally, beneath frontal carinae; frontal carina short, forming vertical flange above antennal insertion; face with longitudinal carinulae between frontal carina and compound eye, grading to foveolate rugulose sculpture posteriorly, grading to smoother on vertex lobes, but vertex lobes still sculptured with faint mesh of foveolae and coarse piligerous puncta; medial area between frontal carinae with a sharp median carina and radiating fan of longitudinal carinulae; head with abundant suberect setae projecting from sides of head in face view; scape curved at base, base slightly flattened, about as wide at curvature as maximum width at apex, with abundant erect setae longer than maximum width of scape; hypostomal margin flat; median tooth absent; inner hypostomal teeth stout, closer to midline than to outer hypostomal teeth; promesonotal groove weakly impressed; pronotal humerus strongly developed as blunt cone; propodeal spines present; anterodorsal pronotum and mesonotum with widely-spaced transverse rugulae; side of pronotum with irregular rugulae; katepisternum smooth and shining; lateral propodeum smooth and shining with 3 strong longitudinal carinae on metapleural bulla; dorsal (outer) margin of hind tibia with abundant suberect setae, some longer than maximum width of tibia; pilosity abundant on mesosomal dorsum; postpetiolar sternite foveolate, with pair of anterolateral gibbosities; first gastral tergite smooth and shining with very faint mesh of foveolae, with abundant flexuous suberect setae; color dark red brown.</paragraph>
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</subSubSection>
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<subSubSection id="B13F9CF64574081D2A9825ED74BE9F7C" type="biology_ecology">
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<paragraph id="620F2A9440B457EE5BBBCF12FBEFFDB6" pageNumber="27">Biology</paragraph>
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<paragraph id="37FBB8BEFA728B5422B67D41B10F9A42" pageNumber="27">
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<taxonomicName id="B5B56FE399681EB68974E3542628BCCD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253257" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole eowilsoni Longino" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="eowilsoni">Pheidole eowilsoni</taxonomicName>
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occurs in mature wet forest. The type series was collected from a nest in a clay bank above a stream. Nocturnal foragers led to a simple hole in a vertical clay bank. The hole led into a tunnel about 1cm diameter. Nest excavation uncovered four chambers containing minor workers, major workers, and brood. One chamber contained a single colony queen. The volume occupied by the nest was about 20cm across and 20cm deep in the bank. One chamber contained what appeared to be an internal trash midden. Among general debris the midden contained nematodes, the remains of a dead
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<taxonomicName id="9566113526188B16E52E65890DD1942C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24785" class="Insecta" family="Formicidae" genus="Myrmelachista" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Myrmelachista Roger" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Myrmelachista</taxonomicName>
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queen, and a few small plant seeds. A second collection was a founding queen with a few minim workers, beneath a stone in a rainforest trail. Minor workers were collected in a pan trap near a stream.
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</paragraph>
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</subSubSection>
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<subSubSection id="AFDD7E0EF8A197924438DAA5059FDA46" type="etymology">
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<paragraph id="B6843DAAA33365E5C42B7E67EA808C78" pageNumber="27">Etymology</paragraph>
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<paragraph id="929728A49AEB5B2FF6E642AA7B145811" pageNumber="27">
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This species is named in honor of E. O. Wilson, whose pioneering work on
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<taxonomicName id="B358D3C58C0B50927865E9910E211BCE" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:24885" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole Westwood" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="genus">Pheidole</taxonomicName>
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has made the present work possible.
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</paragraph>
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</subSubSection>
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<subSubSection id="CE082FEF041A6BC48B94A10C2862D6DD" type="discussion">
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<paragraph id="367968D4FEF3BC6A3F23D51A71035ADA" pageNumber="27">Comments</paragraph>
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<paragraph id="63D7CF5BBC7CFC5C767DB4E6595A97D5" pageNumber="27">
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Two collections from 500m elevation on the Barva Transect in Braulio Carrillo National Park and the type series from 800m in the Penas Blancas Valley have minor workers that are nearly identical to each other. A series of isolated minor workers from 50-500m on the Barva Transect, collected in Winkler and Berlese samples, are tentatively identified as
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<taxonomicName id="4189BE1FF0E0D2F9D57CF6E6B4ADF3D6" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253257" class="Insecta" family="Formicidae" genus="Pheidole" genus.bestMatchDistance="24" genus.bestMatchVote="2" genus.innerRound="1" genus.outerRound="1" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole eowilsoni Longino" order="Hymenoptera" pageNumber="27" phylum="Arthropoda" rank="species" species="eowilsoni">P. eowilsoni</taxonomicName>
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but the clypeus lacks a median carina and the ventral margin of the postpetiole is completely flat.
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</paragraph>
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</subSubSection>
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<subSubSection id="85B8142D4F9799F07B667F24376D089B" type="multiple">
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<caption id="C9C861E4712FE7FA6037AB9C7209CBA0">
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<paragraph id="4FA3C6C7EAEB8CCE2AEEE961BB94A318" pageNumber="28">
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<pageBreakToken id="43C763BE1986660E80DAB96334B649BE" pageNumber="28">FIGURE</pageBreakToken>
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5.
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<taxonomicName id="6E36B03756B03088BD3BAE4CD54DA380" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:253257" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-HNS="Pheidole eowilsoni Longino" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="eowilsoni">Pheidole eowilsoni</taxonomicName>
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. Major worker: A, face view; B, lateral view; C, dorsal view; D, hypostomal margin. Minor worker: E, face view; F, lateral view; G, dorsal view; H, hind tibia. A-C, Holotype major worker; D, non-type major worker; E-H, Paratype minor worker. Scale bars 1mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="890B22BA32E59CB75F249561D287BF5B" type="materials_examined">
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<paragraph id="15CB33A857A9C62100AFF03E1B692D4A" pageNumber="29">
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<pageBreakToken id="1B6194EDE42C7C68205E8719DEBB3B2F" pageNumber="29">Additional</pageBreakToken>
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material examined
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</paragraph>
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<paragraph id="6D53879F7686A9EC066F1760FFD9E1A4" pageNumber="29">
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COSTA RICA: Heredia, La Selva Biological Station,
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<geoCoordinate id="49BE84D49ED0E4594583C42E87EEA8A3" direction="north" orientation="latitude" value="10.416667">10°25'N</geoCoordinate>
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,
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<geoCoordinate id="F9D8010B1EE272C5F545D85974AA4AEB" direction="west" orientation="longitude" value="-84.01667">84°01'W</geoCoordinate>
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, 50m (multiple collectors and collections); 16km SSW Pto. Viejo,
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<geoCoordinate id="3C392E390220C18E344EAF2B5BE5FFCE" direction="north" orientation="latitude" value="10.316667">10°19'N</geoCoordinate>
|
|
,
|
|
<geoCoordinate id="9D5A854864968E0358D697F2E604DC16" direction="west" orientation="longitude" value="-84.05">84°03'W</geoCoordinate>
|
|
, 500m (multiple collectors and collections).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |