484 lines
53 KiB
XML
484 lines
53 KiB
XML
<document ID-DOI="10.3897/zookeys.19.221" ID-GBIF-Dataset="e6ea8eea-7156-4785-8313-f4bd88dadad1" ID-ISSN="1313–2970" ID-Zenodo-Dep="576503" ID-ZooBank="C473F9F6-1AE7-4B3F-B17F-CA1C2709010C" checkinTime="1587956031210" checkinUser="plazi" docAuthor="Wesener, Thomas, Enghoff, Henrik & Sierwald, Petra" docDate="2009" docId="1772122E7822FFEEFF013BD7ACF2E8DA" docLanguage="en" docName="ZK_article_2037.pdf" docOrigin="ZooKeys 19 (19)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@11b73717" docStyleName="zookeys.2008.journal_article" docTitle="Zehntnerobolus rubripes Wesener & Enghoff & Sierwald 2009, comb. n." docType="treatment" docVersion="6" lastPageNumber="41" masterDocId="EB4B6A567807FFC6FF8D3F74AF59EA51" masterDocTitle="Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)" masterLastPageNumber="128" masterPageNumber="1" pageNumber="38" updateTime="1636630017160" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wesener, Thomas</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A. & Zoologisches Forschungsmuseum Alexander Koenig, Museumsmeile Bonn, Adenauerallee 160, D- 53113 Bonn, Germany. & B 54 D 3623309</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Enghoff, Henrik</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Natural History Museum of Denmark, Universitetsparken 15, DK- 2100 Copenhagen, Denmark</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sierwald, Petra</mods:namePart>
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<mods:nameIdentifier type="LSID">urn:lsid:zoobank.org:author:</mods:nameIdentifier>
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<mods:affiliation>Field Museum of Natural History, 1400 S. Lake Shore Drive, Chicago, IL 60605, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2009</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2009-09-04</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>128</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.19.221</mods:identifier>
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<mods:identifier type="GBIF-Dataset">e6ea8eea-7156-4785-8313-f4bd88dadad1</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576503</mods:identifier>
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<mods:identifier type="ZooBank">C473F9F6-1AE7-4B3F-B17F-CA1C2709010C</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3791397" ID-GBIF-Taxon="164184229" ID-Zenodo-Dep="3791397" LSID="urn:lsid:plazi:treatment:1772122E7822FFEEFF013BD7ACF2E8DA" httpUri="http://treatment.plazi.org/id/1772122E7822FFEEFF013BD7ACF2E8DA" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38">
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<subSubSection box="[140,941,1187,1214]" pageId="37" pageNumber="38" type="nomenclature">
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<paragraph blockId="37.[140,941,1187,1214]" box="[140,941,1187,1214]" pageId="37" pageNumber="38">
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<heading box="[140,941,1187,1214]" fontSize="11" level="2" pageId="37" pageNumber="38" reason="3">
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<emphasis bold="true" box="[140,941,1187,1214]" pageId="37" pageNumber="38">
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<taxonomicName authority="(de Saussure & Zehntner, 1897)" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[140,823,1187,1214]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" status="comb. n.">
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<emphasis bold="true" box="[140,433,1187,1214]" italics="true" pageId="37" pageNumber="38">Zehntnerobolus rubripes</emphasis>
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(
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<bibRefCitation author="de Saussure & Zehntner" box="[448,814,1187,1213]" firstAuthor="de Saussure" pageId="37" pageNumber="38" pagination="1 - 12" refId="ref56919" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier (Ed.) Histoire Physique, Naturelle et Politique de Madagascar 27 (53): pl 1 - 12." type="journal article" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel box="[836,941,1187,1213]" pageId="37" pageNumber="38" rank="species">comb. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="37" pageNumber="38" type="reference_group">
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<treatmentCitationGroup pageId="37" pageNumber="38">
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||
<paragraph blockId="37.[140,1108,1258,1672]" box="[140,1087,1258,1284]" pageId="37" pageNumber="38">
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||
<treatmentCitation box="[140,703,1258,1284]" pageId="37" pageNumber="38">
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||
<taxonomicName authority="de Saussure & Zehntner, 1897" authorityName="de Saussure & Zehntner" authorityYear="1897" box="[140,703,1258,1284]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[140,347,1258,1284]" italics="true" pageId="37" pageNumber="38">Spirobolus rubripes</emphasis>
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<bibRefCitation author="de Saussure & Zehntner" box="[354,703,1258,1284]" firstAuthor="de Saussure" pageId="37" pageNumber="38" pagination="1 - 12" refId="ref56919" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier (Ed.) Histoire Physique, Naturelle et Politique de Madagascar 27 (53): pl 1 - 12." type="journal article" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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: plate XI, fig. 48, plate XII, fig. 6.
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</paragraph>
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<paragraph blockId="37.[140,1108,1258,1672]" box="[140,818,1293,1320]" pageId="37" pageNumber="38">
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<taxonomicName authority=", de Saussure and Zehntner 1902: 154" authorityName="de Saussure and Zehntner" authorityPageNumber="154" authorityYear="1902" box="[140,814,1293,1320]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[140,379,1293,1319]" italics="true" pageId="37" pageNumber="38">Spirostrophus rubripes</emphasis>
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,
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<treatmentCitation box="[393,814,1293,1320]" pageId="37" pageNumber="38">
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<bibRefCitation author="de Saussure H" box="[393,814,1293,1320]" pageId="37" pageNumber="38" pagination="1 - 356" refId="ref56994" refString="de Saussure H, Zehntner, L (1902) Myriapodes de Madagascar. In: Grandidier (Ed.), Histoire Physique, Naturelle et Politique de Madagascar 27 (53): 1 - 356, pl 13 - 15." type="journal article" year="1902">de Saussure and Zehntner 1902: 154</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</paragraph>
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</treatmentCitationGroup>
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<paragraph blockId="37.[140,1108,1258,1672]" box="[140,655,1328,1354]" pageId="37" pageNumber="38">
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<treatmentCitationGroup box="[140,655,1328,1354]" pageId="37" pageNumber="38">
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NOT
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<treatmentCitation box="[212,655,1328,1354]" pageId="37" pageNumber="38">
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<taxonomicName authority="Attems, 1927: 64" authorityName="Attems" authorityPageNumber="64" authorityYear="1927" box="[212,655,1328,1354]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[212,451,1328,1354]" italics="true" pageId="37" pageNumber="38">Spirostrophus rubripes</emphasis>
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<bibRefCitation box="[458,655,1328,1354]" pageId="37" pageNumber="38">Attems, 1927: 64</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="37.[140,1108,1258,1672]" box="[140,838,1363,1390]" pageId="37" pageNumber="38">
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<treatmentCitationGroup box="[140,838,1363,1390]" pageId="37" pageNumber="38">
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<treatmentCitation author="Enghoff H" box="[140,609,1363,1390]" page="624" pageId="37" pageNumber="38" year="2003">
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<taxonomicName authority=", Enghoff 2003: 624" authorityName=", Enghoff" authorityPageNumber="624" authorityYear="2003" box="[140,609,1363,1390]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[140,379,1363,1389]" italics="true" pageId="37" pageNumber="38">Spirostrophus rubripes</emphasis>
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,
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<bibRefCitation author="Enghoff" box="[393,609,1363,1390]" firstAuthor="Enghoff" pageId="37" pageNumber="38" pagination="617 - 627" refId="ref57094" refString="Enghoff H (2003) Diplopoda, Millipedes. In: Goodman SM, Benstead JP (Eds) The Natural History of Madagascar. University of Chicago Press, Chicago, 617 - 627." type="book chapter" year="2003">Enghoff 2003: 624</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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(lists species name).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="discussion">
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<paragraph blockId="37.[140,1108,1258,1672]" pageId="37" pageNumber="38">
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||
<emphasis bold="true" box="[185,324,1398,1424]" pageId="37" pageNumber="38">
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<emphasis box="[185,319,1398,1424]" inLineHeading="true" pageId="37" pageNumber="38" reason="1">Comments</emphasis>
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:
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</emphasis>
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drawings of
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[480,745,1399,1425]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[480,745,1399,1425]" italics="true" pageId="37" pageNumber="38">Zehntnerobolus rubripes</emphasis>
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</taxonomicName>
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were first published under the name
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<taxonomicName authority="(de Saussure & Zehntner, 1897)" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[210,797,1434,1460]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[210,418,1434,1460]" italics="true" pageId="37" pageNumber="38">Spirobolus rubripes</emphasis>
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||
(
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<bibRefCitation author="de Saussure & Zehntner" box="[433,789,1434,1460]" firstAuthor="de Saussure" pageId="37" pageNumber="38" pagination="1 - 12" refId="ref56919" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier (Ed.) Histoire Physique, Naturelle et Politique de Madagascar 27 (53): pl 1 - 12." type="journal article" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
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)
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</taxonomicName>
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.
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<taxonomicName authorityName="Brandt" authorityYear="1833" box="[809,921,1434,1460]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis box="[809,921,1434,1460]" italics="true" pageId="37" pageNumber="38">Spirobolus</emphasis>
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</taxonomicName>
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is an East Asian genus. So, when publishing the text description in 1902, de Saussure and Zehntner put
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<taxonomicName baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[186,303,1504,1531]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[186,303,1504,1531]" italics="true" pageId="37" pageNumber="38">S. rubripes</emphasis>
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</taxonomicName>
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in the newly erected genus
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<taxonomicName authorityName="De Saussure & Zehntner" authorityYear="1902" box="[617,761,1504,1530]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis box="[617,761,1504,1530]" italics="true" pageId="37" pageNumber="38">Spirostrophus</emphasis>
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</taxonomicName>
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, which included a non-related assemblage of species. However, they already noted: “Le
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<taxonomicName baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[781,898,1539,1566]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[781,898,1539,1566]" italics="true" pageId="37" pageNumber="38">S. rubripes</emphasis>
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</taxonomicName>
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offre de caractères un peu aberrantes: il est par conséquent difficile à classer. [...]. La position de cette espèce est donc incertaine.” (de Saussure and Zehntner 1902: 158). They, unfortunately, did not put
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<taxonomicName baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[273,388,1645,1671]" class="Diplopoda" family="Spirobolidae" genus="Spirobolus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[273,388,1645,1671]" italics="true" pageId="37" pageNumber="38">S. rubripes</emphasis>
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</taxonomicName>
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into a new genus. The description of
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<taxonomicName authorityName=", Enghoff" authorityYear="2003" box="[804,1042,1645,1671]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species">
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<emphasis box="[804,1042,1645,1671]" italics="true" pageId="37" pageNumber="38">Spirostrophus rubripes</emphasis>
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</taxonomicName>
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by de
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.3767298" ID-Zenodo-Dep="3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="38" pageNumber="39" startId="38.[140,212,1526,1548]" targetBox="[167,1080,166,1506]" targetPageId="38">
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||
<paragraph blockId="38.[140,1108,1526,1676]" pageId="38" pageNumber="39">
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<emphasis bold="true" box="[140,251,1526,1548]" pageId="38" pageNumber="39">Figure 22.</emphasis>
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<taxonomicName authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[257,471,1526,1547]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" status="comb. n.">
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<emphasis box="[257,471,1526,1547]" italics="true" pageId="38" pageNumber="39">Zehntnerobolus rubripes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[477,561,1526,1548]" pageId="38" pageNumber="39" rank="species">comb. n.</taxonomicNameLabel>
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, A, C–G male lectotype, B female paratype,
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||
<emphasis bold="true" box="[975,994,1526,1548]" pageId="38" pageNumber="39">A</emphasis>
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||
telson
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||
<emphasis bold="true" box="[1060,1108,1526,1548]" pageId="38" pageNumber="39">B, C</emphasis>
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||
right midbody leg
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||
<emphasis bold="true" box="[312,331,1558,1580]" pageId="38" pageNumber="39">D</emphasis>
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||
anterior gonopod, anterior view
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||
<emphasis bold="true" box="[638,653,1558,1580]" pageId="38" pageNumber="39">E</emphasis>
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||
anterior gonopod, posterior view
|
||
<emphasis bold="true" box="[969,983,1558,1580]" pageId="38" pageNumber="39">F</emphasis>
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left posterior gonopod, anterior view
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<emphasis bold="true" box="[362,382,1590,1612]" pageId="38" pageNumber="39">G</emphasis>
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left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm.
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</paragraph>
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</caption>
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||
<subSection lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="multiple">
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<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
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||
Saussure and Zehntner was obviously all but forgotten, because Attems also named a non-related species from
|
||
<collectingCountry box="[424,534,202,228]" name="Indonesia" pageId="39" pageNumber="40">Indonesia</collectingCountry>
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<taxonomicName authority="Attems, 1927" authorityName="Attems" authorityYear="1927" box="[541,945,202,228]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species">
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||
<emphasis box="[541,780,202,228]" italics="true" pageId="39" pageNumber="40">Spirostrophus rubripes</emphasis>
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Attems, 1927
|
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</taxonomicName>
|
||
. The
|
||
<typeStatus box="[1006,1054,202,228]" pageId="39" pageNumber="40">type</typeStatus>
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||
species of the genus
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||
<taxonomicName authorityName="De Saussure & Zehntner" authorityYear="1902" box="[333,477,237,263]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[333,477,237,263]" italics="true" pageId="39" pageNumber="40">Spirostrophus</emphasis>
|
||
</taxonomicName>
|
||
is now regarded as a synonym of the
|
||
<typeStatus box="[902,950,238,264]" pageId="39" pageNumber="40">type</typeStatus>
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||
species of the East Indian genus
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||
<taxonomicName authority="Silvestri, 1897" authorityName="Silvestri" authorityYear="1897" box="[349,668,272,298]" class="Diplopoda" family="Trigoniulidae" genus="Leptogoniulus" kingdom="Animalia" order="Spirobolida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[349,499,272,298]" italics="true" pageId="39" pageNumber="40">Leptogoniulus</emphasis>
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||
<bibRefCitation author="Silvestri" box="[505,668,272,298]" firstAuthor="Silvestri" pageId="39" pageNumber="40" pagination="651 - 653" refId="ref57740" refString="Silvestri F (1897) Systema Diplopodum. Annali del Museo civico di storia naturale di Genova 38: 651 - 653." type="journal article" year="1897">Silvestri, 1897</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Hoffman" box="[682,857,272,298]" firstAuthor="Hoffman" pageId="39" pageNumber="40" refId="ref57240" refString="Hoffman RL (1980) Classification of the Diplopoda. Musee d'Histoire Naturelle de Geneve, Geneve, Switzerland, 237 pp." type="book" year="1980">Hoffman 1980</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Shelley and Lehtinen" firstAuthor="Shelley" pageId="39" pageNumber="40" pagination="1379 - 1401" refId="ref57656" refString="Shelley RM, Lehtinen PT (1999) Diagnoses, synonymies and occurrences of the pantropical millipeds, Leptogoniulus sorornus (Butler) and Trigoniulus corallinus (Gervais) (Spirobolida: Pachybolidae: Trigoniulinae). Journal of Natural History 33 (9): 1379 - 1401." type="journal article" year="1999">Shelley and Lehtinen 1999</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Attems" authorityYear="1927" box="[222,460,307,333]" class="Diplopoda" family="Trigoniulidae" genus="Spirostrophus" kingdom="Animalia" order="Spirobolida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[222,460,307,333]" italics="true" pageId="39" pageNumber="40">Spirostrophus rubripes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="de Saussure & Zehntner" box="[475,821,307,334]" firstAuthor="de Saussure" pageId="39" pageNumber="40" pagination="1 - 12" refId="ref56919" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier (Ed.) Histoire Physique, Naturelle et Politique de Madagascar 27 (53): pl 1 - 12." type="journal article" year="1897">de Saussure & Zehntner, 1897</bibRefCitation>
|
||
shares no characters with
|
||
<taxonomicName authorityName="Silvestri" authorityYear="1897" box="[140,290,343,369]" class="Diplopoda" family="Trigoniulidae" genus="Leptogoniulus" kingdom="Animalia" order="Spirobolida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,290,343,369]" italics="true" pageId="39" pageNumber="40">Leptogoniulus</emphasis>
|
||
</taxonomicName>
|
||
, therefore a new genus had to be established to incorporate this species.
|
||
</paragraph>
|
||
<subSubSection pageId="39" pageNumber="40" type="materials_examined">
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis bold="true" box="[185,419,377,403]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,286,377,403]" inLineHeading="true" pageId="39" pageNumber="40" reason="1">Material</emphasis>
|
||
examined:
|
||
</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="2620669322" location="Saussure & Zehntner" municipality="Zehntnerobolus" pageId="39" pageNumber="40" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="lectotype">
|
||
<specimenCount box="[428,476,378,404]" pageId="39" pageNumber="40" type="male">2 ♂</specimenCount>
|
||
,
|
||
<specimenCount box="[491,538,378,404]" pageId="39" pageNumber="40" type="female">2 ♀</specimenCount>
|
||
<emphasis box="[548,654,379,404]" italics="true" pageId="39" pageNumber="40">
|
||
<typeStatus box="[548,648,379,404]" pageId="39" pageNumber="40">Lectotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
designated herewith to fix the concept of
|
||
<taxonomicName authority="(de Saussure & Zehntner, 1897)" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[171,810,413,439]" class="Diplopoda" family="Pachybolidae" genus="Zehntnerobolus" kingdom="Animalia" order="Spirobolida" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species">
|
||
<emphasis box="[171,433,413,439]" italics="true" pageId="39" pageNumber="40">
|
||
<collectingMunicipality box="[171,337,413,439]" pageId="39" pageNumber="40">Zehntnerobolus</collectingMunicipality>
|
||
rubripes
|
||
</emphasis>
|
||
(
|
||
<bibRefCitation author="de Saussure & Zehntner" box="[449,803,413,439]" firstAuthor="de Saussure" pageId="39" pageNumber="40" pagination="1 - 12" refId="ref56919" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier (Ed.) Histoire Physique, Naturelle et Politique de Madagascar 27 (53): pl 1 - 12." type="journal article" year="1897">
|
||
de
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E7822FFEEFF013BD7ACF2E8DA:9A04F5E37820FFE1FE6B3EE9AD83EBE6" box="[486,730,413,439]" municipality="Zehntnerobolus" name="Saussure & Zehntner" pageId="39" pageNumber="40">Saussure & Zehntner</location>
|
||
, 1897
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and to ensure the universal and consistent interpretation of the same
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation ID-GBIF-Occurrence="2620669318" collectorName="Musee & de Geneve" country="Madagascar" determinerName="de Saussure" latitude="-18.938889" location="Foret de Moramanga" longLatPrecision="21" longitude="48.22778" municipality="Province" pageId="39" pageNumber="40" specimenCount="1" stateProvince="Toamasina" typeStatus="lectotype">
|
||
<specimenCount box="[676,698,449,475]" pageId="39" pageNumber="40">♂</specimenCount>
|
||
(dissected,
|
||
<quantity box="[837,923,448,475]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="39" pageNumber="40" unit="mm" value="30.0">30 mm</quantity>
|
||
long), MNHN HB049.
|
||
<collectingCountry box="[239,373,483,509]" name="Madagascar" pageId="39" pageNumber="40">Madagascar</collectingCountry>
|
||
,
|
||
<collectingMunicipality box="[384,482,483,509]" pageId="39" pageNumber="40">Province</collectingMunicipality>
|
||
<collectingRegion box="[487,611,483,509]" country="Madagascar" name="Toamasina" pageId="39" pageNumber="40">Toamasina</collectingRegion>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E7822FFEEFF013BD7ACF2E8DA:9A04F5E37820FFE1FDE33E90AC05EBAF" box="[622,860,483,510]" country="Madagascar" latitude="-18.938889" longLatPrecision="21" longitude="48.22778" municipality="Province" name="Foret de Moramanga" pageId="39" pageNumber="40" stateProvince="Toamasina">Forêt de Moramanga</location>
|
||
, rainforest,
|
||
<geoCoordinate degrees="18" direction="south" minutes="56" orientation="latitude" pageId="39" pageNumber="40" precision="15" seconds="20" value="-18.938889">18°56’20” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[171,316,519,545]" degrees="48" direction="east" minutes="13" orientation="longitude" pageId="39" pageNumber="40" precision="15" seconds="40" value="48.22778">48°13’40” E</geoCoordinate>
|
||
, leg.
|
||
<collectorName box="[379,452,519,545]" pageId="39" pageNumber="40">Musée</collectorName>
|
||
<collectorName box="[462,586,519,545]" pageId="39" pageNumber="40">de Genève</collectorName>
|
||
, det.
|
||
<determinerName box="[652,787,519,545]" pageId="39" pageNumber="40">de Saussure</determinerName>
|
||
</materialsCitation>
|
||
, 1903.
|
||
<materialsCitation ID-GBIF-Occurrence="2620669323" collectorName="Musee & de Geneve" country="Madagascar" determinerName="de Saussure" latitude="-18.938889" location="Foret de Moramanga" longLatPrecision="21" longitude="48.22778" municipality="Province" pageId="39" pageNumber="40" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Toamasina" typeStatus="paratype">
|
||
<emphasis box="[876,983,519,544]" italics="true" pageId="39" pageNumber="40">
|
||
<typeStatus box="[876,978,519,544]" pageId="39" pageNumber="40">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount box="[993,1040,519,545]" pageId="39" pageNumber="40" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount box="[1055,1100,519,545]" pageId="39" pageNumber="40" type="female">2 ♀</specimenCount>
|
||
, MNHN HB049, same data as lectotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis bold="true" box="[185,718,589,615]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,313,589,615]" inLineHeading="true" pageId="39" pageNumber="40" reason="1">Additional</emphasis>
|
||
locality data from the literature:
|
||
</emphasis>
|
||
<materialsCitation ID-GBIF-Occurrence="2620669324" collectingDate="1886" collectingDateMax="1892" collectingDateMin="1886" collectorName="de Saussure & Franz Sikora" country="Madagascar" latitude="-19.033333" location="Zehntner" longLatPrecision="1" longitude="47.916702" municipality="Foret d'Andrangoloaka" pageId="39" pageNumber="40" specimenCount="1">
|
||
<collectorName box="[728,861,589,615]" pageId="39" pageNumber="40">de Saussure</collectorName>
|
||
and
|
||
<location LSID="urn:lsid:plazi:treatment:1772122E7822FFEEFF013BD7ACF2E8DA:9A04F5E37820FFE1FC113D39AB5EE836" box="[924,1031,589,615]" country="Madagascar" latitude="-19.033333" longLatPrecision="1" longitude="47.916702" municipality="Foret d'Andrangoloaka" name="Zehntner" pageId="39" pageNumber="40">Zehntner</location>
|
||
1902:
|
||
<collectingCountry box="[140,274,624,650]" name="Madagascar" pageId="39" pageNumber="40">Madagascar</collectingCountry>
|
||
,
|
||
<collectingMunicipality box="[292,561,624,650]" pageId="39" pageNumber="40">Forêt d’Andrangoloaka</collectingMunicipality>
|
||
, rainforest,
|
||
<geoCoordinate box="[707,861,624,650]" degrees="19" direction="south" minutes="2" orientation="latitude" pageId="39" pageNumber="40" precision="1" seconds="0.00" value="-19.033333">19°2’0.00” S</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[879,1050,624,651]" degrees="47" direction="east" minutes="55" orientation="longitude" pageId="39" pageNumber="40" precision="1" seconds="0.12" value="47.916702">47°55’0.12” E</geoCoordinate>
|
||
, leg.
|
||
<collectorName box="[140,282,659,686]" pageId="39" pageNumber="40">Franz Sikora</collectorName>
|
||
(
|
||
<date box="[297,435,659,686]" pageId="39" pageNumber="40" value="1886" valueMax="1892">
|
||
<collectingDate box="[297,435,659,686]" pageId="39" pageNumber="40" value="1886" valueMax="1892">1886–1892</collectingDate>
|
||
</date>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis bold="true" box="[185,335,694,720]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,330,694,720]" inLineHeading="true" pageId="39" pageNumber="40" reason="1">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
<emphasis box="[342,494,695,720]" italics="true" pageId="39" pageNumber="40">Measurements</emphasis>
|
||
: males with 38 or 39 rings, circa
|
||
<quantity box="[866,947,695,721]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="39" pageNumber="40" unit="mm" value="32.0">32 mm</quantity>
|
||
long,
|
||
<quantity box="[1018,1107,695,721]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="39" pageNumber="40" unit="mm" value="2.9">2.9 mm</quantity>
|
||
wide. Females with 39 or 40 rings,
|
||
<quantity box="[538,664,730,756]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="3.7" metricValueMin="3.6" pageId="39" pageNumber="40" unit="mm" value="36.5" valueMax="37.0" valueMin="36.0">36–37 mm</quantity>
|
||
long,
|
||
<quantity box="[737,827,730,756]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="39" pageNumber="40" unit="mm" value="3.7">3.7 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,293,765,791]" italics="true" pageId="39" pageNumber="40">Coloration</emphasis>
|
||
decolorized in alcohol. Head, antennae and legs red (according to drawing and original description). Body ventrally and dorsally on meso- and metazonites pitchblack. Mesozonites dorsolaterally blood-red (see
|
||
<bibRefCitation author="de Saussure and Zehntner" box="[579,918,835,862]" firstAuthor="de Saussure" pageId="39" pageNumber="40" pagination="1 - 12" refId="ref56919" refString="de Saussure H, Zehntner L (1897) Atlas de l'histoire naturelle des Myriapodes. In: Grandidier (Ed.) Histoire Physique, Naturelle et Politique de Madagascar 27 (53): pl 1 - 12." type="journal article" year="1897">de Saussure and Zehntner 1897</bibRefCitation>
|
||
: plate XI, fig. 48).
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,244,871,897]" italics="true" pageId="39" pageNumber="40">Head</emphasis>
|
||
: each eye with circa 26–28 ocelli arranged in 6 or 7 vertical rows. Labrum with standard three irregular teeth and a single row of 10–12 stout marginal setae. Clypeus with two setiferous foveolae on each side. Antennal cavity absent. Antennae of medium length, protruding back to ring
|
||
<quantity box="[658,707,976,1002]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="39" pageNumber="40" unit="in" value="4.0">4 in</quantity>
|
||
females, ring
|
||
<quantity box="[876,925,976,1003]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="39" pageNumber="40" unit="in" value="5.0">5 in</quantity>
|
||
males. Relative lengths of antennomeres: 1<<2=3=4=5=6. Terminal antennomere with four large sensory cones located together inside a membranous area. Antennomere 5 latero-apically with five rows, antennomere 6 with four rows of sensilla basiconica.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,368,1117,1143]" italics="true" pageId="39" pageNumber="40">Gnathochilarium</emphasis>
|
||
: of usual spirobolidan shape. Lamellae linguales each with two standard setae located behind one another. Stipites each with three apical setae. Basal parts could not be studied because a dissection was not attempted.
|
||
<emphasis box="[811,914,1187,1213]" italics="true" pageId="39" pageNumber="40">Mandible</emphasis>
|
||
not dissected.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" box="[185,823,1223,1249]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,274,1223,1249]" italics="true" pageId="39" pageNumber="40">Collum:</emphasis>
|
||
smooth, laterally not protruding as far as ring 2.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,306,1258,1284]" italics="true" pageId="39" pageNumber="40">Body rings:</emphasis>
|
||
mostly smooth, with some transverse impressions on mesozona. Ozopores starting at ring 6, located closely in front of, but not touching suture between mesozona and metazona.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,251,1363,1389]" italics="true" pageId="39" pageNumber="40">Telson</emphasis>
|
||
: anal valves with neither lips, nor micropunctation. Preanal process sharp, but not protruding above anal valves (
|
||
<figureCitation box="[569,673,1399,1425]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="39" pageNumber="40">Fig. 22A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,229,1435,1460]" italics="true" pageId="39" pageNumber="40">Legs</emphasis>
|
||
: coxae 1 and 2 elongated and fused with sternum, podomeres from prefemur to tarsus in both sexes each with 4–10 ventral/mesal setae. Length of midbody legs circa 0.8 times body diameter in females (
|
||
<figureCitation box="[501,601,1504,1530]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="39" pageNumber="40">Fig. 22B</figureCitation>
|
||
), circa 1.0 times body diameter in males (
|
||
<figureCitation captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="39" pageNumber="40">Fig. 22C</figureCitation>
|
||
). Each podomere ventrally with an apical seta. Coxae 3 and beyond of cylindrical shape. Tarsus with a stout dorso-apical seta and three pairs of regular ventral spines.
|
||
</paragraph>
|
||
<paragraph blockId="39.[140,1108,167,1672]" pageId="39" pageNumber="40">
|
||
<emphasis box="[185,427,1610,1636]" italics="true" pageId="39" pageNumber="40">Male sexual characters</emphasis>
|
||
: tarsal pads absent (
|
||
<figureCitation box="[658,764,1610,1636]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="39" pageNumber="40">Fig. 22C</figureCitation>
|
||
). Coxae 3–5 each with a large conical coxal process, protruding posteriorly to coxa 6.
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,1108,167,651]" pageId="40" pageNumber="41">
|
||
<emphasis box="[185,372,167,193]" italics="true" pageId="40" pageNumber="41">Anterior gonopod</emphasis>
|
||
sternite elongated into a wide, rounded lobe, apically with three very short, well-rounded tips (
|
||
<figureCitation box="[477,584,202,228]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22D</figureCitation>
|
||
). Mesal process of coxite absent. Telopodite on posterior side basally with a circular groove (
|
||
<figureCitation box="[641,742,237,263]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22E</figureCitation>
|
||
). Telopodite process slender and short, curved laterally (
|
||
<figureCitation box="[400,502,272,298]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="40" pageNumber="41">Fig. 23E</figureCitation>
|
||
). Lateral margin elevated (
|
||
<figureCitation box="[800,902,272,298]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,1108,167,651]" pageId="40" pageNumber="41">
|
||
<emphasis box="[185,379,307,333]" italics="true" pageId="40" pageNumber="41">Posterior gonopods</emphasis>
|
||
unique (
|
||
<figureCitation box="[483,619,307,334]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Figs 22F, G</figureCitation>
|
||
). Gonopod pair connected by a sclerotized and visible sternite. Telopodites positioned parallel to one another, sperm canal discharging laterally (
|
||
<figureCitation box="[283,387,378,404]" captionStart="Figure 23" captionStartId="41.[141,213,1526,1548]" captionTargetBox="[140,1109,204,1471]" captionTargetId="figure@41.[140,1109,204,1471]" captionTargetPageId="41" captionText="Figure 23. Flagellobolus pauliani sp. n., male holotype, A telson B left midbody leg C anterior gonopod, anterior view D anterior gonopod, posterior view E posterior gonopods, posterior view, slightly turned F posterior gonopods, posterior view, original position. av = anal valves; cp = coxite process; Cx = coxite; Pre = preanal ring; rp = retrorse process; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767300" httpUri="https://zenodo.org/record/3767300/files/figure.png" pageId="40" pageNumber="41">Fig. 23G</figureCitation>
|
||
). Coxite and telopodite separated by a suture; coxite with a single groove. Telopodite with torsion, tip of telopodite curved laterally, with a sharp-edged tip (
|
||
<figureCitation box="[148,246,448,474]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22F</figureCitation>
|
||
). Sperm canal laterally strongly projecting over telopodite margin (
|
||
<figureCitation box="[990,1095,448,474]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22G</figureCitation>
|
||
). Basally of projection of sperm canal with a row of sclerotized nodules (
|
||
<figureCitation box="[913,1017,483,510]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22G</figureCitation>
|
||
). Sperm canal running through coxite at mesal, through telopodite at lateral margin (
|
||
<figureCitation box="[981,1085,519,545]" captionStart="Figure 22" captionStartId="38.[140,212,1526,1548]" captionTargetBox="[167,1080,166,1506]" captionTargetId="figure@38.[168,1080,165,1506]" captionTargetPageId="38" captionText="Figure 22. Zehntnerobolus rubripes comb. n., A, C–G male lectotype, B female paratype, A telson B, C right midbody leg D anterior gonopod, anterior view E anterior gonopod, posterior view F left posterior gonopod, anterior view G left posterior gonopod, posterior view. Apo = apodeme; av = anal valves; Cx = coxite; Gr = groove; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; Tp = telopodite process. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3767298" httpUri="https://zenodo.org/record/3767298/files/figure.png" pageId="40" pageNumber="41">Fig. 22G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,1108,167,651]" box="[185,752,554,580]" pageId="40" pageNumber="41">
|
||
<emphasis box="[185,446,554,580]" italics="true" pageId="40" pageNumber="41">Female sexual characters</emphasis>
|
||
: vulva simple, bivalve-like.
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,1108,167,651]" pageId="40" pageNumber="41">
|
||
<emphasis bold="true" box="[185,457,589,615]" pageId="40" pageNumber="41">
|
||
<emphasis box="[185,333,589,615]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Intraspecific</emphasis>
|
||
variation:
|
||
</emphasis>
|
||
the measurements of the
|
||
<typeStatus box="[756,804,590,616]" pageId="40" pageNumber="41">type</typeStatus>
|
||
series fit in very well with those recorded in the literature (de Saussure and Zehntner 1902: 159).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph blockId="40.[140,249,745,772]" box="[140,249,745,772]" pageId="40" pageNumber="41">
|
||
<emphasis bold="true" box="[140,249,745,772]" pageId="40" pageNumber="41">Remark</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,1108,800,932]" pageId="40" pageNumber="41">
|
||
The mentum is subdivided by a strong transverse ridge separating a basal, transversely striate part and an apical, smooth part, which is recessed when the gnathochilarium is viewed from below in all the following genera. Such a subdivision is unique within the whole order
|
||
<taxonomicName authorityName="Cook" authorityYear="1895" box="[282,416,906,932]" class="Diplopoda" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="order">Spirobolida</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,534,1011,1037]" pageId="40" pageNumber="41">
|
||
<emphasis bold="true" box="[140,534,1011,1037]" pageId="40" pageNumber="41">
|
||
<emphasis box="[140,226,1011,1037]" inLineHeading="true" pageId="40" pageNumber="41" reason="1">Genera</emphasis>
|
||
with subdivided mentum
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,370,1047,1073]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,278,1047,1073]" class="Diplopoda" family="Pachybolidae" genus="Flagellobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,278,1047,1073]" italics="true" pageId="40" pageNumber="41">Flagellobolus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel box="[293,370,1047,1073]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,368,1082,1108]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,283,1082,1108]" class="Diplopoda" family="Pachybolidae" genus="Riotintobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,283,1082,1108]" italics="true" pageId="40" pageNumber="41">Riotintobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[291,368,1082,1108]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,418,1117,1144]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,333,1117,1143]" class="Diplopoda" family="Pachybolidae" genus="Pseudocentrobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,333,1117,1143]" italics="true" pageId="40" pageNumber="41">Pseudocentrobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[341,418,1118,1144]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,365,1152,1179]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,280,1152,1178]" class="Diplopoda" family="Pachybolidae" genus="Granitobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,280,1152,1178]" italics="true" pageId="40" pageNumber="41">Granitobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[288,365,1153,1179]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,346,1187,1214]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,262,1187,1213]" class="Diplopoda" family="Pachybolidae" genus="Caprobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,262,1187,1213]" italics="true" pageId="40" pageNumber="41">Caprobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[269,346,1188,1214]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,358,1223,1249]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,274,1223,1249]" class="Diplopoda" family="Pachybolidae" genus="Alluviobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,274,1223,1249]" italics="true" pageId="40" pageNumber="41">Alluviobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[281,358,1223,1249]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph blockId="40.[140,534,1011,1284]" box="[140,351,1258,1284]" pageId="40" pageNumber="41">
|
||
<taxonomicName authority="Wesener & Enghoff & Sierwald, 2009" authorityName="Wesener & Enghoff & Sierwald" authorityYear="2009" box="[140,267,1258,1284]" class="Diplopoda" family="Pachybolidae" genus="Ostinobolus" kingdom="Animalia" order="Spirobolida" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. n.">
|
||
<emphasis box="[140,267,1258,1284]" italics="true" pageId="40" pageNumber="41">Ostinobolus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[274,351,1258,1284]" pageId="40" pageNumber="41" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSection>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |